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<document id="9B3EF5D9557A25CE937DEA335ACF5057" ID-DOI="10.1093/zoolinnean/zlad064" ID-ISSN="0024-4082" ID-Zenodo-Dep="10470279" ID-ZooBank="9DCD56DB-3954-47CC-A2C5-483409D3958BC" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1704718537007" checkinUser="plazi" docAuthor="Zelaya, Diego G. &amp; Güller, Marina" docDate="2023" docId="03E587A0FFCBFFC0FC48FC05D93CF850" docLanguage="en" docName="zlad064.pdf" docOrigin="Zoological Journal of the Linnean Society 199 (4)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad064" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Hiatella barnea Zelaya and Marina Güller 2023, comb. nov." docType="treatment" docVersion="1" lastPageNumber="900" masterDocId="FFDCFFD8FFC5FFD2FFD8F867D813FFEC" masterDocTitle="Decrypting the feared genus Hiatella (Bivalvia): South American species" masterLastPageNumber="905" masterPageNumber="882" pageNumber="896" updateTime="1704803696153" updateUser="felipe">
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<mods:title id="AD5C3858DBC6B1F34C62B02B6E606B99">Decrypting the feared genus Hiatella (Bivalvia): South American species</mods:title>
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<mods:namePart id="E73222C11CA036BF2516AE5F4F5F9370">Zelaya, Diego G.</mods:namePart>
<mods:affiliation id="06EA3428D241C4635621294AAC4513DB">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina &amp; Universidad de Buenos Aires, Facultad de Ciencias Exactas y Naturales, Departamento Biodiversidad y Biología Experimental, Ciudad Universitaria, Pabellón 2, 4 ° Piso, Buenos Aires, Argentina</mods:affiliation>
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<mods:namePart id="BF0C2F8136CC2F2ACDF405EB64BCEB52">Güller, Marina</mods:namePart>
<mods:affiliation id="5AC18D9A19BFEAC196C61CD6CC20A183">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Buenos Aires, Argentina &amp; Universidad de Buenos Aires, Facultad de Ciencias Exactas y Naturales, Departamento Biodiversidad y Biología Experimental, Ciudad Universitaria, Pabellón 2, 4 ° Piso, Buenos Aires, Argentina</mods:affiliation>
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<paragraph id="8BF336B6FFCBFFDCFC48FC05DD5EFB90" blockId="14.[912,1357,1122,1149]" box="[912,1357,1122,1149]" pageId="14" pageNumber="896">
<heading id="D0BB81DAFFCBFFDCFC48FC05DD5EFB90" bold="true" box="[912,1357,1122,1149]" centered="true" fontSize="11" level="2" pageId="14" pageNumber="896" reason="2">
<emphasis id="B938EAA4FFCBFFDCFC48FC05DD5EFB90" bold="true" box="[912,1357,1122,1149]" pageId="14" pageNumber="896">
<taxonomicName id="4C4C4D35FFCBFFDCFC48FC05DCC4FB90" ID-CoL="BV45X" authority="(Ihering, 1907)" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[912,1239,1122,1149]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="14" pageNumber="896" phylum="Mollusca" rank="species" species="barnea" status="comb. nov.">
<emphasis id="B938EAA4FFCBFFDCFC48FC05DC3FFB91" bold="true" box="[912,1068,1122,1149]" italics="true" pageId="14" pageNumber="896">Hiatella barnea</emphasis>
(
<bibRefCitation id="EFDD4B47FFCBFFDCFBE4FC05DCDFFB90" author="von Ihering HFA" box="[1084,1228,1122,1148]" pageId="14" pageNumber="896" pagination="1 - 611" refId="ref19407" refString="von Ihering HFA. Les mollusques fossiles du Tertiare et du Cretace Superieur de l'Argentine. Anales del Museo Nacional de Buenos Aires 1907; 14: 1 - 611." type="journal article" year="1907">Ihering, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20B57DFFFCBFFDCFB04FC05DD5EFB90" box="[1244,1357,1122,1148]" pageId="14" pageNumber="896" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356653DFFCBFFDCFC14FCC7DD01FB56" box="[972,1298,1184,1211]" pageId="14" pageNumber="896" type="description">
<paragraph id="8BF336B6FFCBFFDCFC14FCC7DD01FB56" blockId="14.[972,1298,1184,1211]" box="[972,1298,1184,1211]" pageId="14" pageNumber="896">
<emphasis id="B938EAA4FFCBFFDCFC14FCC7DD01FB56" bold="true" box="[972,1298,1184,1211]" pageId="14" pageNumber="896">
(
<figureCitation id="13772A33FFCBFFDCFC0EFCC6DC33FB57" box="[982,1056,1185,1211]" captionStart="Figure 3" captionStartId="5.[129,194,1708,1732]" captionTargetBox="[230,1372,144,1680]" captionTargetId="figure-5@5.[230,1372,144,1680]" captionTargetPageId="5" captionText="Figure 3. Early ontogeny shells of Hiatella meridionalis (AL), Hiatella umbonata (MO) and Hiatella barnea (P). A, C, Saxicava meridionalis: original illustrations (A) and photographs of the lectotype, NHMUK 1854.12.4.772 (C). B, F, G, IK, specimens from Burdwood Bank: MACN-In 44166 (B), MACN-In 44179 (F), MACN-In 44187 (G), MACN-In 44192 (I), MACN-In 44172 (J) and MACN-In 44152 (K). D, Saxicava lebruni, syntype, MNHN-IM-2000-31626. E, specimen from Beagle Channel, MACN-In 10061. L, DNA voucher, H41. H, specimen from Fiordo Ponsonby, MACN-In 44282 (drawn indented lines show the aưachment of the mantle margin to the shell and siphonal muscle scars). M, specimens from Burdwood Bank, MACN-In 44270. N, specimen from Beagle Channel, MACN-In 44235. O, specimen from the Argentine Continental Margin, MACN-In 36596. P, specimens from the Pacific coast, MACN-In 44278. Scale bars: 10 mm in AL; 2 mm in MO; 5 mm in P." figureDoi="http://doi.org/10.5281/zenodo.10470289" httpUri="https://zenodo.org/record/10470289/files/figure.png" pageId="14" pageNumber="896">Figs 3P</figureCitation>
,
<figureCitation id="13772A33FFCBFFDCFBF2FCC6DC99FB57" box="[1066,1162,1185,1211]" captionStart="Figure 6" captionStartId="8.[113,178,1775,1799]" captionTargetBox="[186,1387,145,1746]" captionTargetId="figure-7@8.[184,1388,144,1747]" captionTargetPageId="8" captionText="Figure 6. Anatomy of Hiatella species:Hiatella meridionalis (A, B, G, J) from the Magellan Strait (MACN-In 12414), Hiatella umbonata (C, D, H, K) from Burdwood Bank (MACN-In 44270) and Hiatella barnea (E, F, I, L) from Chilean ḩords (MACN-In 44278). A, C, E, general view with right valve removed.B, D, F, right view with right mantle fold removed.GI, right gill. JL, foot. Abbreviations: aa, anterior adductor muscle; b, byssus; esr, exhalant siphon retractor muscle; f, foot; g, gill; go, gonad; h, heel; id, inner demibranch; isr, inhalant siphon retractor muscle; od, outer demibranch; pa, posterior adductor muscle; t, toe; vm, visceral mass. Scale bars:10 mm in AF; 2 mm in GL." figureDoi="http://doi.org/10.5281/zenodo.10470295" httpUri="https://zenodo.org/record/10470295/files/figure.png" pageId="14" pageNumber="896">6E, F, I, L</figureCitation>
,
<figureCitation id="13772A33FFCBFFDCFB4CFCC6DCA5FB57" box="[1172,1206,1185,1211]" captionStart="Figure 7" captionStartId="9.[129,194,1708,1732]" captionTargetBox="[192,1410,144,1680]" captionTargetId="figure-107@9.[192,1410,144,1680]" captionTargetPageId="9" captionText="Figure 7. Living specimens of Hiatella: Hiatella umbonata (AE) from Burdwood Bank (A, C) and the Argentine continental margin (B, D, E); Hiatella meridionalis (F, G) from Puerto Edén (F) and Magellan Strait (G); and Hiatella barnea (H) from ºuellón. Arrowheads (B, E) point to specimens of Hiatella." figureDoi="http://doi.org/10.5281/zenodo.10470297" httpUri="https://zenodo.org/record/10470297/files/figure.png" pageId="14" pageNumber="896">7H</figureCitation>
,
<figureCitation id="13772A33FFCBFFDCFB19FCC6DD14FB57" box="[1217,1287,1185,1211]" captionStart-0="Figure 11" captionStart-1="Figure 12" captionStart-2="Figure 13" captionStartId-0="15.[129,194,1926,1950]" captionStartId-1="16.[112,177,1708,1732]" captionStartId-2="17.[129,194,1494,1518]" captionTargetBox-0="[225,1377,1438,1898]" captionTargetBox-1="[188,1384,144,1680]" captionTargetBox-2="[209,1393,144,1466]" captionTargetId-0="figure-315@15.[225,1377,1438,1898]" captionTargetId-1="figure-100@16.[188,1384,144,1680]" captionTargetId-2="figure-209@17.[209,1393,144,1466]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionText-0="Figure 11. Saxicava barnea. A, original illustration.B, original label. CF, holotype, MACN-Pi N 510, in right (C), less (D), dorsal (E) and ventral (F) views.Scale bar:5 mm." captionText-1="Figure 12. Shells of Hiatella barnea, external views. A, F, specimens from the Pacific coast, MACN-In 44278. BE, IK, specimens from the Atlantic coast: MACN-In 9175-19 (B, I); MACN-In 9156-3 (C); MACN-In 9032-3 (D, K); MACN-In 9237-8 (E), MACN-In 32620 (J). G, H, DNA vouchers:H43 (G) and H42 (H). Scale bars: 10 mm in AJ; 1 mm in K." captionText-2="Figure 13. Shells of Hiatella barnea, inner and ventral views and hinge teeth. A, specimen from the Pacific coast, MACN-In 44278. B, D, F, G, specimens from the Atlantic coast: MACN-In 9175-19 (B), MACN-In 9156-3 (D), MACN-In 9237-8 (F) and MACN-In 44274 (G). C, DNA voucher, H42 (drawn indented lines show the aưachment of the mantle margin to the shell and siphonal muscle scars). E, specimen from Isla de los Estados, MLP-Ma 4942. Scale bars: 5 mm in AD, F; 500 µm in E; 2 mm in G." figureDoi-0="http://doi.org/10.5281/zenodo.10470309" figureDoi-1="http://doi.org/10.5281/zenodo.10470311" figureDoi-2="http://doi.org/10.5281/zenodo.10470313" httpUri-0="https://zenodo.org/record/10470309/files/figure.png" httpUri-1="https://zenodo.org/record/10470311/files/figure.png" httpUri-2="https://zenodo.org/record/10470313/files/figure.png" pageId="14" pageNumber="896">1113</figureCitation>
)
</emphasis>
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<subSubSection id="C356653DFFCBFFDCFCF2FCAFDCA3FA77" pageId="14" pageNumber="896" type="reference_group">
<paragraph id="8BF336B6FFCBFFDCFCF2FCAFDB7BFB13" blockId="14.[810,1459,1223,1435]" pageId="14" pageNumber="896">
<treatmentCitationGroup id="AB5C1198FFCBFFDCFCF2FCAFDB7BFB13" pageId="14" pageNumber="896">
<taxonomicName id="4C4C4D35FFCBFFDCFCF2FCAFDBD9FB33" box="[810,970,1224,1247]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="14" pageNumber="896" phylum="Mollusca" rank="species" species="carnea">
<emphasis id="B938EAA4FFCBFFDCFCF2FCAFDBD9FB33" box="[810,970,1224,1247]" italics="true" pageId="14" pageNumber="896">Saxicava carnea</emphasis>
</taxonomicName>
(sic)
<treatmentCitation id="0AED10A7FFCBFFDCFBD1FCA0DCDFFB33" author="von Ihering HFA" box="[1033,1228,1223,1247]" page="105" pageId="14" pageNumber="896" year="1907">
<bibRefCitation id="EFDD4B47FFCBFFDCFBD1FCA0DCDFFB33" author="von Ihering HFA" box="[1033,1228,1223,1247]" pageId="14" pageNumber="896" pagination="1 - 611" refId="ref19407" refString="von Ihering HFA. Les mollusques fossiles du Tertiare et du Cretace Superieur de l'Argentine. Anales del Museo Nacional de Buenos Aires 1907; 14: 1 - 611." type="journal article" year="1907">Ihering, 1907: 105</bibRefCitation>
</treatmentCitation>
(
<emphasis id="B938EAA4FFCBFFDCFB07FCAFDD7EFB33" box="[1247,1389,1223,1247]" italics="true" pageId="14" pageNumber="896">nomen nudum</emphasis>
; listed only).
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</paragraph>
<paragraph id="8BF336B6FFCBFFDCFC9DFD61DB98FAD1" blockId="14.[810,1459,1223,1435]" pageId="14" pageNumber="896">
<treatmentCitationGroup id="AB5C1198FFCBFFDCFC9DFD61DB98FAD1" pageId="14" pageNumber="896">
<treatmentCitation id="0AED10A7FFCBFFDCFC9DFD61DCAFFAF2" author="von Ihering HFA" box="[837,1212,1286,1310]" page="325" pageId="14" pageNumber="896" year="1907">
<taxonomicName id="4C4C4D35FFCBFFDCFC9DFD61DCAFFAF2" ID-CoL="BV4LQ" authority="Ihering, 1907: 325" authorityName="Ihering" authorityPageNumber="325" authorityYear="1907" box="[837,1212,1286,1310]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="14" pageNumber="896" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFCBFFDCFC9DFD61DBF8FAF2" box="[837,1003,1286,1310]" italics="true" pageId="14" pageNumber="896">Saxicava barnea</emphasis>
<bibRefCitation id="EFDD4B47FFCBFFDCFC2CFD61DCAFFAF2" author="von Ihering HFA" box="[1012,1212,1286,1310]" pageId="14" pageNumber="896" pagination="1 - 611" refId="ref19407" refString="von Ihering HFA. Les mollusques fossiles du Tertiare et du Cretace Superieur de l'Argentine. Anales del Museo Nacional de Buenos Aires 1907; 14: 1 - 611." type="journal article" year="1907">Ihering, 1907: 325</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 14, fig. 96;
<treatmentCitation id="0AED10A7FFCBFFDCFAB0FD61DB94FAD1" author="von Ihering H" page="63" pageId="14" pageNumber="896" year="1914">
<bibRefCitation id="EFDD4B47FFCBFFDCFAB0FD61DB94FAD1" author="von Ihering H" pageId="14" pageNumber="896" pagination="1 - 148" refId="ref19371" refString="von Ihering H. Catalogo de moluscos Cretaceos e Terciarios da Argentina da collecao do autor. Notas Preliminares da Revista do Museo Paulista (Sao Paulo) 1914; 1: 1 - 148." type="journal article" year="1914">Ihering 1914: 63</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BF336B6FFCBFFDCFC9DFD22DD52FA90" blockId="14.[810,1459,1223,1435]" pageId="14" pageNumber="896">
<treatmentCitationGroup id="AB5C1198FFCBFFDCFC9DFD22DD52FA90" pageId="14" pageNumber="896">
<taxonomicName id="4C4C4D35FFCBFFDCFC9DFD22DBC0FAB1" ID-CoL="3LHVL" baseAuthorityName="G. B. Sowerby I" baseAuthorityYear="1834" box="[837,979,1349,1373]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="14" pageNumber="896" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B938EAA4FFCBFFDCFC9DFD22DBC0FAB1" box="[837,979,1349,1373]" italics="true" pageId="14" pageNumber="896">Hiatella solida</emphasis>
</taxonomicName>
:
<treatmentCitation id="0AED10A7FFCBFFDCFC07FD22DCF3FAB1" author="Carcelles AR" box="[991,1248,1348,1373]" page="222" pageId="14" pageNumber="896" year="1953">
<bibRefCitation id="EFDD4B47FFCBFFDCFC07FD22DCF3FAB1" author="Carcelles AR" box="[991,1248,1348,1373]" pageId="14" pageNumber="896" pagination="155 - 250" refId="ref16576" refString="Carcelles AR. Catalogo de la malacofauna Antartica Argentina. Anales del Museo Nahuel Huapi 1953; 3: 155 - 250." type="journal article" year="1953">Carcelles 1953: 222223</bibRefCitation>
</treatmentCitation>
, pl. 5, fig. 111;
<bibRefCitation id="EFDD4B47FFCBFFDCFAA0FD22DC12FA90" author="Aldea Venegas C &amp; Valdovinos Zarges C" pageId="14" pageNumber="896" pagination="364 - 96" refId="ref16040" refString="Aldea Venegas C, Valdovinos Zarges C. Moluscos del intermareal rocoso del centro-sur de Chile (36 ° - 38 ° S): taxonomia y clave de identificacion. Gayana 2005; 69: 364 - 96. hups: // doi. org / 10.4067 / S 0717 - 65382005000200014" type="journal article" year="2005">Aldea and Valdovinos 2005</bibRefCitation>
: fig. 11I (not Sowerby I, 1834).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BF336B6FFCBFFDCFC9DFDE3DCA3FA77" blockId="14.[810,1459,1223,1435]" box="[837,1200,1411,1435]" pageId="14" pageNumber="896">
<emphasis id="B938EAA4FFCBFFDCFC9DFDE3DBF6FA77" box="[837,997,1412,1435]" italics="true" pageId="14" pageNumber="896">Barnea saxicava</emphasis>
<bibRefCitation id="EFDD4B47FFCBFFDCFC32FDE4DCBFFA77" author="Parodiz JJ" box="[1002,1196,1411,1435]" pageId="14" pageNumber="896" pagination="183 - 296" refId="ref18393" refString="Parodiz JJ. The taxa of fossil Mollusca introduced by Hermann von Ihering. Annals of the Carnegie Museum 1996; 65: 183 - 296." type="journal article" year="1996">Parodiz, 1996: 267</bibRefCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BF336B6FFCBFFDCFCF2FDDADBB3FA18" blockId="14.[810,1459,1467,1524]" pageId="14" pageNumber="896">
<materialsCitation id="3B243CEBFFCBFFDCFCF2FDDADB8FFA18" country="Argentina" latitude="-44.8" location="Chubut Province" longLatPrecision="1134" longitude="-65.7" pageId="14" pageNumber="896" specimenCount="1" stateProvince="Chubut" typeStatus="holotype">
<emphasis id="B938EAA4FFCBFFDCFCF2FDDADBBFFA39" box="[810,940,1469,1493]" italics="true" pageId="14" pageNumber="896">
<typeStatus id="54F78814FFCBFFDCFCF2FDDADB49FA38" box="[810,858,1469,1492]" pageId="14" pageNumber="896">Type</typeStatus>
locality:
</emphasis>
Camarones [
<geoCoordinate id="EE785071FFCBFFDCFBEBFDDADC9FFA39" box="[1075,1164,1467,1493]" degrees="44" direction="south" minutes="48" orientation="latitude" pageId="14" pageNumber="896" precision="925" value="-44.8">
44°48
<emphasis id="B938EAA4FFCBFFDCFBABFDDCDC6AFA38" box="[1139,1145,1467,1492]" italics="true" pageId="14" pageNumber="896">ʹ</emphasis>
S
</geoCoordinate>
,
<geoCoordinate id="EE785071FFCBFFDCFB4EFDDBDCE6FA38" box="[1174,1269,1467,1492]" degrees="65" direction="west" minutes="42" orientation="longitude" pageId="14" pageNumber="896" precision="925" value="-65.7">
65°42
<emphasis id="B938EAA4FFCBFFDCFB0AFDDCDCCBFA38" box="[1234,1240,1467,1492]" italics="true" pageId="14" pageNumber="896">ʹ</emphasis>
W
</geoCoordinate>
,
<collectingRegion id="4988F854FFCBFFDCFB26FDDADDBCFA39" box="[1278,1455,1469,1493]" country="Argentina" name="Chubut" pageId="14" pageNumber="896">Chubut Province</collectingRegion>
,
<collectingCountry id="F35B7626FFCBFFDCFCF2FDBBDB85FA18" box="[810,918,1500,1524]" name="Argentina" pageId="14" pageNumber="896">Argentina</collectingCountry>
]
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336B6FFCBFFDCFCF2FE72DCEBF9A1" blockId="14.[810,1459,1557,1613]" pageId="14" pageNumber="896">
<emphasis id="B938EAA4FFCBFFDCFCF2FE72DBFFF9C1" box="[810,1004,1557,1581]" italics="true" pageId="14" pageNumber="896">Material examined:</emphasis>
<materialsCitation id="3B243CEBFFCBFFDCFC2CFE72DCEBF9A1" collectionCode="ESM" pageId="14" pageNumber="896" specimenCode="ESM1" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F78814FFCBFFDCFC2CFE72DC44F9C1" box="[1012,1111,1557,1581]" pageId="14" pageNumber="896" type="holotype">Holotype</typeStatus>
(
<emphasis id="B938EAA4FFCBFFDCFBB2FE71DD08F9C1" box="[1130,1307,1557,1581]" italics="true" pageId="14" pageNumber="896">MACN-Pi N 510</emphasis>
); and 37 additional lots (Supporting Information,
<specimenCode id="DBEA9ECDFFCBFFDCFB7EFE52DCF4F9A1" box="[1190,1255,1589,1613]" collectionCode="ESM" pageId="14" pageNumber="896">ESM1</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF336B6FFCBFFDCFCF2FE09DD08F949" blockId="14.[810,1458,1644,1701]" pageId="14" pageNumber="896">
<emphasis id="B938EAA4FFCBFFDCFCF2FE09DC0CF96A" box="[810,1055,1646,1670]" italics="true" pageId="14" pageNumber="896">Other published records:</emphasis>
<materialsCitation id="3B243CEBFFCBFFDCFBEAFE09DD04F949" country="Iceland" latitude="-37.04861" location="Santa Maria" longLatPrecision="19" longitude="-73.51139" pageId="14" pageNumber="896" specimenCount="1">
<geoCoordinate id="EE785071FFCBFFDCFBEAFE09DCBEF96A" box="[1074,1197,1644,1670]" degrees="37" direction="south" minutes="02" orientation="latitude" pageId="14" pageNumber="896" precision="15" seconds="55" value="-37.04861">
37°02
<emphasis id="B938EAA4FFCBFFDCFBB6FE0BDC67F969" box="[1134,1140,1644,1669]" italics="true" pageId="14" pageNumber="896">ʹ</emphasis>
55″ S
</geoCoordinate>
,
<geoCoordinate id="EE785071FFCBFFDCFB63FE09DD52F969" box="[1211,1345,1644,1670]" degrees="73" direction="west" minutes="30" orientation="longitude" pageId="14" pageNumber="896" precision="15" seconds="41" value="-73.51139">
73°30
<emphasis id="B938EAA4FFCBFFDCFB2FFE0BDCEEF969" box="[1271,1277,1644,1669]" italics="true" pageId="14" pageNumber="896">ʹ</emphasis>
41″ W
</geoCoordinate>
,
<collectingCountry id="F35B7626FFCBFFDCFA96FE09DD62F96A" box="[1358,1393,1646,1670]" name="Iceland" pageId="14" pageNumber="896">Isla</collectingCountry>
<location id="8E93606DFFCBFFDCFAA2FE09DB74F949" LSID="urn:lsid:plazi:treatment:03E587A0FFCBFFC0FC48FC05D93CF850:8E93606DFFCBFFDCFAA2FE09DB74F949" country="Iceland" latitude="-37.04861" longLatPrecision="19" longitude="-73.51139" name="Santa Maria" pageId="14" pageNumber="896">Santa María</location>
(
<bibRefCitation id="EFDD4B47FFCBFFDCFCAFFEEADC9DF949" author="Aldea Venegas C &amp; Valdovinos Zarges C" box="[887,1166,1677,1701]" pageId="14" pageNumber="896" pagination="364 - 96" refId="ref16040" refString="Aldea Venegas C, Valdovinos Zarges C. Moluscos del intermareal rocoso del centro-sur de Chile (36 ° - 38 ° S): taxonomia y clave de identificacion. Gayana 2005; 69: 364 - 96. hups: // doi. org / 10.4067 / S 0717 - 65382005000200014" type="journal article" year="2005">Aldea and Valdovinos 2005</bibRefCitation>
: as
<taxonomicName id="4C4C4D35FFCBFFDCFB6CFEE9DD18F949" baseAuthorityName="G. B. Sowerby I" baseAuthorityYear="1834" box="[1204,1291,1677,1701]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="14" pageNumber="896" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B938EAA4FFCBFFDCFB6CFEE9DD18F949" box="[1204,1291,1677,1701]" italics="true" pageId="14" pageNumber="896">H. solida</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF336B6FFCBFFDDFCF2FEA0DAB3FE88" blockId="14.[810,1459,1734,1978]" lastBlockId="15.[129,778,144,606]" lastPageId="15" lastPageNumber="897" pageId="14" pageNumber="896">
<emphasis id="B938EAA4FFCBFFDCFCF2FEA0DBB2F932" box="[810,929,1735,1758]" italics="true" pageId="14" pageNumber="896">Description:</emphasis>
Shell ≤
<quantity id="4CB49B53FFCBFFDCFC23FEA1DC4AF932" box="[1019,1113,1734,1758]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" pageId="14" pageNumber="896" unit="mm" value="37.5">37.5 mm</quantity>
long, narrowly elongate, very inflated, with the maximum width at about the anterior third of total shell length; moderately solid (
<figureCitation id="13772A33FFCBFFDCFB46FF62DCF7F8F1" box="[1182,1252,1797,1821]" captionStart="Figure 12" captionStartId="16.[112,177,1708,1732]" captionTargetBox="[188,1384,144,1680]" captionTargetId="figure-100@16.[188,1384,144,1680]" captionTargetPageId="16" captionText="Figure 12. Shells of Hiatella barnea, external views. A, F, specimens from the Pacific coast, MACN-In 44278. BE, IK, specimens from the Atlantic coast: MACN-In 9175-19 (B, I); MACN-In 9156-3 (C); MACN-In 9032-3 (D, K); MACN-In 9237-8 (E), MACN-In 32620 (J). G, H, DNA vouchers:H43 (G) and H42 (H). Scale bars: 10 mm in AJ; 1 mm in K." figureDoi="http://doi.org/10.5281/zenodo.10470311" httpUri="https://zenodo.org/record/10470311/files/figure.png" pageId="14" pageNumber="896">Fig. 12</figureCitation>
). Right valve somewhat larger and ventrally overlapping over less valve. Anterior and posterior ends projected. Posterior area of shell narrow, flat, ill defined. Anterior area of shell with a wide, deep sulcus, radiating from the umbo to the ventral margin. Ventral gape very wide throughout ontogeny; posterior gape small. Anterodorsal margin short, sloping straight. Anterior margin short and rounded. Ventral margin deeply sinuated anteriorly at the gape, rounded to flat posteriorly. Posterior margin widely arched to flaưened, higher than anterior margin. Posterodorsal margin horizontal, straight to slightly convex, longer than the anterodorsal margin. Umbo low, wide, evenly rounded; located at about the anterior third of total shell length. Lunule short, wide, sunken.
</paragraph>
<paragraph id="8BF336B6FFCAFFDDFF44F90CDD28FE4E" blockId="15.[129,778,144,606]" lastBlockId="15.[825,1474,144,418]" pageId="15" pageNumber="897">
Outer shell surface white, chalky (
<figureCitation id="13772A33FFCAFFDDFDDCF90CDA43FE6F" box="[516,592,363,387]" captionStart="Figure 11" captionStartId="15.[129,194,1926,1950]" captionTargetBox="[225,1377,1438,1898]" captionTargetId="figure-315@15.[225,1377,1438,1898]" captionTargetPageId="15" captionText="Figure 11. Saxicava barnea. A, original illustration.B, original label. CF, holotype, MACN-Pi N 510, in right (C), less (D), dorsal (E) and ventral (F) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.10470309" httpUri="https://zenodo.org/record/10470309/files/figure.png" pageId="15" pageNumber="897">Figs 11</figureCitation>
,
<figureCitation id="13772A33FFCAFFDDFD86F90CDA6BFE6F" box="[606,632,363,387]" captionStart="Figure 12" captionStartId="16.[112,177,1708,1732]" captionTargetBox="[188,1384,144,1680]" captionTargetId="figure-100@16.[188,1384,144,1680]" captionTargetPageId="16" captionText="Figure 12. Shells of Hiatella barnea, external views. A, F, specimens from the Pacific coast, MACN-In 44278. BE, IK, specimens from the Atlantic coast: MACN-In 9175-19 (B, I); MACN-In 9156-3 (C); MACN-In 9032-3 (D, K); MACN-In 9237-8 (E), MACN-In 32620 (J). G, H, DNA vouchers:H43 (G) and H42 (H). Scale bars: 10 mm in AJ; 1 mm in K." figureDoi="http://doi.org/10.5281/zenodo.10470311" httpUri="https://zenodo.org/record/10470311/files/figure.png" pageId="15" pageNumber="897">12</figureCitation>
). Dissoconch sculptured with strong, regularly distributed commarginal elements, separated by wide interspaces. Commarginal sculpture consisting of low, wide cords near the umbo, gradually narrowing and increasing in height, originating lamellae ventrally. Lamellae deeply sinuated at anterior sulcus, where they also bear short but stout, triangular, ventrally projected scales. In specimens ≤
<quantity id="4CB49B53FFCAFFDDFF0FFA21D906FDB2" box="[215,277,582,606]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="897" unit="mm" value="5.0">5 mm</quantity>
L, two extremely thin radial ribs bearing small, hollow spines extend posteriorly: one of them running from the prodissoconchdissoconch limit to the junction of posterior and ventral margins, the second to the junction of dorsal and posterior margins. In larger specimens, ribs lack spines distally, and early secreted spines (in the vicinity of the umbo) appear reduced to their bases, appearing as tubercles; the radial ribs are recognizable only by an abrupt change in the orientation of commarginal sculpture. Periostracum thick, yellowish straw coloured to brownish, forming thin periostracal folds.
</paragraph>
<paragraph id="8BF336B6FFCAFFC2FCE1F9A3D94CF82E" blockId="15.[825,1474,452,601]" lastBlockId="16.[113,763,1836,1986]" lastPageId="16" lastPageNumber="898" pageId="15" pageNumber="897">
<emphasis id="B938EAA4FFCAFFDDFCE1F9A3DBF3FE37" box="[825,992,452,476]" italics="true" pageId="15" pageNumber="897">
Hinge (
<figureCitation id="13772A33FFCAFFDDFC5FF9A3DBDEFE30" box="[903,973,452,476]" captionStart="Figure 13" captionStartId="17.[129,194,1494,1518]" captionTargetBox="[209,1393,144,1466]" captionTargetId="figure-209@17.[209,1393,144,1466]" captionTargetPageId="17" captionText="Figure 13. Shells of Hiatella barnea, inner and ventral views and hinge teeth. A, specimen from the Pacific coast, MACN-In 44278. B, D, F, G, specimens from the Atlantic coast: MACN-In 9175-19 (B), MACN-In 9156-3 (D), MACN-In 9237-8 (F) and MACN-In 44274 (G). C, DNA voucher, H42 (drawn indented lines show the aưachment of the mantle margin to the shell and siphonal muscle scars). E, specimen from Isla de los Estados, MLP-Ma 4942. Scale bars: 5 mm in AD, F; 500 µm in E; 2 mm in G." figureDoi="http://doi.org/10.5281/zenodo.10470313" httpUri="https://zenodo.org/record/10470313/files/figure.png" pageId="15" pageNumber="897">Fig. 13</figureCitation>
):
</emphasis>
Small-sized specimens (≤
<quantity id="4CB49B53FFCAFFDDFAD0F9A3DD5BFE37" box="[1288,1352,452,476]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="897" unit="mm" value="8.0">8 mm</quantity>
long) with a delicate, small cardinal tooth at each valve, and the corresponding depression to accommodate the tooth of the opposite valve. Less valve cardinal triangular; depression anterior to cardinal tooth. Right valve cardinal low, peg-like, ventrally directed; depression behind cardinal tooth. Teeth and depressions lost in specimens&gt;
<quantity id="4CB49B53FFD5FFC2FEC5FF2BD97AF88F" box="[285,361,1868,1892]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="898" unit="mm" value="10.0">10 mm</quantity>
long, where valve articulation occurs only by means of the ligament. Nymph short but stout, flaring. Ligament opisthodetic. Pallial sinus deep. Inner shell surface porcellanaceous, white.
</paragraph>
<caption id="DF33663EFFCAFFDDFF58FD5DDA1FFA82" ID-DOI="http://doi.org/10.5281/zenodo.10470307" ID-Zenodo-Dep="10470307" httpUri="https://zenodo.org/record/10470307/files/figure.png" pageId="15" pageNumber="897" startId="15.[128,193,1338,1362]" targetBox="[321,1281,668,1310]" targetPageId="15" targetType="figure">
<paragraph id="8BF336B6FFCAFFDDFF58FD5DDA1FFA82" blockId="15.[128,1443,1338,1391]" pageId="15" pageNumber="897">
<emphasis id="B938EAA4FFCAFFDDFF58FD5DD8F7FABE" bold="true" box="[128,228,1338,1362]" pageId="15" pageNumber="897">Figure 10.</emphasis>
<taxonomicName id="4C4C4D35FFCAFFDDFF31FD5CD967FABE" baseAuthorityName="Sowerby I" baseAuthorityYear="1834" box="[233,372,1338,1362]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="15" pageNumber="897" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B938EAA4FFCAFFDDFF31FD5CD967FABE" box="[233,372,1338,1362]" italics="true" pageId="15" pageNumber="897">Saxicava solida</emphasis>
</taxonomicName>
, lectotype, NHMUK 20220347. A, B, right valve, internal (A) and external (B) views. C, D, less valve, internal (C) and external (D) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="DF33663EFFCAFFDDFF59FFE1D9DAF856" ID-DOI="http://doi.org/10.5281/zenodo.10470309" ID-Zenodo-Dep="10470309" httpUri="https://zenodo.org/record/10470309/files/figure.png" pageId="15" pageNumber="897" startId="15.[129,194,1926,1950]" targetBox="[225,1377,1438,1898]" targetPageId="15" targetType="figure">
<paragraph id="8BF336B6FFCAFFDDFF59FFE1D9DAF856" blockId="15.[129,1419,1926,1979]" pageId="15" pageNumber="897">
<emphasis id="B938EAA4FFCAFFDDFF59FFE1D8F6F873" bold="true" box="[129,229,1926,1951]" pageId="15" pageNumber="897">Figure 11.</emphasis>
<taxonomicName id="4C4C4D35FFCAFFDDFF31FFE0D96DF873" authorityName="Ihering" authorityYear="1907" box="[233,382,1927,1951]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="15" pageNumber="897" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFCAFFDDFF31FFE0D96DF873" box="[233,382,1927,1951]" italics="true" pageId="15" pageNumber="897">Saxicava barnea</emphasis>
</taxonomicName>
. A, original illustration. B, original label. CF, holotype,
<emphasis id="B938EAA4FFCAFFDDFC48FFE0DC3CF873" box="[912,1071,1927,1951]" italics="true" pageId="15" pageNumber="897">MACN-Pi N 510</emphasis>
, in right (C), less (D), dorsal (E) and ventral (F) views. Scale bar: 5 mm.
</paragraph>
</caption>
<caption id="DF33663EFFD5FFC2FFA8FECBDB7DF910" ID-DOI="http://doi.org/10.5281/zenodo.10470311" ID-Zenodo-Dep="10470311" httpUri="https://zenodo.org/record/10470311/files/figure.png" pageId="16" pageNumber="898" startId="16.[112,177,1708,1732]" targetBox="[188,1384,144,1680]" targetPageId="16" targetType="figure">
<paragraph id="8BF336B6FFD5FFC2FFA8FECBDB7DF910" blockId="16.[112,1436,1708,1789]" pageId="16" pageNumber="898">
<emphasis id="B938EAA4FFD5FFC2FFA8FECBD8C7F928" bold="true" box="[112,212,1708,1732]" pageId="16" pageNumber="898">Figure 12.</emphasis>
Shells of
<taxonomicName id="4C4C4D35FFD5FFC2FEF5FECBD9A9F928" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[301,442,1708,1732]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="16" pageNumber="898" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD5FFC2FEF5FECBD9A9F928" box="[301,442,1708,1732]" italics="true" pageId="16" pageNumber="898">Hiatella barnea</emphasis>
</taxonomicName>
, external views. A, F, specimens from the Pacific coast, MACN-In 44278. BE, IK, specimens from the Atlantic coast: MACN-In 9175-19 (B, I); MACN-In 9156-3 (C); MACN-In 9032-3 (D, K); MACN-In 9237-8 (E), MACN-In 32620 (J). G, H, DNA vouchers: H43 (G) and H42 (H). Scale bars: 10 mm in AJ; 1 mm in K.
</paragraph>
</caption>
<paragraph id="8BF336B6FFD5FFC0FCF2FF4AD94DFF2B" blockId="16.[810,1459,1836,1986]" lastBlockId="18.[113,763,144,199]" lastPageId="18" lastPageNumber="900" pageId="16" pageNumber="898">
<emphasis id="B938EAA4FFD5FFC2FCF2FF4ADC25F8A8" box="[810,1078,1836,1860]" italics="true" pageId="16" pageNumber="898">
Anatomy (
<figureCitation id="13772A33FFD5FFC2FC4DFF4ADC37F8A8" box="[917,1060,1836,1860]" captionStart="Figure 6" captionStartId="8.[113,178,1775,1799]" captionTargetBox="[186,1387,145,1746]" captionTargetId="figure-7@8.[184,1388,144,1747]" captionTargetPageId="8" captionText="Figure 6. Anatomy of Hiatella species:Hiatella meridionalis (A, B, G, J) from the Magellan Strait (MACN-In 12414), Hiatella umbonata (C, D, H, K) from Burdwood Bank (MACN-In 44270) and Hiatella barnea (E, F, I, L) from Chilean ḩords (MACN-In 44278). A, C, E, general view with right valve removed.B, D, F, right view with right mantle fold removed.GI, right gill. JL, foot. Abbreviations: aa, anterior adductor muscle; b, byssus; esr, exhalant siphon retractor muscle; f, foot; g, gill; go, gonad; h, heel; id, inner demibranch; isr, inhalant siphon retractor muscle; od, outer demibranch; pa, posterior adductor muscle; t, toe; vm, visceral mass. Scale bars:10 mm in AF; 2 mm in GL." figureDoi="http://doi.org/10.5281/zenodo.10470295" httpUri="https://zenodo.org/record/10470295/files/figure.png" pageId="16" pageNumber="898">Fig. 6E, F, I, L</figureCitation>
):
</emphasis>
Mantle margin fused for most of its length, with a small (anterior) pedal opening and two smaller (posterior) siphonal openings. Suture behind pedal opening extremely short. Anterior and posterior adductor muscles large, ovoid in section, the posterior somewhat dorsally displaced.Gills low (~40% of shell height), elongated, with almost horizontal axis; composed of two complete demibranchs at each side; less and right demibranchs posteriorly fused to each other and to mantle margin. Outer demibranch gradually increasing in height backwards, reaching the maximum size at about two-thirds of its total length. At anterior end, outer demibranch one-third the height of inner demibranch. Ascending and descending lamellae of inner demibranch similar in height throughout their length. Outer demibranch rectangular, anteriorly with ascending lamella higher than descending lamella; similar in size posteriorly. Less and right inner demibranchs ventrally connected by ciliary junctions. Foot medium sized, composed of a compressed stalk and a markedly projected anterior toe. A narrow byssal groove, extending for ~60% of foot sole. Byssus composed of a few, short, flat byssal threads arising from a byssus seam; each byssal thread bearing a distal adhesive disc. Byssal gland remaining functional in adult specimens. Siphons large, fused at the base, distally separated, orange towards the tip in living specimens. Inhalant siphon slightly wider than exhalant siphon; both with numerous series of club-shaped papillae surrounding their openings. Inhalant siphon retractor muscle longer and stronger than exhalant siphon retractor muscle. Anterior and posterior labial palps triangular, similar in size, with ≤ 10 sorting ridges in an 11-mm-long specimen.
</paragraph>
<caption id="DF33663EFFD4FFC3FF59FDB1DB8FF9AF" ID-DOI="http://doi.org/10.5281/zenodo.10470313" ID-Zenodo-Dep="10470313" httpUri="https://zenodo.org/record/10470313/files/figure.png" pageId="17" pageNumber="899" startId="17.[129,194,1494,1518]" targetBox="[209,1393,144,1466]" targetPageId="17" targetType="figure">
<paragraph id="8BF336B6FFD4FFC3FF59FDB1DB8FF9AF" blockId="17.[129,1466,1494,1603]" pageId="17" pageNumber="899">
<emphasis id="B938EAA4FFD4FFC3FF59FDB1D8F6FA02" bold="true" box="[129,229,1494,1518]" pageId="17" pageNumber="899">Figure 13.</emphasis>
Shells of
<taxonomicName id="4C4C4D35FFD4FFC3FEE6FDB1D9D8FA02" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[318,459,1494,1518]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="17" pageNumber="899" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD4FFC3FEE6FDB1D9D8FA02" box="[318,459,1494,1518]" italics="true" pageId="17" pageNumber="899">Hiatella barnea</emphasis>
</taxonomicName>
, inner and ventral views and hinge teeth. A, specimen from the Pacific coast, MACN-In 44278. B, D, F, G, specimens from the Atlantic coast: MACN-In 9175-19 (B), MACN-In 9156-3 (D), MACN-In 9237-8 (F) and MACN-In 44274 (G). C, DNA voucher, H42 (drawn indented lines show the aưachment of the mantle margin to the shell and siphonal muscle scars). E, specimen from Isla de los Estados, MLP-Ma 4942. Scale bars: 5 mm in AD, F; 500 µm in E; 2 mm in G.
</paragraph>
</caption>
<paragraph id="8BF336B6FFD7FFC0FFA9F88ED9DDFED3" blockId="18.[113,762,231,319]" pageId="18" pageNumber="900">
<emphasis id="B938EAA4FFD7FFC0FFA9F88ED8FCFEED" box="[113,239,233,257]" italics="true" pageId="18" pageNumber="900">Distribution:</emphasis>
Along the South American coast, from
<geoCoordinate id="EE785071FFD7FFC0FD5BF88EDAE9FEED" box="[643,762,231,257]" degrees="37" direction="south" minutes="02" orientation="latitude" pageId="18" pageNumber="900" precision="15" seconds="55" value="-37.04861">
37°02
<emphasis id="B938EAA4FFD7FFC0FD67F880DAD6FEEC" box="[703,709,231,256]" italics="true" pageId="18" pageNumber="900">ʹ</emphasis>
55″ S
</geoCoordinate>
in the Pacific to
<geoCoordinate id="EE785071FFD7FFC0FEC1F96FD97CFECC" box="[281,367,262,288]" degrees="41" direction="south" minutes="44" orientation="latitude" pageId="18" pageNumber="900" precision="925" value="-41.733334">
41°44
<emphasis id="B938EAA4FFD7FFC0FE8DF961D948FEF3" box="[341,347,262,287]" italics="true" pageId="18" pageNumber="900">ʹ</emphasis>
S
</geoCoordinate>
in the Atlantic and
<collectingCountry id="F35B7626FFD7FFC0FDE5F96FDA73FECC" box="[573,608,264,288]" name="Iceland" pageId="18" pageNumber="900">Isla</collectingCountry>
de los Estados (
<figureCitation id="13772A33FFD7FFC0FFA4F940D8D5FED3" box="[124,198,295,319]" captionStart="Figure 1" captionStartId="2.[113,178,712,736]" captionTargetBox="[149,1423,147,682]" captionTargetId="figure-646@2.[146,1426,144,684]" captionTargetPageId="2" captionText="Figure 1. Sites in Southern South America (SSA), Prince Edward Islands (PEI) and South Georgia (SG), from where material was studied. A, Hiatella meridionalis. B, Hiatella umbonata. C, Hiatella barnea. Colours represent geographical areas as follows:blue, South Georgia; green, Atlantic coast; light blue, Prince Edward Islands; lilac, Argentine continental margin; orange, Magellan StraitBeagle Channel; purple, Burdwood BankMalvinas/Falkland Islands; red, Pacific coast; yellow, Isla de los Estados." figureDoi="http://doi.org/10.5281/zenodo.10470281" httpUri="https://zenodo.org/record/10470281/files/figure.png" pageId="18" pageNumber="900">Fig. 1C</figureCitation>
). From
<quantity id="4CB49B53FFD7FFC0FEC1F940D994FED3" box="[281,391,295,319]" metricMagnitude="2" metricUnit="m" metricValue="1.545" metricValueMax="3.09" metricValueMin="0.0" pageId="18" pageNumber="900" unit="m" value="154.5" valueMax="309.0" valueMin="0.0">0 to 309 m</quantity>
depth.
</paragraph>
<paragraph id="8BF336B6FFD7FFC0FFA9F903DA05FE90" blockId="18.[113,534,356,380]" box="[113,534,356,380]" pageId="18" pageNumber="900">
<emphasis id="B938EAA4FFD7FFC0FFA9F903D8D5FE90" box="[113,198,356,380]" italics="true" pageId="18" pageNumber="900">Habitat:</emphasis>
Associated with hard substrates.
</paragraph>
<paragraph id="8BF336B6FFD7FFC0FFA9F9C1D925FCB9" blockId="18.[113,763,422,1981]" pageId="18" pageNumber="900">
<emphasis id="B938EAA4FFD7FFC0FFA9F9C1D8C2FE52" box="[113,209,422,446]" italics="true" pageId="18" pageNumber="900">Remarks:</emphasis>
Until this study, and for&gt; 100 years,
<taxonomicName id="4C4C4D35FFD7FFC0FDBEF9C0DADFFE52" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[614,716,422,446]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FDBEF9C0DADFFE52" box="[614,716,422,446]" italics="true" pageId="18" pageNumber="900">H. barnea</emphasis>
</taxonomicName>
had remained known only from its original description, based on a single specimen, inferred to have been collected in Camarones ‘á la base de la formation patagonienne (Tertiary) (
<bibRefCitation id="EFDD4B47FFD7FFC0FD76FA63D8B5FDD7" author="von Ihering HFA" pageId="18" pageNumber="900" pagination="1 - 611" refId="ref19407" refString="von Ihering HFA. Les mollusques fossiles du Tertiare et du Cretace Superieur de l'Argentine. Anales del Museo Nacional de Buenos Aires 1907; 14: 1 - 611." type="journal article" year="1907">Ihering 1907</bibRefCitation>
,
<bibRefCitation id="EFDD4B47FFD7FFC0FF6FFA44D8FFFDD7" author="von Ihering H" box="[183,236,547,571]" pageId="18" pageNumber="900" pagination="1 - 148" refId="ref19371" refString="von Ihering H. Catalogo de moluscos Cretaceos e Terciarios da Argentina da collecao do autor. Notas Preliminares da Revista do Museo Paulista (Sao Paulo) 1914; 1: 1 - 148." type="journal article" year="1914">1914</bibRefCitation>
). The original description is extremely poor in detail and unsatisfactorily figured, and the species was never redescribed. In addition, the
<typeStatus id="54F78814FFD7FFC0FE76FA05D9CEFD96" box="[430,477,610,634]" pageId="18" pageNumber="900">type</typeStatus>
material had not been located subsequently, a fact that has led authors to consider this taxon as a species of questionable status (
<bibRefCitation id="EFDD4B47FFD7FFC0FDEFFAC6DAD7FD55" author="Parodiz JJ" box="[567,708,673,697]" pageId="18" pageNumber="900" pagination="183 - 296" refId="ref18393" refString="Parodiz JJ. The taxa of fossil Mollusca introduced by Hermann von Ihering. Annals of the Carnegie Museum 1996; 65: 183 - 296." type="journal article" year="1996">Parodiz 1996</bibRefCitation>
). To increase the confusion,
<bibRefCitation id="EFDD4B47FFD7FFC0FEB3FAA7DA53FD34" author="von Ihering HFA" box="[363,576,704,728]" pageId="18" pageNumber="900" pagination="1 - 611" refId="ref19407" refString="von Ihering HFA. Les mollusques fossiles du Tertiare et du Cretace Superieur de l'Argentine. Anales del Museo Nacional de Buenos Aires 1907; 14: 1 - 611." type="journal article" year="1907">Ihering (1907: 105)</bibRefCitation>
listed the species as
<taxonomicName id="4C4C4D35FFD7FFC0FF57FA87D920FD1B" box="[143,307,736,759]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="carnea">
<emphasis id="B938EAA4FFD7FFC0FF57FA87D920FD1B" box="[143,307,736,759]" italics="true" pageId="18" pageNumber="900">Saxicava carnea</emphasis>
</taxonomicName>
(indubitably a
<emphasis id="B938EAA4FFD7FFC0FE02FAB8DA71FD1B" box="[474,610,735,759]" italics="true" pageId="18" pageNumber="900">lapsus calami</emphasis>
), and
<bibRefCitation id="EFDD4B47FFD7FFC0FD73FAB8D8E5FCFB" author="Parodiz JJ" pageId="18" pageNumber="900" pagination="183 - 296" refId="ref18393" refString="Parodiz JJ. The taxa of fossil Mollusca introduced by Hermann von Ihering. Annals of the Carnegie Museum 1996; 65: 183 - 296." type="journal article" year="1996">Parodiz (1996: 297)</bibRefCitation>
, in a revision of the taxa introduced by Ihering, referred to it as
<emphasis id="B938EAA4FFD7FFC0FEC8FB78D9A1FCDA" box="[272,434,799,822]" italics="true" pageId="18" pageNumber="900">Barnea saxicava</emphasis>
(instead of
<taxonomicName id="4C4C4D35FFD7FFC0FDF3FB79DADDFCDA" authorityName="Ihering" authorityYear="1907" box="[555,718,798,822]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FDF3FB79DADDFCDA" box="[555,718,798,822]" italics="true" pageId="18" pageNumber="900">Saxicava barnea</emphasis>
</taxonomicName>
, the original binomen).
</paragraph>
<paragraph id="8BF336B6FFD7FFC0FF55FB3AD8D6FA86" blockId="18.[113,763,422,1981]" pageId="18" pageNumber="900">
As part of this study, the
<typeStatus id="54F78814FFD7FFC0FE45FB3BD9EDFC98" box="[413,510,860,884]" pageId="18" pageNumber="900" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4C4D35FFD7FFC0FDF9FB3ADADFFC98" authorityName="Ihering" authorityYear="1907" box="[545,716,860,884]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FDF9FB3ADADFFC98" box="[545,716,860,884]" italics="true" pageId="18" pageNumber="900">Saxicava barnea</emphasis>
</taxonomicName>
was relocated in the palaeoinvertebrate collection of the
<collectionCode id="ED5DAE73FFD7FFC0FD7DFB1BDAE4FC78" box="[677,759,892,916]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="18" pageNumber="900" type="Museum">MACN</collectionCode>
. It consists of an articulated specimen, with the right valve slightly broken dorsally. It is properly figured here, for the first time (
<figureCitation id="13772A33FFD7FFC0FF36FBBDD928FC1E" box="[238,315,986,1010]" captionStart="Figure 11" captionStartId="15.[129,194,1926,1950]" captionTargetBox="[225,1377,1438,1898]" captionTargetId="figure-315@15.[225,1377,1438,1898]" captionTargetPageId="15" captionText="Figure 11. Saxicava barnea. A, original illustration.B, original label. CF, holotype, MACN-Pi N 510, in right (C), less (D), dorsal (E) and ventral (F) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.10470309" httpUri="https://zenodo.org/record/10470309/files/figure.png" pageId="18" pageNumber="900">Fig. 11</figureCitation>
). This specimen is morphologically indistinguishable from present-day specimens collected at the Atlantic and Pacific coasts of Patagonia (
<figureCitation id="13772A33FFD7FFC0FDB4FC7FDAAEFBDD" box="[620,701,1048,1073]" captionStart="Figure 11" captionStartId="15.[129,194,1926,1950]" captionTargetBox="[225,1377,1438,1898]" captionTargetId="figure-315@15.[225,1377,1438,1898]" captionTargetPageId="15" captionText="Figure 11. Saxicava barnea. A, original illustration.B, original label. CF, holotype, MACN-Pi N 510, in right (C), less (D), dorsal (E) and ventral (F) views.Scale bar:5 mm." figureDoi="http://doi.org/10.5281/zenodo.10470309" httpUri="https://zenodo.org/record/10470309/files/figure.png" pageId="18" pageNumber="900">Figs 11</figureCitation>
,
<figureCitation id="13772A33FFD7FFC0FD15FC7EDAFBFBDD" box="[717,744,1049,1073]" captionStart="Figure 12" captionStartId="16.[112,177,1708,1732]" captionTargetBox="[188,1384,144,1680]" captionTargetId="figure-100@16.[188,1384,144,1680]" captionTargetPageId="16" captionText="Figure 12. Shells of Hiatella barnea, external views. A, F, specimens from the Pacific coast, MACN-In 44278. BE, IK, specimens from the Atlantic coast: MACN-In 9175-19 (B, I); MACN-In 9156-3 (C); MACN-In 9032-3 (D, K); MACN-In 9237-8 (E), MACN-In 32620 (J). G, H, DNA vouchers:H43 (G) and H42 (H). Scale bars: 10 mm in AJ; 1 mm in K." figureDoi="http://doi.org/10.5281/zenodo.10470311" httpUri="https://zenodo.org/record/10470311/files/figure.png" pageId="18" pageNumber="900">12</figureCitation>
). Whether
<taxonomicName id="4C4C4D35FFD7FFC0FF01FC5ED952FBBC" authorityName="Ihering" authorityYear="1907" box="[217,321,1080,1104]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FF01FC5ED952FBBC" box="[217,321,1080,1104]" italics="true" pageId="18" pageNumber="900">H. barnea</emphasis>
</taxonomicName>
is a species occurring in Patagonia since the Tertiary or a species from the Present wrongly described from the Tertiary remains uncertain. The exact geological and stratigraphic provenance of the
<typeStatus id="54F78814FFD7FFC0FDD0FCF1DA7AFB42" box="[520,617,1174,1198]" pageId="18" pageNumber="900" type="holotype">holotype</typeStatus>
is unknown. Curiously, a well-preserved system of ºuaternary beach, containing several bivalve species, was described from Camarones by Zanchetta
<emphasis id="B938EAA4FFD7FFC0FE4DFC93D9DBFAE0" box="[405,456,1268,1292]" italics="true" pageId="18" pageNumber="900">et al.</emphasis>
(2012), although
<taxonomicName id="4C4C4D35FFD7FFC0FD4AFC93DAE9FAE0" authorityName="Ihering" authorityYear="1907" box="[658,762,1268,1292]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FD4AFC93DAE9FAE0" box="[658,762,1268,1292]" italics="true" pageId="18" pageNumber="900">H. barnea</emphasis>
</taxonomicName>
was not listed therein. Further palaeontological studies would help to add some precision to the fossil record of this species.
</paragraph>
<paragraph id="8BF336B6FFD7FFC0FF55FD16D934F90D" blockId="18.[113,763,422,1981]" pageId="18" pageNumber="900">
<taxonomicName id="4C4C4D35FFD7FFC0FF55FD16D934FA65" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[141,295,1393,1417]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FF55FD16D934FA65" box="[141,295,1393,1417]" italics="true" pageId="18" pageNumber="900">Hiatella barnea</emphasis>
</taxonomicName>
is an easily recognizable species owing to its elongated and low shell outline, broad, rounded and depressed umbos, wide byssal embayment and prominent commarginal lamellae, which project ventrally into scales. Another distinctive character is the loss of hinge teeth (and the corresponding hinge teeth depressions) through the ontogeny. Anatomically, a peculiar characteristic of this species is the presence of similarly high ascending and descending lamellae of the inner demibranch. The small foot and size of the pedal aperture of this species resemble the characteristics found in
<taxonomicName id="4C4C4D35FFD7FFC0FE52FEEBDA18F94F" authorityName="Zelaya and Marina Güller" authorityYear="2023" box="[394,523,1675,1699]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="umbonata">
<emphasis id="B938EAA4FFD7FFC0FE52FEEBDA18F94F" box="[394,523,1675,1699]" italics="true" pageId="18" pageNumber="900">H. umbonata</emphasis>
</taxonomicName>
, whereas the shape and position of the adductor muscles resemble the condition found in
<taxonomicName id="4C4C4D35FFD7FFC0FF53FEADD932F90E" baseAuthorityName="d'Orbigny" baseAuthorityYear="1846" box="[139,289,1738,1762]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="meridionalis">
<emphasis id="B938EAA4FFD7FFC0FF53FEADD932F90E" box="[139,289,1738,1762]" italics="true" pageId="18" pageNumber="900">H. meridionalis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BF336B6FFD7FFC0FF55FE8ED93CF850" blockId="18.[113,763,422,1981]" pageId="18" pageNumber="900">
Based on the material studied herein,
<taxonomicName id="4C4C4D35FFD7FFC0FDE9FE8DDA8AF8ED" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[561,665,1769,1793]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FDE9FE8DDA8AF8ED" box="[561,665,1769,1793]" italics="true" pageId="18" pageNumber="900">H. barnea</emphasis>
</taxonomicName>
is not a rare species. In fact, it was collected and figured by
<bibRefCitation id="EFDD4B47FFD7FFC0FD45FF6FD8A3F8D3" author="Carcelles AR" pageId="18" pageNumber="900" pagination="155 - 250" refId="ref16576" refString="Carcelles AR. Catalogo de la malacofauna Antartica Argentina. Anales del Museo Nahuel Huapi 1953; 3: 155 - 250." type="journal article" year="1953">Carcelles (1953</bibRefCitation>
: fig. 111) and
<bibRefCitation id="EFDD4B47FFD7FFC0FE93FF4FDA60F8D3" author="Aldea Venegas C &amp; Valdovinos Zarges C" box="[331,627,1831,1856]" pageId="18" pageNumber="900" pagination="364 - 96" refId="ref16040" refString="Aldea Venegas C, Valdovinos Zarges C. Moluscos del intermareal rocoso del centro-sur de Chile (36 ° - 38 ° S): taxonomia y clave de identificacion. Gayana 2005; 69: 364 - 96. hups: // doi. org / 10.4067 / S 0717 - 65382005000200014" type="journal article" year="2005">Aldea and Valdovinos (2005</bibRefCitation>
: fig. 11I), although, in both cases, identified as
<taxonomicName id="4C4C4D35FFD7FFC0FE0EFF2FDA3EF8B3" baseAuthorityName="Sowerby I" baseAuthorityYear="1834" box="[470,557,1863,1887]" class="Bivalvia" family="Hiatellidae" genus="Saxicava" higherTaxonomySource="GBIF" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="solida">
<emphasis id="B938EAA4FFD7FFC0FE0EFF2FDA3EF8B3" box="[470,557,1863,1887]" italics="true" pageId="18" pageNumber="900">H. solida</emphasis>
</taxonomicName>
. Examination of the above-mentioned lots (MACN-In 9032-3 and
<collectionCode id="ED5DAE73FFD7FFC0FD85FF01DAB9F892" box="[605,682,1894,1918]" country="Chile" httpUri="http://grbio.org/cool/70mh-t5r1" name="Museo de Zoologia, Universidad de Concepcion" pageId="18" pageNumber="900">MZUC</collectionCode>
29034, respectively) allows us to confirm that these specimens correspond to
<taxonomicName id="4C4C4D35FFD7FFC0FF1FFFC1D93AF851" authorityName="Zelaya and Marina Güller" authorityYear="2023" baseAuthorityName="Ihering" baseAuthorityYear="1907" box="[199,297,1957,1981]" class="Bivalvia" family="Hiatellidae" genus="Hiatella" kingdom="Animalia" order="Adapedonta" pageId="18" pageNumber="900" phylum="Mollusca" rank="species" species="barnea">
<emphasis id="B938EAA4FFD7FFC0FF1FFFC1D93AF851" box="[199,297,1957,1981]" italics="true" pageId="18" pageNumber="900">H. barnea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>