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<document id="B6BBA88D1F27D27840A675FC953E1586" ID-DOI="10.11646/zootaxa.5415.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10692884" ID-ZooBank="859C14F9-FC21-4076-A212-3B4B6E35E8B7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708603575419" checkinUser="plazi" docAuthor="Mendoza-Penagos, Cristian Camilo, Pérez-Gutiérrez, León A. &amp; Muzón, Javier" docDate="2024" docId="03D88059FF5DFF848FA17C3862D454D8" docLanguage="en" docName="zootaxa.5415.2.1.pdf" docOrigin="Zootaxa 5415 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5415.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lestes helix Ris 1918" docType="treatment" docVersion="2" lastPageNumber="231" masterDocId="FFE1F821FF43FF9A8F367E1B661F5119" masterDocTitle="A revision of Lestidae Calvert, 1901 (Odonata: Zygoptera) from Colombia, with the first record of Lestes dichrostigma Calvert, 1909" masterLastPageNumber="240" masterPageNumber="201" pageNumber="231" updateTime="1708604117699" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6DDA1569C8A7661B3F3D84C5FAE43F0D">A revision of Lestidae Calvert, 1901 (Odonata: Zygoptera) from Colombia, with the first record of Lestes dichrostigma Calvert, 1909</mods:title>
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<mods:namePart id="DDFBA29A24582FAF6B93369BF435242A">Mendoza-Penagos, Cristian Camilo</mods:namePart>
<mods:affiliation id="73A7294A61312BC54C4A2A4B0CC65E97">Laboratório de Ecologia e Conservação-LABECO, Universidade Federal do Pará, Instituto de Ciências Biológicas, Rua Augusto Correia, No. 1 Bairro Guamá, CEP 66.075 - 110 Belém, Pará, Brazil &amp; Laboratorio de Zoología y Ecología Acuática-LAZOEA, Departamentos de Ciencias Biológicas, Universidad de los Andes, Bogotá 111711, Colombia</mods:affiliation>
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<mods:namePart id="8D38B78ACF7F0EA83FF7327BC291FD9F">Pérez-Gutiérrez, León A.</mods:namePart>
<mods:affiliation id="29914B17E75FADE036111054D10EBE6E">Laboratório de Sistemática de Insetos Aquáticos-LABSIA, Universidade Federal do Paraná, Jardim das Américas, Curitiba, Brasi</mods:affiliation>
<mods:nameIdentifier id="DEBE1AD28565A977A6AFC757DDA45FB2" type="email">talysker@gmail.com</mods:nameIdentifier>
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<mods:namePart id="123FB04B7AC37B9F2A2A5BF3A67646F9">Muzón, Javier</mods:namePart>
<mods:affiliation id="74E96094A402DAEF184A1820D00A23A5">Laboratório de Ecologia e Conservação-LABECO, Universidade Federal do Pará, Instituto de Ciências Biológicas, Rua Augusto Correia, No. 1 Bairro Guamá, CEP 66.075 - 110 Belém, Pará, Brazil &amp; Laboratório de Ecologia e Conservação-LABECO, Universidade Federal do Pará, Instituto de Ciências Biológicas, Rua Augusto Correia, No. 1 Bairro Guamá, CEP 66.075 - 110 Belém, Pará, Brazil</mods:affiliation>
<mods:nameIdentifier id="8E39F701670D238CE50C6AB493A07E6D" type="email">jmuzon@gmail.com</mods:nameIdentifier>
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<mods:date id="CB26AAAE130DB7704DB391DF474297CC">2024</mods:date>
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<treatment id="03D88059FF5DFF848FA17C3862D454D8" ID-DOI="http://doi.org/10.5281/zenodo.10692924" ID-Zenodo-Dep="10692924" LSID="urn:lsid:plazi:treatment:03D88059FF5DFF848FA17C3862D454D8" httpUri="http://treatment.plazi.org/id/03D88059FF5DFF848FA17C3862D454D8" lastPageNumber="231" pageId="30" pageNumber="231">
<subSubSection id="C36B62C4FF5DFF848FA17C3867805325" box="[151,415,546,573]" pageId="30" pageNumber="231" type="nomenclature">
<paragraph id="8BCE314FFF5DFF848FA17C3867805325" blockId="30.[151,1437,546,645]" box="[151,415,546,573]" pageId="30" pageNumber="231">
<heading id="D0868623FF5DFF848FA17C3867805325" bold="true" box="[151,415,546,573]" fontSize="11" level="1" pageId="30" pageNumber="231" reason="1">
<taxonomicName id="4C714ACCFF5DFF848FA17C3867805325" ID-CoL="6Q346" authority="Ris, 1918" authorityName="Ris" authorityYear="1918" box="[151,415,546,573]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="helix">
<emphasis id="B905ED5DFF5DFF848FA17C3867805325" bold="true" box="[151,415,546,573]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FA17C38673A5324" bold="true" box="[151,293,546,573]" italics="true" pageId="30" pageNumber="231">Lestes helix</emphasis>
<bibRefCitation id="EFE04CBEFF5DFF848E1B7C3967805325" author="Ris, F." box="[301,415,546,572]" pageId="30" pageNumber="231" pagination="1 - 197" refId="ref21258" refString="Ris, F. (1918) Libellen (Odonata) aus der Regimder amerikanischen Kordilleren von Costaricibis Catamarca. Archiv far Naturgeschichte A, (9), 1 - 197." type="journal article" year="1918">Ris, 1918</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FA17C5C6749539C" pageId="30" pageNumber="231" type="description">
<paragraph id="8BCE314FFF5DFF848FA17C5C6749539C" blockId="30.[151,1437,546,645]" pageId="30" pageNumber="231">
<figureCitation id="134A2DCAFF5DFF848FA17C5C66F35378" box="[151,236,582,609]" captionStart="FIGURE 5" captionStartId="7.[152,255,1378,1402]" captionTargetBox="[239,1354,201,1335]" captionTargetId="figure-19@7.[227,1361,181,1353]" captionTargetPageId="7" captionText="FIGURE 5. Head pattern coloration. Lestes forficula: (A) male, (B) female; L. minutus: (C) male, (D) female; L. tenuatus: (E) male, (F) female; L. helix: (G) male; L. jerrelli: (H) male." figureDoi="http://doi.org/10.5281/zenodo.10692902" httpUri="https://zenodo.org/record/10692902/files/figure.png" pageId="30" pageNumber="231">Fig. 5G</figureCitation>
(♁ head),
<figureCitation id="134A2DCAFF5DFF848E527C5C67C45378" box="[356,475,583,609]" captionStart="FIGURE 8" captionStartId="10.[152,255,1891,1915]" captionTargetBox="[287,1289,183,1865]" captionTargetId="figure-19@10.[287,1300,181,1866]" captionTargetPageId="10" captionText="FIGURE 8. Male pterothorax pattern coloration. Lestes helix: (A) dorsal view, (B) lateral view (C) ventral view; L. jerrelli: (D) dorsal view, (E) lateral view (F) ventral view; L. minutus mature male: (G) dorsal view, (H) lateral view (I) ventral view; L. minutus immature male: (G’) dorsal view, (H’) lateral view (I’) ventral view; L. tenuatus: (J) dorsal view, (K) lateral view (L) ventral view." figureDoi="http://doi.org/10.5281/zenodo.10692912" httpUri="https://zenodo.org/record/10692912/files/figure.png" pageId="30" pageNumber="231">Fig. 8A–C</figureCitation>
(♁ pterothorax),
<figureCitation id="134A2DCAFF5DFF848DA97C5C65045378" box="[671,795,583,609]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[238,1338,184,1882]" captionTargetId="figure-19@11.[237,1350,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Shape of genital ligula. Archilestes chocoanus: (A) ectal view, (B) lateral view; A. grandis: (C) ectal view, (D) lateral view; Lestes apollinaris: (E) ectal view, (F) lateral view; L. curvatus: (G) ectal view, (H) lateral view; L. dicrhostigma: (I) ectal view, (J) lateral view; L. forficula: (K) dorsal view, (L) lateral view; Lestes helix: (M) ectal view, (N) lateral view; L. minutus: (O) ectal view, (P) lateral view; L. tenuatus: (Q) ectal view, (R) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692914" httpUri="https://zenodo.org/record/10692914/files/figure.png" pageId="30" pageNumber="231">Fig. 9M–N</figureCitation>
(♁ genital ligula),
<figureCitation id="134A2DCAFF5DFF848CC77C5C62455378" box="[1009,1114,583,609]" captionStart="FIGURE 11" captionStartId="13.[152,255,1908,1932]" captionTargetBox="[189,1396,188,1883]" captionTargetId="figure-19@13.[189,1398,181,1885]" captionTargetPageId="13" captionText="FIGURE 11. Male cerci. Lestes apollinaris: (A) dorsal view, (B) lateral view; L. curvatus: (C) dorsal view, (D) lateral view; L. dicrhostigma: (E) dorsal view, (F) lateral view; L. forficula: (G) dorsal view, (H) lateral view; Lestes helix: (I) dorsal view, (J) lateral view; L. jerrelli: (K) dorsal view, (L) lateral view; L. minutus: (M) dorsal view, (N) lateral view; L. tenuatus: (O) dorsal view, (P) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692922" httpUri="https://zenodo.org/record/10692922/files/figure.png" pageId="30" pageNumber="231">Fig.11I–J</figureCitation>
(♁ caudal appendages),
<figureCitation id="134A2DCAFF5DFF848A467C5C66AC539C" captionStart="FIGURE 16" captionStartId="19.[152,255,2000,2024]" captionTargetBox="[204,1393,237,1968]" captionTargetId="figure-28@19.[194,1393,237,1976]" captionTargetPageId="19" captionText="FIGURE 16. Distribution of Lestes dichrostigma and L. helix." figureDoi="http://doi.org/10.5281/zenodo.10692936" httpUri="https://zenodo.org/record/10692936/files/figure.png" pageId="30" pageNumber="231">Fig. 16</figureCitation>
(distribution).
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FA17CA862F353D4" box="[151,1260,690,719]" pageId="30" pageNumber="231" type="etymology">
<paragraph id="8BCE314FFF5DFF848FA17CA862F353D4" blockId="30.[151,1437,690,1473]" box="[151,1260,690,719]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FA17CA8673D53D4" bold="true" box="[151,290,691,717]" pageId="30" pageNumber="231">Etymology:</emphasis>
<taxonomicName id="4C714ACCFF5DFF848E1F7CA967AB53D4" authority=", Greek" authorityName="Greek" box="[297,436,690,717]" class="Insecta" family="Lestidae" genus="Lestes" kingdom="Animalia" order="Odonata" pageId="30" pageNumber="231" phylum="Arthropoda" rank="species" species="helix">
<emphasis id="B905ED5DFF5DFF848E1F7CA9677C53D4" bold="true" box="[297,355,690,717]" italics="true" pageId="30" pageNumber="231">helix</emphasis>
, Greek
</taxonomicName>
(ελιξ) spiral, round, curved in reference to the curved apex of male cerci.
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FF17CCC640753E8" box="[199,536,726,753]" pageId="30" pageNumber="231" type="materials_examined">
<paragraph id="8BCE314FFF5DFF848FF17CCC640753E8" blockId="30.[151,1437,690,1473]" box="[199,536,726,753]" pageId="30" pageNumber="231">
<materialsCitation id="3B193B12FF5DFF848FF17CCC640B53E8" box="[199,532,726,753]" country="Peru" location="Pachitea" pageId="30" pageNumber="231" specimenCount="1" typeStatus="holotype">
<emphasis id="B905ED5DFF5DFF848FF17CCC677653E9" bold="true" box="[199,361,726,753]" pageId="30" pageNumber="231">
<typeStatus id="54CA8FEDFF5DFF848FF17CCC671D53E8" box="[199,258,727,753]" pageId="30" pageNumber="231">Type</typeStatus>
locality:
</emphasis>
<location id="8EAE6794FF5DFF848E467CCC67CD53E8" LSID="urn:lsid:plazi:treatment:03D88059FF5DFF848FA17C3862D454D8:8EAE6794FF5DFF848E467CCC67CD53E8" box="[368,466,727,753]" country="Peru" name="Pachitea" pageId="30" pageNumber="231">Pachitea</location>
,
<collectingCountry id="F36671DFFF5DFF848EEB7CCC640B53E8" box="[477,532,727,753]" name="Peru" pageId="30" pageNumber="231">Peru</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FF17CE166FD52BC" pageId="30" pageNumber="231" type="distribution">
<paragraph id="8BCE314FFF5DFF848FF17CE162A6520C" blockId="30.[151,1437,690,1473]" box="[199,1209,762,789]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17CE167C0520D" bold="true" box="[199,479,762,788]" pageId="30" pageNumber="231">
Distribution:
<collectingCountry id="F36671DFFF5DFF848E5C7CE167C0520D" box="[362,479,762,788]" name="Colombia" pageId="30" pageNumber="231">Colombia</collectingCountry>
</emphasis>
(Amazonas,
<collectingRegion id="49B5FFADFF5DFF848D457CE064D0520C" box="[627,719,763,789]" country="Colombia" name="Guainia" pageId="30" pageNumber="231">Guainía</collectingRegion>
,
<collectingRegion id="49B5FFADFF5DFF848DEC7CE0655C520C" box="[730,835,763,789]" country="Colombia" name="Guaviare" pageId="30" pageNumber="231">Guaviare</collectingRegion>
,
<collectingRegion id="49B5FFADFF5DFF848C787CE06595520C" box="[846,906,763,789]" country="Colombia" name="Meta" pageId="30" pageNumber="231">Meta</collectingRegion>
,
<collectingRegion id="49B5FFADFF5DFF848CA37CE065F4520C" box="[917,1003,763,789]" country="Colombia" name="Vaupes" pageId="30" pageNumber="231">Vaupés</collectingRegion>
), Peru, Venezuela.
</paragraph>
<paragraph id="8BCE314FFF5DFF848FF17D0464D85220" blockId="30.[151,1437,690,1473]" box="[199,711,798,825]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17D0467BD5221" bold="true" box="[199,418,798,825]" pageId="30" pageNumber="231">Female described:</emphasis>
No
<emphasis id="B905ED5DFF5DFF848EE47D0464815221" bold="true" box="[466,670,798,825]" pageId="30" pageNumber="231">Larva described:</emphasis>
No
</paragraph>
<paragraph id="8BCE314FFF5DFF848FF17D5962115244" blockId="30.[151,1437,690,1473]" box="[199,1038,834,861]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17D5967805244" bold="true" box="[199,415,834,861]" pageId="30" pageNumber="231">Altitudinal range:</emphasis>
<quantity id="4C899CAAFF5DFF848E907D5964085244" box="[422,535,834,861]" metricMagnitude="2" metricUnit="m" metricValue="2.915" metricValueMax="4.95" metricValueMin="0.88" pageId="30" pageNumber="231" unit="m" value="291.5" valueMax="495.0" valueMin="88.0">88–495 m</quantity>
<emphasis id="B905ED5DFF5DFF848D2B7D5964A45244" bold="true" box="[541,699,834,861]" pageId="30" pageNumber="231">IUCN status:</emphasis>
LC—
<bibRefCitation id="EFE04CBEFF5DFF848C357D5862115244" author="von Ellenrieder, N." box="[771,1038,835,861]" pageId="30" pageNumber="231" refId="ref21850" refString="https: // doi. org / 10.2305 / IUCN. UK. 2009 - 2. RLTS. T 158981 A 5300346. en von Ellenrieder, N. (2009 c) Lestes helix. The IUCN Red List of Threatened Species, 2009, e. T 158971 A 5299010." type="book" year="2009">von Ellenrieder (2009c)</bibRefCitation>
</paragraph>
<paragraph id="8BCE314FFF5DFF848FF17D7C66FD52BC" blockId="30.[151,1437,690,1473]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17D7C64515299" bold="true" box="[199,590,870,897]" pageId="30" pageNumber="231">Records in previous references:</emphasis>
<bibRefCitation id="EFE04CBEFF5DFF848D6C7D7C655F5298" author="Bota-Sierra, C. A." box="[602,832,871,897]" pageId="30" pageNumber="231" pagination="97 - 100" refId="ref18105" refString="Bota-Sierra, C. A. (2014 a) Nine new records of Odonata for Colombia from the Orinoco Basin (Lestidae, Calopterygidae, Heteragrionidae, Coenagrionidae, Libellulidae). Notulae Odonatologicae, 8 (4), 97 - 100." type="journal article" year="2014">Bota-Sierra (2014a)</bibRefCitation>
,
<bibRefCitation id="EFE04CBEFF5DFF848C677D7C626B5298" author="Bota-Sierra, C. A. &amp; Moreno-Arias, C. &amp; Faasen, T." box="[849,1140,870,897]" pageId="30" pageNumber="231" pagination="249 - 268" refId="ref18223" refString="Bota-Sierra, C. A., Moreno-Arias, C. &amp; Faasen, T. (2015) Preliminary list of Odonata from the Colombian Amazon, with descriptions of Inpabasis nigridorsum sp. nov. &amp; Diaphlebia richteri sp. nov. (Coenagrionidae &amp; Gomphidae). International Journal of Odonatology, 18 (3), 249 - 268. https: // doi. org / 10.1080 / 13887890.2015.1081637" type="journal article" year="2015">
Bota-Sierra
<emphasis id="B905ED5DFF5DFF848CE87D7C62045298" box="[990,1051,871,897]" italics="true" pageId="30" pageNumber="231">et al.</emphasis>
(2015)
</bibRefCitation>
,
<bibRefCitation id="EFE04CBEFF5DFF848BB57D7C66FD52BC" author="Aristizabal-Botero, A. &amp; Araque-Orjuela, I. &amp; Cano-Cobos, Y. &amp; Realpe, E." pageId="30" pageNumber="231" pagination="116 - 123" refId="ref17927" refString="Aristizabal-Botero, A., Araque-Orjuela, I., Cano-Cobos, Y. &amp; Realpe, E. (2022) Libelulas: Una conexion entre el agua y la tierra. In: Usma Oviedo, J. S., Trujillo, F. &amp; Naranjo, L. G. (Eds.), Diversidad biologica y cultural del departamento de Guainia. Gobernacion de Guainia, WWF Colombia, Corporacion para el Desarrollo Sostenible del Norte y el Oriente Amazonico- CDA &amp; Instituto Amazonico de Investigaciones Cientificas SINCHI, Bogota, Colombia, pp. 116 - 123." type="book chapter" year="2022">
Aristizabal-Botero
<emphasis id="B905ED5DFF5DFF848A697D7C63825298" box="[1375,1437,871,897]" italics="true" pageId="30" pageNumber="231">et al.</emphasis>
(2022)
</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FF17DB564C9544C" pageId="30" pageNumber="231" type="materials_examined">
<paragraph id="8BCE314FFF5DFF848FF17DB5644C552C" blockId="30.[151,1437,690,1473]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17DB5642952D5" bold="true" box="[199,566,942,972]" pageId="30" pageNumber="231">
Material examined: (2♁♁,
<specimenCount id="9D77FAC6FF5DFF848D3F7DAB643352D5" box="[521,556,944,972]" count="1" pageId="30" pageNumber="231" type="female">1♀</specimenCount>
)
</emphasis>
.
<materialsCitation id="3B193B12FF5DFF848D727DB4628152F4" collectingDate="2014-07-30" collectorName="A. Aristizabal &amp; E. Realpe &amp; J. Sierra &amp; P. Sarmiento" country="Colombia" elevation="83" latitude="3.9075" location="Coco Viejo" longLatPrecision="7" longitude="-67.9089" municipality="Inirida" pageId="30" pageNumber="231" specimenCode="ANDES-E 21579" specimenCount="1" stateProvince="Guainia">
<collectingCountry id="F36671DFFF5DFF848D727DB464AA52D0" box="[580,693,943,969]" name="Colombia" pageId="30" pageNumber="231">Colombia</collectingCountry>
,
<collectingRegion id="49B5FFADFF5DFF848DF67DB4650352D0" box="[704,796,943,969]" country="Colombia" name="Guainia" pageId="30" pageNumber="231">Guainía</collectingRegion>
,
<emphasis id="B905ED5DFF5DFF848C117DB4629652D1" box="[807,1161,943,969]" italics="true" pageId="30" pageNumber="231">
<collectingMunicipality id="6BAAAB35FF5DFF848C117DB4656752D0" box="[807,888,943,969]" pageId="30" pageNumber="231">Inírida</collectingMunicipality>
, comunidad
<location id="8EAE6794FF5DFF848B317DB4629A52D1" LSID="urn:lsid:plazi:treatment:03D88059FF5DFF848FA17C3862D454D8:8EAE6794FF5DFF848B317DB4629A52D1" box="[1031,1157,943,969]" country="Colombia" latitude="3.9075" longLatPrecision="7" longitude="-67.9089" municipality="Inirida" name="Coco Viejo" pageId="30" pageNumber="231" stateProvince="Guainia">Coco Viejo</location>
,
</emphasis>
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,
<geoCoordinate id="EE455788FF5DFF848BC97DB4636A52D0" box="[1279,1397,943,969]" degrees="67.9089" direction="west" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="-67.9089">67.9089W</geoCoordinate>
,
<quantity id="4C899CAAFF5DFF848AB67DB466B052F4" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="30" pageNumber="231" unit="m" value="83.0">
<elevation id="005CD67CFF5DFF848AB67DB466B052F4" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="30" pageNumber="231" unit="m" value="83.0">83 m</elevation>
</quantity>
,
<date id="FFCF178FFF5DFF848F8D7DC8672D52F4" box="[187,306,979,1005]" pageId="30" pageNumber="231" value="2014-07-30">
<collectingDate id="EF8BEE67FF5DFF848F8D7DC8672D52F4" box="[187,306,979,1005]" pageId="30" pageNumber="231" value="2014-07-30">30-7-2014</collectingDate>
</date>
,
<collectorName id="26845499FF5DFF848E0A7DC867C452F4" box="[316,475,979,1005]" pageId="30" pageNumber="231">A. Aristizabal</collectorName>
,
<collectorName id="26845499FF5DFF848ED07DC8644852F4" box="[486,599,979,1005]" pageId="30" pageNumber="231">E. Realpe</collectorName>
,
<collectorName id="26845499FF5DFF848D577DC864DE52F4" box="[609,705,979,1005]" pageId="30" pageNumber="231">J. Sierra</collectorName>
,
<collectorName id="26845499FF5DFF848DFA7DC8654652F4" box="[716,857,979,1005]" pageId="30" pageNumber="231">P. Sarmiento</collectorName>
leg.,
<emphasis id="B905ED5DFF5DFF848CAE7DCF65A452E9" bold="true" box="[920,955,980,1008]" pageId="30" pageNumber="231">1♁</emphasis>
(
<emphasis id="B905ED5DFF5DFF848CFD7DC9628C52F5" bold="true" box="[971,1171,978,1004]" pageId="30" pageNumber="231">
<specimenCode id="DBD79934FF5DFF848CFD7DC9628C52F5" box="[971,1171,978,1004]" collectionCode="ANDES-E" pageId="30" pageNumber="231">ANDES-E 21579</specimenCode>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B193B12FF5DFF848B9C7DC8651B5508" collectionCode="ICN" country="Colombia" county="Amazonas" elevation="103" latitude="1.0471" location="rio Sorotama" longLatPrecision="7" longitude="-70.2241" municipality="Apaporis" pageId="30" pageNumber="231" specimenCount="1" specimenCount-female="1" stateProvince="Guainia">
<collectingCounty id="62AF49C3FF5DFF848B9C7DC8633C52F4" box="[1194,1315,979,1005]" pageId="30" pageNumber="231">Amazonas</collectingCounty>
,
<emphasis id="B905ED5DFF5DFF848A197DC8672B5508" italics="true" pageId="30" pageNumber="231">
<collectingMunicipality id="6BAAAB35FF5DFF848A197DC8638652F4" box="[1327,1433,979,1005]" pageId="30" pageNumber="231">Apaporis</collectingMunicipality>
,
<location id="8EAE6794FF5DFF848FA17DEC672E5508" LSID="urn:lsid:plazi:treatment:03D88059FF5DFF848FA17C3862D454D8:8EAE6794FF5DFF848FA17DEC672E5508" box="[151,305,1015,1041]" country="Colombia" county="Amazonas" latitude="1.0471" longLatPrecision="7" longitude="-70.2241" municipality="Apaporis" name="rio Sorotama" pageId="30" pageNumber="231" stateProvince="Guainia">río Sorotama</location>
,
</emphasis>
<geoCoordinate id="EE455788FF5DFF848E097DEC67BC5508" box="[319,419,1015,1041]" degrees="1.0471" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="1.0471">1.0471N</geoCoordinate>
,
<geoCoordinate id="EE455788FF5DFF848E877DEC64395508" box="[433,550,1015,1041]" degrees="70.2241" direction="west" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="-70.2241">70.2241W</geoCoordinate>
,
<quantity id="4C899CAAFF5DFF848D027DEC649F5508" box="[564,640,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="30" pageNumber="231" unit="m" value="103.0">
<elevation id="005CD67CFF5DFF848D027DEC649F5508" box="[564,640,1015,1041]" metricMagnitude="2" metricUnit="m" metricValue="1.03" pageId="30" pageNumber="231" unit="m" value="103.0">103 m</elevation>
</quantity>
,
<emphasis id="B905ED5DFF5DFF848DB97DE364AD550D" bold="true" box="[655,690,1016,1044]" pageId="30" pageNumber="231">
<specimenCount id="9D77FAC6FF5DFF848DB97DE364AD550D" box="[655,690,1016,1044]" count="1" pageId="30" pageNumber="231" type="female">1♀</specimenCount>
</emphasis>
(
<emphasis id="B905ED5DFF5DFF848DF37DED64E75509" bold="true" box="[709,760,1014,1040]" pageId="30" pageNumber="231">
<collectionCode id="ED60A98AFF5DFF848DF37DED64E75509" box="[709,760,1014,1040]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="30" pageNumber="231">ICN</collectionCode>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="3B193B12FF5DFF848C227DEC6451552C" collectionCode="ICN" country="Colombia" county="Reserva" elevation="153" latitude="2.1771" location="Cerro Moyano" longLatPrecision="7" longitude="-71.1901" municipality="Nukak Maku" pageId="30" pageNumber="231" specimenCount="1" stateProvince="Guaviare">
<collectingRegion id="49B5FFADFF5DFF848C227DEC65625508" box="[788,893,1015,1041]" country="Colombia" name="Guaviare" pageId="30" pageNumber="231">Guaviare</collectingRegion>
,
<emphasis id="B905ED5DFF5DFF848CBD7DEC62C25508" box="[907,1245,1015,1041]" italics="true" pageId="30" pageNumber="231">
<collectingCounty id="62AF49C3FF5DFF848CBD7DEC65FA5509" box="[907,997,1015,1040]" pageId="30" pageNumber="231">Reserva</collectingCounty>
natural
<collectingMunicipality id="6BAAAB35FF5DFF848B7A7DEC62C25508" box="[1100,1245,1015,1041]" pageId="30" pageNumber="231">Nukak Maku</collectingMunicipality>
</emphasis>
,
<emphasis id="B905ED5DFF5DFF848BD87DEC638A5509" box="[1262,1429,1015,1041]" italics="true" pageId="30" pageNumber="231">
<location id="8EAE6794FF5DFF848BD87DEC638A5509" LSID="urn:lsid:plazi:treatment:03D88059FF5DFF848FA17C3862D454D8:8EAE6794FF5DFF848BD87DEC638A5509" box="[1262,1429,1015,1041]" country="Colombia" county="Reserva" latitude="2.1771" longLatPrecision="7" longitude="-71.1901" municipality="Nukak Maku" name="Cerro Moyano" pageId="30" pageNumber="231" stateProvince="Guaviare">Cerro Moyano</location>
</emphasis>
,
<geoCoordinate id="EE455788FF5DFF848FA17A0066E4552C" box="[151,251,1051,1077]" degrees="2.1771" direction="north" orientation="latitude" pageId="30" pageNumber="231" precision="5" value="2.1771">2.1771N</geoCoordinate>
,
<geoCoordinate id="EE455788FF5DFF848E307A006763552C" box="[262,380,1051,1077]" degrees="71.1901" direction="west" orientation="longitude" pageId="30" pageNumber="231" precision="5" value="-71.1901">71.1901W</geoCoordinate>
,
<quantity id="4C899CAAFF5DFF848EB17A0167CF552C" box="[391,464,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="30" pageNumber="231" unit="m" value="153.0">
<elevation id="005CD67CFF5DFF848EB17A0167CF552C" box="[391,464,1050,1077]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="30" pageNumber="231" unit="m" value="153.0">153 m</elevation>
</quantity>
,
<emphasis id="B905ED5DFF5DFF848EEA7A0767E05521" bold="true" box="[476,511,1052,1080]" pageId="30" pageNumber="231">1♁</emphasis>
(
<emphasis id="B905ED5DFF5DFF848D397A01645D552D" bold="true" box="[527,578,1050,1076]" pageId="30" pageNumber="231">
<collectionCode id="ED60A98AFF5DFF848D397A01645D552D" box="[527,578,1050,1076]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="30" pageNumber="231">ICN</collectionCode>
</emphasis>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCE314FFF5DFF848FF17A2564C9544C" blockId="30.[151,1437,690,1473]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17A2564995541" bold="true" box="[199,646,1086,1112]" pageId="30" pageNumber="231">Morphological characteristics (male):</emphasis>
head mostly dark brown except posterior margin, labrum, clypeus, and mandibles pale blue (
<figureCitation id="134A2DCAFF5DFF848EB57A7867FA5564" box="[387,485,1122,1149]" captionStart="FIGURE 5" captionStartId="7.[152,255,1378,1402]" captionTargetBox="[239,1354,201,1335]" captionTargetId="figure-19@7.[227,1361,181,1353]" captionTargetPageId="7" captionText="FIGURE 5. Head pattern coloration. Lestes forficula: (A) male, (B) female; L. minutus: (C) male, (D) female; L. tenuatus: (E) male, (F) female; L. helix: (G) male; L. jerrelli: (H) male." figureDoi="http://doi.org/10.5281/zenodo.10692902" httpUri="https://zenodo.org/record/10692902/files/figure.png" pageId="30" pageNumber="231">Figs. 5G</figureCitation>
,
<figureCitation id="134A2DCAFF5DFF848EC67A7864285564" box="[496,567,1123,1149]" captionStart="FIGURE 8" captionStartId="10.[152,255,1891,1915]" captionTargetBox="[287,1289,183,1865]" captionTargetId="figure-19@10.[287,1300,181,1866]" captionTargetPageId="10" captionText="FIGURE 8. Male pterothorax pattern coloration. Lestes helix: (A) dorsal view, (B) lateral view (C) ventral view; L. jerrelli: (D) dorsal view, (E) lateral view (F) ventral view; L. minutus mature male: (G) dorsal view, (H) lateral view (I) ventral view; L. minutus immature male: (G’) dorsal view, (H’) lateral view (I’) ventral view; L. tenuatus: (J) dorsal view, (K) lateral view (L) ventral view." figureDoi="http://doi.org/10.5281/zenodo.10692912" httpUri="https://zenodo.org/record/10692912/files/figure.png" pageId="30" pageNumber="231">8A–C</figureCitation>
); pterothorax dark brown, black antehumeral stripe with metallic luster; lateral black stripe occupying dorsal 0.5 of mesepimeron and mesinfraepisternum; (
<figureCitation id="134A2DCAFF5DFF848CDE7A9C627D55B8" box="[1000,1122,1159,1185]" captionStart="FIGURE 8" captionStartId="10.[152,255,1891,1915]" captionTargetBox="[287,1289,183,1865]" captionTargetId="figure-19@10.[287,1300,181,1866]" captionTargetPageId="10" captionText="FIGURE 8. Male pterothorax pattern coloration. Lestes helix: (A) dorsal view, (B) lateral view (C) ventral view; L. jerrelli: (D) dorsal view, (E) lateral view (F) ventral view; L. minutus mature male: (G) dorsal view, (H) lateral view (I) ventral view; L. minutus immature male: (G’) dorsal view, (H’) lateral view (I’) ventral view; L. tenuatus: (J) dorsal view, (K) lateral view (L) ventral view." figureDoi="http://doi.org/10.5281/zenodo.10692912" httpUri="https://zenodo.org/record/10692912/files/figure.png" pageId="30" pageNumber="231">Fig. 8A–B</figureCitation>
); pectum pale, metepimeron with two large subtriangular black spots reaching metapleural suture and two smaller at the end of the metathoracic suture (
<figureCitation id="134A2DCAFF5DFF848FD97AD4675455F0" box="[239,331,1231,1257]" captionStart="FIGURE 8" captionStartId="10.[152,255,1891,1915]" captionTargetBox="[287,1289,183,1865]" captionTargetId="figure-19@10.[287,1300,181,1866]" captionTargetPageId="10" captionText="FIGURE 8. Male pterothorax pattern coloration. Lestes helix: (A) dorsal view, (B) lateral view (C) ventral view; L. jerrelli: (D) dorsal view, (E) lateral view (F) ventral view; L. minutus mature male: (G) dorsal view, (H) lateral view (I) ventral view; L. minutus immature male: (G’) dorsal view, (H’) lateral view (I’) ventral view; L. tenuatus: (J) dorsal view, (K) lateral view (L) ventral view." figureDoi="http://doi.org/10.5281/zenodo.10692912" httpUri="https://zenodo.org/record/10692912/files/figure.png" pageId="30" pageNumber="231">Fig. 8C</figureCitation>
). Genital ligula rectangular, internal fold width more than 1/3 of S2 (
<figureCitation id="134A2DCAFF5DFF848B497AD462FD55F0" box="[1151,1250,1231,1257]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[238,1338,184,1882]" captionTargetId="figure-19@11.[237,1350,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Shape of genital ligula. Archilestes chocoanus: (A) ectal view, (B) lateral view; A. grandis: (C) ectal view, (D) lateral view; Lestes apollinaris: (E) ectal view, (F) lateral view; L. curvatus: (G) ectal view, (H) lateral view; L. dicrhostigma: (I) ectal view, (J) lateral view; L. forficula: (K) dorsal view, (L) lateral view; Lestes helix: (M) ectal view, (N) lateral view; L. minutus: (O) ectal view, (P) lateral view; L. tenuatus: (Q) ectal view, (R) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692914" httpUri="https://zenodo.org/record/10692914/files/figure.png" pageId="30" pageNumber="231">Fig. 9M</figureCitation>
), surface entire, reaching anterior fold of S2 (
<figureCitation id="134A2DCAFF5DFF848ED17AE8645B5414" box="[487,580,1267,1293]" captionStart="FIGURE 9" captionStartId="11.[152,255,1908,1932]" captionTargetBox="[238,1338,184,1882]" captionTargetId="figure-19@11.[237,1350,181,1884]" captionTargetPageId="11" captionText="FIGURE 9. Shape of genital ligula. Archilestes chocoanus: (A) ectal view, (B) lateral view; A. grandis: (C) ectal view, (D) lateral view; Lestes apollinaris: (E) ectal view, (F) lateral view; L. curvatus: (G) ectal view, (H) lateral view; L. dicrhostigma: (I) ectal view, (J) lateral view; L. forficula: (K) dorsal view, (L) lateral view; Lestes helix: (M) ectal view, (N) lateral view; L. minutus: (O) ectal view, (P) lateral view; L. tenuatus: (Q) ectal view, (R) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692914" httpUri="https://zenodo.org/record/10692914/files/figure.png" pageId="30" pageNumber="231">Fig. 9N</figureCitation>
). Cerci long, with a bicuspid lobe in basal portion; edge of medial portion irregular; primary curvature
<typeStatus id="54CA8FEDFF5DFF848EEB7B0C64125428" box="[477,525,1303,1329]" pageId="30" pageNumber="231">type</typeStatus>
II (
<figureCitation id="134A2DCAFF5DFF848D0D7B0C64875428" box="[571,664,1303,1329]" captionStart="FIGURE 11" captionStartId="13.[152,255,1908,1932]" captionTargetBox="[189,1396,188,1883]" captionTargetId="figure-19@13.[189,1398,181,1885]" captionTargetPageId="13" captionText="FIGURE 11. Male cerci. Lestes apollinaris: (A) dorsal view, (B) lateral view; L. curvatus: (C) dorsal view, (D) lateral view; L. dicrhostigma: (E) dorsal view, (F) lateral view; L. forficula: (G) dorsal view, (H) lateral view; Lestes helix: (I) dorsal view, (J) lateral view; L. jerrelli: (K) dorsal view, (L) lateral view; L. minutus: (M) dorsal view, (N) lateral view; L. tenuatus: (O) dorsal view, (P) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692922" httpUri="https://zenodo.org/record/10692922/files/figure.png" pageId="30" pageNumber="231">Fig. 11I</figureCitation>
); secondary curvature
<typeStatus id="54CA8FEDFF5DFF848CAB7B0C65D25428" box="[925,973,1303,1329]" pageId="30" pageNumber="231">type</typeStatus>
III (
<figureCitation id="134A2DCAFF5DFF848B327B0C62715428" box="[1028,1134,1303,1329]" captionStart="FIGURE 11" captionStartId="13.[152,255,1908,1932]" captionTargetBox="[189,1396,188,1883]" captionTargetId="figure-19@13.[189,1398,181,1885]" captionTargetPageId="13" captionText="FIGURE 11. Male cerci. Lestes apollinaris: (A) dorsal view, (B) lateral view; L. curvatus: (C) dorsal view, (D) lateral view; L. dicrhostigma: (E) dorsal view, (F) lateral view; L. forficula: (G) dorsal view, (H) lateral view; Lestes helix: (I) dorsal view, (J) lateral view; L. jerrelli: (K) dorsal view, (L) lateral view; L. minutus: (M) dorsal view, (N) lateral view; L. tenuatus: (O) dorsal view, (P) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692922" httpUri="https://zenodo.org/record/10692922/files/figure.png" pageId="30" pageNumber="231">Fig. 11H</figureCitation>
). Apophyses of paraprocts almost 1/3 of cerci length, apices curved (
<figureCitation id="134A2DCAFF5DFF848D5A7B2064D6544C" box="[620,713,1339,1365]" captionStart="FIGURE 11" captionStartId="13.[152,255,1908,1932]" captionTargetBox="[189,1396,188,1883]" captionTargetId="figure-19@13.[189,1398,181,1885]" captionTargetPageId="13" captionText="FIGURE 11. Male cerci. Lestes apollinaris: (A) dorsal view, (B) lateral view; L. curvatus: (C) dorsal view, (D) lateral view; L. dicrhostigma: (E) dorsal view, (F) lateral view; L. forficula: (G) dorsal view, (H) lateral view; Lestes helix: (I) dorsal view, (J) lateral view; L. jerrelli: (K) dorsal view, (L) lateral view; L. minutus: (M) dorsal view, (N) lateral view; L. tenuatus: (O) dorsal view, (P) lateral view." figureDoi="http://doi.org/10.5281/zenodo.10692922" httpUri="https://zenodo.org/record/10692922/files/figure.png" pageId="30" pageNumber="231">Fig. 11J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B62C4FF5DFF848FF17B4462D454D8" pageId="30" pageNumber="231" type="discussion">
<paragraph id="8BCE314FFF5DFF848FF17B4462D454D8" blockId="30.[151,1437,690,1473]" pageId="30" pageNumber="231">
<emphasis id="B905ED5DFF5DFF848FF17B4467225460" bold="true" box="[199,317,1375,1401]" pageId="30" pageNumber="231">Remarks:</emphasis>
This species is known from a few localities in
<collectingCountry id="F36671DFFF5DFF848C407B4465B65460" box="[886,937,1375,1401]" name="Peru" pageId="30" pageNumber="231">Peru</collectingCountry>
(
<bibRefCitation id="EFE04CBEFF5DFF848C8A7B4462365460" author="Ris, F." box="[956,1065,1375,1401]" pageId="30" pageNumber="231" pagination="1 - 197" refId="ref21258" refString="Ris, F. (1918) Libellen (Odonata) aus der Regimder amerikanischen Kordilleren von Costaricibis Catamarca. Archiv far Naturgeschichte A, (9), 1 - 197." type="journal article" year="1918">Ris 1918</bibRefCitation>
;
<bibRefCitation id="EFE04CBEFF5DFF848B0F7B4462C25460" author="Schmidt, E." box="[1081,1245,1375,1401]" pageId="30" pageNumber="231" pagination="225 - 276" refId="ref21399" refString="Schmidt, E. (1942) Odonata nebst Bemerkungen ¸ ber die Anomisma und Chalcopteryx des AmazonasGebiets. In: Titschack, E. (Ed.), Beitrage zur Fauna Perus nach der Ausbeute der Hamburger S ¸ dperu Expedition 1936, 1942, pp. 225 - 276." type="journal article" year="1942">Schmidt 1942</bibRefCitation>
),
<collectingCountry id="F36671DFFF5DFF848BC27B4463785460" box="[1268,1383,1375,1401]" name="Venezuela" pageId="30" pageNumber="231">Venezuela</collectingCountry>
(
<bibRefCitation id="EFE04CBEFF5DFF848A4C7B44679B5484" author="De Marmels, J. &amp; Arias, Q. &amp; Gaiani, M. A." pageId="30" pageNumber="231" refId="ref19193" refString="De Marmels, J., Arias, Q. &amp; Gaiani, M. A. (2022) Odonata of Venezuela. An annotated checklist. Available from: https: // odonates. miza-ucv. org. ve (accessed 3 July 2023)" type="url" year="2022">
De Marmels
<emphasis id="B905ED5DFF5DFF848E327B9867205484" box="[260,319,1411,1437]" italics="true" pageId="30" pageNumber="231">et al.</emphasis>
2022
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), and
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(
<bibRefCitation id="EFE04CBEFF5DFF848D7A7B98653F5484" author="Bota-Sierra, C. A." box="[588,800,1411,1437]" pageId="30" pageNumber="231" pagination="97 - 100" refId="ref18105" refString="Bota-Sierra, C. A. (2014 a) Nine new records of Odonata for Colombia from the Orinoco Basin (Lestidae, Calopterygidae, Heteragrionidae, Coenagrionidae, Libellulidae). Notulae Odonatologicae, 8 (4), 97 - 100." type="journal article" year="2014">Bota-Sierra 2014a</bibRefCitation>
;
<bibRefCitation id="EFE04CBEFF5DFF848C187B9862255484" author="Bota-Sierra, C. A. &amp; Moreno-Arias, C. &amp; Faasen, T." box="[814,1082,1410,1437]" pageId="30" pageNumber="231" pagination="249 - 268" refId="ref18223" refString="Bota-Sierra, C. A., Moreno-Arias, C. &amp; Faasen, T. (2015) Preliminary list of Odonata from the Colombian Amazon, with descriptions of Inpabasis nigridorsum sp. nov. &amp; Diaphlebia richteri sp. nov. (Coenagrionidae &amp; Gomphidae). International Journal of Odonatology, 18 (3), 249 - 268. https: // doi. org / 10.1080 / 13887890.2015.1081637" type="journal article" year="2015">
Bota-Sierra
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2015
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). In
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, it occurs in the Amazon and Orinoquia regions and males are easily recognizable due to the unique cerci shape.
</paragraph>
</subSubSection>
</treatment>
</document>
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