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<document id="C6CCED02CE45E5CCFC85EF8CD7C845F9" ID-CLB-Dataset="56020" ID-DOI="10.5252/geodiversitas2022v44a14" ID-GBIF-Dataset="e0a862de-7b28-4ef1-9861-df72e4dee857" ID-ISSN="1638-9395" ID-Zenodo-Dep="6448632" ID-ZooBank="urn:lsid:zoobank.org:pub:76D6D377-FA63-40C8-9BD1-767B35F1BA09" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649695967684" checkinUser="felipe" docAuthor="Famoso, Nicholas A. &amp; Orcutt, John D." docDate="2022" docId="03D087D5FFB1BD61FF6015E3D8FBB96A" docLanguage="en" docName="Geodiversitas.44.14.427-436.pdf" docOrigin="Geodiversitas 44 (14)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Palaeogale dorothiae MacDonald 1963" docType="treatment" docVersion="5" lastPageNumber="432" masterDocId="FFE9FFADFFB6BD66FF8E1370DD54BE66" masterDocTitle="First occurrences of Palaeogale von Meyer, 1846 in the Pacific Northwest, United States" masterLastPageNumber="436" masterPageNumber="427" pageNumber="432" updateTime="1699319795323" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="625E39AB6C82F3ABD32482D2E0EE421A">First occurrences of Palaeogale von Meyer, 1846 in the Pacific Northwest, United States</mods:title>
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<mods:namePart id="3F9AC08B745BE66DD2498C381492F850">Famoso, Nicholas A.</mods:namePart>
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<mods:namePart id="E62730A40F996AD028768CB7ABB344DF">Orcutt, John D.</mods:namePart>
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<mods:title id="BCC959BDB617C490F8FB223A38E4B91D">Geodiversitas</mods:title>
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<treatment id="03D087D5FFB1BD61FF6015E3D8FBB96A" ID-DOI="http://doi.org/10.5281/zenodo.6458045" ID-GBIF-Taxon="194591545" ID-Zenodo-Dep="6458045" LSID="urn:lsid:plazi:treatment:03D087D5FFB1BD61FF6015E3D8FBB96A" httpUri="http://treatment.plazi.org/id/03D087D5FFB1BD61FF6015E3D8FBB96A" lastPageNumber="432" pageId="7" pageNumber="432">
<subSubSection id="C3636548FFB1BD61FF6015E3DFC9B8CB" box="[238,669,1683,1710]" pageId="7" pageNumber="432" type="nomenclature">
<paragraph id="8BC636C3FFB1BD61FF6015E3DFC9B8CB" blockId="7.[238,669,1683,1742]" box="[238,669,1683,1710]" pageId="7" pageNumber="432">
<taxonomicName id="4C794D40FFB1BD61FF6015E3DFC9B8CB" authority="MacDonald, 1963" authorityName="MacDonald" authorityYear="1963" box="[238,669,1683,1710]" class="Mammalia" family="Odobenidae" genus="Palaeogale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="432" phylum="Chordata" rank="species" species="dorothiae">
<emphasis id="B90DEAD1FFB1BD61FF6015E3DC9AB8C8" bold="true" box="[238,462,1683,1710]" italics="true" pageId="7" pageNumber="432">Palaeogale dorothiae</emphasis>
<bibRefCitation id="EFE84B32FFB1BD61FE5B15E3DFC9B8CB" author="MACDONALD J. R." box="[469,669,1683,1709]" pageId="7" pageNumber="432" pagination="139 - 240" refId="ref7214" refString="MACDONALD J. R. 1963. - The Miocene Faunas from the Wounded Knee Area of Western South Dakota. Bulletin of the American Museum of Natural History 125 (3): 139 - 240. http: // hdl. handle. net / 2246 / 1259" type="journal article" year="1963">MacDonald, 1963</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FEE115C3DF48B8A8" box="[367,540,1715,1742]" pageId="7" pageNumber="432" type="description">
<paragraph id="8BC636C3FFB1BD61FEE115C3DF48B8A8" blockId="7.[238,669,1683,1742]" box="[367,540,1715,1742]" pageId="7" pageNumber="432">
(
<figureCitation id="13422A46FFB1BD61FEF615C3DCE2B8AB" box="[376,438,1715,1741]" captionStart="FIG" captionStartId="8.[132,143,1314,1331]" captionTargetBox="[215,1456,183,1291]" captionTargetId="figure-386@8.[748,1285,716,1286]" captionTargetPageId="8" captionText="FIG. 3. — Dentaries of Palaeogale dorothiae MacDonald, 1963: A, lingual view (JODA 6177);B, occlusal view (JODA 6177);C, buccal view (JODA 6177); D, lingual view (JODA 5893); E, occlusal view (JODA 5893); F, buccal view (JODA 5893). Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6448638" httpUri="https://zenodo.org/record/6448638/files/figure.png" pageId="7" pageNumber="432">Fig. 3</figureCitation>
;
<tableCitation id="C6FB0378FFB1BD61FE4D15C3DF47B8AB" box="[451,531,1715,1741]" captionStart="TABLE" captionStartId="7.[133,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measurements of the lower dentition. Abbreviations: APL, anterior-posterior length; TW, transverse width; , not available; *, type specimen. All measurements are in mm. †, measured scaled figure or image in ImageJ." httpUri="http://table.plazi.org/id/DF06664BFFB1BD61FF0B13ACDFB1BF65" pageId="7" pageNumber="432" tableUuid="DF06664BFFB1BD61FF0B13ACDFB1BF65">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FF0A159EDFE6B9E3" pageId="7" pageNumber="432" type="materials_examined">
<paragraph id="8BC636C3FFB1BD61FF0A159EDFA5B946" blockId="7.[131,774,1773,1824]" pageId="7" pageNumber="432">
REFERRED SPECIMENS. —
<collectionCode id="ED68AE06FFB1BD61FEF6159EDCECB960" box="[376,440,1774,1798]" pageId="7" pageNumber="432">JODA</collectionCode>
6177 left fragmentary dentary with p4-m1.
<collectionCode id="ED68AE06FFB1BD61FF5F1478DC45B946" box="[209,273,1800,1824]" pageId="7" pageNumber="432">JODA</collectionCode>
5893 right fragmentary dentary with p2 and p4.
</paragraph>
<paragraph id="8BC636C3FFB1BD61FF0A1449DFE6B9E3" blockId="7.[131,776,1849,1926]" pageId="7" pageNumber="432">
OCCURRENCE. — JDNM-140, Lonerock, Gilliam County,
<collectingRegion id="49BDF821FFB1BD61FD391449DE50B937" box="[695,772,1849,1873]" country="United States of America" name="Oregon" pageId="7" pageNumber="432">Oregon</collectingRegion>
,
<collectingCountry id="F36E7653FFB1BD61FF0D1423DDE2B90D" box="[131,182,1875,1899]" name="United States of America" pageId="7" pageNumber="432">USA</collectingCountry>
. Kimberly Member, John Day Formation. Precise locality information available to qualified researchers upon request.
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FF0A14EFDF05B98D" pageId="7" pageNumber="432" type="multiple">
<paragraph id="8BC636C3FFB1BD61FF0A14EFDF05B98D" blockId="7.[132,776,1950,2027]" pageId="7" pageNumber="432">
AGE. — Late Oligocene in the Late Arikareean (Ar3) NALMA, between the Tin Roof Tuff (25.3 Ma) and the UNSM JD-BC-3 tuff (23.5-23.8 Ma) (
<bibRefCitation id="EFE84B32FFB1BD61FEDA14A3DF17B98D" author="FREMD T. J. &amp; WHISTLER D. P." box="[340,579,2003,2027]" pageId="7" pageNumber="432" pagination="159 - 169" refId="ref6738" refString="FREMD T. J. &amp; WHISTLER D. P. 2009. - Preliminary description of a new microvertebrate assemblage from the Arikareean (Early Miocene) John Day Formation, Central Oregon. Museum of Northern Arizona Bulletin 65: 159 - 169." type="journal article" year="2009">Fremd &amp; Whistler 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FCA310E4D8FBBDA1" pageId="7" pageNumber="432" type="diagnosis">
<paragraph id="8BC636C3FFB1BD61FCA310E4D8FBBDA1" blockId="7.[811,1456,916,967]" pageId="7" pageNumber="432">
SPECIES DIAGNOSIS. — The p4 is without a posterior accessory cusp and the protoconid of the m1 is not expanded laterally (
<bibRefCitation id="EFE84B32FFB1BD61FB7210DFD8F5BDA1" author="MACDONALD J. R." box="[1276,1441,943,967]" pageId="7" pageNumber="432" pagination="139 - 240" refId="ref7214" refString="MACDONALD J. R. 1963. - The Miocene Faunas from the Wounded Knee Area of Western South Dakota. Bulletin of the American Museum of Natural History 125 (3): 139 - 240. http: // hdl. handle. net / 2246 / 1259" type="journal article" year="1963">MacDonald 1963</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FCA31085D9CCBA88" pageId="7" pageNumber="432" type="description">
<paragraph id="8BC636C3FFB1BD61FCA31085DEE9BA6B" blockId="7.[811,1464,1013,1262]" box="[813,957,1013,1039]" pageId="7" pageNumber="432">DESCRIPTION</paragraph>
<paragraph id="8BC636C3FFB1BD61FCA31764D9CCBA88" blockId="7.[811,1464,1013,1262]" pageId="7" pageNumber="432">
JODA 5893 has no posterior accessory cusp on p4; however, there is a small enamel anomaly on the posterior edge. There is an alveolus for p3. The p2 crown is stretched forward with an upturned posterior heel. JODA 6177 is broken behind the c, and there is no p2 alveoli. There is no p4 accessory cusp. The m1 protoconid is not expanded laterally. See
<tableCitation id="C6FB0378FFB1BD61FAB417C4D8DDBAA8" box="[1338,1417,1204,1230]" captionStart="TABLE" captionStartId="7.[133,143,220,237]" captionTargetPageId="7" captionText="TABLE 2. — Measurements of the lower dentition. Abbreviations: APL, anterior-posterior length; TW, transverse width; , not available; *, type specimen. All measurements are in mm. †, measured scaled figure or image in ImageJ." httpUri="http://table.plazi.org/id/DF06664BFFB1BD61FF0B13ACDFB1BF65" pageId="7" pageNumber="432" tableUuid="DF06664BFFB1BD61FF0B13ACDFB1BF65">Table 2</tableCitation>
for measurements of both specimens.
</paragraph>
</subSubSection>
<subSubSection id="C3636548FFB1BD61FCA31664D8FBB96A" pageId="7" pageNumber="432" type="discussion">
<paragraph id="8BC636C3FFB1BD61FCA31664DEE9BB4A" blockId="7.[810,1457,1300,1805]" box="[813,957,1300,1326]" pageId="7" pageNumber="432">COMPARISON</paragraph>
<paragraph id="8BC636C3FFB1BD61FCA31643D8FBB96A" blockId="7.[810,1457,1300,1805]" pageId="7" pageNumber="432">
Both specimens match the species diagnosis in the absence of the p4 accessory cusp. They are both too large to be
<taxonomicName id="4C794D40FFB1BD61FAAA1624D8DEBB0B" baseAuthorityName="Hayes" baseAuthorityYear="2000" box="[1316,1418,1364,1390]" class="Mammalia" family="Odobenidae" genus="Palaeogale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="432" phylum="Chordata" rank="species" species="minuta">
<emphasis id="B90DEAD1FFB1BD61FAAA1624D867BB0B" box="[1316,1331,1364,1389]" italics="true" pageId="7" pageNumber="432">P</emphasis>
.
<emphasis id="B90DEAD1FFB1BD61FACE1624D8DEBB0B" box="[1344,1418,1364,1389]" italics="true" pageId="7" pageNumber="432">minuta</emphasis>
</taxonomicName>
. In JODA 5893, the morphology of the p4 is similar to
<taxonomicName id="4C794D40FFB1BD61FAC41604D8FABBEB" authorityName="Gervais" authorityYear="1848" box="[1354,1454,1396,1422]" class="Mammalia" family="Odobenidae" genus="Palaeogale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="432" phylum="Chordata" rank="species" species="sectoria">
<emphasis id="B90DEAD1FFB1BD61FAC41604D80DBBEB" box="[1354,1369,1396,1421]" italics="true" pageId="7" pageNumber="432">P</emphasis>
.
<emphasis id="B90DEAD1FFB1BD61FAE81604D8FABBEB" box="[1382,1454,1396,1421]" italics="true" pageId="7" pageNumber="432">sectoria</emphasis>
</taxonomicName>
(Lange-Badre &amp; Dashzeveg 1989;
<bibRefCitation id="EFE84B32FFB1BD61FB1F16E4D825BBC8" author="WANG J. &amp; ZHANG Z. - Q." box="[1169,1393,1427,1454]" pageId="7" pageNumber="432" pagination="310 - 334" refId="ref8205" refString="WANG J. &amp; ZHANG Z. - Q. 2015. - Phylogenetic analysis on Palaeogale (Palaeogalidae, Carnivora) based on specimens from Oligocene strata of Saint-Jacques, Nei Mongol. Vertebrata PalAsiatica 53 (4): 310 - 334. http: // www. vertpala. ac. cn / EN / Y 2015 / V 53 / I 4 / 310" type="journal article" year="2015">Wang &amp; Zhang 2015</bibRefCitation>
). The size of JODA 6177 is comparable to
<taxonomicName id="4C794D40FFB1BD61FB2816C4D85FBBAB" authorityName="Gervais" authorityYear="1848" box="[1190,1291,1459,1485]" class="Mammalia" family="Odobenidae" genus="Palaeogale" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="7" pageNumber="432" phylum="Chordata" rank="species" species="sectoria">
<emphasis id="B90DEAD1FFB1BD61FB2816C4D9E1BBAB" box="[1190,1205,1460,1485]" italics="true" pageId="7" pageNumber="432">P</emphasis>
.
<emphasis id="B90DEAD1FFB1BD61FB4D16C4D85FBBAB" box="[1219,1291,1460,1485]" italics="true" pageId="7" pageNumber="432">sectoria</emphasis>
</taxonomicName>
from
<collectingCountry id="F36E7653FFB1BD61FAC416C3D8E4BBAB" box="[1354,1456,1459,1485]" name="Mongolia" pageId="7" pageNumber="432">Mongolia</collectingCountry>
but the protoconid of the m1 is not expanded (Lange-Badre &amp; Dashzeveg 1989;
<bibRefCitation id="EFE84B32FFB1BD61FC6C1683D997B86B" author="WANG J. &amp; ZHANG Z. - Q." box="[994,1219,1522,1549]" pageId="7" pageNumber="432" pagination="310 - 334" refId="ref8205" refString="WANG J. &amp; ZHANG Z. - Q. 2015. - Phylogenetic analysis on Palaeogale (Palaeogalidae, Carnivora) based on specimens from Oligocene strata of Saint-Jacques, Nei Mongol. Vertebrata PalAsiatica 53 (4): 310 - 334. http: // www. vertpala. ac. cn / EN / Y 2015 / V 53 / I 4 / 310" type="journal article" year="2015">Wang &amp; Zhang 2015</bibRefCitation>
). The
<typeStatus id="54C28861FFB1BD61FA881683D867B86B" box="[1286,1331,1523,1549]" pageId="7" pageNumber="432">type</typeStatus>
specimen is from the Sharps Formation of South Dakota which is 30-28.6 Ma (
<bibRefCitation id="EFE84B32FFB1BD61FCEC1542D970B82A" author="MACDONALD J. R." box="[866,1060,1586,1612]" pageId="7" pageNumber="432" pagination="139 - 240" refId="ref7214" refString="MACDONALD J. R. 1963. - The Miocene Faunas from the Wounded Knee Area of Western South Dakota. Bulletin of the American Museum of Natural History 125 (3): 139 - 240. http: // hdl. handle. net / 2246 / 1259" type="journal article" year="1963">MacDonald 1963</bibRefCitation>
;
<bibRefCitation id="EFE84B32FFB1BD61FBBD1542D8CBB82B" author="MCCONNELL T. H. &amp; DIBENEDETTO J. N." box="[1075,1439,1586,1613]" pageId="7" pageNumber="432" pagination="47759" refId="ref7452" refString="MCCONNELL T. H. &amp; DIBENEDETTO J. N. 2012. - Geology of the Early Arikareean Sharps Formation on the Pine Ridge Indian Reservation and surrounding areas of South Dakota and Nebraska. PloS one 7 (10): e 47759. https: // doi. org / 10.1371 / journal. pone. 0047759" type="journal article" year="2012">McConnell &amp; DiBenedetto 2012</bibRefCitation>
).
<bibRefCitation id="EFE84B32FFB1BD61FCA31522DE9FB80B" author="SWISHER C. C. &amp; III" box="[813,971,1618,1645]" pageId="7" pageNumber="432" refId="ref7834" refString="SWISHER C. C. III 1982. - Stratigraphy and Biostratigraphy of the Eastern Portion of the Wildcat Ridge, Western Nebraska. University of Nebraska-Lincoln, 344 p." type="book" year="1982">Swisher (1982)</bibRefCitation>
states that this species is also known from the Brown Siltstone Beds or Whitney Member of the Brule Formation (
<emphasis id="B90DEAD1FFB1BD61FCBB15E3DE12B8CA" box="[821,838,1683,1708]" italics="true" pageId="7" pageNumber="432">c.</emphasis>
31-30 Ma) and the Unit B Member of the Gering Formation (28.26-28.11 Ma) both of Nebraska (
<bibRefCitation id="EFE84B32FFB1BD61FB6D15C2D8FCB8AA" author="TEDFORD R. H. &amp; SWINEHART J. B. &amp; SWISHER III C. C. &amp; PROTHERO D. R. &amp; KING S. A. &amp; TIERNEY T. E." box="[1251,1448,1714,1740]" pageId="7" pageNumber="432" pagination="312 - 334" refId="ref8070" refString="TEDFORD R. H., SWINEHART J. B., SWISHER III C. C., PROTHERO D. R., KING S. A. &amp; TIERNEY T. E. 1996. - The Whitneyan- Arikareean transition in the High Plains, in PROTHERO D. R. &amp; EMERY R. J. (eds), The Terrestrial Eocene-Oligocene Transition in North America. Cambridge University Press, Cambridge: 312 - 334. https: // doi. org / 10.1017 / CBO 9780511665431.016" type="book chapter" year="1996">
Tedford
<emphasis id="B90DEAD1FFB1BD61FAB215C3D839B8AA" box="[1340,1389,1714,1740]" italics="true" pageId="7" pageNumber="432">et al.</emphasis>
1996
</bibRefCitation>
,
<bibRefCitation id="EFE84B32FFB1BD61FCA315A2DE37B88A" author="TEDFORD R. H. &amp; ALBRIGHT III L. B. &amp; BARNOSKY A. D. &amp; VILLAFRANCA I. &amp; HUNT JR R. M. &amp; STORER J. E. &amp; SWISHER III C. C. &amp; VOORHIES M. R. &amp; WEBB S. D. &amp; WHISTLER D. P." box="[813,867,1746,1772]" pageId="7" pageNumber="432" pagination="169 - 231" refId="ref7939" refString="TEDFORD R. H., ALBRIGHT III L. B., BARNOSKY A. D., FERRUSQUIA- VILLAFRANCA I., HUNT JR R. M., STORER J. E., SWISHER III C. C., VOORHIES M. R., WEBB S. D. &amp; WHISTLER D. P. 2004. - Mammalian biochronology of the Arikareean through Hemphillian interval (late Oligocene through early Pliocene epochs), in WOODBURNE M. O. (ed.), Late Cretaceous and Cenozoic Mammals of North America: Biostratigraphy and Geochronology. Columbia University Press, New York: 169 - 231. http: // www. jstor. org / stable / 10.7312 / wood 13040.12" type="book chapter" year="2004">2004</bibRefCitation>
). This makes the Lonerock specimens, which are between 25.3 and 23.8 Ma, the latest known occurrences of this species.
</paragraph>
</subSubSection>
</treatment>
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