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<mods:title id="FC08ECCAD7FD410ECF2986EB8933B571">The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)</mods:title>
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<treatment id="03D23F44F61D9D2BFF0754E7FADCC5F7" ID-DOI="http://doi.org/10.5281/zenodo.6079495" ID-GBIF-Taxon="137308715" ID-Zenodo-Dep="6079495" LSID="urn:lsid:plazi:treatment:03D23F44F61D9D2BFF0754E7FADCC5F7" httpUri="http://treatment.plazi.org/id/03D23F44F61D9D2BFF0754E7FADCC5F7" lastPageId="16" lastPageNumber="16" pageId="15" pageNumber="16">
<subSubSection id="C361DDD9F61D9D34FF0754E7FE8CC73B" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BC48E52F61D9D34FF0754E7FD4AC71A" blockId="15.[151,745,511,571]" box="[151,745,511,538]" pageId="15" pageNumber="16">
<heading id="D08C393EF61D9D34FF0754E7FD4AC71A" bold="true" box="[151,745,511,538]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<taxonomicName id="4C7BF5D1F61D9D34FF0754E7FD4AC71A" ID-CoL="MVYX" authority="Attems, 1951" authorityName="Attems" authorityYear="1951" box="[151,745,511,538]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pigmentatus">
<emphasis id="B90F5240F61D9D34FF0754E7FD4AC71A" bold="true" box="[151,745,511,538]" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF0754E7FEEAC719" bold="true" box="[151,329,511,537]" italics="true" pageId="15" pageNumber="16">Brachydesmus</emphasis>
(s. str.)
<emphasis id="B90F5240F61D9D34FE3B54E7FD9CC719" bold="true" box="[427,575,511,537]" italics="true" pageId="15" pageNumber="16">pigmentatus</emphasis>
<bibRefCitation id="EFEAF3A3F61D9D34FDD65718FD4AC71A" author="Attems" box="[582,745,512,538]" pageId="15" pageNumber="16" refString="Attems, C. (1951) Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Myriopoden vom Iran gesammelt von der Expedition Hainz Loffler und Genossen 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abt. I, 160, 387 - 426." type="journal article" year="1951">Attems, 1951</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BC48E52F61D9D34FF07573AFE8CC73B" blockId="15.[151,745,511,571]" box="[151,303,546,571]" pageId="15" pageNumber="16">
<figureCitation id="134092D7F61D9D34FF07573AFF7AC73B" box="[151,217,546,571]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
, Map 3
</paragraph>
</subSubSection>
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<paragraph id="8BC48E52F61D9D34FF075772FD3CC780" blockId="15.[151,1435,618,763]" box="[151,671,618,640]" pageId="15" pageNumber="16">
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<taxonomicName id="4C7BF5D1F61D9D34FF075772FDCCC780" ID-CoL="MVYX" authority="Attems, 1951: 407" authorityName="Attems" authorityPageNumber="407" authorityYear="1951" box="[151,623,618,640]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pigmentatus">
<emphasis id="B90F5240F61D9D34FF075772FE0EC780" box="[151,429,618,640]" italics="true" pageId="15" pageNumber="16">Brachydesmus pigmentatus</emphasis>
<treatmentCitation id="0ADAA843F61D9D34FE235773FDCCC780" author="Attems" box="[435,623,618,640]" page="407" pageId="15" pageNumber="16" year="1951">
<bibRefCitation id="EFEAF3A3F61D9D34FE235773FD9CC780" author="Attems" box="[435,575,618,640]" pageId="15" pageNumber="16" refString="Attems, C. (1951) Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Myriopoden vom Iran gesammelt von der Expedition Hainz Loffler und Genossen 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abt. I, 160, 387 - 426." type="journal article" year="1951">Attems, 1951</bibRefCitation>
: 407
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(D).
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<paragraph id="8BC48E52F61D9D34FF075791FBFEC79F" blockId="15.[151,1435,618,763]" box="[151,1117,649,671]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB6BA97CF61D9D34FF075791FBFEC79F" box="[151,1117,649,671]" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF075791FE60C79F" box="[151,451,649,671]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FF075791FE88C79F" ID-CoL="7NTRJ" authorityName="Golovatch &amp; Evsyukov &amp; Reip" authorityYear="2016" box="[151,299,649,671]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Brachydesmus</taxonomicName>
pigmentatus—
</emphasis>
<treatmentCitation id="0ADAA843F61D9D34FE535791FD0FC79F" author="Golovatch" box="[451,684,649,671]" page="426" pageId="15" pageNumber="16" year="1981">
<bibRefCitation id="EFEAF3A3F61D9D34FE535791FDDFC79F" author="Golovatch" box="[451,636,649,671]" pageId="15" pageNumber="16" refString="Golovatch, S. I. (1981 b) Diplopoda from Iran (Glomeridae, Glomeridellidae, Platydesmidae, Polydesmidae). Senckenbergiana biologica, 61 (5 / 6), 421 - 427." type="journal article" year="1981" yearSuffix="b">Golovatch, 1981b</bibRefCitation>
: 426
</treatmentCitation>
(D);
<treatmentCitation id="0ADAA843F61D9D34FD735791FB8DC79F" author="Enghoff" box="[739,1070,649,671]" page="69" pageId="15" pageNumber="16" year="2005">
<bibRefCitation id="EFEAF3A3F61D9D34FD735791FBAFC79F" author="Enghoff" box="[739,1036,649,671]" pageId="15" pageNumber="16" refString="Enghoff, H. &amp; Moravvej, S. A. (2005) A review of the millipede fauna of Iran. Zoology in the Middle East, 35, 61 - 72. http: // dx. doi. org / 10.1080 / 09397140.2005.10638104" type="journal article" year="2005">Enghoff and Moravvej, 2005</bibRefCitation>
: 69
</treatmentCitation>
(R).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC48E52F61D9D34FF0757B0FD4EC7BE" blockId="15.[151,1435,618,763]" box="[151,749,680,702]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB6BA97CF61D9D34FF0757B0FD4EC7BE" box="[151,749,680,702]" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FF0757B0FDD1C7BE" ID-CoL="882TX" authority="Golovatch, 1976: 1567" authorityName="Golovatch" authorityPageNumber="1567" authorityYear="1976" box="[151,626,680,702]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pereliae">
<emphasis id="B90F5240F61D9D34FF0757B0FE20C7BE" box="[151,387,680,702]" italics="true" pageId="15" pageNumber="16">Brachydesmus pereliae</emphasis>
<treatmentCitation id="0ADAA843F61D9D34FE1957B0FDD1C7BE" author="Golovatch" box="[393,626,680,702]" page="1567" pageId="15" pageNumber="16" year="1976">
<bibRefCitation id="EFEAF3A3F61D9D34FE1957B0FD95C7BE" author="Golovatch" box="[393,566,680,702]" pageId="15" pageNumber="16" refString="Golovatch, S. I. (1976) [New species of polydesmidans (Polydesmida, Diplopoda) of the USSR]. Zoologicheskii Zhurnal, 55 (10), 1567 - 1570. [in Russian, English abstract]" type="journal article" year="1976">Golovatch, 1976</bibRefCitation>
: 1567
</treatmentCitation>
</taxonomicName>
(D),
<emphasis id="B90F5240F61D9D34FD3857B0FD4EC7BE" bold="true" box="[680,749,680,702]" pageId="15" pageNumber="16">syn. n.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC48E52F61D9D34FF0757DEFF56C7FB" blockId="15.[151,1435,618,763]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB6BA97CF61D9D34FF0757DEFF56C7FB" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF0757DEFE39C7DC" box="[151,410,710,732]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FF0757DEFE88C7DC" ID-CoL="7NTRJ" authorityName="Golovatch &amp; Evsyukov &amp; Reip" authorityYear="2016" box="[151,299,710,732]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Brachydesmus</taxonomicName>
pereliae—
</emphasis>
<treatmentCitation id="0ADAA843F61D9D34FE0A57DEFD5DC7DC" author="Loksina" box="[410,766,710,732]" page="384" pageId="15" pageNumber="16" year="1979">
<bibRefCitation id="EFEAF3A3F61D9D34FE0A57DEFD6FC7DC" author="Loksina" box="[410,716,710,732]" pageId="15" pageNumber="16" refString="Loksina, I. E. &amp; Golovatch, S. I. (1979) Diplopoda of the USSR fauna. Pedobiologia, 19, 381 - 389." type="journal article" year="1979">Lokšina and Golovatch, 1979</bibRefCitation>
: 384
</treatmentCitation>
(M); Golovatch, 1979: 1323 (M);
<treatmentCitation id="0ADAA843F61D9D34FBF357DEFA38C7DC" author="Read" box="[1123,1435,710,732]" page="65" pageId="15" pageNumber="16" year="1994">
<bibRefCitation id="EFEAF3A3F61D9D34FBF357DEFAD4C7DC" author="Read" box="[1123,1399,710,732]" pageId="15" pageNumber="16" refString="Read, H. &amp; Golovatch, S. I. (1994) A review of the Central Asian millipede fauna. Bulletin of the British Myriapod Group, 10, 59 - 70." type="journal article" year="1994">Read and Golovatch, 1994</bibRefCitation>
: 65
</treatmentCitation>
(M).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61D9D34FF07563FFD83C230" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="8BC48E52F61D9D34FF07563FFDA7C05D" blockId="15.[151,1437,807,2021]" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF07563FFE22C640" bold="true" box="[151,385,807,832]" pageId="15" pageNumber="16">Material examined.</emphasis>
1 male (ZMUM),
<emphasis id="B90F5240F61D9D34FDC6563FFCAFC640" bold="true" box="[598,780,807,832]" pageId="15" pageNumber="16">AZERBAIJAN</emphasis>
, Talysh Mts, Farm Azerbaijan ca
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SE of Lerik,
<quantity id="4C8323B7F61D9D34FAE2563FFF0EC664" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="15" pageNumber="16" unit="m" value="550.0">550 m</quantity>
a.s.l.,
<emphasis id="B90F5240F61D9D34FF6D5655FDB8C663" box="[253,539,844,868]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FF6D5655FEFDC664" box="[253,350,845,868]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FEFC5655FE74C664" box="[364,471,845,868]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FE765654FDB8C663" box="[486,539,844,867]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
</emphasis>
etc. forest along stream, litter and rotten wood,
<date id="FFC5A892F61D9D34FBC65654FB6CC664" box="[1110,1231,844,868]" pageId="15" pageNumber="16" value="1983-10-12">12.X.1983</date>
; 4 juv. (ZMUM), Talysh Mts., Lerik Distr., Zuvand, foot of Mt Kyumyurkey, Oranchay Valley, xerophytous montane grassland,
<quantity id="4C8323B7F61D9D34FF07568CFE90C6AC" box="[151,307,916,941]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="15" pageNumber="16" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">
2100
<emphasis id="B90F5240F61D9D34FF5F568CFF7EC6AD" bold="true" box="[207,221,916,941]" pageId="15" pageNumber="16"></emphasis>
2200 m
</quantity>
a.s.l., under stones,
<date id="FFC5A892F61D9D34FD8A568CFD26C6AC" box="[538,645,916,940]" pageId="15" pageNumber="16" value="1983-10-09">9.X.1983</date>
, all leg. S. Golovatch; 1 juv. (ZMUM), Istisu SW of Astara,
<emphasis id="B90F5240F61D9D34FAA7568DFED3C6CF" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FAA7568DFF5AC6D0" authority=", Parrotia" authorityName="Parrotia" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus, Parrotia</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FE9856A0FED3C6CF" box="[264,368,952,975]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
</emphasis>
,
<taxonomicName id="4C7BF5D1F61D9D34FE1356A0FE1BC6CF" box="[387,440,952,975]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F5240F61D9D34FE1356A0FE1BC6CF" box="[387,440,952,975]" italics="true" pageId="15" pageNumber="16">Acer</emphasis>
</taxonomicName>
etc. forest,
<date id="FFC5A892F61D9D34FDD856A0FD3DC6D0" box="[584,670,952,976]" pageId="15" pageNumber="16" value="1978-10">X.1978</date>
, leg. A. Druk; 1 male (ZMUM), Lenkoran Distr., Hyrcan Nature Reserve, forest, rotten wood,
<date id="FFC5A892F61D9D34FE6056C4FDD3C6F4" box="[496,624,988,1012]" pageId="15" pageNumber="16" value="1971-03-19">19.III.1971</date>
, leg. N. Loginova; 1 male (ZMUM), Lenkoran Distr., Hyrcan Nature Reserve, litter,
<date id="FFC5A892F61D9D34FED656E7FE52C118" box="[326,497,1023,1048]" pageId="15" pageNumber="16" value="1985-01-26" valueMax="1985-02-04" valueMin="1985-01-26">
26.I.
<emphasis id="B90F5240F61D9D34FEEA56E7FE2BC118" bold="true" box="[378,392,1023,1048]" pageId="15" pageNumber="16"></emphasis>
4.II.1985
</date>
; 2 males, 2 females (ZMUM), same locality and data, all leg. A. Druk; 1 female (ZMUM), Azerbaijan, Lenkoran Distr., Hyrcan Nature Reserve, near Alekseevka,
<date id="FFC5A892F61D9D34FBD4513CFB60C13D" box="[1092,1219,1060,1085]" pageId="15" pageNumber="16" value="1985-04-21">21.IV.1985</date>
; 1 male, 2 females (ZMUM), same locality, Khan-Bulan River,
<date id="FFC5A892F61D9D34FD1C515FFCA8C160" box="[652,779,1095,1120]" pageId="15" pageNumber="16" value="1985-04-22">22.IV.1985</date>
; 1 female (ZMUM), same locality, Avrora, lowland forest,
<date id="FFC5A892F61D9D34FF075174FEB6C185" box="[151,277,1132,1157]" pageId="15" pageNumber="16" value="1985-04-21">21.IV.1985</date>
, all leg. E. Kupriyanova; 1 female (ZMUM), same locality, forest, litter,
<date id="FFC5A892F61D9D34FBAF5174FB4AC185" box="[1087,1257,1132,1157]" pageId="15" pageNumber="16" value="1985-01-26" valueMax="1985-02-04" valueMin="1985-01-26">
26.I.
<emphasis id="B90F5240F61D9D34FBE25174FB23C185" bold="true" box="[1138,1152,1132,1157]" pageId="15" pageNumber="16"></emphasis>
4.II.1985
</date>
, leg. A. Druk; 1 female, 2 juv. (ZMUM), same locality,
<quantity id="4C8323B7F61D9D34FDCF5197FD38C1A7" box="[607,667,1167,1192]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="m" value="50.0">50 m</quantity>
a.s.l.,
<emphasis id="B90F5240F61D9D34FD795188FB8CC1A7" box="[745,1071,1168,1192]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FD795188FC1AC1A8" authority=", Parrotia" authorityName="Parrotia" box="[745,953,1168,1192]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus, Parrotia</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FC575188FB8CC1A7" box="[967,1071,1168,1191]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
</emphasis>
etc. forest,
<date id="FFC5A892F61D9D34FB2B5188FA97C1A8" box="[1211,1332,1168,1192]" pageId="15" pageNumber="16" value="1983-10-13">13.X.1983</date>
; 4 males (ZMUM), same locality,
<quantity id="4C8323B7F61D9D34FE2151ACFE49C1CD" box="[433,490,1204,1229]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="km" value="8.0">8 km</quantity>
SW of Alekseevka, Apo,
<emphasis id="B90F5240F61D9D34FC8251ADFB37C1CC" box="[786,1172,1204,1228]" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FC8251ADFCD0C1CC" box="[786,883,1205,1228]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FCEE51ACFC10C1CB" box="[894,947,1204,1227]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FC2F51ADFB37C1CC" authority=", Parrotia" authorityName="Parrotia" box="[959,1172,1205,1228]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Carpinus, Parrotia</taxonomicName>
</emphasis>
etc. forest, litter, under stones and bark,
<date id="FFC5A892F61D9D34FEC851CFFE72C1F0" box="[344,465,1239,1264]" pageId="15" pageNumber="16" value="1983-10-16">16.X.1983</date>
; 4 females (ZMUM), same locality, Apo ca
<quantity id="4C8323B7F61D9D34FC5E51CFFBABC1F0" box="[974,1032,1239,1264]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="16" unit="km" value="6.0">6 km</quantity>
SW of Alekseevka,
<emphasis id="B90F5240F61D9D34FB6551C0FECEC014" italics="true" pageId="15" pageNumber="16">
<taxonomicName id="4C7BF5D1F61D9D34FB6551C0FAF5C1EF" box="[1269,1366,1240,1263]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FAF351C0FA3BC1EF" box="[1379,1432,1240,1263]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61D9D34FF0751E5FECEC014" authority=", Parrotia" authorityName="Parrotia" box="[151,365,1277,1300]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">Carpinus, Parrotia</taxonomicName>
</emphasis>
etc. forest, litter, under stones and bark,
<date id="FFC5A892F61D9D34FCA051E4FC58C015" box="[816,1019,1276,1301]" pageId="15" pageNumber="16" value="1983-10-14" valueMax="1983-10-16" valueMin="1983-10-14">14 and 16.X.1983</date>
; 1 female (ZMUM), Lenkoran Distr., Avrora, Moscow Forest,
<quantity id="4C8323B7F61D9D34FE3F5007FE4BC037" box="[431,488,1311,1336]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="15" pageNumber="16" unit="m" value="50.0">50 m</quantity>
a.s.l.,
<date id="FFC5A892F61D9D34FDA25007FD04C038" box="[562,679,1311,1336]" pageId="15" pageNumber="16" value="1996-06-01">1.VI.1996</date>
; 1 female (ZMUM), Lenkoran, Apo below Bilasar,
<quantity id="4C8323B7F61D9D34FB7F5007FA95C037" box="[1263,1334,1311,1336]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="m" value="350.0">350 m</quantity>
a.s.l.,
<date id="FFC5A892F61D9D34FA105038FEA9C05D" pageId="15" pageNumber="16" value="1996-06-08" valueMax="1996-06-09" valueMin="1996-06-08">
8
<emphasis id="B90F5240F61D9D34FA1E5007FA3FC038" bold="true" box="[1422,1436,1311,1336]" pageId="15" pageNumber="16"></emphasis>
9.VI.1996
</date>
, all leg. S. Golovatch.
</paragraph>
<paragraph id="8BC48E52F61D9D34FF57507FFE4DC0EC" blockId="15.[151,1437,807,2021]" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF57507FFE3EC080" bold="true" box="[199,413,1383,1408]" pageId="15" pageNumber="16">Descriptive notes.</emphasis>
Length ca
<quantity id="4C8323B7F61D9D34FDB15070FD3DC07F" box="[545,670,1383,1408]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="2.1" metricValueMin="1.3" pageId="15" pageNumber="16" unit="mm" value="17.0" valueMax="21.0" valueMin="13.0">
13
<emphasis id="B90F5240F61D9D34FDAD507FFDE8C080" bold="true" box="[573,587,1383,1408]" pageId="15" pageNumber="16"></emphasis>
21 mm
</quantity>
, width of midbody metazonae
<quantity id="4C8323B7F61D9D34FB945070FB2DC07F" box="[1028,1166,1383,1408]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="2.1" metricValueMin="1.2" pageId="15" pageNumber="16" unit="mm" value="1.65" valueMax="2.1" valueMin="1.2">
1.2
<emphasis id="B90F5240F61D9D34FBB7507FFB96C080" bold="true" box="[1063,1077,1383,1408]" pageId="15" pageNumber="16"></emphasis>
2.1 mm
</quantity>
(male, female), but the above samples ranging between 13 and
<quantity id="4C8323B7F61D9D34FDF25094FD10C0A4" box="[610,691,1420,1445]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="16" unit="mm" value="16.0">16 mm</quantity>
in length, and 1.8 and
<quantity id="4C8323B7F61D9D34FC2F5094FBB4C0A4" box="[959,1047,1420,1444]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="16" unit="mm" value="2.1">2.1 mm</quantity>
in width; metazonite to prozonite width ratio &lt;1.6 (
<figureCitation id="134092D7F61D9D34FEFF50B7FE55C0C8" box="[367,502,1455,1480]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">
Fig. 8
<emphasis id="B90F5240F61D9D34FE2550A8FE65C0C7" box="[437,454,1456,1479]" italics="true" pageId="15" pageNumber="16">A</emphasis>
<emphasis id="B90F5240F61D9D34FE5650B7FE77C0C8" bold="true" box="[454,468,1455,1480]" pageId="15" pageNumber="16"></emphasis>
8
<emphasis id="B90F5240F61D9D34FE7250A9FE55C0C8" box="[482,502,1457,1480]" italics="true" pageId="15" pageNumber="16">D</emphasis>
</figureCitation>
). Live coloration usually uniformly grey-brown to brown, in alcohol sometimes faded to light brown (
<figureCitation id="134092D7F61D9D34FE1A50CCFE7EC0EB" box="[394,477,1492,1517]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">
Fig. 8
<emphasis id="B90F5240F61D9D34FE5C50CCFE7EC0EB" box="[460,477,1492,1515]" italics="true" pageId="15" pageNumber="16">A</emphasis>
</figureCitation>
).
</paragraph>
<paragraph id="8BC48E52F61D9D34FF5750EFFD83C230" blockId="15.[151,1437,807,2021]" pageId="15" pageNumber="16">
All characters as in
<taxonomicName id="4C7BF5D1F61D9D34FE3450E1FD87C310" authorityName="Lohmander" authorityYear="1936" box="[420,548,1527,1552]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="assimilis">
<emphasis id="B90F5240F61D9D34FE3450E1FD87C310" box="[420,548,1527,1552]" italics="true" pageId="15" pageNumber="16">B. assimilis</emphasis>
</taxonomicName>
(
<figureCitation id="134092D7F61D9D34FDA350EFFDD5C310" box="[563,630,1527,1552]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
), except as follows. Paraterga subhorizontal to poorly declivous, mostly only slightly rounded laterally; caudolateral corner obtuse-angled only until segment 6, pointed or nearly pointed and increasingly well drawn behind rear tergal margin starting with segment 8 or 9 (
<figureCitation id="134092D7F61D9D34FBDA5327FB77C358" box="[1098,1236,1599,1624]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">
Fig. 8A
<emphasis id="B90F5240F61D9D34FB315327FB0CC358" bold="true" box="[1185,1199,1599,1624]" pageId="15" pageNumber="16"></emphasis>
8D
</figureCitation>
). Fore margins of paraterga mostly straight, anterolateral corner usually angular. Tergal setae a little longer, largely pointed. Male prefemora strongly bulging laterad. Gonopod (
<figureCitation id="134092D7F61D9D34FD3A539FFC8EC3A0" box="[682,813,1671,1696]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B D by S. Dashdamirov, scale bar: 1.0 mm (B D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="15" pageNumber="16">Fig. 8E, 8F</figureCitation>
) subfalcate, bipartite, with a long, very simple, fingershaped, ventrad only slightly and regularly curved, apically acuminate exomere (
<emphasis id="B90F5240F61D9D34FBA153B4FBEFC3C5" bold="true" box="[1073,1100,1708,1733]" pageId="15" pageNumber="16">ex</emphasis>
), this being tightly appressed ventrally to a similar, also finger-shaped, but somewhat shorter endomere (
<emphasis id="B90F5240F61D9D34FC7853D7FBA7C3E8" bold="true" box="[1000,1028,1743,1768]" pageId="15" pageNumber="16">en</emphasis>
), the latter at base carrying a small, medioventral spine (
<emphasis id="B90F5240F61D9D34FE1A53ECFE3AC20D" bold="true" box="[394,409,1780,1805]" pageId="15" pageNumber="16">k</emphasis>
);
<emphasis id="B90F5240F61D9D34FE2853ECFE77C20D" bold="true" box="[440,468,1780,1805]" pageId="15" pageNumber="16">en</emphasis>
representing a solenomere devoid of a hairy pulvillus; a small tooth (
<emphasis id="B90F5240F61D9D34FABB53ECFA97C20D" bold="true" box="[1323,1332,1780,1805]" pageId="15" pageNumber="16">t</emphasis>
) located medioventrally at base of femorite.
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61D9D2BFF575224FADCC5F7" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BC48E52F61D9D34FF575224FC27C29C" blockId="15.[151,1437,807,2021]" pageId="15" pageNumber="16">
<emphasis id="B90F5240F61D9D34FF575224FE98C255" bold="true" box="[199,315,1852,1877]" pageId="15" pageNumber="16">Remarks.</emphasis>
This species has hitherto been considered as subendemic to Hyrcania within N and NW
<collectingCountry id="F36CCEC2F61D9D34FABB5225FAFBC254" box="[1323,1368,1853,1876]" name="Iran" pageId="15" pageNumber="16">Iran</collectingCountry>
alone (
<collectingRegion id="49BF40B0F61D9D34FF0E5247FF4AC278" box="[158,233,1887,1912]" country="Iran" name="Alborz" pageId="15" pageNumber="16">Elburs</collectingRegion>
and Zagros mountains) (
<bibRefCitation id="EFEAF3A3F61D9D34FD965278FD3FC278" author="Attems" box="[518,668,1887,1912]" pageId="15" pageNumber="16" refString="Attems, C. (1951) Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Myriopoden vom Iran gesammelt von der Expedition Hainz Loffler und Genossen 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abt. I, 160, 387 - 426." type="journal article" year="1951">Attems 1951</bibRefCitation>
;
<bibRefCitation id="EFEAF3A3F61D9D34FD395247FCCCC278" author="Golovatch" box="[681,879,1887,1912]" pageId="15" pageNumber="16" refString="Golovatch, S. I. (1981 b) Diplopoda from Iran (Glomeridae, Glomeridellidae, Platydesmidae, Polydesmidae). Senckenbergiana biologica, 61 (5 / 6), 421 - 427." type="journal article" year="1981" yearSuffix="b">Golovatch 1981b</bibRefCitation>
). The above records are thus new to the fauna of the
<collectingCountry id="F36CCEC2F61D9D34FF50529CFE6AC29D" box="[192,457,1924,1949]" name="Azerbaijan" pageId="15" pageNumber="16">
Republic of
<collectingRegion id="49BF40B0F61D9D34FEDB529CFE6AC29D" box="[331,457,1924,1949]" country="Iran" name="Azarbayjan-e Sharqi" pageId="15" pageNumber="16">Azerbaijan</collectingRegion>
</collectingCountry>
, in general also to that of the Caucasus.
</paragraph>
<paragraph id="8BC48E52F61D9D2BFF5752B0FADCC5F7" blockId="15.[151,1437,807,2021]" lastBlockId="16.[151,1436,151,248]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Given such a considerable range of variation in body size in
<taxonomicName id="4C7BF5D1F61D9D34FC1752B1FB92C2C0" authorityName="Attems" authorityYear="1951" box="[903,1073,1961,1984]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pigmentatus">
<emphasis id="B90F5240F61D9D34FC1752B1FB92C2C0" box="[903,1073,1961,1984]" italics="true" pageId="15" pageNumber="16">B. pigmentatus</emphasis>
</taxonomicName>
, we do not hesitate to formally synonymize here this species with
<taxonomicName id="4C7BF5D1F61D9D34FDBC52D5FCD0C2E5" authority="Golovatch, 1976" authorityName="Golovatch" authorityYear="1976" box="[556,883,1996,2021]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="pereliae">
<emphasis id="B90F5240F61D9D34FDBC52D5FD0AC2E5" box="[556,681,1996,2021]" italics="true" pageId="15" pageNumber="16">B. pereliae</emphasis>
<bibRefCitation id="EFEAF3A3F61D9D34FD2352D4FCD0C2E5" author="Golovatch" box="[691,883,1996,2021]" pageId="15" pageNumber="16" refString="Golovatch, S. I. (1976) [New species of polydesmidans (Polydesmida, Diplopoda) of the USSR]. Zoologicheskii Zhurnal, 55 (10), 1567 - 1570. [in Russian, English abstract]" type="journal article" year="1976">Golovatch, 1976</bibRefCitation>
</taxonomicName>
, the latter taxon known from the W Kopetdagh Mountains, SW
<collectingCountry id="F36CCEC2F6029D2BFEDF558FFE4EC5B0" box="[335,493,151,176]" name="Turkmenistan" pageId="16" pageNumber="17">Turkmenistan</collectingCountry>
. Indeed, the
<typeStatus id="54C030F0F6029D2BFDEE558FFD43C5B0" box="[638,736,151,176]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7BF5D1F6029D2BFC965581FC22C5B0" authorityName="Golovatch" authorityYear="1976" box="[774,897,151,176]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pereliae">
<emphasis id="B90F5240F6029D2BFC965581FC22C5B0" box="[774,897,151,176]" italics="true" pageId="16" pageNumber="17">B. pereliae</emphasis>
</taxonomicName>
was said to be
<quantity id="4C8323B7F6029D2BFBA15580FB22C5AF" box="[1073,1153,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="10.0">10 mm</quantity>
long and
<quantity id="4C8323B7F6029D2BFB625580FAF4C5AF" box="[1266,1367,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="16" pageNumber="17" unit="mm" value="1.17">1.17 mm</quantity>
wide, whereas the gonopods of
<taxonomicName id="4C7BF5D1F6029D2BFE2C55A5FDC5C5D4" authorityName="Attems" authorityYear="1951" box="[444,614,189,212]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pigmentatus">
<emphasis id="B90F5240F6029D2BFE2C55A5FDC5C5D4" box="[444,614,189,212]" italics="true" pageId="16" pageNumber="17">B. pigmentatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7BF5D1F6029D2BFD3055A5FCB8C5D5" authorityName="Golovatch" authorityYear="1976" box="[672,795,188,213]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pereliae">
<emphasis id="B90F5240F6029D2BFD3055A5FCB8C5D5" box="[672,795,188,213]" italics="true" pageId="16" pageNumber="17">B. pereliae</emphasis>
</taxonomicName>
have been noted as being virtually identical (
<bibRefCitation id="EFEAF3A3F6029D2BFAB655A4FF70C5F8" author="Golovatch" pageId="16" pageNumber="17" refString="Golovatch, S. I. (1976) [New species of polydesmidans (Polydesmida, Diplopoda) of the USSR]. Zoologicheskii Zhurnal, 55 (10), 1567 - 1570. [in Russian, English abstract]" type="journal article" year="1976">Golovatch 1976</bibRefCitation>
). Hence the new synonymy advanced:
<taxonomicName id="4C7BF5D1F6029D2BFD1855F9FC6EC5F8" authority="Attems, 1951" authorityName="Attems" authorityYear="1951" box="[648,973,223,248]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pigmentatus">
<emphasis id="B90F5240F6029D2BFD1855F9FC93C5F8" box="[648,816,225,248]" italics="true" pageId="16" pageNumber="17">B. pigmentatus</emphasis>
<bibRefCitation id="EFEAF3A3F6029D2BFCA755F8FC6EC5F8" author="Attems" box="[823,973,223,248]" pageId="16" pageNumber="17" refString="Attems, C. (1951) Ergebnisse der Osterreichischen Iran-Expedition 1949 / 50. Myriopoden vom Iran gesammelt von der Expedition Hainz Loffler und Genossen 1949 / 50. Sitzungsberichte der Osterreichischen Akademie der Wissenschaften, Mathematisch-naturwissenschaftliche Klasse, Abt. I, 160, 387 - 426." type="journal article" year="1951">Attems, 1951</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C7BF5D1F6029D2BFC7B55F9FA89C5F8" authority="Golovatch, 1976" authorityName="Golovatch" authorityYear="1976" box="[1003,1322,223,248]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="pereliae">
<emphasis id="B90F5240F6029D2BFC7B55F9FBC6C5F8" box="[1003,1125,223,248]" italics="true" pageId="16" pageNumber="17">B. pereliae</emphasis>
<bibRefCitation id="EFEAF3A3F6029D2BFBFC55C7FA89C5F8" author="Golovatch" box="[1132,1322,223,248]" pageId="16" pageNumber="17" refString="Golovatch, S. I. (1976) [New species of polydesmidans (Polydesmida, Diplopoda) of the USSR]. Zoologicheskii Zhurnal, 55 (10), 1567 - 1570. [in Russian, English abstract]" type="journal article" year="1976">Golovatch, 1976</bibRefCitation>
</taxonomicName>
,
<taxonomicNameLabel id="A23CEF3BF6029D2BFAA555F8FADCC5F7" box="[1333,1407,224,248]" pageId="16" pageNumber="17">syn. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>