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<document id="2EF1F52EF205CBA2E862EBDBA9ABD792" ID-DOI="10.11646/zootaxa.5296.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984131" ID-ZooBank="59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685427549966" checkinUser="plazi" docAuthor="Bartel, Christian, Dunlop, Jason A. &amp; Giribet, Gonzalo" docDate="2023" docId="03D87843394CFD2B8BB5FA93FC83FA0B" docLanguage="en" docName="zootaxa.5296.3.6.pdf" docOrigin="Zootaxa 5296 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stylocellidae Hansen &amp; Sorensen 1904" docType="treatment" docVersion="2" lastPageNumber="432" masterDocId="FFE1003B3945FD208B22FFC5FFC5FFCF" masterDocTitle="An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber" masterLastPageNumber="445" masterPageNumber="421" pageNumber="430" updateTime="1685428803803" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E60E7B2372BF0B0EC35FFAA9DA4EEC40">An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber</mods:title>
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<mods:namePart id="A91B03A642D54B290C58AC3D4B93E7CA">Dunlop, Jason A.</mods:namePart>
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<treatment id="03D87843394CFD2B8BB5FA93FC83FA0B" ID-DOI="http://doi.org/10.5281/zenodo.7984071" ID-Zenodo-Dep="7984071" LSID="urn:lsid:plazi:treatment:03D87843394CFD2B8BB5FA93FC83FA0B" httpUri="http://treatment.plazi.org/id/03D87843394CFD2B8BB5FA93FC83FA0B" lastPageId="11" lastPageNumber="432" pageId="9" pageNumber="430">
<subSubSection id="C36B9ADE394CFD298BB5FA93FE74FABF" box="[151,433,1366,1393]" pageId="9" pageNumber="430" type="nomenclature">
<paragraph id="8BCEC955394CFD298BB5FA93FE74FABF" blockId="9.[151,433,1366,1429]" box="[151,433,1366,1393]" pageId="9" pageNumber="430">
<heading id="D0867E39394CFD298BB5FA93FE74FABF" bold="true" box="[151,433,1366,1393]" fontSize="11" level="1" pageId="9" pageNumber="430" reason="1">
<emphasis id="B9051547394CFD298BB5FA93FE74FABF" bold="true" box="[151,433,1366,1393]" pageId="9" pageNumber="430">
<taxonomicName id="4C71B2D6394CFD298BB5FA93FEF0FABF" ID-CoL="GS8" ID-ENA="57970" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[151,309,1366,1392]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="430" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
gen. sp. 2
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36B9ADE394CFD298BB5FABEFF3AFA5A" box="[151,255,1403,1429]" pageId="9" pageNumber="430" type="description">
<paragraph id="8BCEC955394CFD298BB5FABEFF3AFA5A" blockId="9.[151,433,1366,1429]" box="[151,255,1403,1429]" pageId="9" pageNumber="430">
<figureCitation id="134AD5D0394CFD298BB5FABEFF3AFA5A" box="[151,255,1403,1429]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1909,1933]" captionStartId-1="11.[151,250,696,720]" captionTargetBox-0="[204,1383,181,1873]" captionTargetBox-1="[151,1435,181,672]" captionTargetId-0="figure-20@10.[204,1383,181,1885]" captionTargetId-1="figure-20@11.[151,1435,181,672]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." captionText-1="FIGURE 8. Details of Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Pedipalps and chelicerae in ventral view. B. Close-up of closed gonostome and spiracles (indicated by arrows). Abbreviations: ch, chelicerae; go, closed gonostome; sp, spiracle. Scale bars equal 0.2 mm (A) and 0.1 mm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.7984149" figureDoi-1="http://doi.org/10.5281/zenodo.7984151" httpUri-0="https://zenodo.org/record/7984149/files/figure.png" httpUri-1="https://zenodo.org/record/7984151/files/figure.png" pageId="9" pageNumber="430">Figs. 7–8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36B9ADE394CFD298BB5FA07FDD6F9CE" pageId="9" pageNumber="430" type="materials_examined">
<paragraph id="8BCEC955394CFD298BB5FA07FDD6F9CE" blockId="9.[151,1437,1474,2041]" pageId="9" pageNumber="430">
<emphasis id="B9051547394CFD298BB5FA07FEC3FA13" bold="true" box="[151,262,1474,1500]" pageId="9" pageNumber="430">Material.</emphasis>
Specimen BUB1517, ex coll. Patrick Müller;
<collectingCountry id="F36689C5394CFD298815FA06FC63FA12" box="[823,934,1475,1501]" name="Myanmar" pageId="9" pageNumber="430">Myanmar</collectingCountry>
, Hukawng Valley; Burmese amber, Upper Cretaceous (Lower Cenomanian).
</paragraph>
</subSubSection>
<subSubSection id="C36B9ADE394CFD2B8BE5F9CFFF2FFBDB" lastPageId="11" lastPageNumber="432" pageId="9" pageNumber="430" type="description">
<paragraph id="8BCEC955394CFD298BE5F9CFFE82F836" blockId="9.[151,1437,1474,2041]" pageId="9" pageNumber="430">
<emphasis id="B9051547394CFD298BE5F9CFFE9CF9EB" bold="true" box="[199,345,1546,1572]" pageId="9" pageNumber="430">Description.</emphasis>
Body oval and somewhat very widely subtriangular in profile at anterior end; total L 1.76, maximum prosomal W behind ozophores 1.11, maximum opisthosomal
<geoCoordinate id="EE45AF92394CFD2988CCF9EAFB8CF986" box="[1006,1097,1583,1609]" degrees="1.09" direction="west" orientation="longitude" pageId="9" pageNumber="430" precision="555" value="-1.09">W 1.09</geoCoordinate>
. Prosoma almost completely tuberculate-microgranular, especially near outer edges (
<figureCitation id="134AD5D0394CFD298826F996FCB8F9A2" box="[772,893,1619,1645]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[204,1383,181,1873]" captionTargetId="figure-20@10.[204,1383,181,1885]" captionTargetPageId="10" captionText="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984149" httpUri="https://zenodo.org/record/7984149/files/figure.png" pageId="9" pageNumber="430">Fig. 7A, C</figureCitation>
). Conical ozophores with tubercles and in
<typeStatus id="54CA77F7394CFD298E75F996FA42F9A2" box="[1367,1415,1619,1645]" pageId="9" pageNumber="430">type</typeStatus>
2 position. L of ozophores 0.21, W at base 0.17*, distance between base of ozophores 0.65. Eye lens present anterior to left ozophore. Six opisthosomal tergites visible, each delimited and sparsely granular near left and right edges. Borders delimiting tergites smooth. Chelicerae moderately long, somewhat protruding and completely granular; each cheliceral finger with nine small uniform teeth; basal segment L 0.32, median segment L 0.26, distal segment L 0.22. Pedipalps relatively small and granular, except for tarsus furnished with sensory setae; tarsus with small claw at its tip; pedipalp length: tr 0.14, fe 0.27*, pa 0.21, ti 0.25, ta 0.17, total 1.04. Legs relatively short, robust and strongly granular, especially leg I and II featuring additional setiferous tubercles; coxae not fused; leg tarsi entire, not subdivided, bearing a single smooth claw and numerous setae; tarsus IV without adenostyle; leg lengths: I tr 0.24, fe 0.57, pa 0.24, ti 0.33, mt 0.14, ta 0.44*, total 1.96; II tr 0.21, fe 0.51, pa 0.25, ti 0.33, mt 0.13, ta 0.39, total 1.82; III tr 0.19, fe 0.40, pa 0.21, ti 0.33, mt 0.14, ta 0.32, total 1.59; IV tr 0.22, fe 0.55, pa 0.21, ti 0.21, mt 0.14, ta 0.43, total 1.76.
</paragraph>
<caption id="DF0E99DD394FFD2A8BB5F8B0FD20F837" ID-DOI="http://doi.org/10.5281/zenodo.7984149" ID-Zenodo-Dep="7984149" httpUri="https://zenodo.org/record/7984149/files/figure.png" pageId="10" pageNumber="431" startId="10.[151,250,1909,1933]" targetBox="[204,1383,181,1873]" targetPageId="10" targetType="figure">
<paragraph id="8BCEC955394FFD2A8BB5F8B0FD20F837" blockId="10.[151,1437,1909,2041]" pageId="10" pageNumber="431">
<emphasis id="B9051547394FFD2A8BB5F8B0FED1F842" bold="true" box="[151,276,1909,1933]" pageId="10" pageNumber="431">FIGURE 7.</emphasis>
<taxonomicName id="4C71B2D6394FFD2A8A3EF8B0FE65F842" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[284,416,1909,1933]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="10" pageNumber="431" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing of A. D. Camera lucida drawing of B (closed gonostome and spiracles indicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm.
</paragraph>
</caption>
<caption id="DF0E99DD394EFD2B8BB5FD7DFD07FCD7" ID-DOI="http://doi.org/10.5281/zenodo.7984151" ID-Zenodo-Dep="7984151" httpUri="https://zenodo.org/record/7984151/files/figure.png" pageId="11" pageNumber="432" startId="11.[151,250,696,720]" targetBox="[151,1435,181,672]" targetPageId="11" targetType="figure">
<paragraph id="8BCEC955394EFD2B8BB5FD7DFD07FCD7" blockId="11.[151,1437,696,792]" pageId="11" pageNumber="432">
<emphasis id="B9051547394EFD2B8BB5FD7DFED1FD1F" bold="true" box="[151,276,696,720]" pageId="11" pageNumber="432">FIGURE 8.</emphasis>
Details of
<taxonomicName id="4C71B2D6394EFD2B8AABFD7DFDC8FD1F" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[393,525,696,720]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
gen. sp. 2 (immature specimen BUB1517). A. Pedipalps and chelicerae in ventral view. B. Close-up of closed gonostome and spiracles (indicated by arrows). Abbreviations: ch, chelicerae; go, closed gonostome; sp, spiracle. Scale bars equal 0.2 mm (A) and 0.1 mm (B).
</paragraph>
</caption>
<paragraph id="8BCEC955394EFD2B8BE5FC83FF2FFBDB" blockId="11.[151,1437,838,1477]" pageId="11" pageNumber="432">
Ventral side of opisthosoma covered with small tubercles and setae. Opisthosomal sulci parallel to each other. Sternites 2–3 fused to a subtrapezoidal plate; sternites 4–9 getting increasingly concentrated towards anal plate. Gonostome closed, subhexagonal, covered with numerous small and two larger tubercles proximally,
<geoCoordinate id="EE45AF92394EFD2B8E05FC4AFABAFC67" box="[1319,1407,910,937]" degrees="0.33" direction="west" orientation="longitude" pageId="11" pageNumber="432" precision="555" value="-0.33">W 0.33</geoCoordinate>
, L 0.32. Spiracles C-shaped (consisting of small denticles) and surrounded by smaller tubercles, located on isolated sclerites behind coxae IV, maximum
<geoCoordinate id="EE45AF92394EFD2B891CFC12FD53FC3F" box="[574,662,982,1009]" degrees="0.08" direction="west" orientation="longitude" pageId="11" pageNumber="432" precision="555" value="-0.08">W 0.08</geoCoordinate>
. Anal plate rounded, with few small tubercles,
<geoCoordinate id="EE45AF92394EFD2B8F93FC12FAC2FC3F" box="[1201,1287,982,1009]" degrees="0.20" direction="west" orientation="longitude" pageId="11" pageNumber="432" precision="555" value="-0.2">W 0.20</geoCoordinate>
, L 0.15 (
<figureCitation id="134AD5D0394EFD2B8E4DFC13FF1BFBDB" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[204,1383,181,1873]" captionTargetId="figure-20@10.[204,1383,181,1885]" captionTargetPageId="10" captionText="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984149" httpUri="https://zenodo.org/record/7984149/files/figure.png" pageId="11" pageNumber="432">Fig. 7B, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36B9ADE394EFD2B8BE5FBDBFC83FA0B" pageId="11" pageNumber="432" type="discussion">
<paragraph id="8BCEC955394EFD2B8BE5FBDBFC83FA0B" blockId="11.[151,1437,838,1477]" pageId="11" pageNumber="432">
<emphasis id="B9051547394EFD2B8BE5FBDBFEFEFBF7" bold="true" box="[199,315,1054,1080]" pageId="11" pageNumber="432">Remarks.</emphasis>
Specimen BUB1517 (
<figureCitation id="134AD5D0394EFD2B8963FBDBFD6AFBF7" box="[577,687,1054,1080]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="10.[151,250,1909,1933]" captionStartId-1="11.[151,250,696,720]" captionTargetBox-0="[204,1383,181,1873]" captionTargetBox-1="[151,1435,181,672]" captionTargetId-0="figure-20@10.[204,1383,181,1885]" captionTargetId-1="figure-20@11.[151,1435,181,672]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." captionText-1="FIGURE 8. Details of Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Pedipalps and chelicerae in ventral view. B. Close-up of closed gonostome and spiracles (indicated by arrows). Abbreviations: ch, chelicerae; go, closed gonostome; sp, spiracle. Scale bars equal 0.2 mm (A) and 0.1 mm (B)." figureDoi-0="http://doi.org/10.5281/zenodo.7984149" figureDoi-1="http://doi.org/10.5281/zenodo.7984151" httpUri-0="https://zenodo.org/record/7984149/files/figure.png" httpUri-1="https://zenodo.org/record/7984151/files/figure.png" pageId="11" pageNumber="432">Figs. 7–8</figureCitation>
) is again a juvenile of uncertain gender judging from the closed gonostome and spiracles situated on separate sclerites (
<figureCitation id="134AD5D0394EFD2B8839FB87FC6DFB92" box="[795,936,1090,1117]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[204,1383,181,1873]" captionTargetId="figure-20@10.[204,1383,181,1885]" captionTargetPageId="10" captionText="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984149" httpUri="https://zenodo.org/record/7984149/files/figure.png" pageId="11" pageNumber="432">Figs. 7B, D</figureCitation>
,
<figureCitation id="134AD5D0394EFD2B8895FB87FC1FFB93" box="[951,986,1090,1116]" captionStart="FIGURE 8" captionStartId="11.[151,250,696,720]" captionTargetBox="[151,1435,181,672]" captionTargetId="figure-20@11.[151,1435,181,672]" captionTargetPageId="11" captionText="FIGURE 8. Details of Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Pedipalps and chelicerae in ventral view. B. Close-up of closed gonostome and spiracles (indicated by arrows). Abbreviations: ch, chelicerae; go, closed gonostome; sp, spiracle. Scale bars equal 0.2 mm (A) and 0.1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7984151" httpUri="https://zenodo.org/record/7984151/files/figure.png" pageId="11" pageNumber="432">8B</figureCitation>
). Therefore most diagnostic characters are not fully developed. Nevertheless, granular tegument, C-shaped spiracles,
<typeStatus id="54CA77F7394EFD2B8F2CFBA2FBFBFB4E" box="[1038,1086,1127,1153]" pageId="11" pageNumber="432">type</typeStatus>
2 ozophores, uniform teeth on cheliceral fingers and an ornamented second cheliceral segment (
<figureCitation id="134AD5D0394EFD2B8858FB4FFC16FB6B" box="[890,979,1162,1188]" captionStart="FIGURE 8" captionStartId="11.[151,250,696,720]" captionTargetBox="[151,1435,181,672]" captionTargetId="figure-20@11.[151,1435,181,672]" captionTargetPageId="11" captionText="FIGURE 8. Details of Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Pedipalps and chelicerae in ventral view. B. Close-up of closed gonostome and spiracles (indicated by arrows). Abbreviations: ch, chelicerae; go, closed gonostome; sp, spiracle. Scale bars equal 0.2 mm (A) and 0.1 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7984151" httpUri="https://zenodo.org/record/7984151/files/figure.png" pageId="11" pageNumber="432">Fig. 8A</figureCitation>
) suggest placement in the
<taxonomicName id="4C71B2D6394EFD2B8E21FB4FFA5CFB6B" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[1283,1433,1162,1188]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
. Within
<taxonomicName id="4C71B2D6394EFD2B8BCEFB6BFEBBFB07" authorityName="Hansen &amp; Sorensen" authorityYear="1904" box="[236,382,1198,1224]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
two extant genera feature extensive sculpturing on the second cheliceral segment, similar to that in the fossil:
<taxonomicName id="4C71B2D6394EFD2B8A42FB17FDB7FB23" authority="Thorell, 1890" authorityName="Thorell" authorityYear="1890" box="[352,626,1234,1260]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9051547394EFD2B8A42FB17FE0BFB23" box="[352,462,1234,1260]" italics="true" pageId="11" pageNumber="432">Miopsalis</emphasis>
<bibRefCitation id="EFE0B4A4394EFD2B8AFAFB17FDB7FB23" author="Thorell, T." box="[472,626,1234,1260]" pageId="11" pageNumber="432" pagination="5 - 106" refId="ref14611" refString="Thorell, T. (1890) Aracnidi di Nias e di Sumatra raccolti nel 1886 dal Sig. E. Mondigliani. Annali del Museo Civico di Storia Naturale di Genova, Series 2 a, 10, 5 - 106. https: // doi. org / 10.5962 / bhl. title. 10511" type="journal article" year="1890">Thorell, 1890</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C71B2D6394EFD2B898EFB17FC2BFB23" authority="Westwood, 1874" authorityName="Westwood" authorityYear="1874" box="[684,1006,1234,1260]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9051547394EFD2B898EFB17FCE0FB23" box="[684,805,1234,1260]" italics="true" pageId="11" pageNumber="432">Stylocellus</emphasis>
<bibRefCitation id="EFE0B4A4394EFD2B880CFB17FC2BFB23" author="Westwood, J. O." box="[814,1006,1234,1260]" pageId="11" pageNumber="432" refId="ref14768" refString="Westwood, J. O. (1874) Thesaurus entomologicus oxoniensis; or, illustrations of new, rare, and interesting insects, for the most part contained in the collections presented to the University of Oxford by the Rev. F. W. Hope, M. A., D. C. L., F. R. S., &amp; c. with forty plates from drawings by the author. Clarendon Press, Oxford, 205 pp. https: // doi. org / 10.5962 / bhl. title. 14077" type="book" year="1874">Westwood, 1874</bibRefCitation>
</taxonomicName>
. These can be easily distinguished by the form of the ventral opisthosomal sulci.
<taxonomicName id="4C71B2D6394EFD2B89A4FB33FD3AFADF" authorityName="Westwood" authorityYear="1874" box="[646,767,1270,1296]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9051547394EFD2B89A4FB33FD3AFADF" box="[646,767,1270,1296]" italics="true" pageId="11" pageNumber="432">Stylocellus</emphasis>
</taxonomicName>
features sinoidal ventral opisthosomal sulci and
<taxonomicName id="4C71B2D6394EFD2B8E0CFB33FA59FADF" authorityName="Thorell" authorityYear="1890" box="[1326,1436,1270,1296]" class="Arachnida" family="Stylocellidae" genus="Miopsalis" kingdom="Animalia" order="Opiliones" pageId="11" pageNumber="432" phylum="Arthropoda" rank="genus">
<emphasis id="B9051547394EFD2B8E0CFB33FA59FADF" box="[1326,1436,1270,1296]" italics="true" pageId="11" pageNumber="432">Miopsalis</emphasis>
</taxonomicName>
features straight ventral opisthosomal sulci (Clouse 2012). The latter can also be observed in specimen BUB 1517. However, the fossil also shows a strong concentration of opisthosomal sternites (
<figureCitation id="134AD5D0394EFD2B8F00FAFBFB5EFA97" box="[1058,1179,1342,1369]" captionStart="FIGURE 7" captionStartId="10.[151,250,1909,1933]" captionTargetBox="[204,1383,181,1873]" captionTargetId="figure-20@10.[204,1383,181,1885]" captionTargetPageId="10" captionText="FIGURE 7. Stylocellidae gen. sp. 2 (immature specimen BUB1517). A. Whole body, dorsal view. B. Same, ventral view. C. Camera lucida drawing ofA. D.Camera lucida drawing of B(closed gonostome and spiraclesindicated by arrows).Abbreviations: ap, anal plate; ch, chelicerae; fe, femur; go, closed gonostome; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; legs numbered from I–IV. Scale bars equal 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7984149" httpUri="https://zenodo.org/record/7984149/files/figure.png" pageId="11" pageNumber="432">Fig. 7B, D</figureCitation>
) around the anal plate, a character not known from modern genera. This could either be a juvenile character not present in modern species, or it could be related to a deformation of the opisthosoma as a result of its preservation in amber. For that reason we prefer a cautious approach and leave this specimen unnamed.
</paragraph>
</subSubSection>
</treatment>
</document>
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