treatments-xml/data/03/D1/07/03D1075CB8418657FD5D0D16FE8C4EC8.xml

<document id="C9C994893A345196FCB1C18740F39FA8" ID-DOI="10.5281/zenodo.170073" ID-GBIF-Dataset="a264bf74-a625-4ea9-a290-e34d1bce7500" ID-ISSN="1175­5326" ID-Zenodo-Dep="170073" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459749791144" checkinUser="plazi" docAuthor="Grossi, Paschoal Coelho &amp; Grossi, Everardo José" docDate="2005" docId="03D1075CB8418657FD5D0D16FE8C4EC8" docLanguage="en" docName="zt01054p029.pdf" docOrigin="Zootaxa 1054" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Brachysiderus (Minisiderus) mielkeorum Grossi &amp; Grossi, new species" docType="treatment" docVersion="9" lastPageNumber="28" masterDocId="FFE87F24B8418654FC550B4CFFE84C26" masterDocTitle="A new species of Brachysiderus Waterhouse, 1881 (Coleoptera: Scarabaeidae: Dynastinae) from Maranhão State, Brazil" masterLastPageNumber="29" masterPageNumber="25" pageNumber="25" updateTime="1698211557280" updateUser="plazi">
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<mods:title id="A22C31486FA44D7C190EE4AC5DDC0714">A new species of Brachysiderus Waterhouse, 1881 (Coleoptera: Scarabaeidae: Dynastinae) from Maranhão State, Brazil</mods:title>
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<mods:namePart id="932F4B73DAD02F412E48B3A307B38942">Grossi, Paschoal Coelho</mods:namePart>
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<mods:namePart id="A1FE884500BEA13CDC3E61F9ECFA4322">Grossi, Everardo José</mods:namePart>
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<mods:date id="C0B7A5823D97A5E216C22D7808D5A412">2005</mods:date>
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<subSubSection id="C362E5C1B8418654FD5D0D16FB8A4A52" box="[264,1122,1626,1652]" pageId="0" pageNumber="25" type="nomenclature">
<paragraph id="8BC7B64AB8418654FD5D0D16FB8A4A52" blockId="0.[264,1122,1626,1652]" box="[264,1122,1626,1652]" pageId="0" pageNumber="25">
<heading id="D08F0126B8418654FD5D0D16FB8A4A52" bold="true" box="[264,1122,1626,1652]" fontSize="11" level="1" pageId="0" pageNumber="25" reason="1">
<emphasis id="B90C6A58B8418654FD5D0D16FB8A4A52" bold="true" box="[264,1122,1626,1652]" pageId="0" pageNumber="25">
<taxonomicName id="4C78CDC9B8418654FD5D0D16FC2D4A52" authority="Grossi &amp; Grossi" authorityName="Grossi &amp; Grossi" box="[264,965,1626,1652]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="25" phylum="Arthropoda" rank="species" species="mielkeorum" status="sp. nov." subGenus="Minisiderus">
<emphasis id="B90C6A58B8418654FD5D0D16FE5E4A52" bold="true" box="[264,438,1626,1652]" italics="true" pageId="0" pageNumber="25">Brachysiderus</emphasis>
(
<emphasis id="B90C6A58B8418654FD950D16FDB94A52" bold="true" box="[448,593,1626,1652]" italics="true" pageId="0" pageNumber="25">Minisiderus</emphasis>
) 
<emphasis id="B90C6A58B8418654FE370D16FD1B4A52" bold="true" box="[610,755,1626,1652]" italics="true" pageId="0" pageNumber="25">mielkeorum</emphasis>
Grossi &amp; Grossi
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, 
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</emphasis>
</heading>
</paragraph>
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<subSubSection id="C362E5C1B8418654FD5D0DE9FD7B4B79" pageId="0" pageNumber="25" type="materials_examined">
<paragraph id="8BC7B64AB8418654FD5D0DE9FD4A4B29" blockId="0.[264,1323,1701,1887]" pageId="0" pageNumber="25">
<emphasis id="B90C6A58B8418654FD5D0DE9FD904A99" bold="true" box="[264,632,1701,1727]" pageId="0" pageNumber="25">
Material examined: 
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</emphasis>
,, 
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, MA, Balsas, Serra do Penitente, 
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, 
<date id="FFC6908AB8418654FD5D0D81FE6E4AC1" box="[264,390,1741,1767]" pageId="0" pageNumber="25" value="2004-11-06">6­XI­2004</date>
, C. Mielke leg., Ex Col. E.&amp; P.Grossi, deposited in Museu de Zoologia da Universidade de São Paulo, 
<collectingCountry id="F36FF6DAB8418654FE060DB9FD764B29" box="[595,670,1781,1807]" name="Brazil" pageId="0" pageNumber="25">Brazil</collectingCountry>
.
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<emphasis id="B90C6A58B8418654FD6D0C51FE5D4B11" bold="true" box="[312,437,1821,1847]" pageId="0" pageNumber="25">
<typeStatus id="54C308E8B8418654FD6D0C51FE5D4B11" box="[312,437,1821,1847]" pageId="0" pageNumber="25" type="paratype">Paratypes</typeStatus>
</emphasis>
, 2 ɗ same data as 
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, 1 ɗ, same except 
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13­XI­ 
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</date>
, 500 m, Ex Col. E. Furtado. In col. E. &amp; P. Grossi.
</paragraph>
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<subSubSection id="C362E5C1B8408655FD6D0A57FD3A4F90" pageId="1" pageNumber="26" type="description">
<paragraph id="8BC7B64AB8408655FD6D0A57FD3A4F90" blockId="1.[264,1323,283,950]" pageId="1" pageNumber="26">
<emphasis id="B90C6A58B8408655FD6D0A57FE214D13" bold="true" box="[312,457,283,309]" pageId="1" pageNumber="26">Description</emphasis>
: ɗ 
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in length, 
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in pronotal width (fig. 1). Body elongated oval and convex, color reddish to dark brown, dorsally glabrous and smooth. Clypeus short, as wide as half of head, apically weakly concave on the middle; without sculpturing but with sparse punctures basally. Horns strong, strongly upturned and triangular in cross section, very separated on base from each other and apically acute. Frons with a transversal elevated angled process, strongly punctured on apex and posteriorly, anteriorly with sparse small punctures (fig. 2). Eye canthus with strong coalescent punctures, basally the obtuse angle with sparse small setae (fig. 3). Pronotum bordered all around, very convex and rounded laterally, densely punctured, punctures coalescent anteriorly and on disc. Anteriorly with an elevation with two microgranulated flattened and rounded tubercles, directed forward. Elytral surface completely covered by irregular punctures, with some coalescent punctures and with 5 lines of punctures, humeral punctures smaller than in the remaining elytral disc. Pygidium surface rugulose with very small setae, disc with weak coalescent punctures, basal margin concave. In lateral view, surface weakly convex. Anterior tibiae tridentate externally, basal tooth flattened. Each of meso and metatibiae with 1 strong spine on the external side near middle. Paramera rounded apically, external sides straight (fig. 5).
</paragraph>
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<paragraph id="8BC7B64AB8408655FD5D0C7FFD6C4B48" blockId="1.[264,1322,1843,1902]" pageId="1" pageNumber="26">
<emphasis id="B90C6A58B8408655FD5D0C7FFE514B6D" bold="true" box="[264,441,1843,1867]" pageId="1" pageNumber="26">FIGURES 1–2:</emphasis>
<taxonomicName id="4C78CDC9B8408655FD960C7FFD344B6C" box="[451,732,1843,1866]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="26" phylum="Arthropoda" rank="species" species="miekeorum" status="sp. nov.">
<emphasis id="B90C6A58B8408655FD960C7FFD344B6C" box="[451,732,1843,1866]" italics="true" pageId="1" pageNumber="26">Brachysiderus miekeorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FD723B8408655FEB00C7FFCCA4B6C" box="[741,802,1843,1866]" pageId="1" pageNumber="26" rank="species">n. sp.</taxonomicNameLabel>
holotype ɗ, 1: dorsal view; 2: lateral view of head and anterior part of prothorax.
</paragraph>
</caption>
<caption id="DF07E6C2B8438656FD5D0D9BFCF94B7C" httpUri="https://zenodo.org/record/170075/files/figure.png" pageId="2" pageNumber="27" targetBox="[423,1184,328,1696]" targetPageId="2">
<paragraph id="8BC7B64AB8438656FD5D0D9BFCF94B7C" blockId="2.[264,1323,1751,1882]" pageId="2" pageNumber="27">
<emphasis id="B90C6A58B8438656FD5D0D9BFE5B4AC9" bold="true" box="[264,435,1751,1775]" pageId="2" pageNumber="27">FIGURES 3–6</emphasis>
: 3: 
<taxonomicName id="4C78CDC9B8438656FDB20D94FCEB4AC9" box="[487,771,1752,1775]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="miekeorum" status="sp. nov.">
<emphasis id="B90C6A58B8438656FDB20D94FCEB4AC9" box="[487,771,1752,1775]" italics="true" pageId="2" pageNumber="27">Brachysiderus miekeorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FD723B8438656FF450D9BFCA64AC8" box="[784,846,1751,1774]" pageId="2" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
holotype ɗ, eye canthus (scale 1mm); 4: 
<taxonomicName id="4C78CDC9B8438656FD5D0DB7FDB24B34" authority="Ohaus" authorityName="Ohaus" box="[264,602,1787,1810]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="minicola">
<emphasis id="B90C6A58B8438656FD5D0DB7FDE04B34" box="[264,520,1787,1810]" italics="true" pageId="2" pageNumber="27">Brachysiderus minicola</emphasis>
Ohaus
</taxonomicName>
, ♂ eye canthus (scale 1mm); 5: 
<taxonomicName id="4C78CDC9B8438656FF9E0DB7FB0E4B34" box="[971,1254,1787,1810]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="miekeorum" status="sp. nov.">
<emphasis id="B90C6A58B8438656FF9E0DB7FB0E4B34" box="[971,1254,1787,1810]" italics="true" pageId="2" pageNumber="27">Brachysiderus miekeorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23FD723B8438656F8BA0DB7FAC34B34" box="[1263,1323,1787,1810]" pageId="2" pageNumber="27" rank="species">n. sp.</taxonomicNameLabel>
holotype ɗ, parameres, dorsal and lateral views (scale 1mm); 6: 
<taxonomicName id="4C78CDC9B8438656FF950C6CFAE54B10" authority="Ohaus" authorityName="Ohaus" box="[960,1293,1823,1847]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="minicola">
<emphasis id="B90C6A58B8438656FF950C6CFB574B11" box="[960,1215,1824,1847]" italics="true" pageId="2" pageNumber="27">Brachysiderus minicola</emphasis>
Ohaus
</taxonomicName>
, ♂ parameres, dorsal and lateral views (scale 1mm).
</paragraph>
</caption>
<subSubSection id="C362E5C1B8428657FD6D0A57FEAF4DA0" pageId="3" pageNumber="28" type="etymology">
<paragraph id="8BC7B64AB8428657FD6D0A57FEAF4DA0" blockId="3.[264,1323,283,750]" pageId="3" pageNumber="28">
<emphasis id="B90C6A58B8428657FD6D0A57FE2C4D13" bold="true" box="[312,452,283,309]" pageId="3" pageNumber="28">Etymology.</emphasis>
The specific epithet is a tribute to our friends, Prof. Dr. Olaf Mielke and his son Dr. Carlos Mielke, the later who collected the specimens and kindly provided them to us.
</paragraph>
</subSubSection>
<subSubSection id="C362E5C1B8428657FD6D0ADFFE8C4EC8" pageId="3" pageNumber="28" type="discussion">
<paragraph id="8BC7B64AB8428657FD6D0ADFFCEE4E50" blockId="3.[264,1323,283,750]" pageId="3" pageNumber="28">
<emphasis id="B90C6A58B8428657FD6D0ADFFE2B4D8B" bold="true" box="[312,451,403,429]" pageId="3" pageNumber="28">Discussion.</emphasis>
The new species is distinguished from other 
<taxonomicName id="4C78CDC9B8428657FF800AD8FB964D8B" box="[981,1150,404,429]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B90C6A58B8428657FF800AD8FB964D8B" box="[981,1150,404,429]" italics="true" pageId="3" pageNumber="28">Brachysiderus</emphasis>
</taxonomicName>
species by the transverse angled process on the frons, horns not flattened, eye canthus form, and by the paramera of aedeagus, specially from 
<taxonomicName id="4C78CDC9B8428657FE900AA8FB914DD8" authority="Ohaus, 1930" authorityName="Ohaus" authorityYear="1930" box="[709,1145,484,510]" class="Insecta" family="Dynastidae" genus="Brachysiderus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="minicola">
<emphasis id="B90C6A58B8428657FE900AA8FC354DDB" box="[709,989,484,509]" italics="true" pageId="3" pageNumber="28">Brachysiderus minicola</emphasis>
<bibRefCitation id="EFE9CBBBB8428657FFB10AA8FB914DD8" author="Ohaus" box="[996,1145,484,510]" pageId="3" pageNumber="29" refString="Ohaus, F. (1930) Neue brasilianische Dynastinen. Stettiner Entomlogische Zeitung, 91, 261 - 265." type="journal article" year="1930">Ohaus, 1930</bibRefCitation>
</taxonomicName>
(figs. 4 and 6) by the presence of the tridentate foretibiae that is the characterisc of the 
<emphasis id="B90C6A58B8428657F80C0940FB5F4E03" box="[1113,1207,524,549]" italics="true" pageId="3" pageNumber="28">minicoa</emphasis>
group. In 
<collectingCountry id="F36FF6DAB8428657FD5D0978FEA74E68" box="[264,335,564,590]" name="Brazil" pageId="3" pageNumber="28">Brazil</collectingCountry>
this is the first reported species from the Northeastern Region, all others being from Central­Western and Southeastern Regions.
</paragraph>
<paragraph id="8BC7B64AB8428657FD6D09C8FE8C4EC8" blockId="3.[264,1323,283,750]" pageId="3" pageNumber="28">
Body may be smaller (
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in length, 
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in pronotum width) than the 
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and more strongly punctured, with horns, head process and pronotal tubercles smaller.
</paragraph>
</subSubSection>
</treatment>
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