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<document id="0DED8C185F084FB1DB170A37D1145E57" ID-DOI="10.59893/bjc.23(2).017" ID-ISSN="1407-8619" ID-Zenodo-Dep="13203287" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722657586546" checkinUser="felipe" docAuthor="Rukmane-Bārbale, Anita" docDate="2023" docId="03E587FAB44CB036334A0D13E6E7FA76" docLanguage="en" docName="BalticJColeopterol.23.2.283-294.pdf" docOrigin="Baltic Journal of Coleopterology 23 (2)" docSource="http://dx.doi.org/10.3390/ijms25147806" docStyle="DocumentStyle:514B3A4252435B0AFD496C900C6D7572.1:BalticJColeop.2023-.journal_article" docStyleId="514B3A4252435B0AFD496C900C6D7572" docStyleName="BalticJColeop.2023-.journal_article" docStyleVersion="1" docTitle="Metapocyrtus (Metapocyrtus) tetramaculatus Rukmane-Bārbale, 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="286" masterDocId="FFDCFF82B44DB035313A0C04E449FFA7" masterDocTitle="Four new species of the genus Metapocyrtus Heller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) from Mindanao Island, Philippines" masterLastPageNumber="294" masterPageNumber="283" pageNumber="284" updateTime="1722697247933" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="A0EBD3C7AC852A0626E84D0B612AD295" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3B989FBD28DCBF24C5419653102DF0B2">Four new species of the genus Metapocyrtus Heller, 1912 (Coleoptera: Curculionidae: Pachyrhynchini) from Mindanao Island, Philippines</mods:title>
</mods:titleInfo>
<mods:name id="ACD091AF2EA2F31BE4C33C1ABE6F41B7" type="personal">
<mods:role id="962FE11EB8C362017BD27A83F93DF2FA">
<mods:roleTerm id="711AC306F1DA73C18D0A0C499134914A">Author</mods:roleTerm>
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<mods:namePart id="67753B9930FD5C6A39159ADA9DAE21DC">Rukmane-Bārbale, Anita</mods:namePart>
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<mods:title id="503707FA7E61307AD9ADFDD7C876596F">Baltic Journal of Coleopterology</mods:title>
</mods:titleInfo>
<mods:part id="4CF1270EBEFC4D0796791C495DDC30DE">
<mods:date id="923EA7380BB2B820FED9388E8DFDEE62">2023</mods:date>
<mods:detail id="33C9E76A4600803FA1FF3D3C3042E355" type="volume">
<mods:number id="7EEEFC8F50DD37CD0281B427F202314C">23</mods:number>
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<mods:classification id="98AB417F2EEE8970CB67AAAD9D2684F2">journal article</mods:classification>
<mods:identifier id="DC38F2A2D35361CDC15C214C0E70B6B5" type="DOI">10.59893/bjc.23(2).017</mods:identifier>
<mods:identifier id="CC8302108B027E840DD2916B2B994307" type="ISSN">1407-8619</mods:identifier>
<mods:identifier id="A49CCA201D26ADAA53A04ED80A8D47D3" type="Zenodo-Dep">13203287</mods:identifier>
</mods:mods>
<treatment id="03E587FAB44CB036334A0D13E6E7FA76" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76" httpUri="http://treatment.plazi.org/id/03E587FAB44CB036334A0D13E6E7FA76" lastPageId="3" lastPageNumber="286" pageId="1" pageNumber="284">
<subSubSection id="C3566567B44CB034334A0D13E6A7FEEA" pageId="1" pageNumber="284" type="nomenclature">
<paragraph id="8BF336ECB44CB034334A0D13E6A7FEEA" blockId="1.[624,1097,279,364]" pageId="1" pageNumber="284">
<heading id="D0BB8180B44CB034334A0D13E6A7FEEA" bold="true" centered="true" fontSize="10" level="2" pageId="1" pageNumber="284" reason="4">
<emphasis id="B938EAFEB44CB034334A0D13E6A7FEEA" bold="true" pageId="1" pageNumber="284">
<taxonomicName id="4C4C4D6FB44CB034334A0D13E6D9FEE9" ID-CoL="C3BVK" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB034334A0D13E6D9FEE9" bold="true" italics="true" pageId="1" pageNumber="284">Metapocyrtus (Metapocyrtus) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03433AD0D31E6A7FEEA" box="[663,750,309,333]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44CB034334A0D50E77DFECB" box="[624,820,340,364]" pageId="1" pageNumber="284" type="description">
<paragraph id="8BF336ECB44CB034334A0D50E77DFECB" blockId="1.[624,1097,279,364]" box="[624,820,340,364]" pageId="1" pageNumber="284">
(
<figureCitation id="13772A69B44CB03433420D50E685FECB" box="[632,716,340,364]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A – M. (M.) tetramaculatus sp. nov.; B – M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C – M. (M.) fulvus sp. nov.; D – M. (T.) unalineatus sp. nov.; E – M. (T.) pseudolanaoensis sp. nov.; F – M. (M.) lanaoensis Bollino, 2021." figureDoi="http://doi.org/10.5281/zenodo.13203289" httpUri="https://zenodo.org/record/13203289/files/figure.png" pageId="1" pageNumber="284">Fig. 1A</figureCitation>
,
<figureCitation id="13772A69B44CB03433E20D50E6AEFECB" box="[728,743,340,364]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A – dorsal view, male, holotype; B – dorsal view, female; C – lateral view, male, holotype." figureDoi="http://doi.org/10.5281/zenodo.13203293" httpUri="https://zenodo.org/record/13203293/files/figure.png" pageId="1" pageNumber="284">2</figureCitation>
,
<figureCitation id="13772A69B44CB03433C80D50E765FECB" box="[754,812,340,364]" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E – aedegal body in lateral view; B, D, F – sternite IX; A, B – M. (M.) tetramaculatus sp. nov., C, D – M. (M.) fulvus sp. nov., E, F – M. (T.) unalineatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13203301" httpUri="https://zenodo.org/record/13203301/files/figure.png" pageId="1" pageNumber="284">4A,B</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF336664B44CB03431480813E545FB2C" ID-DOI="http://doi.org/10.5281/zenodo.13203289" ID-Zenodo-Dep="13203289" httpUri="https://zenodo.org/record/13203289/files/figure.png" pageId="1" pageNumber="284" startId="1.[114,153,1047,1071]" targetBox="[114,1096,447,1040]" targetPageId="1" targetType="figure">
<paragraph id="8BF336ECB44CB03431480813E050FB88" blockId="1.[114,1070,1047,1163]" box="[114,1049,1047,1071]" pageId="1" pageNumber="284">
Fig. 1. Dorsal view of rostrum of selected species: A –
<taxonomicName id="4C4C4D6FB44CB03433850813E7FEFB88" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[703,951,1047,1071]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433850813E7FEFB88" box="[703,951,1047,1071]" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03432840813E05CFB88" box="[958,1045,1047,1071]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
;
</paragraph>
<paragraph id="8BF336ECB44CB03431480831E545FB2C" blockId="1.[114,1070,1047,1163]" pageId="1" pageNumber="284">
B –
<taxonomicName id="4C4C4D6FB44CB03431A40832E6ECFBEA" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" box="[158,677,1077,1102]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03431A40832E5D3FBE9" box="[158,410,1078,1102]" italics="true" pageId="1" pageNumber="284">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
; C –
<taxonomicName id="4C4C4D6FB44CB03433E60832E73DFBE9" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[732,884,1078,1102]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="fulvus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433E60832E73DFBE9" box="[732,884,1078,1102]" italics="true" pageId="1" pageNumber="284">M. (M.) fulvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03432410831E79BFBEA" box="[891,978,1077,1101]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; D –
<taxonomicName id="4C4C4D6FB44CB03435300832E555FBCA" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="unalineatus" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44CB03435300832E555FBCA" italics="true" pageId="1" pageNumber="284">M. (T.) unalineatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03430190850E533FBCB" box="[291,378,1108,1132]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; E –
<taxonomicName id="4C4C4D6FB44CB03430950851E6F7FBCA" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[431,702,1109,1133]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="pseudolanaoensis" status="sp. nov." subGenus="Trachycyrtus">
<emphasis id="B938EAFEB44CB03430950851E6F7FBCA" box="[431,702,1109,1133]" italics="true" pageId="1" pageNumber="284">M. (T.) pseudolanaoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44CB03433FF0850E754FBCB" box="[709,797,1108,1132]" pageId="1" pageNumber="284" rank="species">sp. nov.</taxonomicNameLabel>
; F –
<taxonomicName id="4C4C4D6FB44CB034326B0851E54EFB2C" authority="Bollino, 2021" authorityName="Bollino" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="lanaoensis" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB034326B0851E055FBCA" box="[849,1052,1109,1133]" italics="true" pageId="1" pageNumber="284">M. (M.) lanaoensis</emphasis>
Bollino, 2021
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3566567B44CB034314808B4E683FAC5" pageId="1" pageNumber="284" type="materials_examined">
<paragraph id="8BF336ECB44CB034314808B4E644FAC5" blockId="1.[114,593,1200,1378]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB034314808B4E569FB6F" bold="true" box="[114,288,1200,1224]" pageId="1" pageNumber="284">Type material.</emphasis>
<materialsCitation id="3B243CB1B44CB034301608B4E513FA83" collectingDate="2015-09" country="Philippines" county="Sibagat" location="Mindanao" municipality="Agusan" pageId="1" pageNumber="284" specimenCount="1" typeStatus="holotype">
<emphasis id="B938EAFEB44CB034301608B4E5AEFB6F" bold="true" box="[300,487,1200,1224]" pageId="1" pageNumber="284">
<typeStatus id="54F7884EB44CB034301608B4E5DCFB6F" box="[300,405,1200,1224]" pageId="1" pageNumber="284" type="holotype">Holotype</typeStatus>
, male:
</emphasis>
‘’
<collectingCountry id="F35B767CB44CB03430C708B4E49DFB40" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<location id="8E936037B44CB03431D308CBE511FB40" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB03431D308CBE511FB40" box="[233,344,1231,1255]" country="Philippines" county="Sibagat" municipality="Agusan" name="Mindanao" pageId="1" pageNumber="284">Mindanao</location>
,
<collectingMunicipality id="6B97AC96B44CB034305908CBE5FEFB40" box="[355,439,1231,1255]" pageId="1" pageNumber="284">Agusan</collectingMunicipality>
,
<collectingCounty id="62924E60B44CB03430F808CBE65BFB40" box="[450,530,1231,1255]" pageId="1" pageNumber="284">Sibagat</collectingCounty>
/
<date id="FFF2102CB44CB034331D08CBE4E1FAA2" pageId="1" pageNumber="284" value="2015-09">
<collectingDate id="EFB6E9C4B44CB034331D08CBE4E1FAA2" pageId="1" pageNumber="284" value="2015-09">IX. 2015</collectingDate>
</date>
/ local collector leg.’’ (white rectangular label, printed) //
</materialsCitation>
‘’
<typeStatus id="54F7884EB44CB034304B0908E64EFA83" box="[369,519,1292,1316]" pageId="1" pageNumber="284" type="holotype">HOLOTYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44CB03433170909E514FAE4" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB03433170909E514FAE4" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023’’ (red rectangular label, printed).
</paragraph>
<paragraph id="8BF336ECB44CB03431480983E603F9BD" blockId="1.[114,593,1415,1562]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB03431480983E536FA38" bold="true" box="[114,383,1415,1439]" pageId="1" pageNumber="284">
<typeStatus id="54F7884EB44CB03431480983E4A1FA38" box="[114,232,1415,1439]" pageId="1" pageNumber="284" type="paratype">Paratypes</typeStatus>
: (
<specimenCount id="9D4AFD65B44CB03431C50983E53AFA38" box="[255,371,1415,1439]" count="3" pageId="1" pageNumber="284" type="female">3 females</specimenCount>
):
</emphasis>
‘’
<materialsCitation id="3B243CB1B44CB03430A80983E53EFA7B" collectingDate="2015-09" country="Philippines" county="Mindanao" location="Sibagat" municipality="Agusan" pageId="1" pageNumber="284" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F35B767CB44CB03430A80983E671FA38" box="[402,568,1415,1439]" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<collectingCounty id="62924E60B44CB034314809A1E4A8FA1A" box="[114,225,1445,1469]" pageId="1" pageNumber="284">Mindanao</collectingCounty>
,
<collectingMunicipality id="6B97AC96B44CB03431C909A1E50EFA1A" box="[243,327,1445,1469]" pageId="1" pageNumber="284">Agusan</collectingMunicipality>
,
<location id="8E936037B44CB034306009A1E5E3FA1A" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB034306009A1E5E3FA1A" box="[346,426,1445,1469]" country="Philippines" county="Mindanao" municipality="Agusan" name="Sibagat" pageId="1" pageNumber="284">Sibagat</location>
/
<date id="FFF2102CB44CB03430F709A1E67CFA1A" box="[461,565,1445,1469]" pageId="1" pageNumber="284" value="2015-09">
<collectingDate id="EFB6E9C4B44CB03430F709A1E67CFA1A" box="[461,565,1445,1469]" pageId="1" pageNumber="284" value="2015-09">IX. 2015</collectingDate>
</date>
/ local collector leg.’’ //
</materialsCitation>
‘’
<materialsCitation id="3B243CB1B44CB03430B709C0E603F9BD" collectingDate="2015-10" country="Philippines" location="Cabanglasan" municipality="Mindanao" pageId="1" pageNumber="284" specimenCount="1" stateProvince="Bukidnon" typeStatus="paratype">
<collectingCountry id="F35B767CB44CB03430B709C0E67AFA7B" box="[397,563,1476,1500]" name="Philippines" pageId="1" pageNumber="284">PHILIPPINES</collectingCountry>
/
<collectingMunicipality id="6B97AC96B44CB034314809E7E4A8FA5C" box="[114,225,1507,1531]" pageId="1" pageNumber="284">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB44CB03431C209E7E52DFA5C" box="[248,356,1507,1531]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="284">Bukidnon</collectingRegion>
,
<location id="8E936037B44CB034304109E7E64AFA5C" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB034304109E7E64AFA5C" box="[379,515,1507,1531]" country="Philippines" municipality="Mindanao" name="Cabanglasan" pageId="1" pageNumber="284" stateProvince="Bukidnon">Cabanglasan</location>
/
<date id="FFF2102CB44CB034330B09E7E4E1F9BD" pageId="1" pageNumber="284" value="2015-10">
<collectingDate id="EFB6E9C4B44CB034330B09E7E4E1F9BD" pageId="1" pageNumber="284" value="2015-10">X. 2015</collectingDate>
</date>
/ local collector leg.’’// ‘’PHILIP-
</materialsCitation>
</paragraph>
<paragraph id="8BF336ECB44CB03431480AA2E4D3F919" blockId="1.[114,154,1702,1726]" box="[114,154,1702,1726]" pageId="1" pageNumber="284">284</paragraph>
<paragraph id="8BF336ECB44CB034334A08B4E683FAC5" blockId="1.[624,1103,1200,1378]" pageId="1" pageNumber="284">
<materialsCitation id="3B243CB1B44CB034334A08B4E006FAE4" collectingDate="2022-08" collectionCode="PINES" location="Cabanglasan" municipality="Mindanao" pageId="1" pageNumber="284" specimenCount="1" stateProvince="Bukidnon" typeStatus="paratype">
<collectionCode id="ED5DAE29B44CB034334A08B4E6F3FB6F" box="[624,698,1200,1224]" pageId="1" pageNumber="284">PINES</collectionCode>
/
<collectingMunicipality id="6B97AC96B44CB03433F508B4E777FB6F" box="[719,830,1200,1224]" pageId="1" pageNumber="284">Mindanao</collectingMunicipality>
,
<collectingRegion id="4988F80EB44CB034327308B4E7FCFB6F" box="[841,949,1200,1224]" country="Philippines" name="Bukidnon" pageId="1" pageNumber="284">Bukidnon</collectingRegion>
,
<location id="8E936037B44CB03432FA08B4E006FB6F" LSID="urn:lsid:plazi:treatment:03E587FAB44CB036334A0D13E6E7FA76:8E936037B44CB03432FA08B4E006FB6F" box="[960,1103,1200,1224]" municipality="Mindanao" name="Cabanglasan" pageId="1" pageNumber="284" stateProvince="Bukidnon">Cabanglasan</location>
/
<date id="FFF2102CB44CB03433B908CBE6B1FB40" box="[643,760,1231,1255]" pageId="1" pageNumber="284" value="2022-08">
<collectingDate id="EFB6E9C4B44CB03433B908CBE6B1FB40" box="[643,760,1231,1255]" pageId="1" pageNumber="284" value="2022-08">VIII. 2022</collectingDate>
</date>
/ local collector leg.’’ (white rectangular label, printed), with additional red label: ‘’
<typeStatus id="54F7884EB44CB03433CA0908E7CAFA83" box="[752,899,1292,1316]" pageId="1" pageNumber="284" type="paratype">PARATYPE</typeStatus>
/
<taxonomicName id="4C4C4D6FB44CB034329C0909E697FAE4" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="284" phylum="Arthropoda" rank="species" species="tetramaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44CB034329C0909E697FAE4" italics="true" pageId="1" pageNumber="284">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
/ Rukmane-Bārbale, 2023’’ (all in
</materialsCitation>
<collectionCode id="ED5DAE29B44CB034334A094EE6F6FAC5" box="[624,703,1354,1378]" pageId="1" pageNumber="284">DUBC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44CB036334A0983E78DFB7B" lastPageId="3" lastPageNumber="286" pageId="1" pageNumber="284" type="description">
<paragraph id="8BF336ECB44CB034334A0983E71EFA5C" blockId="1.[624,1103,1415,1531]" pageId="1" pageNumber="284">
<emphasis id="B938EAFEB44CB034334A0983E798FA38" bold="true" box="[624,977,1415,1439]" pageId="1" pageNumber="284">
Description.
<typeStatus id="54F7884EB44CB03432310983E73CFA38" box="[779,885,1415,1439]" pageId="1" pageNumber="284" type="holotype">Holotype</typeStatus>
, Male.
</emphasis>
(
<figureCitation id="13772A69B44CB03432D90983E00FFA38" box="[995,1094,1415,1439]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A – dorsal view, male, holotype; B – dorsal view, female; C – lateral view, male, holotype." figureDoi="http://doi.org/10.5281/zenodo.13203293" httpUri="https://zenodo.org/record/13203293/files/figure.png" pageId="1" pageNumber="284">Fig. 2A</figureCitation>
) Dimensions: LB: 6,1, LR: 1,1, WR: 0,9, LP: 2,1, WP: 1,9, LE: 4,0, WE: 2,8. N = 1 for all measurements.
</paragraph>
<paragraph id="8BF336ECB44FB03731FD0C67E0C0FFD1" blockId="2.[199,1161,99,118]" box="[199,1161,99,118]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB03731FD0C67E0C0FFD1" box="[199,1161,99,118]" italics="true" pageId="2" pageNumber="285">
Four new species of the genus
<taxonomicName id="4C4C4D6FB44FB03730F50C67E6FCFFD1" authority="Heller, 1912" authorityName="Heller" authorityYear="1912" box="[463,693,99,118]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="genus">
Metapocyrtus
<bibRefCitation id="EFDD4B1DB44FB03733730C67E6FCFFD1" author="Heller K. M." box="[585,693,99,118]" pageId="2" pageNumber="285" pagination="295 - 402" refId="ref4196" refString="Heller K. M. 1912. Philippinische Russelkafer. The Philippine Journal of Science, 7 (5): 295 - 402." type="journal article" year="1912">Heller, 1912</bibRefCitation>
</taxonomicName>
(
<taxonomicName id="4C4C4D6FB44FB03733FB0C67E76DFFD1" authorityName="Linnaeus" authorityYear="1758" box="[705,804,99,118]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="2" pageNumber="285" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44FB03732140C67E7E2FFD1" box="[814,939,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="family">Curculionidae</taxonomicName>
:
<taxonomicName id="4C4C4D6FB44FB037328F0C67E074FFD1" authorityName="Schoenherr" authorityYear="1826" box="[949,1085,99,118]" class="Insecta" family="Curculionidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="tribe" tribe="Pachyrhynchini">Pachyrhynchini</taxonomicName>
) from …
</emphasis>
</paragraph>
<caption id="DF336664B44FB0373187096EE6B3FA06" ID-DOI="http://doi.org/10.5281/zenodo.13203293" ID-Zenodo-Dep="13203293" httpUri="https://zenodo.org/record/13203293/files/figure.png" pageId="2" pageNumber="285" startId="2.[189,228,1386,1410]" targetBox="[193,1172,171,1380]" targetPageId="2" targetType="figure">
<paragraph id="8BF336ECB44FB0373187096EE6B3FA06" blockId="2.[189,1122,1386,1441]" pageId="2" pageNumber="285">
Fig. 2. Habitus of
<taxonomicName id="4C4C4D6FB44FB03730BA096FE631FA24" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[384,632,1387,1411]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44FB03730BA096FE631FA24" box="[384,632,1387,1411]" italics="true" pageId="2" pageNumber="285">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44FB0373345096EE69DFA25" box="[639,724,1386,1410]" pageId="2" pageNumber="285" rank="species">sp. nov.</taxonomicNameLabel>
A – dorsal view, male, holotype; B – dorsal view, female; C – lateral view, male, holotype.
</paragraph>
</caption>
<paragraph id="8BF336ECB44FB037318709C2E519F9BC" blockId="2.[189,668,1478,1563]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB037318709C2E50FFA79" bold="true" box="[189,326,1478,1502]" pageId="2" pageNumber="285">Coloration.</emphasis>
Integument black, shiny, with markings of round, metallic green to turquoise scales.
</paragraph>
<paragraph id="8BF336ECB44FB03731870A45E0DDF9DF" blockId="2.[189,662,1601,1656]" lastBlockId="2.[699,1178,1478,1656]" pageId="2" pageNumber="285">
<emphasis id="B938EAFEB44FB03731870A45E54EF9FE" bold="true" box="[189,263,1601,1625]" pageId="2" pageNumber="285">Head.</emphasis>
Dorsal surface smooth, with longitudinal line of green to turquoise scales; surface between eyes with sparse white to greenish setae; genae with patch of metallic green to turquoise scales and green hair-like scales; shallow medial longitudinal groov from base of the eyes to base anterior margin of prothorax; transverse, moderate fur-
</paragraph>
<paragraph id="8BF336ECB44FB03735560AA2E0DDF919" blockId="2.[1132,1172,1702,1726]" box="[1132,1172,1702,1726]" pageId="2" pageNumber="285">285</paragraph>
<paragraph id="8BF336ECB44EB03631480CB6E5EEFEA0" blockId="3.[114,592,178,263]" pageId="3" pageNumber="286">row at base of eyes confluent with medial longitudinal groove; eyes large, strongly convex from outline of head.</paragraph>
<paragraph id="8BF336ECB44EB03631480D28E58DFDFF" blockId="3.[114,593,300,600]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480D28E4ADFEE3" bold="true" box="[114,228,300,324]" pageId="3" pageNumber="286">Rostrum.</emphasis>
(
<figureCitation id="13772A69B44EB03631C10D28E512FEE3" box="[251,347,300,324]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A – M. (M.) tetramaculatus sp. nov.; B – M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C – M. (M.) fulvus sp. nov.; D – M. (T.) unalineatus sp. nov.; E – M. (T.) pseudolanaoensis sp. nov.; F – M. (M.) lanaoensis Bollino, 2021." figureDoi="http://doi.org/10.5281/zenodo.13203289" httpUri="https://zenodo.org/record/13203289/files/figure.png" pageId="3" pageNumber="286">Fig. 1A</figureCitation>
) longer than wide (LR/WR: 1,22), smooth at posterior half, slightly punctured at anterior half; green to white setae from apical ½ to base; dorsum with moderate medial longitudinal furrow and corresponding shallow concavity; in dorsal contour straight; in lateral contourraised dorsally at base, then gradually declined to apex; lateral parts covered with long white to shiny green setae.
</paragraph>
<paragraph id="8BF336ECB44EB03631480E79E514FC2C" blockId="3.[114,592,637,907]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480E79E4AFFD32" bold="true" box="[114,230,637,661]" pageId="3" pageNumber="286">Antennae</emphasis>
with antennal scrobe not reaching margin of eye, curved downwards; antennal scape reaching anterior margin of prothorax; funicular segment I nearly three times as long as wide, slightly longer than II, segment II nearly two times longer than III, segments III – VI as long as wide, segment VII larger, as longas wide, club 2,5 times as long as wide.
</paragraph>
<paragraph id="8BF336ECB44EB03631480FB4E4BFFA14" blockId="3.[114,593,944,1459]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB03631480FB4E4A7FC6F" bold="true" box="[114,238,944,968]" pageId="3" pageNumber="286">Prothorax</emphasis>
sub cylindrical, slightly longer than wide (LP/WP: 1,1), strongly punctured, with corresponding, short, white setae; widest just before the middle; in dorsal contour narrowed along anterior margin, widened to just before the middle, gradually narrowed to before posterior margin, straight to posterior margin; with the following markings of round, metallic green to turquoise shales: 1) two spots along anterior margin on disc, each redirected laterally; 2) two spots at basal part, from middle to before posterior margin on disc, each redirected laterally; anterior groove confluent at disc, slightly redirected to middle; posterior groove confluent at disc, curved upwards at lateral parts.
</paragraph>
<paragraph id="8BF336ECB44EB036314809DCE619F9EB" blockId="3.[114,593,1496,1612]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036314809DCE488FA57" bold="true" box="[114,193,1496,1520]" pageId="3" pageNumber="286">Elytra</emphasis>
sub-ellipsoidal, (LE/WE: 1,42), wider than prothorax nearly twice as long as prothorax (WE/WP: 1,47; LE/LP: 1,9); punctured in all length, without expressed
</paragraph>
<paragraph id="8BF336ECB44EB03631480AA2E7D5FE07" blockId="3.[114,154,1702,1726]" lastBlockId="3.[624,1103,178,416]" pageId="3" pageNumber="286">286 puncture rows; in dorsal contour widest just before the middle, moderately convex; each elytron with the following scally markings of round, metallic green to turquoise scales: 1) two sub basal spots; 2) three spots at the medial part; 3) three spots along apical 1/3; 4) longitudinal apical stripe; goldish, thick setae from apical ½ to apex.</paragraph>
<paragraph id="8BF336ECB44EB036334A0DC1E6C1FDBC" blockId="3.[624,1103,453,539]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0DC1E6A0FE7A" bold="true" box="[624,745,453,477]" pageId="3" pageNumber="286">Fore tibia</emphasis>
without denticles along internal margin. Male genitalia as shown in
<figureCitation id="13772A69B44EB03632CD0DE0E6CDFDBC" captionStart="Fig" captionStartId="5.[114,153,1533,1557]" captionTargetBox="[120,1022,855,1503]" captionTargetId="figure-308@5.[114,1032,846,1526]" captionTargetPageId="5" captionText="Fig. 4. Male genitalia of selected species, A, C, E – aedegal body in lateral view; B, D, F – sternite IX; A, B – M. (M.) tetramaculatus sp. nov., C, D – M. (M.) fulvus sp. nov., E, F – M. (T.) unalineatus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13203301" httpUri="https://zenodo.org/record/13203301/files/figure.png" pageId="3" pageNumber="286">Fig.4A, B</figureCitation>
.
</paragraph>
<paragraph id="8BF336ECB44EB036334A0E44E68EFD13" blockId="3.[624,1103,576,692]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0E44E699FDFF" bold="true" box="[624,720,576,600]" pageId="3" pageNumber="286">Female.</emphasis>
Elytra wider with strongly expressed apex, legs not as thick as in males, otherwise as in males; dorsal habitus as in
<figureCitation id="13772A69B44EB036334A0E98E68AFD13" box="[624,707,668,692]" captionStart="Fig" captionStartId="2.[189,228,1386,1410]" captionTargetBox="[193,1172,171,1380]" captionTargetId="figure-114@2.[193,1172,171,1380]" captionTargetPageId="2" captionText="Fig. 2. Habitus of M. (M.) tetramaculatus sp. nov. A – dorsal view, male, holotype; B – dorsal view, female; C – lateral view, male, holotype." figureDoi="http://doi.org/10.5281/zenodo.13203293" httpUri="https://zenodo.org/record/13203293/files/figure.png" pageId="3" pageNumber="286">Fig. 2B</figureCitation>
.
</paragraph>
<paragraph id="8BF336ECB44EB036334A0EDDE78DFB7B" blockId="3.[624,1103,729,1245]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0EDDE723FD56" bold="true" box="[624,874,729,753]" pageId="3" pageNumber="286">Differential analyses.</emphasis>
The new species by black body colour, small size and presence of four scally spots on disc of prothorax, two at basal part and two along apical margin, is similar to
<taxonomicName id="4C4C4D6FB44EB03632050F51E7CDFC2C" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03632050F51E001FCCA" box="[831,1096,853,877]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
, the new species can be easily distinguished by the following characters: 1) large, strongly prominent eyes in
<taxonomicName id="4C4C4D6FB44EB03633E90FCBE794FC40" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[723,989,975,999]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633E90FCBE794FC40" box="[723,989,975,999]" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB03632D60FCBE000FC40" box="[1004,1097,975,999]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
instead of relatively small, weakly prominent eyes in
<taxonomicName id="4C4C4D6FB44EB03633C30809E696FBE4" authority="Cabras &amp; Medina" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633C30809E7BEFB82" box="[761,1015,1037,1061]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina
</taxonomicName>
(
<figureCitation id="13772A69B44EB03633CB082FE70FFBE4" box="[753,838,1067,1091]" captionStart="Fig" captionStartId="1.[114,153,1047,1071]" captionTargetBox="[114,1096,447,1040]" captionTargetId="figure-245@1.[114,1096,447,1040]" captionTargetPageId="1" captionText="Fig. 1. Dorsal view of rostrum of selected species: A – M. (M.) tetramaculatus sp. nov.; B – M. (M.) flavomaculatus Cabras &amp; Medina, 2021; C – M. (M.) fulvus sp. nov.; D – M. (T.) unalineatus sp. nov.; E – M. (T.) pseudolanaoensis sp. nov.; F – M. (M.) lanaoensis Bollino, 2021." figureDoi="http://doi.org/10.5281/zenodo.13203289" httpUri="https://zenodo.org/record/13203289/files/figure.png" pageId="3" pageNumber="286">Fig. 1B</figureCitation>
); 2) scally markings of green to turquoise colour in
<taxonomicName id="4C4C4D6FB44EB0363297084EE697FB27" authorityName="Rukmane-Bārbale" authorityYear="2023" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB0363297084EE697FB27" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB03633DD086CE777FB27" box="[743,830,1128,1152]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
instead of yellow ochre spots in
<taxonomicName id="4C4C4D6FB44EB03633EA0883E741FB1A" authority="Cabras &amp; Medina, 2021" authorityName="Cabras &amp; Medina" authorityYear="2021" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="flavomaculatus" subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB03633EA0883E79DFB38" box="[720,980,1159,1183]" italics="true" pageId="3" pageNumber="286">M. (M.) flavomaculatus</emphasis>
Cabras &amp; Medina, 2021
</taxonomicName>
(
<figureCitation id="13772A69B44EB036322108A1E728FB1A" box="[795,865,1189,1213]" captionStart="Fig" captionStartId="4.[189,228,1083,1107]" captionTargetBox="[189,1091,171,1077]" captionTargetId="figure-205@4.[189,1091,171,1077]" captionTargetPageId="4" captionText="Fig. 3. Habitus of M. (M.) flavomaculatus Cabras &amp; Medina, 2021, female; A – dorsal view; B – lateral view." figureDoi="http://doi.org/10.5281/zenodo.13203297" httpUri="https://zenodo.org/record/13203297/files/figure.png" pageId="3" pageNumber="286">Fig. 3</figureCitation>
); 3) narrow body of
<taxonomicName id="4C4C4D6FB44EB036334A08C1E720FB7A" authorityName="Rukmane-Bārbale" authorityYear="2023" box="[624,873,1221,1245]" class="Insecta" family="Curculionidae" genus="Metapocyrtus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="286" phylum="Arthropoda" rank="species" species="tetramaculatus" status="sp. nov." subGenus="Metapocyrtus">
<emphasis id="B938EAFEB44EB036334A08C1E720FB7A" box="[624,873,1221,1245]" italics="true" pageId="3" pageNumber="286">M. (M.) tetramaculatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5785B44EB036325508C0E78DFB7B" box="[879,964,1220,1244]" pageId="3" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44EB036334A0905E058FA9F" pageId="3" pageNumber="286" type="distribution">
<paragraph id="8BF336ECB44EB036334A0905E058FA9F" blockId="3.[624,1096,1281,1336]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0905E741FABE" bold="true" box="[624,776,1281,1305]" pageId="3" pageNumber="286">Distribution.</emphasis>
North Central Mindanao Island,
<collectingRegion id="4988F80EB44EB03633910924E75DFA9F" box="[683,788,1312,1336]" country="Philippines" name="Bukidnon" pageId="3" pageNumber="286">Bukidnon</collectingRegion>
and Agusan Provinces.
</paragraph>
</subSubSection>
<subSubSection id="C3566567B44EB036334A0959E6E7FA76" pageId="3" pageNumber="286" type="etymology">
<paragraph id="8BF336ECB44EB036334A0959E6E7FA76" blockId="3.[624,1103,1373,1489]" pageId="3" pageNumber="286">
<emphasis id="B938EAFEB44EB036334A0959E6B1FAD2" bold="true" box="[624,760,1373,1397]" pageId="3" pageNumber="286">Etymology.</emphasis>
The specific epithet is a latinised designation of four scally spots on disc of prothorax. Tetra – four, maculata – spots.
</paragraph>
</subSubSection>
</treatment>
</document>
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