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<mods:title id="178AF4B5B7C60558EAA365EBE9C06F6A">A revision of Geostiba of the West Palaearctic region. XXIII. On the Sibiota species of the Caucasus region exclusive of Turkey (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<taxonomicName id="4C794D47FFECFFB6FF7EFDE3DEAF3205" ID-CoL="3FSL6" authority="(EPPELSHEIM, 1878)" baseAuthorityName="EPPELSHEIM" baseAuthorityYear="1878" box="[178,709,584,606]" class="Insecta" family="Staphylinidae" genus="Geostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1106" phylum="Arthropoda" rank="species" species="carinicollis" subGenus="Sibiota">
<emphasis id="B90DEAD6FFECFFB6FF7EFDE3DE153206" bold="true" box="[178,639,584,606]" pageId="9" pageNumber="1106">
<emphasis id="B90DEAD6FFECFFB6FF7EFDE3DD673205" bold="true" box="[178,269,584,606]" italics="true" pageId="9" pageNumber="1106">Geostiba</emphasis>
(
<emphasis id="B90DEAD6FFECFFB6FED0FDE3DD0F3205" bold="true" box="[284,357,584,606]" italics="true" pageId="9" pageNumber="1106">Sibiota</emphasis>
)
<emphasis id="B90DEAD6FFECFFB6FEB8FDE3DD833205" bold="true" box="[372,489,584,606]" italics="true" pageId="9" pageNumber="1106">carinicollis</emphasis>
(
<bibRefCitation id="EFE84B35FFECFFB6FE3BFDE3DE153206" author="EPPELSHEIM E" box="[503,639,584,606]" pageId="9" pageNumber="1106" pagination="3 - 258" refId="ref10056" refString="EPPELSHEIM E. (1878): Staphylinidae; pp. 90 - 131. - In: SCHNEIDER O. &amp; H. LEDER, Beitrage zur Kenntniss der kaukasischen Kaferfauna. Verhandlungen des Naturforschenden Vereines in Brunn 16 (1877): 3 - 258 + 4 plates." type="book chapter" year="1878">EPPELSHEIM</bibRefCitation>
</emphasis>
,
<emphasis id="B90DEAD6FFECFFB6FD46FDE3DEAF3205" bold="true" box="[650,709,584,606]" pageId="9" pageNumber="1106">1878)</emphasis>
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(
<figureCitation id="13422A41FFECFFB6FD1EFDE2DF7F3204" box="[722,789,585,607]" captionStart="Map 2" captionStartId="3.[178,223,1109,1130]" captionTargetBox="[178,1058,212,1100]" captionTargetId="figure-235@3.[178,1058,212,1100]" captionTargetPageId="3" captionText="Map 2: Distributions of the species of the G. carinicollis group: G. articarinata ASSING (black square), G. convergens ASSING (white square), G. uniplicata ASSING (black diamond), G. krzysztofi (ROUBAL) (white diamonds), G. carinicollis (EPPELSHEIM) (white circles), G. unituber nov.sp. (black circle), G. pambakica nov.sp. (white triangles), G. unicuneata nov.sp. (black star), G. meghruica nov.sp. (white star)." figureDoi="http://doi.org/10.5281/zenodo.5183238" httpUri="https://zenodo.org/record/5183238/files/figure.png" pageId="9" pageNumber="1106">Map 2</figureCitation>
)
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</paragraph>
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<treatmentCitation id="0AD810D5FFECFFB6FF7EFDDFDE5132D1" author="EPPELSHEIM E" box="[178,571,628,650]" page="95" pageId="9" pageNumber="1106" year="1878">
<taxonomicName id="4C794D47FFECFFB6FF7EFDDFDE5132D1" authority="EPPELSHEIM, 1878: 95" authorityName="EPPELSHEIM" authorityPageNumber="95" authorityYear="1878" box="[178,571,628,650]" class="Insecta" family="Staphylinidae" genus="Leptusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1106" phylum="Arthropoda" rank="species" species="carinicollis">
<emphasis id="B90DEAD6FFECFFB6FF7EFDDFDD0D32D2" box="[178,359,628,649]" italics="true" pageId="9" pageNumber="1106">Leptusa carinicollis</emphasis>
<bibRefCitation id="EFE84B35FFECFFB6FEA0FDDFDE5132D1" author="EPPELSHEIM E" box="[364,571,628,650]" pageId="9" pageNumber="1106" pagination="3 - 258" refId="ref10056" refString="EPPELSHEIM E. (1878): Staphylinidae; pp. 90 - 131. - In: SCHNEIDER O. &amp; H. LEDER, Beitrage zur Kenntniss der kaukasischen Kaferfauna. Verhandlungen des Naturforschenden Vereines in Brunn 16 (1877): 3 - 258 + 4 plates." type="book chapter" year="1878">
EPPELSHEIM, 1878:
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f.
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</paragraph>
<paragraph id="8BC636C4FFECFFB6FF7EFD33DD853456" blockId="9.[178,1086,664,1038]" pageId="9" pageNumber="1106">
C o m m e n t: The original description is based on an unspecified number of
<typeStatus id="54C28866FFECFFB6FC12FD33D85332F6" box="[990,1081,664,685]" pageId="9" pageNumber="1106" type="syntype">syntypes</typeStatus>
, partly from unspecified localities (&quot;im Kaukasus weiter verbreitet&quot;), but &quot;hauptsächlich bei Mamudly ... und auf den Irgan-Tschaisky-gara&quot; (
<bibRefCitation id="EFE84B35FFECFFB6FD1EFD78DFE432B2" author="EPPELSHEIM E" box="[722,910,723,745]" pageId="9" pageNumber="1106" pagination="3 - 258" refId="ref10056" refString="EPPELSHEIM E. (1878): Staphylinidae; pp. 90 - 131. - In: SCHNEIDER O. &amp; H. LEDER, Beitrage zur Kenntniss der kaukasischen Kaferfauna. Verhandlungen des Naturforschenden Vereines in Brunn 16 (1877): 3 - 258 + 4 plates." type="book chapter" year="1878">EPPELSHEIM 1878</bibRefCitation>
). The type material in the Eppelsheim collection was revised and a male from &quot;Mamudly&quot; was designated as the
<typeStatus id="54C28866FFECFFB6FEFCFCA6DDE73378" box="[304,397,781,803]" pageId="9" pageNumber="1106" type="lectotype">lectotype</typeStatus>
by
<bibRefCitation id="EFE84B35FFECFFB6FE78FCA5DE263378" author="ASSING V" box="[436,588,781,803]" pageId="9" pageNumber="1106" pagination="903 - 1006" refId="ref9754" refString="ASSING V. (2005): A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae). - Linzer Biologische Beitrage 37 (2): 903 - 1006." type="journal article" year="2005">ASSING (2005)</bibRefCitation>
, who identified the type locality as a place in the
<collectingRegion id="49BDF826FFECFFB6FF7EFC80DD5B331A" box="[178,305,811,833]" country="Azerbaijan" name="Baku" pageId="9" pageNumber="1106">Baku region</collectingRegion>
,
<collectingCountry id="F36E7654FFECFFB6FEF0FC80DDC4331A" box="[316,430,811,833]" name="Azerbaijan" pageId="9" pageNumber="1106">Azerbaijan</collectingCountry>
, in the East Caucasus. A re-examination of the data indicated in Leder´s travel reports in SCHNEIDER &amp; LEDER (1878), however, revealed that Mamudly is situated in the south of
<collectingCountry id="F36E7654FFECFFB6FE07FCCEDE753320" box="[459,543,869,891]" name="Georgia" pageId="9" pageNumber="1106">Georgia</collectingCountry>
, close to the border with
<collectingCountry id="F36E7654FFECFFB6FCFEFCCEDFE73320" box="[818,909,869,891]" name="Armenia" pageId="9" pageNumber="1106">Armenia</collectingCountry>
, and most likely identical to Mtisdziri (=Makhmutlo, =Mamudlo, =Mamutlo;
<geoCoordinate id="EE4D5003FFECFFB6FCEFFC28DFF133C3" box="[803,923,899,920]" degrees="41" direction="north" minutes="17" orientation="latitude" pageId="9" pageNumber="1106" precision="15" seconds="12" value="41.286667">41°17'12''N</geoCoordinate>
,
<geoCoordinate id="EE4D5003FFECFFB6FC6BFC28D87133C2" box="[935,1051,899,921]" degrees="44" direction="east" minutes="10" orientation="longitude" pageId="9" pageNumber="1106" precision="15" seconds="08" value="44.16889">44°10'08''E</geoCoordinate>
) in Kvemo Kartli province. The second specified locality (&quot;Irgan-Tschaisky-gara&quot;) are probably the mountains (gara) near Irgani (=Irganchay, =Irganchai, =Irganchaj;
<geoCoordinate id="EE4D5003FFECFFB6FC0DFC15D8533388" box="[961,1081,958,979]" degrees="41" direction="north" minutes="12" orientation="latitude" pageId="9" pageNumber="1106" precision="15" seconds="24" value="41.20667">41°12'24''N</geoCoordinate>
,
<geoCoordinate id="EE4D5003FFECFFB6FF7EFC70DD4233AA" box="[178,296,987,1009]" degrees="44" direction="east" minutes="08" orientation="longitude" pageId="9" pageNumber="1106" precision="15" seconds="24" value="44.140003">44°08'24''E</geoCoordinate>
), which are not far away from the type locality. The revised, currently known distribution is shown in
<figureCitation id="13422A41FFECFFB6FE6BFC53DD813456" box="[423,491,1016,1037]" captionStart="Map 2" captionStartId="3.[178,223,1109,1130]" captionTargetBox="[178,1058,212,1100]" captionTargetId="figure-235@3.[178,1058,212,1100]" captionTargetPageId="3" captionText="Map 2: Distributions of the species of the G. carinicollis group: G. articarinata ASSING (black square), G. convergens ASSING (white square), G. uniplicata ASSING (black diamond), G. krzysztofi (ROUBAL) (white diamonds), G. carinicollis (EPPELSHEIM) (white circles), G. unituber nov.sp. (black circle), G. pambakica nov.sp. (white triangles), G. unicuneata nov.sp. (black star), G. meghruica nov.sp. (white star)." figureDoi="http://doi.org/10.5281/zenodo.5183238" httpUri="https://zenodo.org/record/5183238/files/figure.png" pageId="9" pageNumber="1106">Map 2</figureCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BC636C4FFECFFB6FF7EFBB5DE963435" blockId="9.[178,1085,1053,1134]" pageId="9" pageNumber="1106">
As can be inferred from the morphology of the median lobe of the aedeagus (base abruptly convex in lateral view; see figure
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<bibRefCitation id="EFE84B35FFECFFB6FD67FB90DF59340A" author="ASSING V" box="[683,819,1083,1105]" pageId="9" pageNumber="1106" pagination="903 - 1006" refId="ref9754" refString="ASSING V. (2005): A revision of the species of Geostiba THOMSON and Tropimenelytron PACE of the Eastern Mediterranean, the Caucasus, and adjacent regions (Coleoptera: Staphylinidae, Aleocharinae). - Linzer Biologische Beitrage 37 (2): 903 - 1006." type="journal article" year="2005">ASSING 2005</bibRefCitation>
),
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<emphasis id="B90DEAD6FFECFFB6FC89FB90DFBD340B" box="[837,983,1083,1104]" italics="true" pageId="9" pageNumber="1106">G. carinicollis</emphasis>
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is closely allied to the four species described from
<collectingCountry id="F36E7654FFECFFB6FD83FBF3DEC23435" box="[591,680,1112,1134]" name="Armenia" pageId="9" pageNumber="1106">Armenia</collectingCountry>
below).
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