106 lines
14 KiB
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106 lines
14 KiB
XML
<document id="7C8D235A3273C4454AD94FAF1F582CB2" ID-DOI="10.11646/zootaxa.5296.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984131" ID-ZooBank="59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685427549966" checkinUser="plazi" docAuthor="Bartel, Christian, Dunlop, Jason A. & Giribet, Gonzalo" docDate="2023" docId="03D87843394EFD2C8BB5F9D3FA5EFDD6" docLanguage="en" docName="zootaxa.5296.3.6.pdf" docOrigin="Zootaxa 5296 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stylocellidae Hansen & Sorensen 1904" docType="treatment" docVersion="2" lastPageNumber="433" masterDocId="FFE1003B3945FD208B22FFC5FFC5FFCF" masterDocTitle="An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber" masterLastPageNumber="445" masterPageNumber="421" pageNumber="432" updateTime="1685428803803" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1A0F3F1BECB45CAD8FF4AC7DCB98609C">An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber</mods:title>
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<mods:namePart id="64B9AFF0EF103928E88B132E83C2B4B2">Bartel, Christian</mods:namePart>
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<mods:namePart id="14A2F35A351BFE2A0158DB4CD5AD15BB">Dunlop, Jason A.</mods:namePart>
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<mods:namePart id="A2E25F01C1835AC562880D08207080EC">Giribet, Gonzalo</mods:namePart>
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<mods:title id="5580F97566F92CFC9A72C81171C2FFB7">Zootaxa</mods:title>
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<mods:date id="544668A56785ACA1699CCA374284A10D">2023</mods:date>
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<mods:number id="BF8B614156DEFC1F655249C6154D264D">2023-05-29</mods:number>
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<mods:number id="A9502EEEF943C422FC7C24DB6CC58345">5296</mods:number>
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<treatment id="03D87843394EFD2C8BB5F9D3FA5EFDD6" LSID="urn:lsid:plazi:treatment:03D87843394EFD2C8BB5F9D3FA5EFDD6" httpUri="http://treatment.plazi.org/id/03D87843394EFD2C8BB5F9D3FA5EFDD6" lastPageId="12" lastPageNumber="433" pageId="11" pageNumber="432">
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<subSubSection id="C36B9ADE394EFD2B8BB5F9D3FE74F9FF" box="[151,433,1558,1584]" pageId="11" pageNumber="432" type="nomenclature">
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<paragraph id="8BCEC955394EFD2B8BB5F9D3FE74F9FF" blockId="11.[151,433,1558,1620]" box="[151,433,1558,1584]" pageId="11" pageNumber="432">
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<heading id="D0867E39394EFD2B8BB5F9D3FE74F9FF" bold="true" box="[151,433,1558,1584]" fontSize="11" level="1" pageId="11" pageNumber="432" reason="1">
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<emphasis id="B9051547394EFD2B8BB5F9D3FE74F9FF" bold="true" box="[151,433,1558,1584]" pageId="11" pageNumber="432">
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<taxonomicName id="4C71B2D6394EFD2B8BB5F9D3FEF0F9FF" ID-CoL="GS8" ID-ENA="57970" authorityName="Hansen & Sorensen" authorityYear="1904" box="[151,309,1558,1584]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
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gen. sp. 3
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE394EFD2B8BB5F9FFFF1CF99B" box="[151,217,1594,1620]" pageId="11" pageNumber="432" type="description">
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<paragraph id="8BCEC955394EFD2B8BB5F9FFFF1CF99B" blockId="11.[151,433,1558,1620]" box="[151,217,1594,1620]" pageId="11" pageNumber="432">
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<figureCitation id="134AD5D0394EFD2B8BB5F9FFFF1CF99B" box="[151,217,1594,1620]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="11" pageNumber="432">Fig. 9</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE394EFD2B8BB5F947FDD6F90F" pageId="11" pageNumber="432" type="materials_examined">
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<paragraph id="8BCEC955394EFD2B8BB5F947FDD6F90F" blockId="11.[151,1436,1666,2017]" pageId="11" pageNumber="432">
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<emphasis id="B9051547394EFD2B8BB5F947FEC3F953" bold="true" box="[151,262,1666,1692]" pageId="11" pageNumber="432">Material.</emphasis>
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Specimen BUB3653, ex coll. Patrick Müller;
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<collectingCountry id="F36689C5394EFD2B8815F946FC63F952" box="[823,934,1667,1693]" name="Myanmar" pageId="11" pageNumber="432">Myanmar</collectingCountry>
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, Hukawng Valley; Burmese amber, Upper Cretaceous (Lower Cenomanian).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE394EFD2C8BE5F90FFB92FF1A" lastPageId="12" lastPageNumber="433" pageId="11" pageNumber="432" type="description">
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<paragraph id="8BCEC955394EFD2C8BE5F90FFB92FF1A" blockId="11.[151,1436,1666,2017]" lastBlockId="12.[151,1437,150,537]" lastPageId="12" lastPageNumber="433" pageId="11" pageNumber="432">
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<emphasis id="B9051547394EFD2B8BE5F90FFE9CF92B" bold="true" box="[199,345,1738,1764]" pageId="11" pageNumber="432">Description.</emphasis>
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Body oval, total L 2.32, maximum opisthosomal
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<geoCoordinate id="EE45AF92394EFD2B88A1F90EFC12F92B" box="[899,983,1738,1765]" degrees="1.23" direction="west" orientation="longitude" pageId="11" pageNumber="432" precision="555" value="-1.23">W 1.23</geoCoordinate>
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. Ozophores equivocal. Chelicerae rather short, its distal segment ornamented with small tubercles (
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<figureCitation id="134AD5D0394EFD2B8804F92BFC67F8C7" box="[806,930,1774,1800]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="11" pageNumber="432">Fig. 9A–B</figureCitation>
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); basal segment L?, median segment L 0.36, distal segment L 0.18. Pedipalps moderately long; tarsus bearing a small claw on its tip and furnished with a few sensory setae; pedipalp length: tr?, fe?, pa?, ti 0.34, ta 0.27. Legs long, robust and finely granular; tarsi with a single smooth claw and additional setae; tarsus IV not subdivided, bearing a lamelliform proximal adenostyle; leg lengths: I tr 0.21, fe 0.43*, pa 0.23, ti 0.49*, mt 0.30, ta 0.41, total 2.07; II tr 0.20, fe 0.46, pa 0.14, ti 0.37, mt 0.30, ta 0.43, total 1.90; III tr 0.16, fe 0.41, pa 0.18, ti 0.42*, mt 0.28, ta 0.37, total 1.82; IV tr 0.23, fe 0.52, pa 0.18, ti 0.34, mt 0.36, ta 0.34, total 1.97. Gonostome equivocal. Opisthosomal sternites completely granular. Sternites 2–3 fused into a plate; sternites 4–8 free, their widths: 4, 0.98; 5, 0.93; 6, 0.82; 7, 0.64; 8, 0.52. Opisthosomal sternal sulci straight and parallel to each other. Anal plate rounded and unmodified (
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<figureCitation id="134AD5D03949FD2C88D3FF7EFB8FFF1A" box="[1009,1098,187,213]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="12" pageNumber="433">Fig. 9B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B9ADE3949FD2C8BE5FF1AFA5EFDD6" pageId="12" pageNumber="433" type="discussion">
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<paragraph id="8BCEC9553949FD2C8BE5FF1AFA5EFDD6" blockId="12.[151,1437,150,537]" pageId="12" pageNumber="433">
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<emphasis id="B90515473949FD2C8BE5FF1AFEFEFF36" bold="true" box="[199,315,223,249]" pageId="12" pageNumber="433">Remarks.</emphasis>
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Specimen BUB3653 (
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<figureCitation id="134AD5D03949FD2C891FFF1AFD47FF36" box="[573,642,223,249]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="12" pageNumber="433">Fig. 9</figureCitation>
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) is interpreted as an adult male as it bears an adenostyle on tarsus IV and seems heavily sclerotized. The habitus, in combination with an ornamented second cheliceral segment and free sternites, indicates stylocellid affinities. Additionally, typical pettalid or ogoveid characters, like a modified anal region or opisthosomal apophyses, are missing. The proximal position of the adenostyle (
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<figureCitation id="134AD5D03949FD2C8FA5FE8EFAC4FEAA" box="[1159,1281,331,357]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="12" pageNumber="433">Fig. 9A–B</figureCitation>
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) is somewhat similar to that observed in specimen GPIH05128 and is thus more typical for modern sironids or neogoveids. However, specimen GPIH05128 features a much larger and lamelliform adenostyle and a longer tarsus IV. Due to the lack of important dorsal and ventral characteristics which could not be reconstructed with the help of micro-CT (
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<figureCitation id="134AD5D03949FD2C8BBDFE1EFF33FE3A" box="[159,246,475,501]" captionStart="FIGURE 9" captionStartId="12.[151,250,1088,1112]" captionTargetBox="[151,1435,561,1065]" captionTargetId="figure-207@12.[151,1435,561,1065]" captionTargetPageId="12" captionText="FIGURE 9. Stylocellidae gen. sp. 3 (male specimen BUB3653). A. Whole body, ventral view. B. Camera lucida drawing of A. C. 3D reconstruction of whole body, ventral view. Abbreviations: ad, adenostyle; ch, chelicerae; pp, pedipalp; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984153" httpUri="https://zenodo.org/record/7984153/files/figure.png" pageId="12" pageNumber="433">Fig. 9C</figureCitation>
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), this fossil cannot be placed any further in the family
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<taxonomicName id="4C71B2D63949FD2C8868FE1EFC25FE3A" authorityName="Hansen & Sorensen" authorityYear="1904" box="[842,992,475,501]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="433" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
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. Nevertheless, this is another interesting specimen showing a possible combination of sironid and stylocellid affinities and it is available for further studies.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |