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<mods:title id="712C98F9B0FC8B6733FF8C5F37CF210C">The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)</mods:title>
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<mods:namePart id="3DE1E4CF0819FB94EBCA3A39BA3BDD6E">Reip, Hans</mods:namePart>
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<treatment id="03D23F44F61B9D31FF07531BFE95C718" ID-DOI="http://doi.org/10.5281/zenodo.6079488" ID-GBIF-Taxon="137308726" ID-Zenodo-Dep="6079488" LSID="urn:lsid:plazi:treatment:03D23F44F61B9D31FF07531BFE95C718" httpUri="http://treatment.plazi.org/id/03D23F44F61B9D31FF07531BFE95C718" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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<paragraph id="8BC48E52F61B9D32FF07531BFD5DC31E" blockId="9.[151,766,1539,1599]" box="[151,766,1539,1566]" pageId="9" pageNumber="10">
<heading id="D08C393EF61B9D32FF07531BFD5DC31E" bold="true" box="[151,766,1539,1566]" fontSize="11" level="1" pageId="9" pageNumber="10" reason="1">
<emphasis id="B90F5240F61B9D32FF07531BFD5DC31E" bold="true" box="[151,766,1539,1566]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF07531BFE85C31D" bold="true" box="[151,294,1539,1565]" italics="true" pageId="9" pageNumber="10">Polydesmus</emphasis>
<taxonomicName id="4C7BF5D1F61B9D32FEBD531CFD5DC31E" authority="Lohmander, 1936" authorityName="Lohmander" authorityYear="1936" box="[301,766,1539,1566]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="muralewiczi">
(s. str.)
<emphasis id="B90F5240F61B9D32FE19531BFDBFC31D" bold="true" box="[393,540,1539,1565]" italics="true" pageId="9" pageNumber="10">muralewiczi</emphasis>
<bibRefCitation id="EFEAF3A3F61B9D32FDB3531CFD5DC31E" author="Lohmander" box="[547,766,1540,1566]" pageId="9" pageNumber="10" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungl. Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B, 5 (1), 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC48E52F61B9D32FF07533EFE5FC33F" blockId="9.[151,766,1539,1599]" box="[151,508,1574,1599]" pageId="9" pageNumber="10">
<figureCitation id="134092D7F61B9D32FF07533EFEEAC33F" box="[151,329,1574,1599]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="9" pageNumber="10">
Figs 3A, 3H
<emphasis id="B90F5240F61B9D32FEB0533EFE8DC33F" bold="true" box="[288,302,1574,1599]" pageId="9" pageNumber="10"></emphasis>
3J
</figureCitation>
,
<figureCitation id="134092D7F61B9D32FEC5533FFE0AC33F" box="[341,425,1575,1599]" captionStart="FIGURE 4" captionStartId="8.[151,249,1796,1818]" captionTargetBox="[198,1381,227,1769]" captionTargetId="figure@8.[198,1388,193,1774]" captionTargetPageId="8" captionText="FIGURE 4. Left gonopods of Polydesmus abchasius Attems, 1898, P. lignaui Lohmander, 1936 and P. muralewiczi Lohmander, 1936 (A C), and right gonopod of P. muralewiczi Lohmander, 1936 (D). A: male from near Lake Ritsa, Abkhazia, mesal view. B: male from Sukhumi Botanical Garden, Abkhazia, mesal view. C: male from near Kursavka, Stavropol Prov., Russia, mesal view. D: male lectotype from near Psebai, Krasnodar Prov., Russia (after Lohmander 1936). Designations: en, endomere; ex, exomere; d, exomere tooth; pu, hairy pulvillus; r, endomere rod. Scale bar: 0.2 mm (A C), not to scale (D)." httpUri="https://zenodo.org/record/1052389/files/figure.png" pageId="9" pageNumber="10">4C, 4D</figureCitation>
, Map 1
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61B9D32FF07537FFD11C37D" box="[151,690,1639,1661]" pageId="9" pageNumber="10" type="description">
<paragraph id="8BC48E52F61B9D32FF07537FFD11C37D" blockId="9.[151,1435,1639,1784]" box="[151,690,1639,1661]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB6BA97CF61B9D32FF07537FFD11C37D" box="[151,690,1639,1661]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF07537FFE13C37D" box="[151,432,1639,1661]" italics="true" pageId="9" pageNumber="10">
Polydesmus
<taxonomicName id="4C7BF5D1F61B9D32FE86537FFE34C37D" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[278,407,1639,1661]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="complanatus">complanatus</taxonomicName>
<emphasis id="B90F5240F61B9D32FE075370FE13C37D" bold="true" box="[407,432,1640,1661]" italics="true" pageId="9" pageNumber="10"></emphasis>
</emphasis>
<treatmentCitation id="0ADAA843F61B9D32FE21537FFD20C37D" author="Muralewicz" box="[433,643,1639,1661]" page="7" pageId="9" pageNumber="10" year="1927">
<bibRefCitation id="EFEAF3A3F61B9D32FE21537FFDCEC37D" author="Muralewicz" box="[433,621,1639,1661]" pageId="9" pageNumber="10" refString="Muralewicz, W. S. (1927) [To the fauna of Myriopoda of the Caucasus]. Acta Societatis Entomologicae Stauropolitanae [Izvestiya Stavropolskogo entomologicheskogo obshchestva], 3 (1), 1 - 7. [Stavropol, in Russian]" type="journal article" year="1927">Muralewicz, 1927</bibRefCitation>
: 7
</treatmentCitation>
(R).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61B9D32FF07539EFCC5C3BB" pageId="9" pageNumber="10" type="reference_group">
<paragraph id="8BC48E52F61B9D32FF07539EFC86C39C" blockId="9.[151,1435,1639,1784]" box="[151,805,1670,1692]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB6BA97CF61B9D32FF07539EFC86C39C" box="[151,805,1670,1692]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF07539EFEB3C39C" box="[151,272,1670,1692]" italics="true" pageId="9" pageNumber="10">Polydesmus</emphasis>
<taxonomicName id="4C7BF5D1F61B9D32FE86539EFD56C39C" authority="Lohmander, 1936: 9" authorityName="Lohmander" authorityPageNumber="9" authorityYear="1936" box="[278,757,1670,1692]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="muralewiczi" subGenus="Polydesmus">
(
<emphasis id="B90F5240F61B9D32FE8E539EFE34C39C" box="[286,407,1670,1692]" italics="true" pageId="9" pageNumber="10">Polydesmus</emphasis>
)
<emphasis id="B90F5240F61B9D32FE36539EFD81C39C" box="[422,546,1670,1692]" italics="true" pageId="9" pageNumber="10">muralewiczi</emphasis>
<treatmentCitation id="0ADAA843F61B9D32FDB8539EFD56C39C" author="Lohmander" box="[552,757,1670,1692]" page="9" pageId="9" pageNumber="10" year="1936">
<bibRefCitation id="EFEAF3A3F61B9D32FDB8539EFD7CC39C" author="Lohmander" box="[552,735,1670,1692]" pageId="9" pageNumber="10" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungl. Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B, 5 (1), 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
(D).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC48E52F61B9D32FF0753BDFCC5C3BB" blockId="9.[151,1435,1639,1784]" box="[151,870,1701,1723]" pageId="9" pageNumber="10">
<treatmentCitationGroup id="AB6BA97CF61B9D32FF0753BDFCC5C3BB" box="[151,870,1701,1723]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF0753BDFEB3C3BB" box="[151,272,1701,1723]" italics="true" pageId="9" pageNumber="10">Polydesmus</emphasis>
<taxonomicName id="4C7BF5D1F61B9D32FE8653BDFE4DC3BB" authorityName="Lohmander" authorityYear="1936" box="[278,494,1701,1723]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="lignaui" subGenus="Polydesmus">
(
<emphasis id="B90F5240F61B9D32FE8E53BDFE34C3BB" box="[286,407,1701,1723]" italics="true" pageId="9" pageNumber="10">Polydesmus</emphasis>
)
<emphasis id="B90F5240F61B9D32FE3653BDFE4DC3BB" box="[422,494,1701,1723]" italics="true" pageId="9" pageNumber="10">lignaui</emphasis>
</taxonomicName>
<emphasis id="B90F5240F61B9D32FE6453BDFD25C3BB" box="[500,646,1701,1723]" italics="true" pageId="9" pageNumber="10">muralewiczi—</emphasis>
<treatmentCitation id="0ADAA843F61B9D32FD1653BEFC96C3BB" author="Attems" box="[646,821,1701,1723]" page="21" pageId="9" pageNumber="10" year="1940">
<bibRefCitation id="EFEAF3A3F61B9D32FD1653BEFCB1C3BB" author="Attems" box="[646,786,1701,1723]" pageId="9" pageNumber="10" refString="Attems, C. (1940) Myriapoda 3. Polydesmoidea III. Fam. Polydesmidae, Vanhoeffeniidae, Cryptodesmidae, Oniscodesmidae, Sphaerotrichopidae, Peridontodesmidae, Rhachidesmidae, Macellolophidae, Pandirodesmidae. Das Tierreich, 70, i - xvii + 1 - 577. http: // dx. doi. org / 10.1515 / 9783111609645.1" type="journal article" year="1940">Attems, 1940</bibRefCitation>
: 21
</treatmentCitation>
(D).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61B9D32FF0753DCFD67C3F8" pageId="9" pageNumber="10" type="description">
<paragraph id="8BC48E52F61B9D32FF0753DCFD67C3F8" blockId="9.[151,1435,1639,1784]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF0753DCFE0DC3DA" box="[151,430,1732,1754]" italics="true" pageId="9" pageNumber="10">Polydesmus muralewiczi—</emphasis>
<bibRefCitation id="EFEAF3A3F61B9D32FE3E53DDFD88C3DA" author="Lang" box="[430,555,1732,1754]" pageId="9" pageNumber="10" refString="Lang, J. (1959) [To the knowledge of millipedes (Diplopoda) of the USSR territory] // Zoologicheskii Zhurnal, 38 (12), 1790 - 1796. [in Russian, a summary in English]" type="journal article" year="1959">Lang, 1959</bibRefCitation>
: 1790 (M);
<bibRefCitation id="EFEAF3A3F61B9D32FD2353DCFCD6C3DA" author="Kobakhidze" box="[691,885,1732,1754]" pageId="9" pageNumber="10" refString="Kobakhidze, D. N. (1965) [A list of millipedes (Diplopoda) of SSR Georgia]. Fragmenta Faunistica, 11 (21), 390 - 398. [in Russian, summaries in Polish and German]" type="journal article" year="1965">Kobakhidze, 1965</bibRefCitation>
: 391 (R);
<bibRefCitation id="EFEAF3A3F61B9D32FC7B53DCFA88C3DA" author="Loksina" box="[1003,1323,1732,1754]" pageId="9" pageNumber="10" refString="Loksina, I. E. &amp; Golovatch, S. I. (1979) Diplopoda of the USSR fauna. Pedobiologia, 19, 381 - 389." type="journal article" year="1979">Lokšina and Golovatch, 1979</bibRefCitation>
: 384 (M);
<bibRefCitation id="EFEAF3A3F61B9D32FF5753FAFEC8C3F8" author="Talikadze" box="[199,363,1762,1784]" pageId="9" pageNumber="10" refString="Talikadze, D. A. (1984) [On the millipede fauna (Diplopoda) of the Colchidan Province of the Caucasus]. Zoologicheskii Zhurnal, 63 (1), 124 - 145. [in Russian, a summary in English]" type="journal article" year="1984">Talikadze, 1984</bibRefCitation>
: 143 (R);
<bibRefCitation id="EFEAF3A3F61B9D32FE4153FBFDE4C3F8" author="Zuev" box="[465,583,1762,1784]" pageId="9" pageNumber="10" refString="Zuev, R. V. (2014) Preliminary data on the millipedes (Diplopoda) of the Stavropol Territory, northern Caucasus, Russia. Arthropoda Selecta, 23 (4), 347 - 354." type="journal article" year="2014">Zuev, 2014</bibRefCitation>
: 348 (R, D).
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F61B9D31FF07523CFDB9C488" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="materials_examined">
<paragraph id="8BC48E52F61B9D32FF07523CFBCEC2F2" blockId="9.[151,1436,1828,2034]" pageId="9" pageNumber="10">
<emphasis id="B90F5240F61B9D32FF07523CFE24C23D" bold="true" box="[151,391,1828,1853]" pageId="9" pageNumber="10">Material examined.</emphasis>
1 male (lectotype, designated herewith), 1 female (paralectotype, designated herewith), “Typen” (ZISP),
<emphasis id="B90F5240F61B9D32FEC85251FE1CC262" bold="true" box="[344,447,1865,1890]" pageId="9" pageNumber="10">RUSSIA</emphasis>
, Kuban Region, Station Psebai, under stones during ascent to Hill Gerisgel,
<date id="FFC5A892F61B9D32FAB35251FA3BC261" box="[1315,1432,1865,1889]" pageId="9" pageNumber="10" value="1911-05-20">20.V.1911</date>
, leg. D. Volnukhin; 4 juv. (ZMUM), Krasnodar Prov., Mostovskoy Distr., Shedok, N of Psebai, karst funnel, wet
<emphasis id="B90F5240F61B9D32FF07528AFE2DC2A9" box="[151,398,1938,1961]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C7BF5D1F61B9D32FF07528AFF43C2A9" box="[151,224,1938,1961]" class="Magnoliopsida" family="Fagaceae" genus="Fagus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">Fagus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61B9D32FF7D528AFE81C2A9" box="[237,290,1938,1961]" class="Magnoliopsida" family="Sapindaceae" genus="Acer" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sapindales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">Acer</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61B9D32FEA0528AFE2DC2A9" box="[304,398,1938,1961]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
</emphasis>
etc. forest,
<quantity id="4C8323B7F61B9D32FD895289FDC1C2A9" box="[537,610,1937,1962]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="9" pageNumber="10" unit="m" value="650.0">650 m</quantity>
a.s.l.,
<date id="FFC5A892F61B9D32FD205289FC9BC2AA" box="[688,824,1937,1962]" pageId="9" pageNumber="10" value="1986-08-04">4.VIII.1986</date>
; 1 male (ZMUM), Stavropol Prov., E of Georgievsk,
<emphasis id="B90F5240F61B9D32FF0752AEFECDC2CD" box="[151,366,1974,1997]" italics="true" pageId="9" pageNumber="10">
<taxonomicName id="4C7BF5D1F61B9D32FF0752AEFF5BC2CD" box="[151,248,1974,1997]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
,
<taxonomicName id="4C7BF5D1F61B9D32FE9652AEFECDC2CD" box="[262,366,1974,1997]" class="Magnoliopsida" family="Betulaceae" genus="Carpinus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">Carpinus</taxonomicName>
</emphasis>
etc. forest, litter and under bark,
<date id="FFC5A892F61B9D32FD6C52ADFC68C2CD" box="[764,971,1973,1998]" pageId="9" pageNumber="10" value="1982-05-28" valueMax="1982-05-31" valueMin="1982-05-28">28 and 31.V.1982</date>
; 1 male, 2 females (ZMUM), Stavropol Prov., Kursavska,
<taxonomicName id="4C7BF5D1F61B9D32FEF352C1FE1FC2F2" box="[355,444,2009,2034]" class="Magnoliopsida" family="Fabaceae" genus="Robinia" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fabales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="genus">
<emphasis id="B90F5240F61B9D32FEF352C1FE1FC2F2" box="[355,444,2009,2034]" italics="true" pageId="9" pageNumber="10">Robinia</emphasis>
</taxonomicName>
and
<emphasis id="B90F5240F61B9D32FE6252C2FD89C2F1" box="[498,554,2010,2033]" italics="true" pageId="9" pageNumber="10">Rosa</emphasis>
hedge along road,
<date id="FFC5A892F61B9D32FC9052C1FCD6C2F1" box="[768,885,2009,2033]" pageId="9" pageNumber="10" value="1982-05-30">30.V.1982</date>
, all leg. S. Golovatch.
</paragraph>
<paragraph id="8BC48E52F6189D31FF57558FFD22C5F8" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF57558FFE00C5B0" bold="true" box="[199,419,151,176]" pageId="10" pageNumber="11">Descriptive notes.</emphasis>
Length ca
<quantity id="4C8323B7F6189D31FDA85580FD19C5AF" box="[568,698,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="10" pageNumber="11" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">
11
<emphasis id="B90F5240F6189D31FDC3558FFDC2C5B0" bold="true" box="[595,609,151,176]" pageId="10" pageNumber="11"></emphasis>
12 mm
</quantity>
, width of midbody metazonae
<quantity id="4C8323B7F6189D31FBAE5580FB78C5AF" box="[1086,1243,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="10" pageNumber="11" unit="mm" value="1.225" valueMax="1.25" valueMin="1.2">
1.2
<emphasis id="B90F5240F6189D31FBF0558FFBCDC5B0" bold="true" box="[1120,1134,151,176]" pageId="10" pageNumber="11"></emphasis>
1.25 mm
</quantity>
(male, female), metazonite to prozonite width ratio &lt;1.7 (
<figureCitation id="134092D7F6189D31FD1655A4FC9FC5D4" box="[646,828,188,213]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="10" pageNumber="11">
Fig. 3A, 3H
<emphasis id="B90F5240F6189D31FC8355A4FC82C5D5" bold="true" box="[787,801,188,213]" pageId="10" pageNumber="11"></emphasis>
3J
</figureCitation>
). Live coloration usually uniformly light brown, not reddish, about same as in alcohol (
<figureCitation id="134092D7F6189D31FD8A55C7FDD7C5F8" box="[538,628,223,248]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="10" pageNumber="11">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8BC48E52F6189D31FF57541CFDB9C488" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="11">
All characters as in
<taxonomicName id="4C7BF5D1F6189D31FE3B541DFD9BC41D" authorityName="Attems" authorityYear="1898" box="[427,568,260,285]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="abchasius">
<emphasis id="B90F5240F6189D31FE3B541DFD9BC41D" box="[427,568,260,285]" italics="true" pageId="10" pageNumber="11">P. abchasius</emphasis>
</taxonomicName>
(
<figureCitation id="134092D7F6189D31FDD8541CFD5CC41C" box="[584,767,260,285]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="10" pageNumber="11">
Figs 3A, 3H
<emphasis id="B90F5240F6189D31FD46541CFD47C41D" bold="true" box="[726,740,260,285]" pageId="10" pageNumber="11"></emphasis>
3J
</figureCitation>
,
<figureCitation id="134092D7F6189D31FC9D541CFCC1C41C" box="[781,866,260,284]" captionStart="FIGURE 4" captionStartId="8.[151,249,1796,1818]" captionTargetBox="[198,1381,227,1769]" captionTargetId="figure@8.[198,1388,193,1774]" captionTargetPageId="8" captionText="FIGURE 4. Left gonopods of Polydesmus abchasius Attems, 1898, P. lignaui Lohmander, 1936 and P. muralewiczi Lohmander, 1936 (A C), and right gonopod of P. muralewiczi Lohmander, 1936 (D). A: male from near Lake Ritsa, Abkhazia, mesal view. B: male from Sukhumi Botanical Garden, Abkhazia, mesal view. C: male from near Kursavka, Stavropol Prov., Russia, mesal view. D: male lectotype from near Psebai, Krasnodar Prov., Russia (after Lohmander 1936). Designations: en, endomere; ex, exomere; d, exomere tooth; pu, hairy pulvillus; r, endomere rod. Scale bar: 0.2 mm (A C), not to scale (D)." httpUri="https://zenodo.org/record/1052389/files/figure.png" pageId="10" pageNumber="11">4C, 4D</figureCitation>
), except as follows. Collum clearly narrower than head (
<figureCitation id="134092D7F6189D31FF70543FFED7C440" box="[224,372,295,320]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="10" pageNumber="11">Fig. 3A, 3H</figureCitation>
). Paraterga not upturned, subhorizontal, mostly acute-angled and drawn behind rear tergal margin starting with segment 14 or 15 (
<figureCitation id="134092D7F6189D31FDDF5454FD5CC464" box="[591,767,332,357]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H J), P. lignaui Lohmander, 1936 (D G) and P. abchasius Attems, 1898 (B, C). A, H J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="10" pageNumber="11">
Fig. 3A, 3H
<emphasis id="B90F5240F6189D31FD455454FD40C465" bold="true" box="[725,739,332,357]" pageId="10" pageNumber="11"></emphasis>
3J
</figureCitation>
). Gonopod (
<figureCitation id="134092D7F6189D31FC1A5454FBB2C464" box="[906,1041,332,357]" captionStart="FIGURE 4" captionStartId="8.[151,249,1796,1818]" captionTargetBox="[198,1381,227,1769]" captionTargetId="figure@8.[198,1388,193,1774]" captionTargetPageId="8" captionText="FIGURE 4. Left gonopods of Polydesmus abchasius Attems, 1898, P. lignaui Lohmander, 1936 and P. muralewiczi Lohmander, 1936 (A C), and right gonopod of P. muralewiczi Lohmander, 1936 (D). A: male from near Lake Ritsa, Abkhazia, mesal view. B: male from Sukhumi Botanical Garden, Abkhazia, mesal view. C: male from near Kursavka, Stavropol Prov., Russia, mesal view. D: male lectotype from near Psebai, Krasnodar Prov., Russia (after Lohmander 1936). Designations: en, endomere; ex, exomere; d, exomere tooth; pu, hairy pulvillus; r, endomere rod. Scale bar: 0.2 mm (A C), not to scale (D)." httpUri="https://zenodo.org/record/1052389/files/figure.png" pageId="10" pageNumber="11">
Fig. 4C
<emphasis id="B90F5240F6189D31FC4F5454FC4EC465" bold="true" box="[991,1005,332,357]" pageId="10" pageNumber="11"></emphasis>
4D
</figureCitation>
) with a shorter endomere rod and a less heavily spinulate exomere tip.
</paragraph>
</subSubSection>
<subSubSection id="C361DDD9F6189D31FF57548CFE95C718" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BC48E52F6189D31FF57548CFE95C718" blockId="10.[151,1436,151,536]" pageId="10" pageNumber="11">
<emphasis id="B90F5240F6189D31FF57548CFE98C4AD" bold="true" box="[199,315,404,429]" pageId="10" pageNumber="11">Remarks.</emphasis>
Although the original description was clearly based only on the male (
<bibRefCitation id="EFEAF3A3F6189D31FBF0548CFA8EC4AD" author="Lohmander" box="[1120,1325,404,429]" pageId="10" pageNumber="11" refString="Lohmander, H. (1936) Uber die Diplopoden des Kaukasusgebietes. Goteborgs Kungl. Vetenskaps- och Vitterhets-Samhalles Handlingar, Series 5 B, 5 (1), 1 - 196." type="journal article" year="1936">Lohmander, 1936</bibRefCitation>
), the
<typeStatus id="54C030F0F6189D31FAFC548DFA3FC4AD" box="[1388,1436,405,429]" pageId="10" pageNumber="11">type</typeStatus>
series, “
<typeStatus id="54C030F0F6189D31FF6054A0FE9FC4D0" box="[240,316,440,464]" pageId="10" pageNumber="11">Typen</typeStatus>
”, actually consists of two specimens (male and female) which are to be considered as
<typeStatus id="54C030F0F6189D31FA9E54A0FAD7C4D0" box="[1294,1396,440,464]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
. To secure that the species is based on male material, the male is designated herewith as
<typeStatus id="54C030F0F6189D31FB1754C4FB53C4F5" box="[1159,1264,476,501]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
, the female as
<typeStatus id="54C030F0F6189D31FF0754E7FE91C718" box="[151,306,511,536]" pageId="10" pageNumber="11" type="paralectotype">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>