treatments-xml/data/03/D1/07/03D1075CB8428650FD5D0800FAC34F90.xml

<document id="D6DBC6CF6A18C683F43C9ED06C7D8E85" ID-DOI="10.5281/zenodo.170073" ID-GBIF-Dataset="a264bf74-a625-4ea9-a290-e34d1bce7500" ID-ISSN="1175­5326" ID-Zenodo-Dep="170073" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749791144" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho &amp; Grossi, Everardo José" docDate="2005" docId="03D1075CB8428650FD5D0800FAC34F90" docLanguage="en" docName="zt01054p029.pdf" docOrigin="Zootaxa 1054" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Brachysiderus Waterhouse" docType="key" docVersion="9" lastPageNumber="29" masterDocId="FFE87F24B8418654FC550B4CFFE84C26" masterDocTitle="A new species of Brachysiderus Waterhouse, 1881 (Coleoptera: Scarabaeidae: Dynastinae) from Maranhão State, Brazil" masterLastPageNumber="29" masterPageNumber="25" pageNumber="28" updateTime="1698211557280" updateUser="plazi">
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<mods:title id="ABA842A411ABD735F0B56B33275FF554">A new species of Brachysiderus Waterhouse, 1881 (Coleoptera: Scarabaeidae: Dynastinae) from Maranhão State, Brazil</mods:title>
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<mods:namePart id="EF3C1B6636A60C26D6F1047765190943">Grossi, Paschoal Coelho</mods:namePart>
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<mods:namePart id="F3AF35BB45FB454C3CC813A1CDEB6043">Grossi, Everardo José</mods:namePart>
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<mods:date id="83E5359F7F0CBF270464C9D5389AABB5">2005</mods:date>
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<paragraph id="8BC7B64AB8428657FD5D0800FC014F40" blockId="3.[264,1001,844,905]" box="[264,1001,844,870]" pageId="3" pageNumber="28">
<heading id="D08F0126B8428657FD5D0800FC014F40" bold="true" box="[264,1001,844,870]" fontSize="11" level="1" pageId="3" pageNumber="28" reason="1">
<emphasis id="B90C6A58B8428657FD5D0800FC014F40" bold="true" box="[264,1001,844,870]" pageId="3" pageNumber="28">
Key to subgenera and species of 
<taxonomicName id="4C78CDC9B8428657FEC80800FC014F40" authority="Waterhouse" authorityName="Waterhouse" box="[669,1001,844,870]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B90C6A58B8428657FEC80800FCA34F40" bold="true" box="[669,843,844,870]" italics="true" pageId="3" pageNumber="28">Brachysiderus</emphasis>
Waterhouse
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC7B64AB8428657FD5D0823FD9C4FAF" blockId="3.[264,1001,844,905]" box="[264,628,879,905]" pageId="3" pageNumber="28">
(modified from 
<bibRefCitation id="EFE9CBBBB8428657FD960823FD844FAF" author="Endrodi" box="[451,620,879,905]" pageId="3" pageNumber="29" refString="Endrodi, S. (1985) The Dynastinae of the World. Dr. W. Junk Publ., Dordrecht. 800 pp., 46 plates. Endrodi, S. (1970) Monographie der Dynastinae (Coleoptera). 3. Tribus: Agaocephalini. Acta Zoologica Academiae Scientarium Hungaricae, 16, 27 - 96." type="book chapter" year="1985">Endrödi, 1985</bibRefCitation>
)
</paragraph>
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<paragraph id="8BC7B64AB8428657FD5D08F5FAC3486D" blockId="3.[264,1325,953,1899]" pageId="3" pageNumber="28">
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1.­ Body flattened, brownish­yellow with black spots on elytra and pronotum; eye canthus in males produced into a small forward and upward directed horn. Male protarsi strongly thickened, inner claw on its under edge with strong tooth. Female head with 2 transverse tubercles ....................................... Subgenus 
<taxonomicName id="4C78CDC9B8428657FFEA0F7DFAEB486D" authority="Waterhouse" authorityName="Waterhouse" box="[959,1283,1073,1099]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B90C6A58B8428657FFEA0F7DFB86486D" bold="true" box="[959,1134,1073,1099]" italics="true" pageId="3" pageNumber="28">Brachysiderus</emphasis>
Waterhouse
</taxonomicName>
…2
</keyLead>
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<keyLead id="3089157FB8428657FD5D0F15FAC348E5" pageId="3" pageNumber="28">
1’­ Body convex, dorsally reddish to darkish­brown, eye canthus simple or somewhat projected outside. Male protarsi not thickened. Female head without tubercles. Subgenus 
<taxonomicName id="4C78CDC9B8428657FD6D0FE5FDC648E5" ID-CoL="5TCZ" authority="Endrodi" authorityName="Endrodi" box="[312,558,1193,1219]" class="Insecta" family="Scarabaeidae" genus="Minisiderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B90C6A58B8428657FD6D0FE5FE2148E5" bold="true" box="[312,457,1193,1219]" italics="true" pageId="3" pageNumber="28">Minisiderus</emphasis>
Endrödi
</taxonomicName>
..................................................................................................... 3
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2.­ Head of males with long, upturned, bifurcated horn. Elytra finely reticulated, shiny, strongly and densely punctate. Northwestern 
<collectingCountry id="F36FF6DAB8428657FF1E0FB5FC7B4935" box="[843,915,1273,1299]" name="Brazil" pageId="3" pageNumber="28">Brazil</collectingCountry>
and Eastern 
<collectingCountry id="F36FF6DAB8428657F87E0FB5FB8A4935" box="[1067,1122,1273,1299]" name="Peru" pageId="3" pageNumber="28">Peru</collectingCountry>
........................... ........................................................................................ 
<taxonomicName id="4C78CDC9B8428657FF990E6DFAC3491D" authority="Waterhouse" authorityName="Waterhouse" box="[972,1323,1313,1339]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="quadrimaculatus">
<emphasis id="B90C6A58B8428657FF990E6DFB71491D" bold="true" box="[972,1177,1313,1339]" italics="true" pageId="3" pageNumber="28">quadrimaculatus</emphasis>
Waterhouse
</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="3089157FB8428657FD5D0E05FAC24995" pageId="3" pageNumber="28">
2’­ Head of males with long, forward­directed, bifurcated horn, with a middle­apical acute process. Elytra strongly reticulated, oily, more sparsely and finely punctate than previous species. Western 
<collectingCountry id="F36FF6DAB8428657FE670ED5FD7E4995" box="[562,662,1433,1459]" name="Ecuador" pageId="3" pageNumber="28">Ecuador</collectingCountry>
, Western 
<collectingCountry id="F36FF6DAB8428657FF520ED5FCD54995" box="[775,829,1433,1459]" name="Peru" pageId="3" pageNumber="28">Peru</collectingCountry>
and 
<collectingCountry id="F36FF6DAB8428657FF230ED5FC254995" box="[886,973,1433,1459]" name="Bolivia" pageId="3" pageNumber="28">Bolivia</collectingCountry>
................. 
<emphasis id="B90C6A58B8428657F8070ED5FB394995" bold="true" box="[1106,1233,1433,1459]" italics="true" pageId="3" pageNumber="28">tridentiger</emphasis>
(Prell)
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<keyLead id="3089157FB8428657FD5D0E8DFAC549FD" box="[264,1325,1473,1499]" pageId="3" pageNumber="28">3.­ Protibiae externally tridentate ....................................................................................... 4</keyLead>
</paragraph>
<paragraph id="8BC7B64AB8428657FD5D0EA5FAC54A25" blockId="3.[264,1325,953,1899]" box="[264,1325,1513,1539]" pageId="3" pageNumber="28">
<keyLead id="3089157FB8428657FD5D0EA5FAC54A25" box="[264,1325,1513,1539]" pageId="3" pageNumber="28">3’­ Protibiae externally quadridentate................................................................................. 6</keyLead>
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4 ­ Male conjoint frontal horn basis much narrower than head, anterior angle of eye canthus acute and slightly upturned, frons smooth. Apical margin of pronotum strongly lobed at middle, posterior margin nearly straight. Pygidium without setae. Elytra sparsely punctate. 
<collectingCountry id="F36FF6DAB8428657FE440DC5FDB14A85" box="[529,601,1673,1699]" name="Brazil" pageId="3" pageNumber="28">Brazil</collectingCountry>
(Paraná, São Paulo, Minas Gerais) .......... 
<taxonomicName id="4C78CDC9B8428657F8180DC5FAC34A85" authority="Arrow." authorityName="Arrow." box="[1101,1323,1673,1699]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="paranensis">
<emphasis id="B90C6A58B8428657F8180DC5FB394A85" bold="true" box="[1101,1233,1673,1699]" italics="true" pageId="3" pageNumber="28">paranensis</emphasis>
Arrow.
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</paragraph>
<paragraph id="8BC7B64AB8428657FD5D0DFDFAC34B3D" blockId="3.[264,1325,953,1899]" pageId="3" pageNumber="28">
<keyLead id="3089157FB8428657FD5D0DFDFAC34B3D" pageId="3" pageNumber="28">4’­ Male conjoint frontal horn basis more wide than half of head, eye canthus simple and not upturned, frons punctate. Elytra densely punctate. Pygidium with many small setae ....................................................................................................................................... 5</keyLead>
</paragraph>
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<keyLead id="3089157FB8428657FD5D0C65FAC34B4D" pageId="3" pageNumber="28">5 ­ Horns triangular in cross section, frons with transverse process. Pronotal tubercles large and pointed forward, pronotum strongly punctured on disc. Paramera slightly</keyLead>
</paragraph>
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curved with outer edges straight. Female unknown. 
<collectingCountry id="F36FF6DAB8458650FFC20A50FC374D10" box="[919,991,284,310]" name="Brazil" pageId="4" pageNumber="29">Brazil</collectingCountry>
(Maranhão) ........................ ................................................................................ 
<taxonomicName id="4C78CDC9B8458650FFDB0A08FB0C4D78" authority="Grossi &amp; Grossi" authorityName="Grossi &amp; Grossi" box="[910,1252,324,350]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="mielkeorum" status="sp. nov.">
<emphasis id="B90C6A58B8458650FFDB0A08FBF74D78" bold="true" box="[910,1055,324,350]" italics="true" pageId="4" pageNumber="29">mielkeorum</emphasis>
Grossi &amp; Grossi
</taxonomicName>
<taxonomicNameLabel id="A23FD723B8458650F8B90A08FAC24D78" box="[1260,1322,324,350]" pageId="4" pageNumber="29" rank="species">n. sp.</taxonomicNameLabel>
<keyLead id="3089157FB8458650FD5D0A20FAC34DD8" pageId="4" pageNumber="29">
5’­ Horns flattened, frons simple, without transversal process, strongly punctured. Pronotal tubercles small, pointed upwards, disc with sparse punctures. Paramera strongly curved with abrupt straightening on apex. 
<collectingCountry id="F36FF6DAB8458650FF780AF0FC9C4DF0" box="[813,884,444,470]" name="Brazil" pageId="4" pageNumber="29">Brazil</collectingCountry>
(Minas Gerais, Mato Grosso, Goiás, Distrito Federal) .................................................................................. 
<taxonomicName id="4C78CDC9B8458650F8320AA8FAC34DD8" authority="Ohaus." authorityName="Ohaus." box="[1127,1323,484,510]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="minicola">
<emphasis id="B90C6A58B8458650F8320AA8FB394DD8" bold="true" box="[1127,1233,484,510]" italics="true" pageId="4" pageNumber="29">minicola</emphasis>
Ohaus.
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<keyLead id="3089157FB8458650FD5D0940FAC34F30" pageId="4" pageNumber="29">
6­ Protibial teeth unequally spaced from each other, 2 apical and 2 basal pairs placed far from each other. Pronotum of male with erect small horn at mid­apical margin with a fovea behind, that of female with traces of 2 tubercles without a fovea. Elytra finely reticulated, more finely so than pronotum, irregularly punctate, with traces of double rows. Pygidium smooth and shiny, in female densely punctate. Paramera slightly assymetrical, outer edge rounded, apices sharp. 
<collectingCountry id="F36FF6DAB8458650FF630998FC964EC8" box="[822,894,724,750]" name="Brazil" pageId="4" pageNumber="29">Brazil</collectingCountry>
(Mato Grosso, Goiás)...................... ................................................................................................. 
<taxonomicName id="4C78CDC9B8458650F84C09B0FAC34F30" authority="Ohaus." authorityName="Ohaus." box="[1049,1323,764,790]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="matogrossensis">
<emphasis id="B90C6A58B8458650F84C09B0FB394F30" bold="true" box="[1049,1233,764,790]" italics="true" pageId="4" pageNumber="29">matogrossensis</emphasis>
Ohaus.
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC7B64AB8458650FD5D0868FAC34F90" blockId="4.[264,1323,284,950]" pageId="4" pageNumber="29">
<keyLead id="3089157FB8458650FD5D0868FAC34F90" pageId="4" pageNumber="29">
6’­ The four protibial teeth placed equidistant from each other, basal tooth small and acuminate. Pronotum behind apical margin with 2 small tubercles, without a fovea. Paramera broad, asymmetrical, outer edge of apices toothed. Female unknown. 
<collectingCountry id="F36FF6DAB8458650F8B40838FAC14FA8" box="[1249,1321,884,910]" name="Brazil" pageId="4" pageNumber="29">Brazil</collectingCountry>
(Goiás).................................................................................................. 
<taxonomicName id="4C78CDC9B8458650F83008D0FAC34F90" authority="Ohaus." authorityName="Ohaus." box="[1125,1323,924,950]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="29" phylum="Arthropoda" rank="species" species="goyanus">
<emphasis id="B90C6A58B8458650F83008D0FB224F90" bold="true" box="[1125,1226,924,950]" italics="true" pageId="4" pageNumber="29">goyanus</emphasis>
Ohaus.
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