480 lines
52 KiB
XML
480 lines
52 KiB
XML
<document id="03F71D5CFCEB035332573BD8D58870B2" ID-DOI="10.5281/zenodo.6391953" ID-ISSN="1942-1354" ID-Zenodo-Dep="6391953" ID-ZooBank="B6862B03-E882-437A-B9E0-3B5FB1443602" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648504685522" checkinUser="felipe" docAuthor="Skelley, Paul E. & Keller, Oliver" docDate="2022" docId="03D5014CFFF3FFF7FF1EC9A33EB967AD" docLanguage="en" docName="InsectaMundi.2022.0916.1-11.pdf" docOrigin="Insecta Mundi 2022 (916)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Haroldiellus sallei" docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="FFEC7934FFF7FFF2FFDECD463F236519" masterDocTitle="A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="3" updateTime="1699308862729" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="1EE212A661B2B84F6D5E44A63519AE62">
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<mods:title id="6A5E041BE447914668C593323ECB4FF4">A third species of Haroldiellus Gordon and Skelley, 2007 from Mexico and Central America (Coleoptera: Scarabaeidae: Aphodiinae: Aphodiini)</mods:title>
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<mods:namePart id="C4779CE11E76A76ADB1615A01F777174">Skelley, Paul E.</mods:namePart>
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<mods:affiliation id="A54EDCAF40AB5D2DE9B5403DFEBD8F19">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:namePart id="2CECA78802DE352360358BF9A0A8913B">Keller, Oliver</mods:namePart>
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<mods:affiliation id="9FEED48936A93B6133AEB47A05BE0C06">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:title id="FADD030E41013E88AAB22C6804942837">Insecta Mundi</mods:title>
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<mods:part id="3AAC71D48A3B66085FB040CE42CE62A6">
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<mods:date id="5294FCF70BC662E5438EE7FFF0B8DFAF">2022</mods:date>
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<mods:number id="2F319177500B74AC17A7532B2D0D306D">2022-02-24</mods:number>
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<mods:number id="443F8A5613DC1506005AC92C92A6B326">2022</mods:number>
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<mods:identifier id="630AEF64381A344FE93D625A8DBC268B" type="DOI">10.5281/zenodo.6391953</mods:identifier>
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<mods:identifier id="F2C62F3D5353AA0C3703B60815EC141D" type="ISSN">1942-1354</mods:identifier>
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<treatment id="03D5014CFFF3FFF7FF1EC9A33EB967AD" LSID="urn:lsid:plazi:treatment:03D5014CFFF3FFF7FF1EC9A33EB967AD" httpUri="http://treatment.plazi.org/id/03D5014CFFF3FFF7FF1EC9A33EB967AD" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
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<subSubSection id="C366E3D1FFF3FFF6FF1EC9A33D3F601A" box="[192,540,1253,1283]" pageId="4" pageNumber="3" type="nomenclature">
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<paragraph id="8BC3B05AFFF3FFF6FF1EC9A33D3F601A" blockId="4.[192,540,1253,1316]" box="[192,540,1253,1283]" pageId="4" pageNumber="3">
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<heading id="D08B0736FFF3FFF6FF1EC9A33D3F601A" box="[192,540,1253,1283]" fontSize="12" level="2" pageId="4" pageNumber="3" reason="2">
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<taxonomicName id="4C7CCBD9FFF3FFF6FF1EC9A33D3F601A" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1863" box="[192,540,1253,1283]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallei">
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<emphasis id="B9086C48FFF3FFF6FF1EC9A33D3F601A" bold="true" box="[192,540,1253,1283]" pageId="4" pageNumber="3">
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<emphasis id="B9086C48FFF3FFF6FF1EC9A33E86601B" bold="true" box="[192,421,1253,1282]" italics="true" pageId="4" pageNumber="3">Haroldiellus sallei</emphasis>
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(Harold)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C366E3D1FFF3FFF6FF1EC84C3EA1603D" box="[192,386,1289,1316]" pageId="4" pageNumber="3" type="description">
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<paragraph id="8BC3B05AFFF3FFF6FF1EC84C3EA1603D" blockId="4.[192,540,1253,1316]" box="[192,386,1289,1316]" pageId="4" pageNumber="3">
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<figureCitation id="1347ACDFFFF3FFF6FF1EC84C3E61603D" box="[192,322,1290,1316]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="4" pageNumber="3">Figures 1, 4</figureCitation>
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,
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<figureCitation id="1347ACDFFFF3FFF6FE93C84F3E7F603A" box="[333,348,1289,1315]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="4" pageNumber="3">6</figureCitation>
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,
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<figureCitation id="1347ACDFFFF3FFF6FEB9C84C3EA1603D" box="[359,386,1290,1316]" captionStart="Figure 11" captionStartId="10.[192,268,1675,1701]" captionTargetBox="[192,1440,1021,1644]" captionTargetId="figure-87@10.[192,1440,1021,1644]" captionTargetPageId="10" captionText="Figure 11. Distribution map of Haroldiellus sallei (blue) as currently known from specimen labels and literature records. Map made with SimpleMappr (Shorthouse 2010)." figureDoi="http://doi.org/10.5281/zenodo.6391965" httpUri="https://zenodo.org/record/6391965/files/figure.png" pageId="4" pageNumber="3">11</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C366E3D1FFF3FFF6FF3EC8743C1D631B" pageId="4" pageNumber="3" type="reference_group">
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<paragraph id="8BC3B05AFFF3FFF6FF3EC8743D106070" blockId="4.[224,1440,1330,1538]" pageId="4" pageNumber="3">
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<treatmentCitation id="0ADD964BFFF3FFF6FF3EC8743D0E6053" author="Harold E." box="[224,557,1330,1354]" page="336" pageId="4" pageNumber="3" year="1863">
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<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC8743D0E6053" authority="Harold 1863: 336" authorityName="Harold" authorityPageNumber="336" authorityYear="1863" box="[224,557,1330,1354]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallei">
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<emphasis id="B9086C48FFF3FFF6FF3EC8743E546053" box="[224,375,1330,1354]" italics="true" pageId="4" pageNumber="3">Aphodius sallei</emphasis>
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<bibRefCitation id="EFEDCDABFFF3FFF6FEA0C8743D0E6053" author="Harold E." box="[382,557,1330,1354]" pageId="4" pageNumber="3" pagination="327 - 389" refId="ref5343" refString="Harold E. 1863. Beitrage zur Kenntnis einiger coprophagen Lamellicornien. (Funftes Stuck). Berliner Entomologische Zeitschrift 7: 327 - 389." type="journal article" year="1863">Harold 1863: 336</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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~
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<treatmentCitation id="0ADD964BFFF3FFF6FD97C8743C0D6053" author="Blackwelder RE" box="[585,814,1330,1354]" page="213" pageId="4" pageNumber="3" year="1944">
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<bibRefCitation id="EFEDCDABFFF3FFF6FD97C8743C0D6053" author="Blackwelder RE" box="[585,814,1330,1354]" pageId="4" pageNumber="3" pagination="189 - 341" refId="ref4964" refString="Blackwelder RE. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part II. Bulletin of the United States National Museum 185: i - iii + 189 - 341." type="journal article" year="1944">Blackwelder 1944: 213</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="0ADD964BFFF3FFF6FCE7C8743B1B6053" author="Dellacasa M & Gordon RD & Dellacasa G." box="[825,1080,1330,1354]" page="159" pageId="4" pageNumber="3" year="1998">
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<bibRefCitation id="EFEDCDABFFF3FFF6FCE7C8743B1B6053" author="Dellacasa M & Gordon RD & Dellacasa G." box="[825,1080,1330,1354]" pageId="4" pageNumber="3" pagination="139 - 162" refId="ref5126" refString="Dellacasa M, Gordon RD, Dellacasa G. 1998. The types of Aphodius species described by Harold from Mexico with description of a new genus. Acta Zoologica Mexicana 74: 139 - 162." type="journal article" year="1998">Dellacasa et al. 1998: 159</bibRefCitation>
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</treatmentCitation>
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(habitus, epipharynx and aedeagus figured;
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<typeStatus id="54C70EF8FFF3FFF6FE8BC8173E926070" box="[341,433,1361,1385]" pageId="4" pageNumber="3" type="lectotype">lectotype</typeStatus>
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designated).
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</paragraph>
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<paragraph id="8BC3B05AFFF3FFF6FF3EC8293CE6609E" blockId="4.[224,1440,1330,1538]" box="[224,965,1391,1415]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3EC8293CE6609E" box="[224,965,1391,1415]" pageId="4" pageNumber="3">
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<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC8293EC2609E" authority="(Harold)" baseAuthorityName="Harold" box="[224,481,1391,1415]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallaei">
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<emphasis id="B9086C48FFF3FFF6FF3EC8293EA1609E" box="[224,386,1391,1415]" italics="true" pageId="4" pageNumber="3">Aphodius sallaei</emphasis>
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||
(Harold)
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||
</taxonomicName>
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||
~
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||
<treatmentCitation id="0ADD964BFFF3FFF6FE25C8363DA9609E" author="Bates HW" box="[507,650,1391,1415]" page="83" pageId="4" pageNumber="3" year="1887">
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<bibRefCitation id="EFEDCDABFFF3FFF6FE25C8363DA9609E" author="Bates HW" box="[507,650,1391,1415]" pageId="4" pageNumber="3" pagination="25 - 160" refId="ref4902" refString="Bates HW. 1887. Biologia Centrali-Americana. Insecta. Coleoptera: Lamellicornia. (Copridae, Aphodiidae, Orphnidae, Hybosoridae, Geotrupidae, Trogidae, Aclopidae, Chasmatopteridae, Melolonthidae) 2: 25 - 160." type="journal article" year="1887">Bates 1887: 83</bibRefCitation>
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</treatmentCitation>
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(incorrect spelling); 1889: 391.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC3B05AFFF3FFF6FF3EC8C83B7360BF" blockId="4.[224,1440,1330,1538]" box="[224,1104,1422,1446]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3EC8C83B7360BF" box="[224,1104,1422,1446]" pageId="4" pageNumber="3">
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<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC8C83D1560BF" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1863" box="[224,566,1422,1446]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallei" subGenus="Bodilus">
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<emphasis id="B9086C48FFF3FFF6FF3EC8C83E1D60BF" box="[224,318,1422,1446]" italics="true" pageId="4" pageNumber="3">Aphodius</emphasis>
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(
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<emphasis id="B9086C48FFF3FFF6FE93C8C83EB460BF" box="[333,407,1422,1446]" italics="true" pageId="4" pageNumber="3">Bodilus</emphasis>
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)
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<emphasis id="B9086C48FFF3FFF6FE7BC8C83EF460BF" box="[421,471,1422,1446]" italics="true" pageId="4" pageNumber="3">sallei</emphasis>
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(Harold)
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</taxonomicName>
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~
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<treatmentCitation id="0ADD964BFFF3FFF6FD8EC8C83C2E60BF" author="Schmidt A." box="[592,781,1422,1446]" page="166" pageId="4" pageNumber="3" year="1913">
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<bibRefCitation id="EFEDCDABFFF3FFF6FD8EC8C83C2E60BF" author="Schmidt A." box="[592,781,1422,1446]" pageId="4" pageNumber="3" pagination="117 - 178" refId="ref5655" refString="Schmidt A. 1913. Erster Versuch einer Einteilung der exotischen Aphodien in Subgenera und als Anhang einige Neubeschreibungen. Archiv fur Naturgeschichte, Abteilung A 79: 117 - 178." type="journal article" year="1913">Schmidt 1913: 166</bibRefCitation>
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</treatmentCitation>
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||
;
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||
<treatmentCitation id="0ADD964BFFF3FFF6FCC9C8C83C5960BF" author="Schmidt A." box="[791,890,1422,1446]" page="304" pageId="4" pageNumber="3" year="1922">
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<bibRefCitation id="EFEDCDABFFF3FFF6FCC9C8C83C5960BF" author="Schmidt A." box="[791,890,1422,1446]" pageId="4" pageNumber="3" refId="ref5687" refString="Schmidt A. 1922. Coleoptera Aphodiinae. Das Tierreich, Vol. 45. Walter de Gruyter and Co.; Berlin and Leipzig, Germany. 614 p." type="book" year="1922">1922: 304</bibRefCitation>
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</treatmentCitation>
|
||
;
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||
<treatmentCitation id="0ADD964BFFF3FFF6FC5DC8C83B6F60BF" author="Dellacasa M." box="[899,1100,1422,1446]" page="373" pageId="4" pageNumber="3" year="1988">
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<bibRefCitation id="EFEDCDABFFF3FFF6FC5DC8C83B6F60BF" author="Dellacasa M." box="[899,1100,1422,1446]" pageId="4" pageNumber="3" pagination="1 - 455" refId="ref5091" refString="Dellacasa M. 1988 [1987]. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae Part I. Memorie della Societa Entomologica Italiana 66: 1 - 455." type="journal article" year="1988">Dellacasa 1988: 373</bibRefCitation>
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||
</treatmentCitation>
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||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC3B05AFFF3FFF6FF3EC8EA3DCD60DD" blockId="4.[224,1440,1330,1538]" box="[224,750,1452,1477]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3EC8EA3DCD60DD" box="[224,750,1452,1477]" pageId="4" pageNumber="3">
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<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC8EA3EF260DD" authority="(Harold)" baseAuthorityName="Harold" box="[224,465,1452,1476]" class="Insecta" family="Aphodiidae" genus="Agrilinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallei">
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||
<emphasis id="B9086C48FFF3FFF6FF3EC8EA3E5060DD" box="[224,371,1452,1476]" italics="true" pageId="4" pageNumber="3">Agrilinus sallei</emphasis>
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||
(Harold)
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||
</taxonomicName>
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||
~
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||
<treatmentCitation id="0ADD964BFFF3FFF6FE35C8EA3DC960DD" author="Dellacasa M & Gordon RD & Dellacasa G." box="[491,746,1452,1477]" page="159" pageId="4" pageNumber="3" year="2002">
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||
<bibRefCitation id="EFEDCDABFFF3FFF6FE35C8EA3DC960DD" author="Dellacasa M & Gordon RD & Dellacasa G." box="[491,746,1452,1477]" pageId="4" pageNumber="3" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. 2002: 159</bibRefCitation>
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</treatmentCitation>
|
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.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="8BC3B05AFFF3FFF6FF3EC88D3A1260FA" blockId="4.[224,1440,1330,1538]" box="[224,1329,1483,1507]" pageId="4" pageNumber="3">
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<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3EC88D3A1260FA" box="[224,1329,1483,1507]" pageId="4" pageNumber="3">
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<treatmentCitation id="0ADD964BFFF3FFF6FF3EC88D3D8E60FA" author="Balthasar V." box="[224,685,1483,1507]" page="164" pageId="4" pageNumber="3" year="1941">
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||
<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC88D3D8E60FA" authority="Balthasar 1941: 164" authorityName="Balthasar" authorityPageNumber="164" authorityYear="1941" box="[224,685,1483,1507]" class="Insecta" family="Aphodiidae" genus="Aphodius" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="freyi" subGenus="Agrilinus">
|
||
<emphasis id="B9086C48FFF3FFF6FF3EC88D3E1D60FA" box="[224,318,1483,1507]" italics="true" pageId="4" pageNumber="3">Aphodius</emphasis>
|
||
(
|
||
<emphasis id="B9086C48FFF3FFF6FE93C88D3E8B60FA" box="[333,424,1483,1507]" italics="true" pageId="4" pageNumber="3">Agrilinus</emphasis>
|
||
)
|
||
<emphasis id="B9086C48FFF3FFF6FE68C88D3EC060FA" box="[438,483,1483,1507]" italics="true" pageId="4" pageNumber="3">freyi</emphasis>
|
||
<bibRefCitation id="EFEDCDABFFF3FFF6FE36C88D3D8E60FA" author="Balthasar V." box="[488,685,1483,1507]" pageId="4" pageNumber="3" pagination="161 - 171" refId="ref4878" refString="Balthasar V. 1941. Neue Arten der coprophagen Scarabaeiden aus dem Hamburger Zoologischen Museum. Zoologischer Anzeiger 133: 161 - 171." type="journal article" year="1941">Balthasar 1941: 164</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
~ M.
|
||
<treatmentCitation id="0ADD964BFFF3FFF6FD37C88D3C9260FA" author="Dellacasa M." box="[745,945,1483,1507]" page="366" pageId="4" pageNumber="3" year="1988">
|
||
<bibRefCitation id="EFEDCDABFFF3FFF6FD37C88D3C9260FA" author="Dellacasa M." box="[745,945,1483,1507]" pageId="4" pageNumber="3" pagination="1 - 455" refId="ref5091" refString="Dellacasa M. 1988 [1987]. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae Part I. Memorie della Societa Entomologica Italiana 66: 1 - 455." type="journal article" year="1988">Dellacasa 1988: 366</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0ADD964BFFF3FFF6FC65C88D3B8960FA" author="Dellacasa M & Gordon RD & Dellacasa G." box="[955,1194,1483,1507]" page="15" pageId="4" pageNumber="3" year="2000">
|
||
<bibRefCitation id="EFEDCDABFFF3FFF6FC65C88D3B8960FA" author="Dellacasa M & Gordon RD & Dellacasa G." box="[955,1194,1483,1507]" pageId="4" pageNumber="3" pagination="11 - 15" refId="ref5163" refString="Dellacasa M, Gordon RD, Dellacasa G. 2000. Aphodius (Platyderides) pullatus A. Schmidt, 1913, another Mexican taxon to be moved into the genus Cephalocycluys and remarks on Cephalocyclus mexicanus (Harold, 1862) and Aphodius freyi Balthasar, 1941. Acta Zoologica Mexicana 79: 11 - 15." type="journal article" year="2000">Dellacasa et al. 2000: 15</bibRefCitation>
|
||
</treatmentCitation>
|
||
(synonymy).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC3B05AFFF3FFF6FF3EC8AC3C1D631B" blockId="4.[224,1440,1330,1538]" box="[224,830,1514,1538]" pageId="4" pageNumber="3">
|
||
<treatmentCitationGroup id="AB6C9774FFF3FFF6FF3EC8AC3C1D631B" box="[224,830,1514,1538]" pageId="4" pageNumber="3">
|
||
<taxonomicName id="4C7CCBD9FFF3FFF6FF3EC8AC3ECC631B" authority="(Harold)" baseAuthorityName="Harold" baseAuthorityYear="1863" box="[224,495,1514,1538]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B9086C48FFF3FFF6FF3EC8AC3EB3631B" box="[224,400,1514,1538]" italics="true" pageId="4" pageNumber="3">Haroldiellus sallei</emphasis>
|
||
(Harold)
|
||
</taxonomicName>
|
||
~
|
||
<treatmentCitation id="0ADD964BFFF3FFF6FDD7C8AC3C1A631B" author="Gordon RD & Skelley PE" box="[521,825,1514,1538]" page="270" pageId="4" pageNumber="3" year="2007">
|
||
<bibRefCitation id="EFEDCDABFFF3FFF6FDD7C8AC3C1A631B" author="Gordon RD & Skelley PE" box="[521,825,1514,1538]" pageId="4" pageNumber="3" pagination="1 - 580" refId="ref5257" refString="Gordon RD, Skelley PE. 2007. A monograph of the Aphodiini inhabiting the United States and Canada (Coleoptera: Scarabaeidae: Aphodiinae). Memoirs of the American Entomological Institute (Gainesville) 79: i - iv + 1 - 580." type="journal article" year="2007">Gordon and Skelley 2007: 270</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366E3D1FFF3FFF6FF1ECB543D9D6388" pageId="4" pageNumber="3" type="diagnosis">
|
||
<paragraph id="8BC3B05AFFF3FFF6FF1ECB543D9D6388" blockId="4.[191,1441,1554,1682]" pageId="4" pageNumber="3">
|
||
<emphasis id="B9086C48FFF3FFF6FF1ECB543E1F6335" bold="true" box="[192,316,1554,1580]" pageId="4" pageNumber="3">Diagnosis.</emphasis>
|
||
Length
|
||
<quantity id="4C841DBFFFF3FFF6FE46CB543D036335" box="[408,544,1554,1581]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.5" metricValueMin="3.5" pageId="4" pageNumber="3" unit="mm" value="4.5" valueMax="5.5" valueMin="3.5">3.5–5.5 mm</quantity>
|
||
, width 1.8–2.0. mm. A member of
|
||
<taxonomicName id="4C7CCBD9FFF3FFF6FC76CB543B0E6335" authorityName="Gordon and Skelley" authorityYear="2007" box="[936,1069,1554,1580]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9086C48FFF3FFF6FC76CB543B0E6335" box="[936,1069,1554,1580]" italics="true" pageId="4" pageNumber="3">Haroldiellus</emphasis>
|
||
</taxonomicName>
|
||
distinguished from other species by: Elytral striae deeply impressed, intervals convex; interval color pale lacking distinct marks (
|
||
<figureCitation id="1347ACDFFFF3FFF6FB29CB723A146357" box="[1271,1335,1588,1614]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
|
||
). Female lacking median frontal tubercle; male median frontal tubercle conical, not transversely widened (
|
||
<figureCitation id="1347ACDFFFF3FFF6FB30CB133A136369" box="[1262,1328,1621,1648]" captionStart="Figures 6–9" captionStartId="9.[195,282,992,1018]" captionTargetBox="[192,1440,213,962]" captionTargetId="figure-11@9.[192,1440,213,962]" captionTargetPageId="9" captionText="Figures 6–9. Heads of Haroldiellus species. 6) H. sallei. 7) H. lansbergei. 8) H. woodruffi female allotype. 9) H. woodruffi male holotype. Scale line = 0.5 mm (all figures)." figureDoi="http://doi.org/10.5281/zenodo.6391959" httpUri="https://zenodo.org/record/6391959/files/figure.png" pageId="4" pageNumber="3">Fig. 6</figureCitation>
|
||
). Protibia with dorsal surface lacking punctures (
|
||
<figureCitation id="1347ACDFFFF3FFF6FDB0CB313D936388" box="[622,688,1655,1681]" captionStart="Figures 1–5" captionStartId="8.[192,279,1279,1305]" captionTargetBox="[192,1440,213,1249]" captionTargetId="figure-66@8.[192,1440,213,1249]" captionTargetPageId="8" captionText="Figures 1–5. Haroldiellus species. 1) H. sallei dorsal habitus. 2) H. lansbergei dorsal habitus. 3) H. woodruffi, holotype, dorsal habitus. 4) H. sallei protibia dorsal surface with punctures. 5) H. woodruffi protibia dorsal surface without punctures. Scale line = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6391957" httpUri="https://zenodo.org/record/6391957/files/figure.png" pageId="4" pageNumber="3">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366E3D1FFF3FFF7FF1ECBE73CC96706" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="distribution">
|
||
<paragraph id="8BC3B05AFFF3FFF7FF1ECBE73D08644C" blockId="4.[192,1441,1697,1927]" lastBlockId="5.[191,1442,213,543]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">
|
||
<emphasis id="B9086C48FFF3FFF6FF1ECBE73E7963A2" bold="true" box="[192,346,1697,1723]" pageId="4" pageNumber="3">Distribution.</emphasis>
|
||
(
|
||
<figureCitation id="1347ACDFFFF3FFF6FEB3CBE73E9C63A2" box="[365,447,1697,1723]" captionStart="Figure 11" captionStartId="10.[192,268,1675,1701]" captionTargetBox="[192,1440,1021,1644]" captionTargetId="figure-87@10.[192,1440,1021,1644]" captionTargetPageId="10" captionText="Figure 11. Distribution map of Haroldiellus sallei (blue) as currently known from specimen labels and literature records. Map made with SimpleMappr (Shorthouse 2010)." figureDoi="http://doi.org/10.5281/zenodo.6391965" httpUri="https://zenodo.org/record/6391965/files/figure.png" pageId="4" pageNumber="3">Fig. 11</figureCitation>
|
||
).
|
||
<emphasis id="B9086C48FFF3FFF6FE09CBE73D2E63A2" bold="true" box="[471,525,1697,1723]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FE09CBE73D2E63A2" box="[471,525,1697,1723]" name="United States of America" pageId="4" pageNumber="3">USA</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDC1CBE73D5D63A2" box="[543,638,1697,1723]" country="United States of America" name="Arizona" pageId="4" pageNumber="3">Arizona</collectingRegion>
|
||
, south central
|
||
<collectingRegion id="49B87EB8FFF3FFF6FCF2CBE43C4C63A5" box="[812,879,1698,1724]" country="United States of America" name="Texas" pageId="4" pageNumber="3">Texas</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FC5ACBE73CFA63A2" bold="true" box="[900,985,1697,1723]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FC5ACBE73CFA63A2" box="[900,985,1697,1723]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC35CBE73B6A63A2" box="[1003,1097,1697,1723]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB86CBE43B9D63A5" box="[1112,1214,1698,1724]" country="Mexico" name="Durango" pageId="4" pageNumber="3">Durango</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB12CBE73ABF63A2" box="[1228,1436,1697,1723]" country="Mexico" name="Mexico" pageId="4" pageNumber="3">Estado de México</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FF1ECB853E0A63C4" box="[192,297,1731,1757]" country="Mexico" name="Guerrero" pageId="4" pageNumber="3">Guerrero</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FEE9CB853EB063C4" box="[311,403,1731,1757]" country="Mexico" name="Hidalgo" pageId="4" pageNumber="3">Hidalgo</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE7FCB853EC863C4" box="[417,491,1731,1757]" country="Mexico" name="Jalisco" pageId="4" pageNumber="3">Jalisco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE26CB853D5463C4" box="[504,631,1731,1757]" country="Mexico" name="Michoacan de Ocampo" pageId="4" pageNumber="3">Michoacán</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD58CB853DC763C4" box="[646,740,1731,1757]" country="Mexico" name="Morelos" pageId="4" pageNumber="3">Morelos</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD2CCB853C6463C4" box="[754,839,1731,1757]" country="Mexico" name="Nayarit" pageId="4" pageNumber="3">Nayarit</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC88CB853CC663C4" box="[854,997,1731,1757]" country="Mexico" name="Nuevo Leon" pageId="4" pageNumber="3">Nuevo León</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC2DCB853B6B63C4" box="[1011,1096,1731,1757]" country="Mexico" name="Oaxaca" pageId="4" pageNumber="3">Oaxaca</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB88CB853B8763C4" box="[1110,1188,1731,1757]" country="Mexico" name="Puebla" pageId="4" pageNumber="3">Puebla</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB6DCB853A0463C4" box="[1203,1319,1731,1757]" country="Mexico" name="Queretaro de Arteaga" pageId="4" pageNumber="3">Querétaro</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FAEBCB853FD363E6" country="Mexico" name="Quintana Roo" pageId="4" pageNumber="3">Quintana Roo</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FF23CBA33E8E63E6" box="[253,429,1765,1791]" country="Mexico" name="San Luis Potosi" pageId="4" pageNumber="3">San Luis Potosí</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE64CBA33D2F63E6" box="[442,524,1765,1791]" country="Mexico" name="Sonora" pageId="4" pageNumber="3">Sonora</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDC7CBA33D5663E6" box="[537,629,1765,1791]" country="Mexico" name="Tabasco" pageId="4" pageNumber="3">Tabasco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD5FCBA33C2063E6" box="[641,771,1765,1791]" country="Mexico" name="Tamaulipas" pageId="4" pageNumber="3">Tamaulipas</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FCCECBA33C5463E6" box="[784,887,1765,1791]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="4" pageNumber="3">Veracruz</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC5ACBA33CC163E6" box="[900,994,1765,1791]" country="Mexico" name="Yucatan" pageId="4" pageNumber="3">Yucatán</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FC29CBA23B5463E7" bold="true" box="[1015,1143,1764,1790]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FC29CBA23B5463E7" box="[1015,1143,1764,1790]" name="Guatemala" pageId="4" pageNumber="3">Guatemala</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB58CBA33A3E63E6" box="[1158,1309,1765,1791]" country="Guatemala" name="Alta Verapaz" pageId="4" pageNumber="3">Alta Verapaz</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FAF4CBA33FCA6238" country="Guatemala" name="Baja Verapaz" pageId="4" pageNumber="3">Baja Verapaz</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FF2ACA403E546238" box="[244,375,1798,1825]" country="Guatemala" name="El Progreso" pageId="4" pageNumber="3">El Progreso</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE5CCA403ECF6239" box="[386,492,1798,1824]" country="Guatemala" name="Escuintla" pageId="4" pageNumber="3">Escuintla</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE29CA403D506239" box="[503,627,1798,1824]" country="Guatemala" name="Guatemala" pageId="4" pageNumber="3">Guatemala</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDA3CA403DE16239" box="[637,706,1798,1824]" country="Guatemala" name="Izabal" pageId="4" pageNumber="3">Izabal</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD13CA403CA36239" box="[717,896,1798,1824]" country="Guatemala" name="Quetzaltenango" pageId="4" pageNumber="3">Quetzaltenango</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC54CA413B016238" box="[906,1058,1799,1825]" country="Guatemala" name="Sacatepequez" pageId="4" pageNumber="3">Sacatepéquez</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FBF2CA413B8A6238" box="[1068,1193,1799,1825]" country="Guatemala" name="Santa Rosa" pageId="4" pageNumber="3">Santa Rosa</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB6DCA413A2B6238" box="[1203,1288,1799,1825]" country="Guatemala" name="Zacapa" pageId="4" pageNumber="3">Zacapa</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FAC5CA413A826239" bold="true" box="[1307,1441,1798,1825]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FAC5CA413A826239" box="[1307,1441,1798,1825]" name="El Salvador" pageId="4" pageNumber="3">El Salvador</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FF16CA6E3E45625B" box="[200,358,1832,1858]" country="El Salvador" name="Chalatenango" pageId="4" pageNumber="3">Chalatenango</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FEAECA6E3EC2625B" box="[368,481,1832,1858]" country="El Salvador" name="Cuscatlan" pageId="4" pageNumber="3">Cuscatlán</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE35CA6F3D4D625B" box="[491,622,1832,1859]" country="El Salvador" name="La Libertad" pageId="4" pageNumber="3">La Libertad</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDA6CA6F3C2B625B" box="[632,776,1832,1859]" country="El Salvador" name="San Salvador" pageId="4" pageNumber="3">San Salvador</collectingRegion>
|
||
, San Vincente,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC6ECA6F3B05625A" box="[944,1062,1833,1859]" country="El Salvador" name="Santa Ana" pageId="4" pageNumber="3">Santa Ana</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FBE6CA6F3B8D625A" bold="true" box="[1080,1198,1833,1859]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FBE6CA6F3B8D625A" box="[1080,1198,1833,1859]" name="Honduras" pageId="4" pageNumber="3">Honduras</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB65CA6F3A67625A" box="[1211,1348,1833,1859]" country="Honduras" name="Comayagua" pageId="4" pageNumber="3">Comayagua</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FA90CA6E3AB8625B" box="[1358,1435,1832,1858]" country="Honduras" name="Copan" pageId="4" pageNumber="3">Copán</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FF1ECA0C3E2F627D" box="[192,268,1866,1892]" country="Honduras" name="Cortes" pageId="4" pageNumber="3">Cortés</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FEC8CA0C3EA4627D" box="[278,391,1866,1892]" country="Honduras" name="El Paraiso" pageId="4" pageNumber="3">El Paraíso</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FE4FCA0C3D4A627D" box="[401,617,1866,1892]" country="Honduras" name="Francisco Morazan" pageId="4" pageNumber="3">Francisco Morazán</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDAACA0C3DDE627D" box="[628,765,1866,1892]" country="Honduras" name="Ocotepeque" pageId="4" pageNumber="3">Ocotepeque</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FCD9CA0C3C48627C" box="[775,875,1866,1893]" country="Honduras" name="Olancho" pageId="4" pageNumber="3">Olancho</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FCA3CA0C3CD5627D" bold="true" box="[893,1014,1866,1892]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FCA3CA0C3CD5627D" box="[893,1014,1866,1892]" name="Nicaragua" pageId="4" pageNumber="3">Nicaragua</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FBDACA0C3B6E627D" box="[1028,1101,1866,1892]" country="Nicaragua" name="Boaco" pageId="4" pageNumber="3">Boaco</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB86CA0C3B96627D" box="[1112,1205,1866,1892]" country="Nicaragua" name="Jinotega" pageId="4" pageNumber="3">Jinotega</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB61CA0C3BD9627D" box="[1215,1274,1866,1892]" country="Nicaragua" name="Leon" pageId="4" pageNumber="3">León</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FADACA0D3A4C627C" box="[1284,1391,1867,1893]" country="Nicaragua" name="Managua" pageId="4" pageNumber="3">Managua</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FAA7CA0C3E05629E" country="Nicaragua" name="Rio San Juan" pageId="4" pageNumber="3">Río San Juan</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FEEDCA2A3E57629E" box="[307,372,1900,1927]" country="Nicaragua" name="Rivas" pageId="4" pageNumber="3">Rivas</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FE54CA2A3D29629F" bold="true" box="[394,522,1900,1926]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FE54CA2A3D29629F" box="[394,522,1900,1926]" name="Costa Rica" pageId="4" pageNumber="3">Costa Rica</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF3FFF6FDC6CA2A3D5B629F" box="[536,632,1900,1926]" country="Costa Rica" name="Alajuela" pageId="4" pageNumber="3">Alajuela</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD5ACA2A3DFC629F" box="[644,735,1900,1926]" country="Costa Rica" name="Cartago" pageId="4" pageNumber="3">Cartago</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FD32CA2A3C4C629F" box="[748,879,1900,1926]" country="Costa Rica" name="Guanacaste" pageId="4" pageNumber="3">Guanacaste</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FCA2CA2A3CFB629F" box="[892,984,1900,1926]" country="Costa Rica" name="Heredia" pageId="4" pageNumber="3">Heredia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FC3ACA2A3B12629F" box="[996,1073,1900,1926]" country="Costa Rica" name="Limon" pageId="4" pageNumber="3">Limón</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FBE0CA2B3B9D629E" box="[1086,1214,1901,1927]" country="Costa Rica" name="Puntarenas" pageId="4" pageNumber="3">Puntarenas</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF3FFF6FB15CA2A3A0D629F" box="[1227,1326,1900,1926]" country="Costa Rica" name="San Jose" pageId="4" pageNumber="3">San José</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF3FFF6FA9CCA2A3A83629F" bold="true" box="[1346,1440,1900,1926]" pageId="4" pageNumber="3">
|
||
<collectingCountry id="F36BF0CAFFF3FFF6FA9CCA2A3A83629F" box="[1346,1440,1900,1926]" name="Panama" pageId="4" pageNumber="3">Panama</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF2FFF7FF16CD903E0F65E9" box="[200,300,214,240]" country="Panama" name="Chiriqui" pageId="5" pageNumber="4">Chiriqui</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FEE6CD903E5965E9" box="[312,378,214,240]" country="Panama" name="Cocle" pageId="5" pageNumber="4">Coclé</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FE59CD903EC065E9" box="[391,483,214,240]" country="Panama" name="Panama" pageId="5" pageNumber="4">Panama</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FE31CD903D7A65E9" box="[495,601,214,240]" country="Panama" name="Veraguas" pageId="5" pageNumber="4">Veraguas</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF2FFF7FDB1CD933DC065F6" bold="true" box="[623,739,213,239]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FDB1CD933DC065F6" box="[623,739,213,239]" name="Colombia" pageId="5" pageNumber="4">Colombia</collectingCountry>
|
||
</emphasis>
|
||
(Valle de
|
||
<collectingRegion id="49B87EB8FFF2FFF7FC88CD903C8365E9" box="[854,928,214,240]" country="Colombia" name="Cauca" pageId="5" pageNumber="4">Cauca</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF2FFF7FC68CD933B3065F6" bold="true" box="[950,1043,213,239]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FC68CD933B3065F6" box="[950,1043,213,239]" name="Jamaica" pageId="5" pageNumber="4">Jamaica</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF2FFF7FBFFCD903BBF65E9" box="[1057,1180,214,240]" country="Jamaica" name="Clarendon" pageId="5" pageNumber="4">Clarendon</collectingRegion>
|
||
, Manchester,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FAE6CD903AB865E9" box="[1336,1435,214,240]" country="Jamaica" name="Portland" pageId="5" pageNumber="4">Portland</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FF1CCDB13E786408" box="[194,347,247,273]" country="Jamaica" name="Saint Andrew" pageId="5" pageNumber="4">Saint Andrew</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FEBACDB13EF6640B" box="[356,469,247,274]" country="Jamaica" name="Saint Ann" pageId="5" pageNumber="4">Saint Ann</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FE01CDB13DAD6408" box="[479,654,247,273]" country="Jamaica" name="Saint Catherine" pageId="5" pageNumber="4">Saint Catherine</collectingRegion>
|
||
, Saint Thomas,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FCE4CDB13C86640B" box="[826,933,247,274]" country="Jamaica" name="Trelawny" pageId="5" pageNumber="4">Trelawny</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF2FFF7FC6DCDB13CCC6408" bold="true" box="[947,1007,247,273]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FC6DCDB13CCC6408" box="[947,1007,247,273]" name="Haiti" pageId="5" pageNumber="4">Haiti</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF2FFF7FC23CDB13B556408" box="[1021,1142,247,273]" country="Haiti" name="Artibonite" pageId="5" pageNumber="4">Artibonite</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FB5ECDB13BE66408" box="[1152,1221,247,273]" country="Haiti" name="Ouest" pageId="5" pageNumber="4">Ouest</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FB10CDB13A09640B" box="[1230,1322,247,274]" country="Haiti" name="Sud-Est" pageId="5" pageNumber="4">Sud-Est</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF2FFF7FAE4CDB13E75642A" bold="true" pageId="5" pageNumber="4">
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FAE4CDB13E75642A" name="Dominican Republic" pageId="5" pageNumber="4">Dominican Republic</collectingCountry>
|
||
</emphasis>
|
||
(
|
||
<collectingRegion id="49B87EB8FFF2FFF7FEBDCC5F3E82642A" box="[355,417,281,307]" country="Dominican Republic" name="Azua" pageId="5" pageNumber="4">Azua</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FE74CC5F3D3B642A" box="[426,536,281,307]" country="Dominican Republic" name="Barahona" pageId="5" pageNumber="4">Barahona</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FDFCCC5F3D9B642A" box="[546,696,281,307]" country="Dominican Republic" name="La Altagracia" pageId="5" pageNumber="4">La Altagracia</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FD1FCC5F3C38642A" box="[705,795,281,307]" country="Dominican Republic" name="La Vega" pageId="5" pageNumber="4">La Vega</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FCFACC5F3C90642A" box="[804,947,281,307]" country="Dominican Republic" name="Monte Cristi" pageId="5" pageNumber="4">Monte Cristi</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FC62CC5F3B14642A" box="[956,1079,281,307]" country="Dominican Republic" name="Pedernales" pageId="5" pageNumber="4">Pedernales</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FB9FCC5F3BF4642A" box="[1089,1239,281,307]" country="Dominican Republic" name="San Cristobal" pageId="5" pageNumber="4">San Cristobal</collectingRegion>
|
||
,
|
||
<collectingRegion id="49B87EB8FFF2FFF7FB3FCC5F3AB7642D" box="[1249,1428,281,308]" country="Dominican Republic" name="Santo Domingo" pageId="5" pageNumber="4">Santo Domingo</collectingRegion>
|
||
);
|
||
<emphasis id="B9086C48FFF2FFF7FF61CC7C3EAE644C" bold="true" box="[191,397,314,341]" pageId="5" pageNumber="4">
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FF61CC7C3EAE644C" box="[191,397,314,341]" name="US Virgin Islands" pageId="5" pageNumber="4">US Virgin Islands</collectingCountry>
|
||
</emphasis>
|
||
(St. Thomas).
|
||
</paragraph>
|
||
<paragraph id="8BC3B05AFFF2FFF7FEDECC1A3CC96706" blockId="5.[191,1442,213,543]" pageId="5" pageNumber="4">
|
||
These are records repeated from
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FDB3CC1B3DC4646F" author="Bates HW" box="[621,743,348,375]" pageId="5" pageNumber="4" pagination="25 - 160" refId="ref4902" refString="Bates HW. 1887. Biologia Centrali-Americana. Insecta. Coleoptera: Lamellicornia. (Copridae, Aphodiidae, Orphnidae, Hybosoridae, Geotrupidae, Trogidae, Aclopidae, Chasmatopteridae, Melolonthidae) 2: 25 - 160." type="journal article" year="1887">Bates 1887</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FD2FCC1A3C0A646F" author="Bates HW" box="[753,809,348,374]" pageId="5" pageNumber="4" pagination="337 - 416" refId="ref4941" refString="Bates HW. 1889. Biologia Centrali-Americana. Insecta. Coleoptera: Lamellicornia (Suppl.) 2: 337 - 416." type="journal article" year="1889">1889</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FCEACC1A3CF8646F" author="Balthasar V." box="[820,987,348,374]" pageId="5" pageNumber="4" pagination="161 - 171" refId="ref4878" refString="Balthasar V. 1941. Neue Arten der coprophagen Scarabaeiden aus dem Hamburger Zoologischen Museum. Zoologischer Anzeiger 133: 161 - 171." type="journal article" year="1941">Balthasar 1941</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FC38CC1A3BE9646F" author="Dellacasa M & Gordon RD & Dellacasa G." box="[998,1226,348,375]" pageId="5" pageNumber="4" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. 2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FB0ACC1A3ABF646F" author="Skelley PE & Dellacasa M & Dellacasa G & Gordon RD" box="[1236,1436,348,375]" pageId="5" pageNumber="4" pagination="1 - 14" refId="ref5812" refString="Skelley PE, Dellacasa M, Dellacasa G, Gordon RD. 2007. Checklist of the Aphodiini of Mexico, Central and South America (Coleoptera: Scarabaeidae: Aphodiinae). Insecta Mundi 0014: 1 - 14." type="book chapter" year="2007">Skelley et al. 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FF1ECC383EC76481" author="Ramirez-Ponce A & Allende-Canseco J & Moron MA" box="[192,484,382,408]" pageId="5" pageNumber="4" pagination="323 - 343" refId="ref5576" refString="Ramirez-Ponce A, Allende-Canseco J, Moron MA. 2009. Fauna de Coleopteros Lamellicornios de Santiago Xiacui, Sierra Norte, Oaxaca, Mexico. Acta Zoologica Mexicana (nueva serie) 25 (2): 323 - 343." type="journal article" year="2009">Ramírez-Ponce et al. 2009</bibRefCitation>
|
||
; Cruz R. et al. 2012;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FD19CC383CB56481" author="Halffter G & Verdu JR & Moreno CE & Halffter V." box="[711,918,382,408]" pageId="5" pageNumber="4" pagination="54 - 60" refId="ref5303" refString="Halffter G, Verdu JR, Moreno CE, Halffter V. 2012. Historical and ecological determinants of dung beetle assemblages in two arid zones of central Mexico. Journal of Arid Environments 76: 54 - 60." type="journal article" year="2012">Halffter et al. 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FC7ECC383B9C6481" author="Lopez-Collado J. & Cruz-Rosales M & Vilaboa-Arroniz J & Martinez-Morales I & Gonzalez-Hernandez H." box="[928,1215,382,408]" pageId="5" pageNumber="4" pagination="78 - 87" refId="ref5395" refString="Lopez-Collado J. Cruz-Rosales M, Vilaboa-Arroniz J, Martinez-Morales I, Gonzalez-Hernandez H. 2017. Contributions of dung beetles to cattle productivity in the tropics: A stochastic-dynamic modeling approach. Agricultural Systems 155: 78 - 87." type="journal article" year="2017">López-Collado et al. 2017</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FB17CC383A6C6481" author="Minor P." box="[1225,1359,382,408]" pageId="5" pageNumber="4" pagination="493 - 514" refId="ref5476" refString="Minor P. 2017. New state records and updated checklist of Aphodiini and Eupariini (Coleoptera: Scarabaeidae: Aphodiinae) from Mexico. Zootaxa 4244 (4): 493 - 514." type="journal article" year="2017">Minor 2017</bibRefCitation>
|
||
; Perez- Gelabert 2020;
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FEB4CCD93C5564A0" author="Rivera-Gasperin SL & Escobar-Hernandez F." box="[362,886,415,441]" pageId="5" pageNumber="4" pagination="1 - 19" refId="ref5618" refString="Rivera-Gasperin SL, Escobar-Hernandez F. 2020. Especies de Scarabaeoidea (Coleoptera) del CICOLMA, Veracruz, Mexico. Acta Zoologica Mexicana (nueva serie) 36 (1): 1 - 19." type="journal article" year="2020">Rivera-Gasperín and Escobar-Hernández 2020</bibRefCitation>
|
||
; and
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FC6FCCE63B6164A0" author="Warner WB" box="[945,1090,415,442]" pageId="5" pageNumber="4" refId="ref5853" refString="Warner WB. 2022. Some new USA country and state records of Scarabaeidae (Coleoptera). The Coleopterists Bulletin (in press)." type="book" year="2022">
|
||
Warner
|
||
<quantity id="4C841DBFFFF2FFF7FBD2CCD93B6164A0" box="[1036,1090,415,441]" metricMagnitude="1" metricUnit="m" metricValue="5.135879999999999" pageId="5" pageNumber="4" unit="in" value="2022.0">2022</quantity>
|
||
</bibRefCitation>
|
||
<emphasis id="B9086C48FFF2FFF7FB96CCE63BB964A0" box="[1096,1178,416,441]" italics="true" pageId="5" pageNumber="4">in press</emphasis>
|
||
. Additional records are from museum specimens deposited in CMN; FSCA; LSAM; TAMU. In the
|
||
<collectingCountry id="F36BF0CAFFF2FFF7FBC2CC873B9764C2" box="[1052,1204,449,475]" name="United States of America" pageId="5" pageNumber="4">United States</collectingCountry>
|
||
, the only records we were able to confirm are from southern
|
||
<collectingRegion id="49B87EB8FFF2FFF7FDA8CCA53D9764E4" box="[630,692,483,509]" country="United States of America" name="Texas" pageId="5" pageNumber="4">Texas</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B87EB8FFF2FFF7FD36CCA43C6564E5" box="[744,838,482,508]" country="United States of America" name="Arizona" pageId="5" pageNumber="4">Arizona</collectingRegion>
|
||
. The distributional comments in
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FB6FCCA43A8164E4" author="Dellacasa M & Gordon RD & Dellacasa G." box="[1201,1442,482,509]" pageId="5" pageNumber="4" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. (2002)</bibRefCitation>
|
||
of “southern coast, from
|
||
<collectingRegion id="49B87EB8FFF2FFF7FE0DCF423D5F6707" box="[467,636,516,542]" country="United States of America" name="South Carolina" pageId="5" pageNumber="4">South Carolina</collectingRegion>
|
||
and
|
||
<collectingRegion id="49B87EB8FFF2FFF7FD6DCF423C276707" box="[691,772,516,542]" country="United States of America" name="Florida" pageId="5" pageNumber="4">Florida</collectingRegion>
|
||
to
|
||
<collectingRegion id="49B87EB8FFF2FFF7FCF6CF423C446707" box="[808,871,516,542]" country="United States of America" name="Texas" pageId="5" pageNumber="4">Texas</collectingRegion>
|
||
” is in error.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366E3D1FFF2FFF7FF1ECF6B3A496770" pageId="5" pageNumber="4" type="materials_examined">
|
||
<paragraph id="8BC3B05AFFF2FFF7FF1ECF6B3A496770" blockId="5.[192,1439,557,618]" pageId="5" pageNumber="4">
|
||
<emphasis id="B9086C48FFF2FFF7FF1ECF6B3E8E675E" bold="true" box="[192,429,557,583]" pageId="5" pageNumber="4">Materials examined.</emphasis>
|
||
Over the years, we have examined hundreds of specimens in the museums cited above plus many more. These data are too voluminous to present in this paper. Their distribution is abbreviated above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C366E3D1FFF2FFF7FF1ECF3F3EB967AD" pageId="5" pageNumber="4" type="discussion">
|
||
<paragraph id="8BC3B05AFFF2FFF7FF1ECF3F3EB967AD" blockId="5.[192,1440,633,693]" pageId="5" pageNumber="4">
|
||
<emphasis id="B9086C48FFF2FFF7FF1ECF3F3E0C678A" bold="true" box="[192,303,633,659]" pageId="5" pageNumber="4">Remarks.</emphasis>
|
||
As noted by
|
||
<bibRefCitation id="EFEDCDABFFF2FFF7FE1DCF3F3D9F678A" author="Dellacasa M & Gordon RD & Dellacasa G." box="[451,700,633,659]" pageId="5" pageNumber="4" pagination="155 - 223" refId="ref5221" refString="Dellacasa M, Gordon RD, Dellacasa G. 2002. Aphodiinae described or recorded by Bates in Biologia Centrali-Americana (Coleoptera Scarabaeoidea: Aphodiidae). Acta Zoologica Mexicana 86: 155 - 223." type="journal article" year="2002">Dellacasa et al. (2002)</bibRefCitation>
|
||
,
|
||
<taxonomicName id="4C7CCBD9FFF2FFF7FD16CF3C3C02678A" baseAuthorityName="Harold" baseAuthorityYear="1863" box="[712,801,633,659]" class="Insecta" family="Aphodiidae" genus="Haroldiellus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="sallei">
|
||
<emphasis id="B9086C48FFF2FFF7FD16CF3C3DFF678A" box="[712,732,634,659]" italics="true" pageId="5" pageNumber="4">H</emphasis>
|
||
.
|
||
<emphasis id="B9086C48FFF2FFF7FD34CF3F3C02678A" box="[746,801,633,659]" italics="true" pageId="5" pageNumber="4">sallei</emphasis>
|
||
</taxonomicName>
|
||
is a widespread common lowland species. It is often collected in cow dung.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |