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<document id="92E871A2957646241A977307E36084FB" ID-DOI="10.11646/zootaxa.4996.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="5074778" ID-ZooBank="50282C10-075C-4B2C-B61C-3A772D5BC2F2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625557780024" checkinUser="plazi" docAuthor="Hsueh, Pan-Wen" docDate="2021" docId="03E587C6FFA5FF8D85A0FADAFE1BF380" docLanguage="en" docName="zootaxa.4996.3.4.pdf" docOrigin="Zootaxa 4996 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudonereis kihawensis Hsueh 2021, n. sp." docType="treatment" docUuid="A3DBE7D1-9A60-4949-8174-BA12A1BC6174" docUuidSource="ZooBank" docVersion="3" lastPageNumber="502" masterDocId="FFDCFFBEFFA3FF878537FFA5FFB0F06D" masterDocTitle="New species and record of Pseudonereis (Annelida, Polychaeta, Nereididae) from Taiwan" masterLastPageNumber="512" masterPageNumber="492" pageNumber="498" updateTime="1698992174721" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="0EE0830F5B64B0B85F19680E8F837B3F">New species and record of Pseudonereis (Annelida, Polychaeta, Nereididae) from Taiwan</mods:title>
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<mods:namePart id="7015FB03D29D96E80E9E4ECBD28A7991">Hsueh, Pan-Wen</mods:namePart>
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<mods:title id="030740B8B8F2BBBA0A471BCC9E9A3F93">Zootaxa</mods:title>
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<mods:date id="CFF3236292CF783F1A5291D4CE5B7ECD">2021</mods:date>
<mods:detail id="BD3CEC22FCB11B62001D5DBE48C50732" type="pubDate">
<mods:number id="B38577FE422474B11F552973F79B4E04">2021-07-06</mods:number>
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<mods:number id="64FD98F8F29A498E94EC9A1D0ECDF4B3">4996</mods:number>
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<treatment id="03E587C6FFA5FF8D85A0FADAFE1BF380" LSID="urn:lsid:zoobank.org:act:A3DBE7D1-9A60-4949-8174-BA12A1BC6174" httpUri="http://treatment.plazi.org/id/03E587C6FFA5FF8D85A0FADAFE1BF380" lastPageId="10" lastPageNumber="502" pageId="6" pageNumber="498">
<subSubSection id="C356655BFFA5FF8185A0FADAFDBFF5F4" box="[151,527,1407,1434]" pageId="6" pageNumber="498" type="nomenclature">
<paragraph id="8BF336D0FFA5FF8185A0FADAFDBFF5F4" blockId="6.[151,527,1407,1470]" box="[151,527,1407,1434]" pageId="6" pageNumber="498">
<heading id="D0BB81BCFFA5FF8185A0FADAFDBFF5F4" bold="true" box="[151,527,1407,1434]" fontSize="11" level="1" pageId="6" pageNumber="498" reason="1">
<emphasis id="B938EAC2FFA5FF8185A0FADAFDBFF5F4" bold="true" box="[151,527,1407,1434]" pageId="6" pageNumber="498">
<taxonomicName id="4C4C4D53FFA5FF8185A0FADAFE76F5F7" ID-CoL="8M35G" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[151,454,1407,1434]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="6" pageNumber="498" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFA5FF8185A0FADAFE76F5F7" bold="true" box="[151,454,1407,1434]" italics="true" pageId="6" pageNumber="498">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57B9FFA5FF8184FAFADAFDBFF5F4" box="[461,527,1407,1433]" pageId="6" pageNumber="498" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFA5FF8185A0FA06FE1BF5D0" box="[151,427,1443,1470]" pageId="6" pageNumber="498" type="description">
<paragraph id="8BF336D0FFA5FF8185A0FA06FE1BF5D0" blockId="6.[151,527,1407,1470]" box="[151,427,1443,1470]" pageId="6" pageNumber="498">
<figureCitation id="13772A55FFA5FF8185A0FA06FEA4F5D3" box="[151,276,1443,1470]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="6" pageNumber="498">Figs 5A–C</figureCitation>
,
<figureCitation id="13772A55FFA5FF818428FA06FED0F5D0" box="[287,352,1443,1469]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="6" pageNumber="498">6A–F</figureCitation>
,
<figureCitation id="13772A55FFA5FF81845CFA06FE1BF5D0" box="[363,427,1443,1469]" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="6" pageNumber="498">7A–F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFA5FF8185A0FA43FDD2F649" pageId="6" pageNumber="498" type="materials_examined">
<paragraph id="8BF336D0FFA5FF8185A0FA43FDD2F649" blockId="6.[151,1437,1510,2041]" pageId="6" pageNumber="498">
<emphasis id="B938EAC2FFA5FF8185A0FA43FE31F66D" bold="true" box="[151,385,1510,1536]" pageId="6" pageNumber="498">Material examined.</emphasis>
<materialsCitation id="3B243C8DFFA5FF8184BFFA43FDD2F649" collectingDate="2014-03-27" collectionCode="NMNS" country="Taiwan" latitude="23.116167" location="Jihuei" longLatPrecision="12" longitude="121.40517" pageId="6" pageNumber="498" specimenCode="NMNS 8383-25" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
<typeStatus id="54F78872FFA5FF8184BFFA43FE40F66D" box="[392,496,1510,1536]" pageId="6" pageNumber="498" type="holotype">Holotype</typeStatus>
(
<specimenCode id="DBEA9EABFFA5FF818737FA43FD0BF66C" box="[512,699,1510,1537]" collectionCode="NMNS" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="6" pageNumber="498">NMNS 8383-25</specimenCode>
), atoke,
<location id="8E93600BFFA5FF81862DFA43FCEDF66D" LSID="urn:lsid:plazi:treatment:03E587C6FFA5FF8D85A0FADAFE1BF380:8E93600BFFA5FF81862DFA43FCEDF66D" box="[794,861,1510,1536]" country="Taiwan" latitude="23.116167" longLatPrecision="12" longitude="121.40517" name="Jihuei" pageId="6" pageNumber="498" stateProvince="Taitung">Jihuei</location>
(
<geoCoordinate id="EE785017FFA5FF81865BFA43FC6FF66D" box="[876,991,1510,1536]" degrees="23" direction="north" minutes="6.97" orientation="latitude" pageId="6" pageNumber="498" precision="9" value="23.116167">23°6.97'N</geoCoordinate>
<geoCoordinate id="EE785017FFA5FF8186D1FA43FBC4F66D" box="[998,1140,1510,1536]" degrees="121" direction="east" minutes="24.31" orientation="longitude" pageId="6" pageNumber="498" precision="9" value="121.40517">121°24.31'E</geoCoordinate>
),
<collectingRegion id="4988F832FFA5FF8181BFFA43FA89F66D" box="[1160,1337,1510,1536]" country="Taiwan" name="Taitung" pageId="6" pageNumber="498">Taitung County</collectingRegion>
,
<collectingCountry id="F35B7640FFA5FF818074FA43FA28F66D" box="[1347,1432,1510,1536]" name="Taiwan" pageId="6" pageNumber="498">Taiwan</collectingCountry>
, rocky intertidal habitats,
<date id="FFF21010FFA5FF818498F9AFFDEEF649" box="[431,606,1546,1572]" pageId="6" pageNumber="498" value="2014-03-27">
<collectingDate id="EFB6E9F8FFA5FF818498F9AFFDEEF649" box="[431,606,1546,1572]" pageId="6" pageNumber="498" value="2014-03-27">27 March 2014</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFA5FF8F85F0F98BFC7EF75F" lastPageId="8" lastPageNumber="500" pageId="6" pageNumber="498" type="description">
<paragraph id="8BF336D0FFA5FF8185F0F98BFCCCF704" blockId="6.[151,1437,1510,2041]" pageId="6" pageNumber="498">
<emphasis id="B938EAC2FFA5FF8185F0F98BFEE9F625" bold="true" box="[199,345,1582,1608]" pageId="6" pageNumber="498">Description.</emphasis>
<typeStatus id="54F78872FFA5FF818468F98BFE7AF625" box="[351,458,1582,1608]" pageId="6" pageNumber="498" type="holotype">Holotype</typeStatus>
, complete, body length
<quantity id="4CB49B35FFA5FF8187E6F98BFC84F625" box="[721,820,1582,1609]" metricMagnitude="-2" metricUnit="m" metricValue="4.95" pageId="6" pageNumber="498" unit="mm" value="49.5">49.5 mm</quantity>
with 95 chaetigers, chaetiger 10 width
<quantity id="4CB49B35FFA5FF8181D0F98BFA8EF625" box="[1255,1342,1582,1609]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="498" unit="mm" value="1.5">1.5 mm</quantity>
, excluding parapodia; beige in alcohol (
<figureCitation id="13772A55FFA5FF818734F9F7FDEEF600" box="[515,606,1618,1645]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="6" pageNumber="498">Fig. 5A</figureCitation>
). Prostomium wider than long, lateral antennae antero-lateral, longer than palps, palpophores globose, palpostyles conical. Four pairs of tentacular cirri, longest one reaching chaetiger 3. Two pairs of eyes, in trapezoidal arrangement, subequal in size. Apodous anterior segment about 1.3 times longer than chaetiger 1. Pharynx with dark brown jaws, each with 3 lateral teeth; paragnath pattern: I=1; II=35 (left), 33 (right), in 5 comb-like rows; III=58, in 4 transverse comb-like rows; IV=71 (left), 67 (right), in 5 comb-like rows with addi- tional cones and p-bars in sigmoid towards the jaws; V=1; VI=1 (left), 1 (right), crescent-shaped bars; VII–VIII=19, p-bars and cones interspaced, p-bar row slightly behind cone row, one paragnath on each furrow and ridge regions (
<figureCitation id="13772A55FFA5FF818597F8EBFEA9F704" box="[160,281,1870,1897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="6" pageNumber="498">Fig. 5B–C</figureCitation>
). Ridge pattern of areas VI–V–VI, λ-shaped (
<figureCitation id="13772A55FFA5FF818620F8EBFCDFF704" box="[791,879,1870,1897]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="6" pageNumber="498">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA5FF8185F0F8D7FA2CF794" blockId="6.[151,1437,1510,2041]" pageId="6" pageNumber="498">
Dorsal cirri digitiform, basally attached to dorsal ligule on anterior chaetigers, about as long as dorsal ligule (
<figureCitation id="13772A55FFA5FF818597F833FEA9F7DC" box="[160,281,1942,1969]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="6" pageNumber="498">Fig. 6A–B</figureCitation>
), subdistally attached to dorsal ligule on mid-body to posterior chaetigers, about 0.8 times as long as dorsal ligule on mid-body chaetigers (
<figureCitation id="13772A55FFA5FF818729F81FFD29F7B8" box="[542,665,1978,2005]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="6" pageNumber="498">Fig. 6C–D</figureCitation>
), about 0.4–0.5 times as long as dorsal ligule on posterior chaetigers (
<figureCitation id="13772A55FFA5FF818597F87BFF47F794" box="[160,247,2014,2041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="6" pageNumber="498">Fig. 6E</figureCitation>
), distally attached to dorsal ligule on last 13 chaetigers, about 0.5 times as long as dorsal ligule (
<figureCitation id="13772A55FFA5FF81800EF87BFA3FF794" box="[1337,1423,2014,2041]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="6" pageNumber="498">Fig. 6F</figureCitation>
).
</paragraph>
<caption id="DF336658FFA4FF8085A0F831FBE7F7A2" ID-DOI="http://doi.org/10.5281/zenodo.5074788" ID-Zenodo-Dep="5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="7" pageNumber="499" startId="7.[151,250,1940,1964]" targetBox="[185,1404,199,1916]" targetPageId="7">
<paragraph id="8BF336D0FFA4FF8085A0F831FBE7F7A2" blockId="7.[151,1436,1940,2000]" pageId="7" pageNumber="499">
<emphasis id="B938EAC2FFA4FF8085A0F831FEA4F7C1" bold="true" box="[151,276,1940,1964]" pageId="7" pageNumber="499">FIGURE 5.</emphasis>
<taxonomicName id="4C4C4D53FFA4FF80842BF831FDABF7C1" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[284,539,1940,1964]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="7" pageNumber="499" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFA4FF80842BF831FDABF7C1" box="[284,539,1940,1964]" italics="true" pageId="7" pageNumber="499">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFA4FF808714F831FDEDF7C1" bold="true" box="[547,605,1940,1964]" pageId="7" pageNumber="499">
<taxonomicNameLabel id="A20B57B9FFA4FF808714F831FDEDF7C1" box="[547,605,1940,1964]" pageId="7" pageNumber="499" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm.
</paragraph>
</caption>
<caption id="DF336658FFABFF8F85A0FBA7FB12F4EB" ID-DOI="http://doi.org/10.5281/zenodo.5074790" ID-Zenodo-Dep="5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500" startId="8.[151,250,1026,1050]" targetBox="[177,1410,181,1001]" targetPageId="8">
<paragraph id="8BF336D0FFABFF8F85A0FBA7FB12F4EB" blockId="8.[151,1437,1026,1158]" pageId="8" pageNumber="500">
<emphasis id="B938EAC2FFABFF8F85A0FBA7FEA4F477" bold="true" box="[151,276,1026,1050]" pageId="8" pageNumber="500">FIGURE 6.</emphasis>
<taxonomicName id="4C4C4D53FFABFF8F842CFBA7FDA8F477" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[283,536,1026,1050]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="8" pageNumber="500" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFABFF8F842CFBA7FDA8F477" box="[283,536,1026,1050]" italics="true" pageId="8" pageNumber="500">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFABFF8F8717FBA7FDEAF477" bold="true" box="[544,602,1026,1050]" pageId="8" pageNumber="500">
<taxonomicNameLabel id="A20B57B9FFABFF8F8717FBA7FDEAF477" box="[544,602,1026,1050]" pageId="8" pageNumber="500" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BF336D0FFABFF8F85F0FB11FE28F557" blockId="8.[151,1437,1204,1987]" pageId="8" pageNumber="500">
Dorsal ligules subconical on anterior chaetigers (
<figureCitation id="13772A55FFABFF8F87C5FB11FCDEF4A2" box="[754,878,1204,1231]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6A–B</figureCitation>
), becoming small pennant-like lobe on middle of mid-body to anterior half of posterior chaetigers (
<figureCitation id="13772A55FFABFF8F87E7FB7DFCFAF49E" box="[720,842,1240,1267]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6C–E</figureCitation>
); base of dorsal ligule slightly elongate and broader on mid-body chaetigers (
<figureCitation id="13772A55FFABFF8F8486FB59FDBBF57A" box="[433,523,1276,1303]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6D</figureCitation>
), markedly elongate and broader on posterior chaetigers (
<figureCitation id="13772A55FFABFF8F81A2FB59FABBF57A" box="[1173,1291,1276,1303]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6E–F</figureCitation>
). Notopodial prechaetal lobe absent.
</paragraph>
<paragraph id="8BF336D0FFABFF8F85F0FAE1FC80F5EE" blockId="8.[151,1437,1204,1987]" pageId="8" pageNumber="500">
Median ligule conical on anterior-most chaetigers, shorter than neuroacicular ligule (
<figureCitation id="13772A55FFABFF8F81A5FAE1FABEF532" box="[1170,1294,1348,1375]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6A–B</figureCitation>
), subconical thereafter and longer than neuroacicular ligule (
<figureCitation id="13772A55FFABFF8F879AFACDFC94F5EE" box="[685,804,1384,1411]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6C–F</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFABFF8F85F0FA29FE94F612" blockId="8.[151,1437,1204,1987]" pageId="8" pageNumber="500">
Neuroacicular ligule with predominant inferior lobe on anterior to anterior half of posterior chaetigers (
<figureCitation id="13772A55FFABFF8F8058FA29FF6CF5A6" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6A–E</figureCitation>
), inferior and superior lobes subequal in length on last 13 chaetigers (
<figureCitation id="13772A55FFABFF8F86CEFA15FBFFF5A6" box="[1017,1103,1456,1483]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6F</figureCitation>
), about 1.3 times longer than ventral ligule on anterior to mid-body chaetigers, about as long as ventral ligule on posterior chaetigers (
<figureCitation id="13772A55FFABFF8F802EFA71FA21F582" box="[1305,1425,1492,1519]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6A–F</figureCitation>
). Neuropodial postchaetal lobe present throughout. Ventral ligule subconical throughout. Ventral cirri mid-ventrally attached to ventral edge of parapodia, about 0.8 times as long as ventral ligule on anterior chaetigers, about 0.7 times as long as ventral ligule on mid-body chaetigers, about 0.5 times as long as ventral ligule on posterior chaetigers (
<figureCitation id="13772A55FFABFF8F8597F9C1FEA8F612" box="[160,280,1636,1663]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="8" pageNumber="500">Fig. 6A–F</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFABFF8F85F0F92DFB67F762" blockId="8.[151,1437,1204,1987]" pageId="8" pageNumber="500">
Notochaetae present from chaetiger 3 to posterior chaetigers, homogomph spinigers present throughout (
<figureCitation id="13772A55FFABFF8F8058F92DFF0DF6AA" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="8" pageNumber="500">Fig. 7A</figureCitation>
). Supra-acicular fascicle of neuropodia: homogomph spinigers and short-bladed heterogomph falcigers with serrations present throughout (
<figureCitation id="13772A55FFABFF8F84DAF975FDD8F687" box="[493,616,1744,1770]" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="8" pageNumber="500">Fig. 7B–C</figureCitation>
). Subacicular fascicle of neuropodia: short-bladed heterogomph falcigers with serrations present throughout (
<figureCitation id="13772A55FFABFF8F8711F951FD30F762" box="[550,640,1780,1807]" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="8" pageNumber="500">Fig. 7D</figureCitation>
), heterogomph spinigers absent throughout (
<figureCitation id="13772A55FFABFF8F8144F951FB7BF762" box="[1139,1227,1780,1807]" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="8" pageNumber="500">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BF336D0FFABFF8F85F0F8BDFC7EF75F" blockId="8.[151,1437,1204,1987]" box="[199,974,1816,1843]" pageId="8" pageNumber="500">Pygidium crenulated; anal cirri cirriform, as long as last 2 chaetigers.</paragraph>
</subSubSection>
<subSubSection id="C356655BFFABFF8F85F0F899FE83F717" pageId="8" pageNumber="500" type="etymology">
<paragraph id="8BF336D0FFABFF8F85F0F899FE83F717" blockId="8.[151,1437,1204,1987]" pageId="8" pageNumber="500">
<emphasis id="B938EAC2FFABFF8F85F0F899FEFEF73B" bold="true" box="[199,334,1852,1878]" pageId="8" pageNumber="500">Etymology.</emphasis>
The name is derived from “Kihaw”, the local Amis aboriginal language for the location of the sampling site.
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFABFF8F85F0F821FD61F7F3" box="[199,721,1924,1950]" pageId="8" pageNumber="500" type="materials_examined">
<paragraph id="8BF336D0FFABFF8F85F0F821FD61F7F3" blockId="8.[151,1437,1204,1987]" box="[199,721,1924,1950]" pageId="8" pageNumber="500">
<materialsCitation id="3B243C8DFFABFF8F85F0F821FD7DF7F3" box="[199,717,1924,1950]" country="Taiwan" location="Jihuei" pageId="8" pageNumber="500" specimenCount="1" stateProvince="Taitung" typeStatus="holotype">
<emphasis id="B938EAC2FFABFF8F85F0F821FED6F7F3" bold="true" box="[199,358,1924,1950]" pageId="8" pageNumber="500">
<typeStatus id="54F78872FFABFF8F85F0F821FEB2F7F3" box="[199,258,1924,1950]" pageId="8" pageNumber="500">Type</typeStatus>
locality.
</emphasis>
<location id="8E93600BFFABFF8F845BF821FE02F7F3" LSID="urn:lsid:plazi:treatment:03E587C6FFA5FF8D85A0FADAFE1BF380:8E93600BFFABFF8F845BF821FE02F7F3" box="[364,434,1924,1950]" country="Taiwan" name="Jihuei" pageId="8" pageNumber="500" stateProvince="Taitung">Jihuei</location>
,
<collectingRegion id="4988F832FFABFF8F848AF821FDDDF7F3" box="[445,621,1924,1950]" country="Taiwan" name="Taitung" pageId="8" pageNumber="500">Taitung County</collectingRegion>
,
<collectingCountry id="F35B7640FFABFF8F8740F821FD7DF7F3" box="[631,717,1924,1950]" name="Taiwan" pageId="8" pageNumber="500">Taiwan</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356655BFFABFF8F85F0F80DFD78F7AF" box="[199,712,1960,1987]" pageId="8" pageNumber="500" type="distribution">
<paragraph id="8BF336D0FFABFF8F85F0F80DFD78F7AF" blockId="8.[151,1437,1204,1987]" box="[199,712,1960,1987]" pageId="8" pageNumber="500">
<emphasis id="B938EAC2FFABFF8F85F0F80DFED1F7AF" bold="true" box="[199,353,1960,1986]" pageId="8" pageNumber="500">Distribution.</emphasis>
Known only from
<typeStatus id="54F78872FFABFF8F8700F80CFDD7F7AE" box="[567,615,1961,1987]" pageId="8" pageNumber="500">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<caption id="DF336658FFAAFF8E85A0FB4EFE1DF5FE" ID-DOI="http://doi.org/10.5281/zenodo.5074794" ID-Zenodo-Dep="5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="9" pageNumber="501" startId="9.[151,250,1259,1283]" targetBox="[177,1410,181,1235]" targetPageId="9">
<paragraph id="8BF336D0FFAAFF8E85A0FB4EFE1DF5FE" blockId="9.[151,1437,1259,1428]" pageId="9" pageNumber="501">
<emphasis id="B938EAC2FFAAFF8E85A0FB4EFEA5F569" bold="true" box="[151,277,1259,1284]" pageId="9" pageNumber="501">FIGURE 7.</emphasis>
<taxonomicName id="4C4C4D53FFAAFF8E842AFB49FDACF569" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[285,540,1260,1284]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E842AFB49FDACF569" box="[285,540,1260,1284]" italics="true" pageId="9" pageNumber="501">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E8713FB49FDEFF569" bold="true" box="[548,607,1260,1284]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E8713FB49FDEFF569" box="[548,607,1260,1284]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm.
</paragraph>
</caption>
<subSubSection id="C356655BFFAAFF8D85F0FA67FE1BF380" lastPageId="10" lastPageNumber="502" pageId="9" pageNumber="501" type="discussion">
<paragraph id="8BF336D0FFAAFF8D85F0FA67FCDAF1E4" blockId="9.[151,1437,1474,2040]" lastBlockId="10.[151,1437,151,1005]" lastPageId="10" lastPageNumber="502" pageId="9" pageNumber="501">
<emphasis id="B938EAC2FFAAFF8E85F0FA67FE8BF5B1" bold="true" box="[199,315,1474,1500]" pageId="9" pageNumber="501">Remarks.</emphasis>
Of all known
<taxonomicName id="4C4C4D53FFAAFF8E84DDFA67FD30F5B1" authorityName="Kinberg" authorityYear="1865" box="[490,640,1474,1500]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="genus">
<emphasis id="B938EAC2FFAAFF8E84DDFA67FD30F5B1" box="[490,640,1474,1500]" italics="true" pageId="9" pageNumber="501">Pseudonereis</emphasis>
</taxonomicName>
, only
<taxonomicName id="4C4C4D53FFAAFF8E87F9FA66FCFCF5B1" authorityName="Kinberg" authorityYear="1865" box="[718,844,1474,1500]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="formosa">
<emphasis id="B938EAC2FFAAFF8E87F9FA66FD6FF5B1" box="[718,735,1475,1500]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E87C7FA67FCFCF5B1" box="[752,844,1474,1500]" italics="true" pageId="9" pageNumber="501">formosa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C4C4D53FFAAFF8E866AFA66FC48F5B1" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[861,1016,1474,1500]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="jihueiensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E866AFA66FCDEF5B1" box="[861,878,1475,1500]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E86B7FA67FC48F5B1" box="[896,1016,1474,1500]" italics="true" pageId="9" pageNumber="501">jihueiensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E8135FA67FBF4F5B1" bold="true" box="[1026,1092,1474,1500]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E8135FA67FBF4F5B1" box="[1026,1092,1474,1500]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C4C4D53FFAAFF8E8161FA66FB43F5B1" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[1110,1267,1474,1500]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E8161FA66FBD7F5B1" box="[1110,1127,1475,1500]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E814FFA67FB43F5B1" box="[1144,1267,1474,1500]" italics="true" pageId="9" pageNumber="501">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E81C9FA67FAF0F5B1" bold="true" box="[1278,1344,1474,1500]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E81C9FA67FAF0F5B1" box="[1278,1344,1474,1500]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C4C4D53FFAAFF8E80B3FA66FEB7F66D" authorityName="" baseAuthorityName="Schmarda" baseAuthorityYear="1861" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="podocirra">
<emphasis id="B938EAC2FFAAFF8E80B3FA66FA25F5B1" box="[1412,1429,1475,1500]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E85A0FA43FEB7F66D" box="[151,263,1510,1536]" italics="true" pageId="9" pageNumber="501">podocirra</emphasis>
</taxonomicName>
are noted without heterogomph spinigers in the subacicular fascicle of the neuropodia (
<figureCitation id="13772A55FFAAFF8E81CEFA43FAE1F66D" box="[1273,1361,1510,1536]" captionStart="FIGURE 7" captionStartId="9.[151,250,1259,1283]" captionTargetBox="[177,1410,181,1235]" captionTargetId="figure-33@9.[177,1410,181,1235]" captionTargetPageId="9" captionText="FIGURE 7. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, notochaeta, homogomph spiniger, chaetiger 72; B, supra-acicular homogomph spiniger, chaetiger 72; C, supra-acicular short-bladed heterogomph falciger, chaetiger 72; D, subacicular heterogomph falciger, chaetiger 72; E, parapodia, posterior view, chaetiger 88. Abbreviation: SubHeF, subacicu- lar heterogomph falciger; SupHeF, supra-acicular heterogomph falciger; SupHoS, supra-acicular homogomph spiniger. Scale: A–D, 0.01 mm; E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074794" httpUri="https://zenodo.org/record/5074794/files/figure.png" pageId="9" pageNumber="501">Fig. 7E</figureCitation>
,
<tableCitation id="C6CE036BFFAAFF8E8069FA43FF14F649" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="9" pageNumber="501">Table 1</tableCitation>
). However,
<taxonomicName id="4C4C4D53FFAAFF8E8406F9AEFE7CF649" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[305,460,1546,1572]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E8406F9AEFEF2F649" box="[305,322,1547,1572]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E8466F9AFFE7CF649" box="[337,460,1546,1572]" italics="true" pageId="9" pageNumber="501">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E84E3F9AFFDA3F649" bold="true" box="[468,531,1546,1572]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E84E3F9AFFDA3F649" box="[468,531,1546,1572]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be readily distinguished from the other three congeners by having crescentshaped bars in Area VI of the pharynx (
<figureCitation id="13772A55FFAAFF8E877EF98BFD2FF624" box="[585,671,1582,1609]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="9" pageNumber="501">Fig. 5B</figureCitation>
,
<tableCitation id="C6CE036BFFAAFF8E879EF98BFD4AF625" box="[681,762,1582,1608]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="9" pageNumber="501">Table 1</tableCitation>
).
<taxonomicName id="4C4C4D53FFAAFF8E8625F98BFB98F625" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[786,1064,1582,1608]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E8625F98BFB98F625" box="[786,1064,1582,1608]" italics="true" pageId="9" pageNumber="501">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E8119F98BFBDCF625" bold="true" box="[1070,1132,1582,1608]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E8119F98BFBDCF625" box="[1070,1132,1582,1608]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
has additional features that are not seen in
<taxonomicName id="4C4C4D53FFAAFF8E8477F9F6FE0CF601" authorityName="Kinberg" authorityYear="1865" box="[320,444,1618,1644]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="formosa">
<emphasis id="B938EAC2FFAAFF8E8477F9F6FEE1F601" box="[320,337,1619,1644]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E8457F9F7FE0CF601" box="[352,444,1618,1644]" italics="true" pageId="9" pageNumber="501">formosa</emphasis>
</taxonomicName>
, which are having: 1) smaller ratio of dorsal cirrus to dorsal ligule on anterior chaetigers (about 1.0 versus 2.0); 2) tip of dorsal ligule becoming small pennant-like lobe on mid-body to the anterior half of posterior chaetigers (versus rounded); 3) greater length to width ratio of dorsal ligule on chaetigers 30, 50, and 70 (2.1, 1.8, 2.3 versus 1.3, 1.4 and 1.4, respectively); and 4) greater number of chaetigers with dorsal cirri distally attached to notopodial dorsal ligules (last 13 chaetigers versus posterior-most chaetigers only) (
<bibRefCitation id="EFDD4B21FFAAFF8E81F2F947FA27F690" author="Bakken, T." box="[1221,1431,1762,1789]" pageId="9" pageNumber="501" pagination="145 - 176" refId="ref11104" refString="Bakken, T. (2007) Revision of Pseudonereis (Polychaeta, Nereididae). Zoological Journal of the Linnean Society, 150 (1), 145 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00289. x" type="journal article" year="2007">Bakken 2007: 158</bibRefCitation>
, fig. 7A–E;
<bibRefCitation id="EFDD4B21FFAAFF8E842EF8A3FDADF74D" author="Conde-Vela, V. M." box="[281,541,1798,1824]" pageId="9" pageNumber="501" pagination="245 - 278" refId="ref11340" refString="Conde-Vela, V. M. (2018) New species of Pseudonereis Kinberg, 1865 (Polychaeta: Nereididae) from the Atlantic Ocean, and a review of paragnath morphology and methodology. Zootaxa, 4471 (2), 245 - 278. https: // doi. org / 10.11646 / zootaxa. 4471.2.2" type="journal article" year="2018">Conde-Vela 2018: 270</bibRefCitation>
;
<figureCitation id="13772A55FFAAFF8E871CF8A3FD12F74D" box="[555,674,1798,1824]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="9" pageNumber="501">Fig. 6A–F</figureCitation>
,
<tableCitation id="C6CE036BFFAAFF8E8798F8A3FCB3F74D" box="[687,771,1798,1824]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="9" pageNumber="501">Table 1</tableCitation>
). Also,
<taxonomicName id="4C4C4D53FFAAFF8E865CF8A2FBB8F74D" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[875,1032,1798,1824]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E865CF8A2FCCCF74D" box="[875,892,1799,1824]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E86BAF8A3FBB8F74D" box="[909,1032,1798,1824]" italics="true" pageId="9" pageNumber="501">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E8126F8A3FBE2F74D" bold="true" box="[1041,1106,1798,1824]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E8126F8A3FBE2F74D" box="[1041,1106,1798,1824]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be further differentiated from
<taxonomicName id="4C4C4D53FFAAFF8E85E1F88EFEDFF729" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[214,367,1834,1860]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="jihueiensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E85E1F88EFF57F729" box="[214,231,1835,1860]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E85C0F88FFEDFF729" box="[247,367,1834,1860]" italics="true" pageId="9" pageNumber="501">jihueiensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E8440F88FFE07F729" bold="true" box="[375,439,1834,1860]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E8440F88FFE07F729" box="[375,439,1834,1860]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by having: 1) three lateral teeth on cutting edge of the jaws (versus none); 2) λ-shaped ridge pattern of areas VI–V–VI (versus χ-shaped); 3) smaller ratio of dorsal cirrus to dorsal ligule on anterior and mid-body chaetigers (1.0 and 0.8 versus 1.3 and 1.0, respectively); 4) greater length to width ratio of dorsal ligule on chaetigers 30, 50, and 70 (2.1, 1.8, 2.3 versus 1.3, 1.4 and 1.4, respectively); and 5) shorter anal cirri (as long as last 2 chaetigers versus as long as 4–7 chaetigers) (
<figureCitation id="13772A55FFAAFF8E87E3F81FFCE0F7B9" box="[724,848,1978,2004]" captionStart="FIGURE 1" captionStartId="2.[151,250,1148,1172]" captionTargetBox="[151,1436,179,1122]" captionTargetId="figure-15@2.[151,1436,179,1122]" captionTargetPageId="2" captionText="FIGURE 1. Pseudonereis jihueiensis n. sp.; A, paratype (NMNS 8383-16); B, holotype (NMNS 8383-1); C, paratype (NMNS 8383-18); D, paratype (NMNS 8383-5); E–F, paratype (NMNS 8383-11): A–C, anterior body, dorsal view; D, jaws; E, parag- naths of the pharynx, dorsal view; F, paragnaths of the pharynx, frontal view. Scale: A–C, 1.0 mm; D–F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074780" httpUri="https://zenodo.org/record/5074780/files/figure.png" pageId="9" pageNumber="501">Figs 1D–E</figureCitation>
,
<figureCitation id="13772A55FFAAFF8E866BF81FFC2DF7B9" box="[860,925,1978,2004]" captionStart="FIGURE 2" captionStartId="3.[151,250,1148,1172]" captionTargetBox="[153,1435,273,1124]" captionTargetId="figure-78@3.[153,1435,273,1124]" captionTargetPageId="3" captionText="FIGURE 2. Pseudonereis jihueiensis n. sp.; holotype (NMNS 8383-1): A, left parapodium, anterior view, chaetiger 10; B, left parapodium, posterior view, chaetiger 10; C, left parapodium, chaetiger 30; D, left parapodium, chaetiger 43; E, left parapo- dium, chaetiger 64; F, left parapodium, chaetiger 74. Abbreviation: DC, dorsal cirrus; NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe; NeVL, neuropodial ventral ligule; NoDL, notopodial dorsal ligule; NoVL, notopodial ventral ligule; VC, ventral cirrus. Scale: A–F, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5074782" httpUri="https://zenodo.org/record/5074782/files/figure.png" pageId="9" pageNumber="501">2A–F</figureCitation>
,
<figureCitation id="13772A55FFAAFF8E869FF81FFC5EF7B8" box="[936,1006,1978,2005]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="9" pageNumber="501">5B–D</figureCitation>
,
<figureCitation id="13772A55FFAAFF8E86CEF81FFB8AF7B9" box="[1017,1082,1978,2004]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="9" pageNumber="501">6A–F</figureCitation>
,
<tableCitation id="C6CE036BFFAAFF8E8172F81FFB27F7B9" box="[1093,1175,1978,2004]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="9" pageNumber="501">Table 1</tableCitation>
). Lastly,
<taxonomicName id="4C4C4D53FFAAFF8E8035F81EFA2CF7B9" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[1282,1436,1978,2004]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFAAFF8E8035F81EFAA3F7B9" box="[1282,1299,1979,2004]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E8016F81FFA2CF7B9" box="[1313,1436,1978,2004]" italics="true" pageId="9" pageNumber="501">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFAAFF8E85A0F87BFF67F795" bold="true" box="[151,215,2014,2040]" pageId="9" pageNumber="501">
<taxonomicNameLabel id="A20B57B9FFAAFF8E85A0F87BFF67F795" box="[151,215,2014,2040]" pageId="9" pageNumber="501" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be further distinguished from
<taxonomicName id="4C4C4D53FFAAFF8E8756F87AFD42F795" authorityName="" baseAuthorityName="Schmarda" baseAuthorityYear="1861" box="[609,754,2014,2040]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="9" pageNumber="501" phylum="Annelida" rank="species" species="podocirra">
<emphasis id="B938EAC2FFAAFF8E8756F87AFDC2F795" box="[609,626,2015,2040]" italics="true" pageId="9" pageNumber="501">P</emphasis>
.
<emphasis id="B938EAC2FFAAFF8E87B5F87BFD42F795" box="[642,754,2014,2040]" italics="true" pageId="9" pageNumber="501">podocirra</emphasis>
</taxonomicName>
by having: 1) fewer number and arrangement of paragnaths in Areas VII–VIII (
<quantity id="4CB49B35FFA9FF8D8459FF32FE17F0DC" box="[366,423,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="4.826" pageId="10" pageNumber="502" unit="in" value="19.0">19 in</quantity>
two rows with one paragnath on each furrow and ridge regions versus
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2–4 rows with 2–3 paragnaths on each furrow and ridge regions in Area VIII); 3) smaller ratio of dorsal cirrus to notopodial dorsal ligule on chaetigers 30, 50 but a larger ratio of dorsal cirrus to notopodial dorsal ligule on chaetiger 70 (1.0, 0.8, 0.4 and 0.5 versus 1.7, 2.5, 0.8 and 0.4, respectively); 4) greater length to width ratio of dorsal ligule on chaetigers 30, 50, and 70 (2.1, 1.8, and 2.3 versus 0.9, 1.3 and 2.2, respectively); and 5) greater number of chaetigers with dorsal cirri distally attached to notopodial dorsal ligules (last 13 chaetigers versus the entire posterior chaetigers) (
<bibRefCitation id="EFDD4B21FFA9FF8D807EFEEEFE33F1E4" author="Kara, J. &amp; Macdonald, A. H. H. &amp; Simon, C. A." pageId="10" pageNumber="502" pagination="1282 - 1297" refId="ref11582" refString="Kara, J., Macdonald, A. H. H. &amp; Simon, C. A. (2018) Integrative taxonomic methods reveal an incorrect synonymisation of the South African Pseudonereis podocirra (Schmarda) as the widespread Pseudonereis variegata (Grube) from Chile. Invertebrate Systematics, 32 (6), 1282 - 1297. https: // doi. org / 10.1071 / IS 18016" type="journal article" year="2018">
Kara
<emphasis id="B938EAC2FFA9FF8D80BFFEEEFF1DF1E4" italics="true" pageId="10" pageNumber="502">et al</emphasis>
. 2018: 1287–1288
</bibRefCitation>
, figs 3C–I, 4A–C;
<figureCitation id="13772A55FFA9FF8D8761FECAFD00F1E4" box="[598,688,366,393]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="10" pageNumber="502">Figs 5B</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D878BFECAFD4DF1E4" box="[700,765,367,393]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="10" pageNumber="502">6A–F</figureCitation>
,
<tableCitation id="C6CE036BFFA9FF8D8630FECAFCE9F1E4" box="[775,857,367,393]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="10" pageNumber="502">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BF336D0FFA9FF8D85F0FE36FE1BF380" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="502">
Of the 20 known species of the genus, only
<taxonomicName id="4C4C4D53FFA9FF8D87FBFE36FC50F1C0" authority="(Hansen, 1882)" baseAuthorityName="Hansen" baseAuthorityYear="1882" box="[716,992,403,429]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="ferox">
<emphasis id="B938EAC2FFA9FF8D87FBFE36FD6DF1C1" box="[716,733,403,428]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D87D9FE36FC97F1C0" box="[750,807,403,429]" italics="true" pageId="10" pageNumber="502">ferox</emphasis>
(Hansen, 1882)
</taxonomicName>
<emphasis id="B938EAC2FFA9FF8D86DDFE36FBA4F1C0" box="[1002,1044,403,429]" italics="true" pageId="10" pageNumber="502">and</emphasis>
<taxonomicName id="4C4C4D53FFA9FF8D812AFE36FB1AF1C1" baseAuthorityName="Grube" baseAuthorityYear="1857" box="[1053,1194,403,429]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="variegata">
<emphasis id="B938EAC2FFA9FF8D812AFE36FB9EF1C1" box="[1053,1070,403,428]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D8108FE36FB1AF1C1" box="[1087,1194,403,428]" italics="true" pageId="10" pageNumber="502">variegata</emphasis>
</taxonomicName>
(Grube &amp; Kröyer in Grube, 1858) have crescent-shaped paragnaths in Area VI of the pharynx as in
<taxonomicName id="4C4C4D53FFA9FF8D8116FE12FB0DF1BC" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[1057,1213,439,465]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFA9FF8D8116FE12FB82F1BD" box="[1057,1074,439,464]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D8175FE12FB0DF1BC" box="[1090,1213,439,465]" italics="true" pageId="10" pageNumber="502">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFA9FF8D81F2FE12FAB6F1BC" bold="true" box="[1221,1286,439,465]" pageId="10" pageNumber="502">
<taxonomicNameLabel id="A20B57B9FFA9FF8D81F2FE12FAB6F1BC" box="[1221,1286,439,465]" pageId="10" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(
<bibRefCitation id="EFDD4B21FFA9FF8D8021FE12FEF4F198" author="Conde-Vela, V. M." pageId="10" pageNumber="502" pagination="245 - 278" refId="ref11340" refString="Conde-Vela, V. M. (2018) New species of Pseudonereis Kinberg, 1865 (Polychaeta: Nereididae) from the Atlantic Ocean, and a review of paragnath morphology and methodology. Zootaxa, 4471 (2), 245 - 278. https: // doi. org / 10.11646 / zootaxa. 4471.2.2" type="journal article" year="2018">Conde-Vela 2018: 268–269</bibRefCitation>
, fig. 10D; Fig, 5B,
<tableCitation id="C6CE036BFFA9FF8D8714FE7EFDC6F198" box="[547,630,475,501]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="10" pageNumber="502">Table 1</tableCitation>
). However,
<taxonomicName id="4C4C4D53FFA9FF8D8632FE7EFC10F198" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[773,928,475,501]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFA9FF8D8632FE7EFCA6F199" box="[773,790,475,500]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D8612FE7EFC10F198" box="[805,928,475,501]" italics="true" pageId="10" pageNumber="502">kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFA9FF8D869FFE7EFC58F198" bold="true" box="[936,1000,475,501]" pageId="10" pageNumber="502">
<taxonomicNameLabel id="A20B57B9FFA9FF8D869FFE7EFC58F198" box="[936,1000,475,501]" pageId="10" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="4C4C4D53FFA9FF8D8028FE7EFAC8F198" baseAuthorityName="Hansen" baseAuthorityYear="1882" box="[1311,1400,475,501]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="ferox">
<emphasis id="B938EAC2FFA9FF8D8028FE7EFA80F199" box="[1311,1328,475,500]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D8008FE7EFAC8F198" box="[1343,1400,475,501]" italics="true" pageId="10" pageNumber="502">ferox</emphasis>
</taxonomicName>
by having: 1) more rows of comb-like paragnaths in Areas II &amp; IV (5 and 5 versus 3 and 4, respectively); 2) λ-shaped ridge pattern of areas VI–V–VI (versus ɔc-shaped); 3) dorsal ligule longer than median ligule on anterior chaetigers (versus subequal); 4) dorsal ligule becoming small pennant-like lobe on mid-body to the anterior half of posterior chaetigers (versus rounded); 5) dorsal ligule broaden basally on posterior-most chaetigers (versus broaden in the middle); 6) greater number of dorsal cirrus distally attached to dorsal ligule (last 13 chaetigers versus posterior-most chaetigers); and 7) no heterogomph spinigers in the subacicular fascicle of the neuropodia (versus present from about chaetiger 40) (
<bibRefCitation id="EFDD4B21FFA9FF8D84B3FD72FD20F29C" author="Bakken, T." box="[388,656,727,753]" pageId="10" pageNumber="502" pagination="145 - 176" refId="ref11104" refString="Bakken, T. (2007) Revision of Pseudonereis (Polychaeta, Nereididae). Zoological Journal of the Linnean Society, 150 (1), 145 - 176. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00289. x" type="journal article" year="2007">Bakken 2007: 116–170</bibRefCitation>
, figs 14A, 15A–F;
<bibRefCitation id="EFDD4B21FFA9FF8D8647FD72FBC2F29C" author="Conde-Vela, V. M." box="[880,1138,727,753]" pageId="10" pageNumber="502" pagination="245 - 278" refId="ref11340" refString="Conde-Vela, V. M. (2018) New species of Pseudonereis Kinberg, 1865 (Polychaeta: Nereididae) from the Atlantic Ocean, and a review of paragnath morphology and methodology. Zootaxa, 4471 (2), 245 - 278. https: // doi. org / 10.11646 / zootaxa. 4471.2.2" type="journal article" year="2018">Conde-Vela 2018: 269</bibRefCitation>
;
<figureCitation id="13772A55FFA9FF8D81B7FD72FB6CF29C" box="[1152,1244,726,753]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="10" pageNumber="502">Figs 5B</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D81DDFD72FA9BF29C" box="[1258,1323,727,753]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="10" pageNumber="502">6A–F</figureCitation>
,
<tableCitation id="C6CE036BFFA9FF8D8000FD72FA3BF29C" box="[1335,1419,727,753]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="10" pageNumber="502">Table 1</tableCitation>
).
<taxonomicName id="4C4C4D53FFA9FF8D85A0FD5EFE1FF378" authority="Hsueh, 2021" authorityName="Hsueh" authorityYear="2021" box="[151,431,763,789]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="kihawensis" status="sp. nov.">
<emphasis id="B938EAC2FFA9FF8D85A0FD5EFE1FF378" box="[151,431,763,789]" italics="true" pageId="10" pageNumber="502">Pseudonereis kihawensis</emphasis>
</taxonomicName>
<emphasis id="B938EAC2FFA9FF8D8482FD5EFE44F378" bold="true" box="[437,500,763,789]" pageId="10" pageNumber="502">
<taxonomicNameLabel id="A20B57B9FFA9FF8D8482FD5EFE44F378" box="[437,500,763,789]" pageId="10" pageNumber="502" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs from by
<taxonomicName id="4C4C4D53FFA9FF8D879DFD5EFC84F379" baseAuthorityName="Grube" baseAuthorityYear="1857" box="[682,820,763,789]" class="Polychaeta" family="Nereididae" genus="Pseudonereis" kingdom="Animalia" order="Phyllodocida" pageId="10" pageNumber="502" phylum="Annelida" rank="species" species="variegata">
<emphasis id="B938EAC2FFA9FF8D879DFD5EFD0BF379" box="[682,699,763,788]" italics="true" pageId="10" pageNumber="502">P</emphasis>
.
<emphasis id="B938EAC2FFA9FF8D87FEFD5EFC84F379" box="[713,820,763,788]" italics="true" pageId="10" pageNumber="502">variegata</emphasis>
</taxonomicName>
by having: 1) fewer number of paragnaths and a single band arrangement of paragnaths in Area VII–VIII (versus 43 and 2 bands, respectively); 2) λ-shaped ridge pattern of areas VI–V–VI (versus ɔc-shaped); 3) dorsal cirrus as long as dorsal ligule on anterior chaetigers (versus longer than dorsal ligule); 4) dorsal ligule longer than median ligule on anterior chaetigers (versus subequal); 5) dorsal cirrus distally attached to dorsal ligule from last 13 chaetigers (versus posterior-most chaetigers); and 6) no heterogomph spinigers in the subacicular fascicle of the neuropodia (versus present) (
<bibRefCitation id="EFDD4B21FFA9FF8D86FAFC0AFAB9F3A4" author="Conde-Vela, V. M." box="[973,1289,943,969]" pageId="10" pageNumber="502" pagination="245 - 278" refId="ref11340" refString="Conde-Vela, V. M. (2018) New species of Pseudonereis Kinberg, 1865 (Polychaeta: Nereididae) from the Atlantic Ocean, and a review of paragnath morphology and methodology. Zootaxa, 4471 (2), 245 - 278. https: // doi. org / 10.11646 / zootaxa. 4471.2.2" type="journal article" year="2018">Conde-Vela 2018: 267–269</bibRefCitation>
, fig. 10C–I;
<figureCitation id="13772A55FFA9FF8D85A0FC76FF41F380" box="[151,241,978,1005]" captionStart="FIGURE 5" captionStartId="7.[151,250,1940,1964]" captionTargetBox="[185,1404,199,1916]" captionTargetId="figure-33@7.[183,1404,199,1916]" captionTargetPageId="7" captionText="FIGURE 5. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, anterior body, dorsal view; B, paragnaths of the pharynx; C, Jaws; D, posterior body, dorsal view. Scale: A, 1.0 mm; B, D, 0.5 mm; C, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074788" httpUri="https://zenodo.org/record/5074788/files/figure.png" pageId="10" pageNumber="502">Figs 5B</figureCitation>
,
<figureCitation id="13772A55FFA9FF8D85CAFC76FE8EF380" box="[253,318,979,1005]" captionStart="FIGURE 6" captionStartId="8.[151,250,1026,1050]" captionTargetBox="[177,1410,181,1001]" captionTargetId="figure-15@8.[177,1410,181,1001]" captionTargetPageId="8" captionText="FIGURE 6. Pseudonereis kihawensis n. sp.; holotype (NMNS 8383-25): A, right parapodium, anterior view, chaetiger 10; B, right parapodium, posterior view, chaetiger 10; C, right parapodium, chaetiger 30; D, right parapodium, chaetiger 50; E, right parapodium, anterior view, chaetiger 71; F, right parapodium, chaetiger 88. Abbreviation: NeIA, neuroacicular inferior lobe; NePL, neuropodial postchaetal lobe; NeSA, neuroacicular superior acicular lobe. Scale: A–F, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5074790" httpUri="https://zenodo.org/record/5074790/files/figure.png" pageId="10" pageNumber="502">6A–F</figureCitation>
,
<tableCitation id="C6CE036BFFA9FF8D847FFC76FE2AF380" box="[328,410,979,1005]" captionStart="TABLE 1" captionStartId="16.[151,239,500,526]" captionText="TABLE 1. Key morphological characters of all known Pseudonereis to the world. Data in the table is largely based on Bakken (2007) with additional information from new described, new combined and resurrected species since 2007 by Glasby et al. (2013), Conde-Vela (2018), Kara et al. (2018) and the present study. Abbreviation: HeF, heterogomph falciger; HeS, heterogomph, spiniger; HoF, homogomph falciger; HoS, homogomph spiniger; LPTC, chaetigers reached by longest tentacular cirri; NTJ, number of teeth on the jaws. na = not available. * Conde-Vela (personal communicate)." pageId="10" pageNumber="502">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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