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<mods:title id="81AA27B12773D938EA67973280976596">Revisionary checklist of the Southern African Sesiini (Lepidoptera: Sesiidae) with description of new species</mods:title>
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<subSubSection id="C356657AFFCB440EFF4FF8D5FDFBD813" box="[151,606,1878,1904]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BF336F1FFCB440EFF4FF8D5FDFBD813" blockId="4.[151,606,1878,1904]" box="[151,606,1878,1904]" pageId="4" pageNumber="5">
<heading id="D0BB819DFFCB440EFF4FF8D5FDFBD813" bold="true" box="[151,606,1878,1904]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B938EAE3FFCB440EFF4FF8D5FDFBD813" bold="true" box="[151,606,1878,1904]" pageId="4" pageNumber="5">
<taxonomicName id="4C4C4D72FFCB440EFF4FF8D5FE4BD813" ID-CoL="B8XNC" authority="Holland, 1893" authorityName="Holland" authorityYear="1893" box="[151,494,1878,1904]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCB440EFF4FF8D5FE9DD813" bold="true" box="[151,312,1878,1904]" italics="true" pageId="4" pageNumber="5">Cicinnoscelis</emphasis>
<bibRefCitation id="EFDD4B00FFCB440EFEE7F8D5FE4BD813" author="Holland" box="[319,494,1878,1904]" firstAuthor="Holland" pageId="4" pageNumber="5" pagination="181 - 184" refId="ref28768" refString="Holland, W. J. (1893) Four new genera and species of West African Sesiidae. Journal of the New York Entomological Society, 1, 181 - 184. http: // dx. doi. org / 10.4039 / ent 251 - 1" type="journal article" year="1893">Holland, 1893</bibRefCitation>
</taxonomicName>
rev. stat.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFCB440FFF4FF81DFB1FDD74" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BF336F1FFCB440EFF4FF81DFC69D8D0" blockId="4.[151,1436,1949,2032]" box="[151,972,1949,1971]" pageId="4" pageNumber="5">
<typeStatus id="54F78853FFCB440EFF4FF81DFF6CD8D0" box="[151,201,1950,1971]" pageId="4" pageNumber="5">Type</typeStatus>
species:
<taxonomicName id="4C4C4D72FFCB440EFEFEF81EFD74D8D0" authority="Holland 1893: 183" authorityName="Holland" authorityPageNumber="183" authorityYear="1893" box="[294,721,1949,1971]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B938EAE3FFCB440EFEFEF81EFDAFD8D0" box="[294,522,1949,1971]" italics="true" pageId="4" pageNumber="5">Cicinnoscelis longipes</emphasis>
<bibRefCitation id="EFDD4B00FFCB440EFDC9F81EFD74D8D0" author="Holland" box="[529,721,1949,1971]" firstAuthor="Holland" page="183" pageId="4" pageNumber="5" pagination="181 - 184" refId="ref28768" refString="Holland, W. J. (1893) Four new genera and species of West African Sesiidae. Journal of the New York Entomological Society, 1, 181 - 184. http: // dx. doi. org / 10.4039 / ent 251 - 1" type="journal article" year="1893">Holland 1893: 183</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BF336F1FFCB440EFF4FF83FFB25D893" blockId="4.[151,1436,1949,2032]" pageId="4" pageNumber="5">
Literature.
<bibRefCitation id="EFDD4B00FFCB440EFED0F83FFE6FD8B1" author="Hampson" box="[264,458,1980,2002]" firstAuthor="Hampson" page="78" pageId="4" pageNumber="5" pagination="46 - 119" refId="ref28526" refString="Hampson, G. F. (1919) A classification of the Aegeriadae of the Oriental and Ethiopian Regions. Novitates Zoologicae, 26 (1), 46 - 119." type="journal article" year="1919">Hampson 1919: 78</bibRefCitation>
(as
<taxonomicName id="4C4C4D72FFCB440EFE2FF83FFDD6D8B1" authorityName="von Dalla Torre &amp; Strand" authorityYear="1925" box="[503,627,1980,2002]" class="Insecta" family="Sesiidae" genus="Cicinoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCB440EFE2FF83FFDD6D8B1" box="[503,627,1980,2002]" italics="true" pageId="4" pageNumber="5">Cicinoscelis</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFDD4B00FFCB440EFD66F83FFBA0D8B1" author="Dalla Torre &amp; Strand" box="[702,1029,1980,2002]" firstAuthor="Dalla Torre" page="120" pageId="4" pageNumber="5" refId="ref27751" refString="Dalla Torre, K. W. &amp; Strand, E. (1925) Aegeriidae. Lepidopterorum Catalogus, 31. W. Junk, Berlin, 202 pp." type="book" year="1925">Dalla Torre &amp; Strand 1925: 120</bibRefCitation>
(as
<taxonomicName id="4C4C4D72FFCB440EFBEAF83FFB0BD8B1" authorityName="von Dalla Torre &amp; Strand" authorityYear="1925" box="[1074,1198,1980,2002]" class="Insecta" family="Sesiidae" genus="Cicinoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCB440EFBEAF83FFB0BD8B1" box="[1074,1198,1980,2002]" italics="true" pageId="4" pageNumber="5">Cicinoscelis</emphasis>
</taxonomicName>
[sic]);
<bibRefCitation id="EFDD4B00FFCB440EFB20F83FFF46D893" author="Naumann" firstAuthor="Naumann" page="14" pageId="4" pageNumber="5" refId="ref29205" refString="Naumann, C. M. (1971) Untersuchungen zur Systematik und Phylogenese der holarktischen Sesiiden (Insecta, Lepidoptera). Bonner Zoologische Monographien, 1, 190 pp." type="book" year="1971">Naumann 1971: 14</bibRefCitation>
;
<bibRefCitation id="EFDD4B00FFCB440EFF35F858FD9CD893" author="Heppner &amp; Duckworth" box="[237,569,2010,2032]" firstAuthor="Heppner" page="42" pageId="4" pageNumber="5" pagination="1 - 144" refId="ref28716" refString="Heppner, J. B. &amp; Duckworth, W. D. (1981) Classification of the Superfamily Sesioidea (Lepidoptera, Ditrysia). Smithsonian Contributions to Zoology, 314, 1 - 144. http: // dx. doi. org / 10.5479 / si. 00810282.314" type="journal article" year="1981">Heppner &amp; Duckworth 1981: 42</bibRefCitation>
; Fletcher &amp; Nye 1982: 38;
<bibRefCitation id="EFDD4B00FFCB440EFC96F859FBD9D893" author="Puhringer &amp; Kallies" box="[846,1148,2010,2032]" firstAuthor="Puhringer" page="43" pageId="4" pageNumber="5" pagination="185" refId="ref29350" refString="Puhringer, F. &amp; Kallies, A. (2004) Provisional checklist of the Sesiidae of the world (Lepidoptera: Ditrysia). Mitteilungen der Entomologischen Arbeitsgemeinschaft Salzkammergut, 4, 185. Available from: http: // www. sesiidae. net / Checklst. htm (accessed 30 October 2013)" type="journal article" year="2004">Pühringer &amp; Kallies 2004: 43</bibRefCitation>
.
</paragraph>
<paragraph id="8BF336F1FFCA440FFF4FFF14FB1FDD74" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
The poorly known genus
<taxonomicName id="4C4C4D72FFCA440FFE6DFF14FDE9DFD3" authorityName="Holland" authorityYear="1893" box="[437,588,151,176]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFE6DFF14FDE9DFD3" box="[437,588,151,176]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
was considered synonymous with
<taxonomicName id="4C4C4D72FFCA440FFC0BFF14FB8EDFD3" authorityName="Walker" authorityYear="1856" box="[979,1067,151,176]" class="Insecta" family="Sesiidae" genus="Alonina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFC0BFF14FB8EDFD3" box="[979,1067,151,176]" italics="true" pageId="5" pageNumber="6">Alonina</emphasis>
</taxonomicName>
for many years (sensu
<bibRefCitation id="EFDD4B00FFCA440FFAE9FF1BFF76DFB6" author="Hampson" firstAuthor="Hampson" pageId="5" pageNumber="6" pagination="46 - 119" refId="ref28526" refString="Hampson, G. F. (1919) A classification of the Aegeriadae of the Oriental and Ethiopian Regions. Novitates Zoologicae, 26 (1), 46 - 119." type="journal article" year="1919">Hampson 1919</bibRefCitation>
). However, significant differences of the antenna, the wing venation and the structure of the genitalia do not support a close relationship to
<taxonomicName id="4C4C4D72FFCA440FFE21FF5CFDF7DF9B" authorityName="Walker" authorityYear="1856" box="[505,594,223,248]" class="Insecta" family="Sesiidae" genus="Alonina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFE21FF5CFDF7DF9B" box="[505,594,223,248]" italics="true" pageId="5" pageNumber="6">Alonina</emphasis>
</taxonomicName>
. The
<typeStatus id="54F78853FFCA440FFD4FFF63FD62DF9B" box="[663,711,224,248]" pageId="5" pageNumber="6">type</typeStatus>
species
<taxonomicName id="4C4C4D72FFCA440FFCF3FF5CFB70DF9B" authority="Holland, 1893" authorityName="Holland" authorityYear="1893" box="[811,1237,223,248]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B938EAE3FFCA440FFCF3FF5CFB8FDF9B" box="[811,1066,223,248]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis longipes</emphasis>
<bibRefCitation id="EFDD4B00FFCA440FFBEBFF5CFB70DF9B" author="Holland" box="[1075,1237,223,248]" firstAuthor="Holland" pageId="5" pageNumber="6" pagination="181 - 184" refId="ref28768" refString="Holland, W. J. (1893) Four new genera and species of West African Sesiidae. Journal of the New York Entomological Society, 1, 181 - 184. http: // dx. doi. org / 10.4039 / ent 251 - 1" type="journal article" year="1893">Holland, 1893</bibRefCitation>
</taxonomicName>
<emphasis id="B938EAE3FFCA440FFB07FF5CFAF9DF9B" bold="true" box="[1247,1372,223,248]" pageId="5" pageNumber="6">rev. comb.</emphasis>
from
<collectingCountry id="F35B7661FFCA440FFF4FFE87FF47DE7E" box="[151,226,260,285]" name="Gabon" pageId="5" pageNumber="6">Gabon</collectingCountry>
is only known in the male sex. It seems closely related to and may well be congeneric or even conspecific with
<taxonomicName id="4C4C4D72FFCA440FFF09FEA4FD3ADE23" authority="Le Cerf, 1916" authorityName="Le Cerf" authorityYear="1916" box="[209,671,295,320]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gigantipes">
<emphasis id="B938EAE3FFCA440FFF09FEA4FE5CDE5C" box="[209,505,295,320]" italics="true" pageId="5" pageNumber="6">Megalosphecia gigantipes</emphasis>
<bibRefCitation id="EFDD4B00FFCA440FFDD8FEABFD3ADE23" author="Le Cerf" box="[512,671,295,320]" firstAuthor="Le Cerf" pageId="5" pageNumber="6" pagination="7 - 14" refId="ref28989" refString="Le Cerf, F. (1916) Explication des planches. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 12 (1), 7 - 14, pls. 373 - 381." type="journal article" year="1916">Le Cerf, 1916</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F35B7661FFCA440FFD3DFEA4FCF9DE23" box="[741,860,295,320]" name="Cameroon" pageId="5" pageNumber="6">Cameroon</collectingCountry>
, the
<typeStatus id="54F78853FFCA440FFC4BFEABFC66DE23" box="[915,963,296,320]" pageId="5" pageNumber="6">type</typeStatus>
species of
<taxonomicName id="4C4C4D72FFCA440FFB9CFEA4FA3DDE23" authority="Le Cerf, 1916" authorityName="Le Cerf" authorityYear="1916" box="[1092,1432,295,320]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFB9CFEA4FB4ADE23" box="[1092,1263,295,320]" italics="true" pageId="5" pageNumber="6">Megalosphecia</emphasis>
<bibRefCitation id="EFDD4B00FFCA440FFB2EFEABFA3DDE23" author="Le Cerf" box="[1270,1432,295,320]" firstAuthor="Le Cerf" pageId="5" pageNumber="6" pagination="7 - 14" refId="ref28989" refString="Le Cerf, F. (1916) Explication des planches. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 12 (1), 7 - 14, pls. 373 - 381." type="journal article" year="1916">Le Cerf, 1916</bibRefCitation>
</taxonomicName>
, which is solely represented by females. Both species share important structural characters, namely the large size, the long, clavate antennae, the very long and tufted hind legs, the opaque, blackish wings and the rather similar wing venation. The exact relationship of
<taxonomicName id="4C4C4D72FFCA440FFDBAFE17FD5CDECE" authorityName="Holland" authorityYear="1893" box="[610,761,404,429]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFDBAFE17FD5CDECE" box="[610,761,404,429]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D72FFCA440FFCF7FE17FC7FDECE" authorityName="Le Cerf" authorityYear="1916" box="[815,986,404,429]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFCF7FE17FC7FDECE" box="[815,986,404,429]" italics="true" pageId="5" pageNumber="6">Megalosphecia</emphasis>
</taxonomicName>
could not be definitively clarified. Both genera are here placed in
<taxonomicName id="4C4C4D72FFCA440FFE6BFE34FE5ADEB3" authorityName="Boisduval" authorityYear="1828" box="[435,511,439,464]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
for the first time. The following redescription of
<taxonomicName id="4C4C4D72FFCA440FFBFCFE34FB1EDEB3" authorityName="Holland" authorityYear="1893" box="[1060,1211,439,464]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFBFCFE34FB1EDEB3" box="[1060,1211,439,464]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
based on
<specimenCount id="9D4AFD78FFCA440FFAF2FE3BFA39DEB3" box="[1322,1436,439,464]" pageId="5" pageNumber="6" type="male">two males</specimenCount>
of
<taxonomicName id="4C4C4D72FFCA440FFF6CFE5FFE14DE96" authorityName="Holland" authorityYear="1893" box="[180,433,476,501]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B938EAE3FFCA440FFF6CFE5FFE14DE96" box="[180,433,476,501]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis longipes</emphasis>
</taxonomicName>
from Eala (
<figureCitation id="13772A74FFCA440FFDECFE5FFDD1DE96" box="[564,628,476,501]" captionStart="FIGURES 112" captionStartId="30.[151,264,1789,1812]" captionTargetBox="[153,1436,201,1762]" captionTargetId="figure-114@30.[151,1436,195,1765]" captionTargetPageId="30" captionText="FIGURES 112. Specimens of Cicinnoscelis Holland, 1893; Megalosphecia Le Cerf, 1916 and Alonina Walker, 1856: (1) Cicinnoscelis longipes Holland, 1893, ♂, Congo, Eala. (23) C. flavipes new sp., ♀, holotype. (45) C. krooni new sp., ♀, holotype. (6) Megalosphecia callosoma Hampson, 1919 ♀, holotype. 712. Alonina rygchiiformis Walker, 1856, (7) ♂ holotype; (8) ♂, syntype of A. difformis Hampson, 1919 new syn.; (9) ♀, syntype of A. difformis Hampson, 1919 new syn.; (1011) ♂, Kenya, Eastern Province, road Meru to Nyambeni Hills, 10 km N/E Meru; (12) ♀ South Africa [unknown locality]. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10099868" httpUri="https://zenodo.org/record/10099868/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
) and Bamanya,
<collectingCountry id="F35B7661FFCA440FFCF4FE5FFCDCDE96" box="[812,889,476,501]" name="Democratic Republic of the Congo" pageId="5" pageNumber="6">Congo</collectingCountry>
, which are deposited in MRAC, and
<specimenCount id="9D4AFD78FFCA440FFACCFE5FFA39DE96" box="[1300,1436,476,501]" pageId="5" pageNumber="6" type="female">two females</specimenCount>
from southern Africa, below described as
<taxonomicName id="4C4C4D72FFCA440FFDB5FE7CFCC7DD7B" authorityName="Bartsch" authorityYear="2013" box="[621,866,511,536]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="flavipes" status="sp. nov.">
<emphasis id="B938EAE3FFCA440FFDB5FE7CFCC7DD7B" box="[621,866,511,536]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis flavipes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5798FFCA440FFCB1FD83FC1ADD74" box="[873,959,512,535]" pageId="5" pageNumber="6" rank="species">new sp.</taxonomicNameLabel>
and
<taxonomicName id="4C4C4D72FFCA440FFC2DFD83FBFBDD7B" authorityName="Bartsch" authorityYear="2013" box="[1013,1118,511,536]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="krooni" status="sp. nov.">
<emphasis id="B938EAE3FFCA440FFC2DFD83FBFBDD7B" box="[1013,1118,511,536]" italics="true" pageId="5" pageNumber="6">C. krooni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B5798FFCA440FFBBDFD83FB1FDD74" box="[1125,1210,512,535]" pageId="5" pageNumber="6" rank="species">new sp.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFCA440FFF1FFDA7FCEFDA36" pageId="5" pageNumber="6" type="description">
<paragraph id="8BF336F1FFCA440FFF1FFDA7FBA5DCC6" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FFDA7FED1DD5E" bold="true" box="[199,372,548,573]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">Redescription.</emphasis>
Very large clearwing moths with long, slender, tufted hind legs, wingspan varies from
<quantity id="4CB49B14FFCA440FFA98FDA7FF63DD3C" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="6.4" metricValueMin="4.5" pageId="5" pageNumber="6" unit="mm" value="54.5" valueMax="64.0" valueMin="45.0">45 to 64 mm</quantity>
. Head with labial palpus moderately upturned, covered with short and ventrally rough scales, third palpomere smooth, second palpomere about three times the length of other palpomeres; proboscis reduced; eyes very large, width of frons about equal the diameter of the eye; antenna spindle-shaped, in male bipectinate, rami covered with minute ciliae, in female simple or bipectinate, without ciliae. Thorax and abdomen smooth, in males particularly dorsally sparsely mottled with hair-like scales; abdomen long, rather slender, cylindrical. Fore- and mid leg strong with long and slender fore coxa; hind leg extremely long; tibia and first tarsomere of all legs, most conspicuous in hind tibia, dorsally and apically with tufts of rough, hair-like scales; spurs of mid leg rather short, of hind leg long and strong, lateral spur half as long as mesal one on all spur pairs. Wings rather narrow, almost opaque; hindwing at dorsum partially hyaline or semihyaline. Wing venation (
<figureCitation id="13772A74FFCA440FFCC4FCE4FCCBDCE3" box="[796,878,871,896]" captionStart="FIGURES 8491" captionStartId="38.[151,264,1763,1786]" captionTargetBox="[160,1428,202,1727]" captionTargetId="figure-70@38.[151,1436,193,1742]" captionTargetPageId="38" captionText="FIGURES 8491. Wing venations: (84) Cicinnoscelis longipes Holland, 1893; (85) Alonina rygchiiformis Walker, 1856; (86) Anaudia felderi Wallengren, 1863, syntype; (87) Austrosetia semirufa Felder &amp; Felder, 1874; (88) Felderiola candescens (Felder &amp; Felder, 1874); (89) Monopetalotaxis doleriformis (Walker, 1856); (90) Hagnogyna sanguicosta (Hampson, 1919); (91) Vespanthedon cerceris Le Cerf, 1917, holotype." figureDoi="http://doi.org/10.5281/zenodo.10099887" httpUri="https://zenodo.org/record/10099887/files/figure.png" pageId="5" pageNumber="6">Fig. 84</figureCitation>
) of forewing with common stalk of veins R3 and R5, and common origin of M2 and M3; hindwing with stalked M3 and CuA1.
</paragraph>
<paragraph id="8BF336F1FFCA440FFF1FFC2CFF70DBC3" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FFC2CFE59DCAB" box="[199,508,943,968]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">
<emphasis id="B938EAE3FFCA440FFF1FFC2CFED3DCAB" bold="true" box="[199,374,943,968]" pageId="5" pageNumber="6">Male genitalia</emphasis>
(
<figureCitation id="13772A74FFCA440FFE51FC2CFE55DCAB" box="[393,496,943,968]" captionStart="FIGURES 107108" captionStartId="41.[151,264,1784,1807]" captionTargetBox="[163,1426,210,1763]" captionTargetId="figure-33@41.[151,1436,193,1763]" captionTargetPageId="41" captionText="FIGURES 107108. Genitalia of Cicinnoscelis Holland, 1893: (107) Cicinnoscelis longipes Holland, 1893; (108) C. flavipes new sp, holotype." figureDoi="http://doi.org/10.5281/zenodo.10099893" httpUri="https://zenodo.org/record/10099893/files/figure.png" pageId="5" pageNumber="6">Fig. 107</figureCitation>
).
</emphasis>
Strongly asymmetric; tegumen very broad, gnathos medially located, long and narrow, bifurcate; uncus extremely broad and flat, caudally bilobed, lobes extraordinaryly wide apart, each caudally with strong thorn-like, ventrad pointing sparse setae; valva short and broad, somewhat quadratic, distal margin with irregular protrusions, dorsally pointed and covered with rows of strong thorn-like setae, ventrally rounded; vinculum small, forming a long and narrow saccus; phallus long and slender, slightly s-curved; coecum penis short and round; vesica with enlarged basal part ovoid, dorso-distally covered with numerous sclerotized teeth.
</paragraph>
<paragraph id="8BF336F1FFCA440FFF1FFB2FFCEFDA36" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FFB2FFDA8DBA6" box="[199,525,1196,1221]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">
<emphasis id="B938EAE3FFCA440FFF1FFB2FFE28DBA6" bold="true" box="[199,397,1196,1221]" pageId="5" pageNumber="6">Female genitalia</emphasis>
(
<figureCitation id="13772A74FFCA440FFE45FB2FFDA4DBA6" box="[413,513,1196,1221]" captionStart="FIGURES 107108" captionStartId="41.[151,264,1784,1807]" captionTargetBox="[163,1426,210,1763]" captionTargetId="figure-33@41.[151,1436,193,1763]" captionTargetPageId="41" captionText="FIGURES 107108. Genitalia of Cicinnoscelis Holland, 1893: (107) Cicinnoscelis longipes Holland, 1893; (108) C. flavipes new sp, holotype." figureDoi="http://doi.org/10.5281/zenodo.10099893" httpUri="https://zenodo.org/record/10099893/files/figure.png" pageId="5" pageNumber="6">Fig. 108</figureCitation>
).
</emphasis>
Papillae anales very narrow; segment eight and lamella postvaginalis fused to a well sclerotized, compact ring, whose diameter is distinctly larger than width of papillae anales; posterior apophyses about twice the length of anterior pair; ostium wide, round, located basally of segment eight; antrum strong sclerotized, smooth, cylindrical, cranially narrowed; ductus bursae short and broad, gradually enlarged to form a ovoid bursa copulatrix with large longitudinal signum.
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFCA440FFF1FFADCFEA3D9BC" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BF336F1FFCA440FFF1FFADCFEF0DA86" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FFADCFEE4DA1B" bold="true" box="[199,321,1375,1400]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">Diagnosis.</emphasis>
The male genitalia are characterized as follows: (1) tegumen including gnathos, uncus and valva strongly asymmetric; (2) uncus large, extremely broad and flat, of nearly the same dimensions as valva; (3) valva almost quadratic with irregular shape; (4) vesica with proximal part bulbous, dorso-distally covered with numerous sclerotized teeth.
</paragraph>
<paragraph id="8BF336F1FFCA440FFF1FFA6CFB50D9FB" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<taxonomicName id="4C4C4D72FFCA440FFF1FFA6CFEFBD96B" authorityName="Holland" authorityYear="1893" box="[199,350,1519,1544]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFF1FFA6CFEFBD96B" box="[199,350,1519,1544]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
was previously considered synonymous with
<taxonomicName id="4C4C4D72FFCA440FFC59FA6CFC7FD96B" authorityName="Walker" authorityYear="1856" box="[897,986,1519,1544]" class="Insecta" family="Sesiidae" genus="Alonina" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFC59FA6CFC7FD96B" box="[897,986,1519,1544]" italics="true" pageId="5" pageNumber="6">Alonina</emphasis>
</taxonomicName>
, but species of the latter genus differ clearly by the well-developed haustellum, the much shorter second segment of the labial palpus and the shorter hind legs. Males differ further by the serrate, not bipectinate antenna and the completely different, symmetric structure of the genitalia. The assignment of the following two species in
<taxonomicName id="4C4C4D72FFCA440FFC2CF9DFFB2ED916" authorityName="Holland" authorityYear="1893" box="[1012,1163,1628,1653]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFC2CF9DFFB2ED916" box="[1012,1163,1628,1653]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
, each described from a single female from Southern Africa, should be considered provisionally until males become known.
</paragraph>
<paragraph id="8BF336F1FFCA440FFF1FF927FEA3D9BC" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FF927FE16D9DE" bold="true" box="[199,435,1700,1725]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">Genus composition.</emphasis>
Currently
<taxonomicName id="4C4C4D72FFCA440FFDF4F927FD66D9DE" authorityName="Holland" authorityYear="1893" box="[556,707,1700,1725]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFCA440FFDF4F927FD66D9DE" box="[556,707,1700,1725]" italics="true" pageId="5" pageNumber="6">Cicinnoscelis</emphasis>
</taxonomicName>
includes the following three species:
<emphasis id="B938EAE3FFCA440FFBB4F927FAC9D9DE" box="[1132,1388,1700,1725]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C4C4D72FFCA440FFBB4F927FB49D9DE" authorityName="Holland" authorityYear="1893" box="[1132,1260,1700,1725]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longipes">C. longipes</taxonomicName>
<taxonomicName id="4C4C4D72FFCA440FFB2CF927FAC9D9DE" authorityName="Bartsch" authorityYear="2013" box="[1268,1388,1700,1725]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="flavipes">C. flavipes</taxonomicName>
</emphasis>
and
<taxonomicName id="4C4C4D72FFCA440FFF4FF94BFF5AD983" authorityName="Bartsch" authorityYear="2013" box="[151,255,1735,1760]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="krooni">
<emphasis id="B938EAE3FFCA440FFF4FF94BFF5AD983" box="[151,255,1735,1760]" italics="true" pageId="5" pageNumber="6">C. krooni</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFCA440FFF1FF96FFBABD8BE" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BF336F1FFCA440FFF1FF96FFBABD8BE" blockId="5.[151,1436,151,2013]" pageId="5" pageNumber="6">
<emphasis id="B938EAE3FFCA440FFF1FF96FFE95D866" bold="true" box="[199,304,1772,1797]" inLineHeading="true" pageId="5" pageNumber="6" reason="1">Remark.</emphasis>
One species, the enigmatic
<taxonomicName id="4C4C4D72FFCA440FFDA7F96FFBCFD866" authority="Hampson, 1919" authorityName="Hampson" authorityYear="1919" box="[639,1130,1772,1797]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="callosoma">
<emphasis id="B938EAE3FFCA440FFDA7F96FFC0CD866" box="[639,937,1772,1797]" italics="true" pageId="5" pageNumber="6">Megalosphecia callosoma</emphasis>
<bibRefCitation id="EFDD4B00FFCA440FFC6CF96FFBCFD866" author="Hampson" box="[948,1130,1772,1797]" firstAuthor="Hampson" pageId="5" pageNumber="6" pagination="46 - 119" refId="ref28526" refString="Hampson, G. F. (1919) A classification of the Aegeriadae of the Oriental and Ethiopian Regions. Novitates Zoologicae, 26 (1), 46 - 119." type="journal article" year="1919">Hampson, 1919</bibRefCitation>
</taxonomicName>
(
<figureCitation id="13772A74FFCA440FFBA6F96FFB63D866" box="[1150,1222,1772,1797]" captionStart="FIGURES 112" captionStartId="30.[151,264,1789,1812]" captionTargetBox="[153,1436,201,1762]" captionTargetId="figure-114@30.[151,1436,195,1765]" captionTargetPageId="30" captionText="FIGURES 112. Specimens of Cicinnoscelis Holland, 1893; Megalosphecia Le Cerf, 1916 and Alonina Walker, 1856: (1) Cicinnoscelis longipes Holland, 1893, ♂, Congo, Eala. (23) C. flavipes new sp., ♀, holotype. (45) C. krooni new sp., ♀, holotype. (6) Megalosphecia callosoma Hampson, 1919 ♀, holotype. 712. Alonina rygchiiformis Walker, 1856, (7) ♂ holotype; (8) ♂, syntype of A. difformis Hampson, 1919 new syn.; (9) ♀, syntype of A. difformis Hampson, 1919 new syn.; (1011) ♂, Kenya, Eastern Province, road Meru to Nyambeni Hills, 10 km N/E Meru; (12) ♀ South Africa [unknown locality]. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10099868" httpUri="https://zenodo.org/record/10099868/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
), differs distinctly from both
<taxonomicName id="4C4C4D72FFCA440FFEC8F892FE09D844" authorityName="Le Cerf" authorityYear="1916" box="[272,428,1808,1832]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gigantipes">
<emphasis id="B938EAE3FFCA440FFEC8F892FE09D844" box="[272,428,1808,1832]" italics="true" pageId="5" pageNumber="6">M. gigantipes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D72FFCA440FFE3FF893FDC2D84B" authorityName="Holland" authorityYear="1893" box="[487,615,1807,1832]" class="Insecta" family="Sesiidae" genus="Cicinnoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="B938EAE3FFCA440FFE3FF893FDC2D84B" box="[487,615,1807,1832]" italics="true" pageId="5" pageNumber="6">C. longipes</emphasis>
</taxonomicName>
(
<figureCitation id="13772A74FFCA440FFDA0F88CFD1FD84B" box="[632,698,1807,1832]" captionStart="FIGURES 112" captionStartId="30.[151,264,1789,1812]" captionTargetBox="[153,1436,201,1762]" captionTargetId="figure-114@30.[151,1436,195,1765]" captionTargetPageId="30" captionText="FIGURES 112. Specimens of Cicinnoscelis Holland, 1893; Megalosphecia Le Cerf, 1916 and Alonina Walker, 1856: (1) Cicinnoscelis longipes Holland, 1893, ♂, Congo, Eala. (23) C. flavipes new sp., ♀, holotype. (45) C. krooni new sp., ♀, holotype. (6) Megalosphecia callosoma Hampson, 1919 ♀, holotype. 712. Alonina rygchiiformis Walker, 1856, (7) ♂ holotype; (8) ♂, syntype of A. difformis Hampson, 1919 new syn.; (9) ♀, syntype of A. difformis Hampson, 1919 new syn.; (1011) ♂, Kenya, Eastern Province, road Meru to Nyambeni Hills, 10 km N/E Meru; (12) ♀ South Africa [unknown locality]. Scale bars 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10099868" httpUri="https://zenodo.org/record/10099868/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
) by its external appearance, particularly by the stalked forewing veins R3 and R4/R5. It is so far only known from
<specimenCount id="9D4AFD78FFCA440FFD1AF8B7FC9AD82E" box="[706,831,1844,1869]" pageId="5" pageNumber="6" type="female">two female</specimenCount>
specimens, the
<typeStatus id="54F78853FFCA440FFC28F8B7FBF7D82E" box="[1008,1106,1844,1869]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
and a
<typeStatus id="54F78853FFCA440FFB42F8B7FB5FD82F" box="[1178,1274,1844,1868]" pageId="5" pageNumber="6" type="paratype">paratype</typeStatus>
from Kashitu,
<collectingCountry id="F35B7661FFCA440FFF4FF8D4FF4AD813" box="[151,239,1879,1904]" name="Zambia" pageId="5" pageNumber="6">Zambia</collectingCountry>
, in BMNH. This species is immediately identified by the beautiful red abdominal segments with broad black and narrow yellow posterior margin, and by the large transparent areas of the wings. According to the wing venation,
<taxonomicName id="4C4C4D72FFCA440FFEDDF822FE38D8DA" authorityName="Hampson" authorityYear="1919" box="[261,413,1952,1977]" class="Insecta" family="Sesiidae" genus="Megalosphecia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="callosoma">
<emphasis id="B938EAE3FFCA440FFEDDF822FE38D8DA" box="[261,413,1952,1977]" italics="true" pageId="5" pageNumber="6">M. callosoma</emphasis>
</taxonomicName>
represents most probably an undescribed genus of the
<taxonomicName id="4C4C4D72FFCA440FFBDCF823FBF4D8DA" authorityName="Boisduval" authorityYear="1828" box="[1028,1105,1952,1977]" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Sesiini">Sesiini</taxonomicName>
or perhaps of the
<taxonomicName id="4C4C4D72FFCA440FFAC3F823FE77D8BE" authority="Duckworth &amp; Eichlin, 1977" authorityName="Duckworth &amp; Eichlin" authorityYear="1977" class="Insecta" family="Sesiidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="tribe" tribe="Cissuvorini">Cissuvorini Duckworth &amp; Eichlin, 1977</taxonomicName>
, a tribe, which is currently not known from Africa.
</paragraph>
</subSubSection>
</treatment>
</document>