136 lines
16 KiB
XML
136 lines
16 KiB
XML
<document id="EB626A6F6111EB8F37FB66F27B704FF9" ID-CLB-Dataset="53499" ID-DOI="10.11646/zootaxa.5296.3.6" ID-GBIF-Dataset="2825cf24-234a-4b85-b997-673ee08019bf" ID-ISSN="1175-5326" ID-Zenodo-Dep="7984131" ID-ZooBank="59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1685427549966" checkinUser="plazi" docAuthor="Bartel, Christian, Dunlop, Jason A. & Giribet, Gonzalo" docDate="2023" docId="03D878433947FD228BB5FAD0FB66F835" docLanguage="en" docName="zootaxa.5296.3.6.pdf" docOrigin="Zootaxa 5296 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.3.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sirocellus Bartel & Dunlop & Giribet 2023, gen. nov." docType="treatment" docUuid="EC139A74-F787-4577-9A7D-823A662E238F" docUuidSource="ZooBank" docVersion="6" lastPageNumber="423" masterDocId="FFE1003B3945FD208B22FFC5FFC5FFCF" masterDocTitle="An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber" masterLastPageNumber="445" masterPageNumber="421" pageNumber="423" updateTime="1699510734804" updateUser="plazi" zenodo-license-document="CLOSED">
|
||
<mods:mods id="82E43E917269B57C9EE4FA26B4BEF2B0" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1CE8033E5890D11FB9DA48F798BA14E9">
|
||
<mods:title id="FD2886A22EBB622779C99F6089D74BE2">An unexpected diversity of Cyphophthalmi (Arachnida: Opiliones) in Upper Cretaceous Burmese amber</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="FE25104EC9F9313C6A2F774405C0BA72" type="personal">
|
||
<mods:role id="C5179F56AFB293628EFB4C109AC7ABDF">
|
||
<mods:roleTerm id="9D9289A9E2D8C24E26DAE060D5EC3263">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E8E7C58A689A0B03BC17B22E055DDB79">Bartel, Christian</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="CB79A73AEDAABB79506AAF3277B5032E" type="personal">
|
||
<mods:role id="088C463345646320DA58F87DF45A6F0A">
|
||
<mods:roleTerm id="CB540E8A9473CD84BD0DA97C85B0A05A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="72403C5BD8B819595DCC23EA5220BCDE">Dunlop, Jason A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="4FC72048289071C07C59575F2D15042C" type="personal">
|
||
<mods:role id="BA37157A73C93A8F7DFBD880BD628752">
|
||
<mods:roleTerm id="B24602ABC8CFF3D96D0360FF92D63F7D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="DE70192120F4B41808EA7EF248D4AFAD">Giribet, Gonzalo</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="975745A87ECEDBA3E576CF30E035F4CB">text</mods:typeOfResource>
|
||
<mods:relatedItem id="47B070F6BA1B8F6041831BB4CEE486A1" type="host">
|
||
<mods:titleInfo id="707F3DFBAC999BD3F04D1C3DDFDAF37C">
|
||
<mods:title id="389C6E0C93C99D006A6564ABE2788F8F">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="8CCF74006390AAB6DDB041C454726852">
|
||
<mods:date id="7BF908867619544ADE11E273A5ABCB73">2023</mods:date>
|
||
<mods:detail id="A916C36B7D8E69D02A9D30A171D25817" type="pubDate">
|
||
<mods:number id="6311D692DC60EC49D5832A360F91E088">2023-05-29</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="30B708C691588179FF085FF0219C597C" type="volume">
|
||
<mods:number id="653CF7B8B49BF3333E2A7D1569FFB3A2">5296</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A02479CC1B4D956BC3EB6027FC67C327" type="issue">
|
||
<mods:number id="F188B17F17A1E95140425A2D1055483C">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="B9FB05987359774192339F38206EC2FF" unit="page">
|
||
<mods:start id="438FBA61B31DA459569AD025B852FC32">421</mods:start>
|
||
<mods:end id="7FC859A2692D3BED7B95CAD74EC113A3">445</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="4C667017DE45803F18B49B5DCDE1743D">
|
||
<mods:url id="1F6B68989FC0D922EE427DC7F7FA4519">http://dx.doi.org/10.11646/zootaxa.5296.3.6</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="00B876399259A27918545DA0AAF7CE15">journal article</mods:classification>
|
||
<mods:identifier id="46249004010A2FB2B4832540217BCDB1" type="CLB-Dataset">53499</mods:identifier>
|
||
<mods:identifier id="9CFF7A2DBFE9D4B473667C7EB46AED52" type="DOI">10.11646/zootaxa.5296.3.6</mods:identifier>
|
||
<mods:identifier id="8A40B9C8E30DA7DD513B4B6C4C23581B" type="GBIF-Dataset">2825cf24-234a-4b85-b997-673ee08019bf</mods:identifier>
|
||
<mods:identifier id="B0EA1973B9D5BC01FB567329A4D2F7FA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="DC671673B71AC496BA45B3F1C59703AE" type="Zenodo-Dep">7984131</mods:identifier>
|
||
<mods:identifier id="02D8273BAF20717212BFA4F08E6D57C5" type="ZooBank">59AD1B4F-15B2-4DC0-A57E-2F6B57539D1A</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D878433947FD228BB5FAD0FB66F835" ID-DOI="http://doi.org/10.5281/zenodo.7984033" ID-GBIF-Taxon="209663743" ID-Zenodo-Dep="7984033" LSID="urn:lsid:zoobank.org:act:EC139A74-F787-4577-9A7D-823A662E238F" httpUri="http://treatment.plazi.org/id/03D878433947FD228BB5FAD0FB66F835" lastPageNumber="423" pageId="2" pageNumber="423">
|
||
<subSubSection id="C36B9ADE3947FD228BB5FAD0FCAAFABD" pageId="2" pageNumber="423" type="nomenclature">
|
||
<paragraph id="8BCEC9553947FD228BB5FAD0FE19FAE0" blockId="2.[151,476,1301,1328]" box="[151,476,1301,1328]" pageId="2" pageNumber="423">
|
||
<heading id="D0867E393947FD228BB5FAD0FE19FAE0" bold="true" box="[151,476,1301,1328]" fontSize="11" level="1" pageId="2" pageNumber="423" reason="1">
|
||
<emphasis id="B90515473947FD228BB5FAD0FE19FAE0" bold="true" box="[151,476,1301,1328]" pageId="2" pageNumber="423">
|
||
Genus
|
||
<taxonomicName id="4C71B2D63947FD228BCCFAD0FEA2FAFF" ID-CoL="BHZMB" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[238,359,1301,1328]" class="Arachnida" family="Stylocellidae" genus="Sirocellus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B90515473947FD228BCCFAD0FEA2FAFF" bold="true" box="[238,359,1301,1328]" italics="true" pageId="2" pageNumber="423">Sirocellus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A236A83C3947FD228A4CFAD0FE19FAE0" box="[366,476,1301,1327]" pageId="2" pageNumber="423" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCEC9553947FD228BB5FA9FFCAAFABD" blockId="2.[151,879,1370,1394]" box="[151,879,1370,1394]" pageId="2" pageNumber="423">
|
||
<uri id="FFE0C5573947FD228BB5FA9FFCAAFABD" box="[151,879,1370,1394]" pageId="2" pageNumber="423">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFD7F3803947FD228AACFA9FFCAAFABD" box="[398,879,1370,1394]" pageId="2" pageNumber="423">EC139A74-F787-4577-9A7D-823A662E238F</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B9ADE3947FD228BB5FA65FD9BFA74" box="[151,606,1440,1467]" pageId="2" pageNumber="423" type="type_taxon">
|
||
<paragraph id="8BCEC9553947FD228BB5FA65FD9BFA74" blockId="2.[151,1437,1440,2043]" box="[151,606,1440,1467]" pageId="2" pageNumber="423">
|
||
<emphasis id="B90515473947FD228BB5FA65FEF7FA75" bold="true" box="[151,306,1440,1466]" pageId="2" pageNumber="423">
|
||
<typeStatus id="54CA77F73947FD228BB5FA65FF17FA75" box="[151,210,1440,1466]" pageId="2" pageNumber="423">Type</typeStatus>
|
||
species.
|
||
</emphasis>
|
||
<taxonomicName id="4C71B2D63947FD228A1BFA65FDC5FA75" ID-CoL="BHZMC" authority="Bartel & Dunlop & Giribet, 2023" authorityName="Bartel & Dunlop & Giribet" authorityYear="2023" box="[313,512,1440,1466]" class="Arachnida" family="Stylocellidae" genus="Sirocellus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="iunctus" status="sp. nov.">
|
||
<emphasis id="B90515473947FD228A1BFA65FDC5FA75" box="[313,512,1440,1466]" italics="true" pageId="2" pageNumber="423">Sirocellus iunctus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A236A83C3947FD228925FA64FD9BFA74" box="[519,606,1441,1467]" pageId="2" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B9ADE3947FD228BE5FA01FAA7FA11" box="[199,1378,1476,1503]" pageId="2" pageNumber="423" type="etymology">
|
||
<paragraph id="8BCEC9553947FD228BE5FA01FAA7FA11" blockId="2.[151,1437,1440,2043]" box="[199,1378,1476,1503]" pageId="2" pageNumber="423">
|
||
<emphasis id="B90515473947FD228BE5FA01FE8BFA11" bold="true" box="[199,334,1476,1502]" pageId="2" pageNumber="423">Etymology.</emphasis>
|
||
A combination of the names of the modern genera
|
||
<taxonomicName id="4C71B2D63947FD2288AEFA01FC7FFA11" authorityName="Latreille" authorityYear="1795" box="[908,954,1476,1502]" class="Arachnida" family="Sironidae" genus="Siro" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90515473947FD2288AEFA01FC7FFA11" box="[908,954,1476,1502]" italics="true" pageId="2" pageNumber="423">Siro</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C71B2D63947FD2288D2FA01FBACFA11" authorityName="Westwood" authorityYear="1874" box="[1008,1129,1476,1502]" class="Arachnida" family="Stylocellidae" genus="Stylocellus" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90515473947FD2288D2FA01FBACFA11" box="[1008,1129,1476,1502]" italics="true" pageId="2" pageNumber="423">Stylocellus</emphasis>
|
||
</taxonomicName>
|
||
. Masculine in gender.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B9ADE3947FD228BE5FA2DFD6BF9A1" pageId="2" pageNumber="423" type="diagnosis">
|
||
<paragraph id="8BCEC9553947FD228BE5FA2DFD6BF9A1" blockId="2.[151,1437,1440,2043]" pageId="2" pageNumber="423">
|
||
<emphasis id="B90515473947FD228BE5FA2DFE87F9CD" bold="true" box="[199,322,1512,1538]" pageId="2" pageNumber="423">Diagnosis.</emphasis>
|
||
Body oval, completely granular dorsally as well as ventrally. Almost conical ozophores in the
|
||
<typeStatus id="54CA77F73947FD228E4EFA2CFA59F9CC" box="[1388,1436,1513,1539]" pageId="2" pageNumber="423">type</typeStatus>
|
||
2 position. Chelicerae long and mostly smooth, with just a few granules or tubercles. Pedipalps long and rather thin. Legs robust and with a granular surface. Leg tarsi not subdivided, bearing a single smooth claw. Tarsus IV in males proximally with a large lamelliform adenostyle.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36B9ADE3947FD228BE5F9BDFB66F835" pageId="2" pageNumber="423" type="discussion">
|
||
<paragraph id="8BCEC9553947FD228BE5F9BDFB66F835" blockId="2.[151,1437,1440,2043]" pageId="2" pageNumber="423">
|
||
<emphasis id="B90515473947FD228BE5F9BDFEFEF95D" bold="true" box="[199,315,1656,1682]" pageId="2" pageNumber="423">Remarks.</emphasis>
|
||
Specimen GPIH05128 (
|
||
<figureCitation id="134AD5D03947FD228969F9BDFD4FF95D" box="[587,650,1656,1682]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1852]" captionTargetBox="[151,1435,943,1796]" captionTargetId="figure-293@3.[151,1435,943,1802]" captionTargetPageId="3" captionText="FIGURE 1. Sirocellus iunctus gen. et sp. nov., male holotype (GPIH05128). A. Whole specimen, dorsolateral view. B. Same, ventrolateral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ad, adenostyle; ch, chelicerae; fe, femur; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; tr, trochanter; legs numbered from I–IV. Scale bars equal 1 mm (A, C, D) and 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7984133" httpUri="https://zenodo.org/record/7984133/files/figure.png" pageId="2" pageNumber="423">Fig. 1</figureCitation>
|
||
) is recognized as an adult male due to the presence of an adenostyle on tarsus IV (
|
||
<figureCitation id="134AD5D03947FD228A29F959FEA5F979" box="[267,352,1692,1718]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1852]" captionTargetBox="[151,1435,943,1796]" captionTargetId="figure-293@3.[151,1435,943,1802]" captionTargetPageId="3" captionText="FIGURE 1. Sirocellus iunctus gen. et sp. nov., male holotype (GPIH05128). A. Whole specimen, dorsolateral view. B. Same, ventrolateral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ad, adenostyle; ch, chelicerae; fe, femur; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; tr, trochanter; legs numbered from I–IV. Scale bars equal 1 mm (A, C, D) and 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7984133" httpUri="https://zenodo.org/record/7984133/files/figure.png" pageId="2" pageNumber="423">Fig. 1C</figureCitation>
|
||
) and due to the high degree of body sclerotization. The families
|
||
<taxonomicName id="4C71B2D63947FD228F12F959FB6CF979" authorityName="Shear" authorityYear="1980" box="[1072,1193,1692,1718]" class="Arachnida" family="Ogoveidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="family">Ogoveidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C71B2D63947FD228FFDF959FA88F979" authorityName="Shear" authorityYear="1980" box="[1247,1357,1692,1718]" class="Arachnida" family="Pettalidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="family">Pettalidae</taxonomicName>
|
||
can be excluded, in this case due to the absence of a large opisthosomal apophysis on the second sternite and of a modified anal region. The subtriangular anterior part of the body in combination with
|
||
<typeStatus id="54CA77F73947FD2288DBF920FBECF930" box="[1017,1065,1765,1791]" pageId="2" pageNumber="423">type</typeStatus>
|
||
2 ozophores (
|
||
<figureCitation id="134AD5D03947FD228FE8F921FA81F931" box="[1226,1348,1764,1790]" captionStart="FIGURE 1" captionStartId="3.[151,250,1828,1852]" captionTargetBox="[151,1435,943,1796]" captionTargetId="figure-293@3.[151,1435,943,1802]" captionTargetPageId="3" captionText="FIGURE 1. Sirocellus iunctus gen. et sp. nov., male holotype (GPIH05128). A. Whole specimen, dorsolateral view. B. Same, ventrolateral view. C. Camera lucida drawing of A. D. Same of B. Abbreviations: ad, adenostyle; ch, chelicerae; fe, femur; mt, metatarsus; oz, ozophore; pa, patella; pp, pedipalp; ta, tarsus; ti, tibia; tr, trochanter; legs numbered from I–IV. Scale bars equal 1 mm (A, C, D) and 0.5 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.7984133" httpUri="https://zenodo.org/record/7984133/files/figure.png" pageId="2" pageNumber="423">Fig. 1C, D</figureCitation>
|
||
) is also found in modern members of the family
|
||
<taxonomicName id="4C71B2D63947FD22897BF8CDFD2AF8ED" authorityName="Hansen & Sorensen" authorityYear="1904" box="[601,751,1800,1826]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
|
||
. However, a large lamelliform adenostyle situated proximally is not known from modern stylocellid genera. There it is often described as fringed or plumose and it is situated near the middle of the tarsus. The form and position of the adenostyle is in fact more alike those of most modern sironids or neogoveids, but this could be a plesiomorphic character for
|
||
<taxonomicName id="4C71B2D63947FD22887AF8B1FBC3F841" authorityName="Simon" authorityYear="1879" box="[856,1030,1908,1934]" class="Arachnida" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="444" phylum="Arthropoda" rank="subOrder" subOrder="Cyphophthalmi">Cyphophthalmi</taxonomicName>
|
||
not found in extant stylocellids. We choose to place this fossil in the family
|
||
<taxonomicName id="4C71B2D63947FD228977F85DFD2FF87D" authorityName="Hansen & Sorensen" authorityYear="1904" box="[597,746,1944,1970]" class="Arachnida" family="Stylocellidae" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="423" phylum="Arthropoda" rank="family">Stylocellidae</taxonomicName>
|
||
, which better fits the biogeographic distribution of the family and corresponds with the overall habitus of the specimen. This fossil is, however, placed in a new genus due to its unique adenostyle in combination with other characters which are otherwise stylocellid-like.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |