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<document id="E32D81A1B6C0B420014F5549FFE8A701" ID="10.11646/zootaxa.4144.1.2" ID-DOI="10.11646/zootaxa.4144.1.2" ID-GBIF-Dataset="96118837-83f9-42e9-8426-5e03a819aa5a" ID-ISSN="1175-5326" ID-Zenodo-Dep="267229" ID-ZooBank="F1A6E9DE-2CF3-4C4E-8E7F-29F44476CE15" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1469513822376" checkinUser="plazi" docAuthor="Freidberg, Amnon" docDate="2016" docId="03D087C843119D3E5FB3FC1BFE9AFF60" docLanguage="en" docName="zootaxa.4144.1.2.pdf" docOrigin="Zootaxa 4144 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Goniglossum Rondani" docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFE9FFB043129D3B5F24FFC9FFFAFFB4" masterDocTitle="New taxa of Carpomyini, with special emphasis on Goniglossum (Diptera: Tephritidae: Trypetinae)" masterLastPageNumber="70" masterPageNumber="54" pageNumber="57" updateTime="1698366019578" updateUser="plazi">
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<mods:title id="78FC79BC7947EA6A6E13FAAFC8AAC667">New taxa of Carpomyini, with special emphasis on Goniglossum (Diptera: Tephritidae: Trypetinae)</mods:title>
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<mods:namePart id="1440C388711FCE7F0FACC1BF4BFF53A4">Freidberg, Amnon</mods:namePart>
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<mods:title id="2177B2EFAEF8F2DC95A2A83F2FD9043B">Zootaxa</mods:title>
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<mods:date id="83E92E40D6F43BFDDAFF4EE6E97BCC7B">2016</mods:date>
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<treatment id="03D087C843119D3E5FB3FC1BFE9AFF60" ID-DOI="http://doi.org/10.5281/zenodo.6086595" ID-GBIF-Taxon="123207571" ID-Zenodo-Dep="6086595" LSID="urn:lsid:plazi:treatment:03D087C843119D3E5FB3FC1BFE9AFF60" httpUri="http://treatment.plazi.org/id/03D087C843119D3E5FB3FC1BFE9AFF60" lastPageId="5" lastPageNumber="58" pageId="3" pageNumber="57">
<subSubSection id="C363655543119D385FB3FC1BFE53FC58" box="[151,425,978,1004]" pageId="3" pageNumber="57" type="nomenclature">
<paragraph id="8BC636DE43119D385FB3FC1BFE53FC58" blockId="3.[151,425,978,1004]" box="[151,425,978,1004]" pageId="3" pageNumber="57">
<heading id="D08E81B243119D385FB3FC1BFE53FC58" bold="true" box="[151,425,978,1004]" fontSize="11" level="1" pageId="3" pageNumber="57" reason="1">
<taxonomicName id="4C794D5D43119D385FB3FC1BFE53FC58" authority="Rondani" authorityName="Rondani" box="[151,425,978,1004]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385FB3FC1BFE53FC58" bold="true" box="[151,425,978,1004]" pageId="3" pageNumber="57">
<emphasis id="B90DEACC43119D385FB3FC1BFEC2FC58" bold="true" box="[151,312,978,1004]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
Rondani
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363655543119D385FB3FBD0FB6FFB3F" pageId="3" pageNumber="57" type="reference_group">
<paragraph id="8BC636DE43119D385FB3FBD0FB5FFB9B" blockId="3.[151,1436,1049,1163]" box="[151,1189,1049,1071]" pageId="3" pageNumber="57">
<treatmentCitationGroup id="AB6911F043119D385FB3FBD0FB5FFB9B" box="[151,1189,1049,1071]" pageId="3" pageNumber="57">
<taxonomicName id="4C794D5D43119D385FB3FBD0FE17FB9B" authority="Rondani, 1856: 110" authorityName="Rondani" authorityPageNumber="110" authorityYear="1856" box="[151,493,1049,1071]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385FB3FBD0FEE7FB9B" box="[151,285,1049,1071]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
<treatmentCitation id="0AD810CF43119D385E07FBD0FE17FB9B" author="Rondani" box="[291,493,1049,1071]" page="110" pageId="3" pageNumber="57" year="1856">
<bibRefCitation id="EFE84B2F43119D385E07FBD0FE41FB9B" author="Rondani" box="[291,443,1049,1071]" pageId="3" pageNumber="57" refString="Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. 1: Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familiae et strips agregata. Parmae [Parma], 228 pp." type="book" year="1856">Rondani, 1856</bibRefCitation>
: 110
</treatmentCitation>
</taxonomicName>
. Type species:
<taxonomicName id="4C794D5D43119D385DA2FBD3FC50FB9B" authority="Meigen" authorityName="Meigen" box="[646,938,1049,1071]" class="Insecta" family="Lauxaniidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="wiedemanni">
<emphasis id="B90DEACC43119D385DA2FBD3FCAEFB9B" box="[646,852,1049,1071]" italics="true" pageId="3" pageNumber="57">Trypeta wiedemanni</emphasis>
Meigen
</taxonomicName>
, by original designation.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC636DE43119D385FB3FBF1FB6FFB3F" blockId="3.[151,1436,1049,1163]" pageId="3" pageNumber="57">
Partial synonymy:
<bibRefCitation id="EFE84B2F43119D385E73FBF1FE1EFBFA" author="Hendel" box="[343,484,1080,1102]" pageId="3" pageNumber="57" refString="Hendel, F. (1927) Trypetidae. In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. Vol. 5, 221 pp." type="book" year="1927">Hendel, 1927</bibRefCitation>
: 88 (Palaearctic fauna [
<taxonomicName id="4C794D5D43119D385DF0FBF1FC9CFBFA" ID-CoL="87G7Q" box="[724,870,1080,1102]" class="Insecta" family="Tephritidae" genus="Gonioglossum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385DF0FBF1FC9CFBFA" box="[724,870,1080,1102]" italics="true" pageId="3" pageNumber="57">Gonioglossum</emphasis>
</taxonomicName>
, sic]).
<bibRefCitation id="EFE84B2F43119D385C8AFBF1FBCAFBFA" author="Seguy" box="[942,1072,1080,1102]" pageId="3" pageNumber="57" refString="Seguy, E. (1934) Faune de France. 28. Dipteres (Brachyceres). Paul Lechevalier, Paris. 832 pp + 27 plates." type="book" year="1934">Séguy, 1934</bibRefCitation>
: 118 (French fauna [
<taxonomicName id="4C794D5D43119D385A27FBF1FA6FFBFA" ID-CoL="87G7Q" box="[1283,1429,1080,1102]" class="Insecta" family="Tephritidae" genus="Gonioglossum" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385A27FBF1FA6FFBFA" box="[1283,1429,1080,1102]" italics="true" pageId="3" pageNumber="57">Gonioglossum</emphasis>
</taxonomicName>
, sic]).
<bibRefCitation id="EFE84B2F43119D385E25FB9FFDDAFBD8" author="Freidberg" box="[257,544,1110,1132]" pageId="3" pageNumber="57" refString="Freidberg, A. &amp; Kugler, J. (1989) Fauna Palaestina. Insecta IV. Diptera: Tephritidae. The Israel Academy of Sciences and Humanities, Jerusalem, 212 pp + 8 plates + 1 map." type="book" year="1989">Freidberg and Kugler, 1989</bibRefCitation>
: 188 (Israel fauna, redescription).
<bibRefCitation id="EFE84B2F43119D385CA4FB9EFC00FBD8" author="Merz" box="[896,1018,1110,1132]" pageId="3" pageNumber="57" refString="Merz, B. (1994) Insecta Helvetica Fauna. Diptera Tephritidae, 10, 1 - 198." type="journal article" year="1994">Merz, 1994</bibRefCitation>
: 103 (Swiss fauna).
<bibRefCitation id="EFE84B2F43119D385BEEFB9FFA96FBD8" author="Norrbom" box="[1226,1388,1110,1132]" pageId="3" pageNumber="57" refString="Norrbom, A. L. (1997) The genus Carpomya Costa (Diptera: Tephritidae): New synonymy, description of first American species, and phylogenetic analysis. Proceedings of the Entomological Society of Washington, 99 (2), 338 - 347." type="journal article" year="1997">Norrbom, 1997</bibRefCitation>
: 340 (key, cladistics and synonymy).
<bibRefCitation id="EFE84B2F43119D385D29FBBCFD77FB3F" author="White" box="[525,653,1141,1163]" pageId="3" pageNumber="57" refString="White, I. M. (1998) Tephritid flies Diptera Tephritidae. Handbooks for the Identification of British insects 10 (5 a), Royal Entomological Society of London., 134 pp." type="book" year="1998">White, 1998</bibRefCitation>
: 37 (British fauna).
<bibRefCitation id="EFE84B2F43119D385C7CFBBCFC31FB3F" author="Smit" box="[856,971,1141,1163]" pageId="3" pageNumber="57" refString="Smit, J. T. (2010) De Nederlandse boorvliegen (Tephritidae). Entomologischen Tabellen 5. Groen Media Services, Leiderdorp. 159 pp. [in Dutch]" type="book" year="2010">Smit, 2010</bibRefCitation>
: 128 (Dutch fauna).
</paragraph>
</subSubSection>
<subSubSection id="C363655543119D385FB3FB7EFD90FAA3" pageId="3" pageNumber="57" type="description">
<paragraph id="8BC636DE43119D385FB3FB7EFD90FAA3" blockId="3.[151,1436,1207,1883]" pageId="3" pageNumber="57">
<bibRefCitation id="EFE84B2F43119D385FB3FB7EFEB5FB64" author="Norrbom" box="[151,335,1207,1232]" pageId="3" pageNumber="57" refString="Norrbom, A. L. (1997) The genus Carpomya Costa (Diptera: Tephritidae): New synonymy, description of first American species, and phylogenetic analysis. Proceedings of the Entomological Society of Washington, 99 (2), 338 - 347." type="journal article" year="1997">Norrbom (1997)</bibRefCitation>
synonymized
<taxonomicName id="4C794D5D43119D385ED6FB7EFD7CFB64" box="[498,646,1207,1232]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385ED6FB7EFD7CFB64" box="[498,646,1207,1232]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
under
<taxonomicName id="4C794D5D43119D385DF0FB71FCB0FB7B" box="[724,842,1208,1231]" class="Insecta" family="Tephritidae" genus="Carpomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385DF0FB71FCB0FB7B" box="[724,842,1208,1231]" italics="true" pageId="3" pageNumber="57">Carpomya</emphasis>
</taxonomicName>
, and transferred
<taxonomicName id="4C794D5D43119D385B23FB71FB57FB64" box="[1031,1197,1207,1232]" class="Insecta" family="Lauxaniidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="wiedemanni">
<emphasis id="B90DEACC43119D385B23FB71FB57FB64" box="[1031,1197,1207,1232]" italics="true" pageId="3" pageNumber="57">G. wiedemanni</emphasis>
</taxonomicName>
to
<taxonomicName id="4C794D5D43119D385BF5FB71FABDFB7B" box="[1233,1351,1208,1231]" class="Insecta" family="Tephritidae" genus="Carpomya" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385BF5FB71FABDFB7B" box="[1233,1351,1208,1231]" italics="true" pageId="3" pageNumber="57">Carpomya</emphasis>
</taxonomicName>
. Here I reinstate
<taxonomicName id="4C794D5D43119D385FDBFB15FE68FB41" box="[255,402,1244,1269]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385FDBFB15FE68FB41" box="[255,402,1244,1269]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
as a valid genus (new status), return
<taxonomicName id="4C794D5D43119D385C63FB15FC34FB41" box="[839,974,1244,1269]" class="Insecta" family="Lauxaniidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="wiedemanni">
<emphasis id="B90DEACC43119D385C63FB15FC34FB41" box="[839,974,1244,1269]" italics="true" pageId="3" pageNumber="57">wiedemanni</emphasis>
</taxonomicName>
to its former genus (
<taxonomicName id="4C794D5D43119D385BE2FB15FAA0FB41" box="[1222,1370,1244,1269]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385BE2FB15FAA0FB41" box="[1222,1370,1244,1269]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
), and describe a new species,
<taxonomicName id="4C794D5D43119D385E87FB36FD99FAAC" box="[419,611,1279,1304]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="species" species="liat">
<emphasis id="B90DEACC43119D385E87FB36FD99FAAC" box="[419,611,1279,1304]" italics="true" pageId="3" pageNumber="57">Goniglossum liat</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C363655543119D385FE3FAEDFD7EF9A1" pageId="3" pageNumber="57" type="diagnosis">
<paragraph id="8BC636DE43119D385FE3FAEDFD7EF9A1" blockId="3.[151,1436,1207,1883]" pageId="3" pageNumber="57">
<emphasis id="B90DEACC43119D385FE3FAEDFEBBFA89" bold="true" box="[199,321,1316,1341]" pageId="3" pageNumber="57">Diagnosis.</emphasis>
Although clearly a carpomyine,
<taxonomicName id="4C794D5D43119D385DEEFAEDFCA4FA89" box="[714,862,1316,1341]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385DEEFAEDFCA4FA89" box="[714,862,1316,1341]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
is not only easily distinguishable from all other genera of the subtribe but is also unique among all trypetines and related taxa in possessing a long proboscis. A long proboscis is encountered occasionally among the
<taxonomicName id="4C794D5D43119D385C30FAA5FC69FA31" box="[788,915,1388,1413]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="subFamily" subFamily="Tephritinae">Tephritinae</taxonomicName>
but is entirely absent in the other groups of
<taxonomicName id="4C794D5D43119D385FB3FA46FEE0FA1C" box="[151,282,1423,1448]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
. In
<taxonomicName id="4C794D5D43119D385E67FA46FE2DFA1C" box="[323,471,1423,1448]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385E67FA46FE2DFA1C" box="[323,471,1423,1448]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
the long proboscis is apparently associated with the peculiar shape of the head which is extended in an unusual way along a posterodorsal-anteroventral axis. Other characters, unique to
<taxonomicName id="4C794D5D43119D385A2CFA7DFA66FA79" box="[1288,1436,1460,1485]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="B90DEACC43119D385A2CFA7DFA66FA79" box="[1288,1436,1460,1485]" italics="true" pageId="3" pageNumber="57">Goniglossum</emphasis>
</taxonomicName>
when compared to other
<taxonomicName id="4C794D5D43119D385E88FA1EFDC0FA44" box="[428,570,1495,1520]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="57" phylum="Arthropoda" rank="subTribe" subTribe="Carpomyina">Carpomyina</taxonomicName>
, are the lack of a posterodistal lobe in cell cup and the arrow-shaped, apically serrated aculeus (with 515 marginal teeth).
</paragraph>
</subSubSection>
<subSubSection id="C363655543119D3F5FE3F9D6FCD0F80E" lastPageId="4" lastPageNumber="58" pageId="3" pageNumber="57" type="description">
<paragraph id="8BC636DE43119D385FE3F9D6FE18F8EF" blockId="3.[151,1436,1207,1883]" pageId="3" pageNumber="57">
<emphasis id="B90DEACC43119D385FE3F9D6FE8EF98C" bold="true" box="[199,372,1567,1592]" pageId="3" pageNumber="57">Redescription.</emphasis>
<emphasis id="B90DEACC43119D385E5FF9D6FE42F98C" box="[379,440,1567,1592]" italics="true" pageId="3" pageNumber="57">Head</emphasis>
(
<figureCitation id="13422A5B43119D385EE2F9D6FDF0F98C" box="[454,522,1567,1592]" captionStart="FIGURE 2" captionStartId="4.[151,250,1121,1143]" captionTargetBox="[483,1100,364,966]" captionTargetId="figure@4.[473,1114,193,1100]" captionTargetPageId="4" captionText="FIGURE 2. Goniglossum liat, head, lateral view. 2 a. photo. 2 b. drawing (drawing, reproduced by permission, from: A. Freidberg &amp; J. Kugler, Diptera: Tephritidae (Fauna Palaestina: Insecta, IV), Jerusalem: The Israel Academy of Sciences and Humanities, 1989, fig. 201 (by Walter Ferguson), p. 190 © The Israel Academy of Sciences and Humanities)." httpUri="https://zenodo.org/record/267231/files/figure.png" pageId="3" pageNumber="57">Fig. 2</figureCitation>
): Frons flat, more than twice as wide as eye; frontal vitta with fine setulae; frontofacial angle about 135o; face with prominent carina; ventral part of carina broader and protuberant; angle between parafacial and gena markedly protuberant, with coarse setulae; occiput strongly convex; eye rather oblique, longest diameter about 2.5 times as long as shortest diameter. First flagellomere about 2.5 times as long as high, with or without distinct apicodorsal point; arista almost bare. Proboscis geniculate; haustellum about as long as longest diameter of eye; labella slightly longer than haustellum; palpus about as long as antenna. Setae mostly black; postocellar seta sometimes yellowish; both vertical setae black, ocellar seta about as long as lateral vertical seta; 2 orbital setae, anterior orbital seta longer than postocellar seta; 3 frontal setae; genal seta usually indistinct; postocullar setae fine, brown.
</paragraph>
<caption id="DF06665643169D3F5FB3FBA8FB17FB00" httpUri="https://zenodo.org/record/267231/files/figure.png" pageId="4" pageNumber="58" targetBox="[483,1100,364,966]" targetPageId="4">
<paragraph id="8BC636DE43169D3F5FB3FBA8FB17FB00" blockId="4.[151,1436,1121,1204]" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FB3FBA8FEEDFBC2" bold="true" box="[151,279,1121,1143]" pageId="4" pageNumber="58">FIGURE 2.</emphasis>
<taxonomicName id="4C794D5D43169D3F5E07FBA8FE2FFBC3" box="[291,469,1121,1143]" class="Insecta" family="Tephritidae" genus="Goniglossum" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="species" species="liat">
<emphasis id="B90DEACC43169D3F5E07FBA8FE2FFBC3" box="[291,469,1121,1143]" italics="true" pageId="4" pageNumber="58">Goniglossum liat</emphasis>
</taxonomicName>
, head, lateral view. 2a. photo. 2b. drawing (drawing, reproduced by permission, from: A. Freidberg &amp; J. Kugler,
<emphasis id="B90DEACC43169D3F5EACFB48FDA1FB22" box="[392,603,1152,1174]" italics="true" pageId="4" pageNumber="58">
<taxonomicName id="4C794D5D43169D3F5EACFB48FE20FB22" box="[392,474,1153,1174]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="order">Diptera</taxonomicName>
:
<taxonomicName id="4C794D5D43169D3F5EC3FB49FDA1FB22" box="[487,603,1152,1174]" class="Insecta" family="Tephritidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="family">Tephritidae</taxonomicName>
</emphasis>
(Fauna
<taxonomicName id="4C794D5D43169D3F5D95FB49FCE6FB22" box="[689,796,1152,1174]" class="Arachnida" family="Zodariidae" genus="Palaestina" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="58" phylum="Arthropoda" rank="genus">Palaestina</taxonomicName>
: Insecta, IV), Jerusalem: The Israel Academy of Sciences and Humanities, 1989, fig. 201 (by Walter Ferguson), p. 190 © The Israel Academy of Sciences and Humanities).
</paragraph>
</caption>
<paragraph id="8BC636DE43169D3F5FE3FB28FD73FA3E" blockId="4.[151,1436,1249,2014]" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FE3FB28FEE2FB4E" box="[199,280,1249,1274]" italics="true" pageId="4" pageNumber="58">Thorax</emphasis>
: Mesonotum about 1.01.1 times as long as wide; scutellum strongly convex. Chaetotaxy normal for the tribe; setae mostly black; scapulars vary from yellow to blackish; dorsocentral seta aligned with or slightly anterior to anterior supra-alar seta; 2 pairs of equal scutellar setae and 2 anepisternal setae present; Mesonotum and pleuron with elaborate dark (brown or black) pattern over contrasting paler (white or yellow) background. See species descriptions and
<figureCitation id="13422A5B43169D3F5E89FAB8FDEFFA3E" box="[429,533,1393,1418]" captionStart-0="FIGURES 3 5" captionStart-1="FIGURES 6 8" captionStartId-0="6.[151,264,1228,1251]" captionStartId-1="7.[151,264,855,878]" captionTargetBox-0="[475,1111,309,1203]" captionTargetBox-1="[430,1161,199,825]" captionTargetId-0="figure@6.[473,1114,301,1208]" captionTargetId-1="figure@7.[340,1247,193,834]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 3 5. Goniglossum, thorax. 3. G. liat, mesonotum, dorsal view, drawing. 4. G. liat, thorax, posterodorsal view, photo. 5. G. wiedemanni, thorax, posterodorsal view, photo. (drawing reproduced, by permission, from: A. Freidberg &amp; J. Kugler, Diptera: Tephritidae (Fauna Palaestina: Insecta, IV), Jerusalem: The Israel Academy of Sciences and Humanities, 1989, fig. 202 (by Walter Ferguson), p. 190 © The Israel Academy of Sciences and Humanities)." captionText-1="FIGURES 6 8. Goniglossum, thorax. 6. G. liat, thorax, lateral view. 7. G. wiedemanni, thorax, lateral view. 8. G. liat, wing." httpUri-0="https://zenodo.org/record/267232/files/figure.png" httpUri-1="https://zenodo.org/record/267233/files/figure.png" pageId="4" pageNumber="58">Figs. 37</figureCitation>
for detail.
</paragraph>
<paragraph id="8BC636DE43169D3F5FE3FA5FFC4DFA66" blockId="4.[151,1436,1249,2014]" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FE3FA5FFF06FA19" box="[199,252,1430,1453]" italics="true" pageId="4" pageNumber="58">Legs</emphasis>
: Mostly yellow; femora sometimes brownish. setae brown to black; Setulae yellowish; midfemur and hindfemur in male with comb of rather short, fine, erect setae ventrally.
</paragraph>
<paragraph id="8BC636DE43169D3F5FE3FA17FD5EF8B6" blockId="4.[151,1436,1249,2014]" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FE3FA17FF05FA41" box="[199,255,1502,1525]" italics="true" pageId="4" pageNumber="58">Wing</emphasis>
(
<figureCitation id="13422A5B43169D3F5E34FA14FEAFFA42" box="[272,341,1501,1526]" captionStart="FIGURES 6 8" captionStartId="7.[151,264,855,878]" captionTargetBox="[430,1161,199,825]" captionTargetId="figure@7.[340,1247,193,834]" captionTargetPageId="7" captionText="FIGURES 6 8. Goniglossum, thorax. 6. G. liat, thorax, lateral view. 7. G. wiedemanni, thorax, lateral view. 8. G. liat, wing." httpUri="https://zenodo.org/record/267233/files/figure.png" pageId="4" pageNumber="58">Fig. 8</figureCitation>
): Enlarged costal setae (near costal break) short, not much longer than adjacent setulae; vein R4+5 dorsally with 16 setulae, including 13 (usually 23) setulae at node, rarely with some (13) setulae along proximal section, ventrally with 03 setulae at node (usually 0). Vein R2+3 with stump vein extending anteriorly, approximately aligned with crossvein DM-Cu, usually extending to 0.500.66 of width of cell r2+3; crossvein R-M at mid-length of cell dm; veins R4+5 and M somewhat divergent towards wing apex; vein A1+Cu2 slightly curved near vein Cu2; cell bcu without or with indistinct posterodistal point. Wing pattern banded, with bands partly yellow, partly dark brown or blackish. Basal band united with 2nd band between Sc and vein M, and 3rd band united with apical band between vein C and vein R4+5.
</paragraph>
<paragraph id="8BC636DE43169D3F5FE3F8D8FCD0F80E" blockId="4.[151,1436,1249,2014]" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FE3F8D8FECBF89E" box="[199,305,1809,1834]" italics="true" pageId="4" pageNumber="58">Abdomen</emphasis>
: Coloration strongly variable both inter- and intra-specifically, with dark pattern usually less contrasted than on thorax and less obvious in
<taxonomicName id="4C794D5D43169D3F5D88F8FFFCAEF8FA" box="[684,852,1845,1870]" class="Insecta" family="Lauxaniidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="species" species="wiedemanni">
<emphasis id="B90DEACC43169D3F5D88F8FFFCAEF8FA" box="[684,852,1845,1870]" italics="true" pageId="4" pageNumber="58">G. wiedemanni</emphasis>
</taxonomicName>
than in
<emphasis id="B90DEACC43169D3F5C93F8FFFC00F8F9" box="[951,1018,1845,1870]" italics="true" pageId="4" pageNumber="58">G. l i a t</emphasis>
. Epandrium with elongate surstylus (
<figureCitation id="13422A5B43169D3F5FBBF890FEE0F8C5" box="[159,282,1881,1906]" captionStart="FIGURES 9 12" captionStartId="7.[151,264,1855,1878]" captionTargetBox="[484,1099,934,1832]" captionTargetId="figure@7.[473,1114,927,1834]" captionTargetPageId="7" captionText="FIGURES 9 12. Goniglossum spp., male terminalia. 9. G. liat, epandrium, posterior view. 10. G. liat, epandrium, lateral view. 11. G. liat, phallus. 12. G. wiedemanni, phallus (Solid bar indicates the length of the “ bubble-like structure ”; dashed bar indicates the length of the distal part of the sclerotization)." httpUri="https://zenodo.org/record/267234/files/figure.png" pageId="4" pageNumber="58">Figs. 9, 10</figureCitation>
), similar to
<taxonomicName id="4C794D5D43169D3F5E80F893FC9CF8C5" authority="Norrbom" authorityName="Norrbom" authorityYear="1997" box="[420,870,1881,1906]" class="Insecta" family="Tephritidae" genus="Carpomya" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="species" species="tica">
<emphasis id="B90DEACC43169D3F5E80F893FE0BF8C5" box="[420,497,1882,1905]" italics="true" pageId="4" pageNumber="58">C. tica</emphasis>
<bibRefCitation id="EFE84B2F43169D3F5EDCF890FC9CF8C5" author="Norrbom" box="[504,870,1881,1906]" pageId="4" pageNumber="58" refString="Norrbom, A. L. (1997) The genus Carpomya Costa (Diptera: Tephritidae): New synonymy, description of first American species, and phylogenetic analysis. Proceedings of the Entomological Society of Washington, 99 (2), 338 - 347." type="journal article" year="1997">
Norrbom (1997: 343,
<figureCitation id="13422A5B43169D3F5DD4F890FCA4F8C5" box="[752,862,1881,1906]" captionStart="FIGURES 3 5" captionStartId="6.[151,264,1228,1251]" captionTargetBox="[475,1111,309,1203]" captionTargetId="figure@6.[473,1114,301,1208]" captionTargetPageId="6" captionText="FIGURES 3 5. Goniglossum, thorax. 3. G. liat, mesonotum, dorsal view, drawing. 4. G. liat, thorax, posterodorsal view, photo. 5. G. wiedemanni, thorax, posterodorsal view, photo. (drawing reproduced, by permission, from: A. Freidberg &amp; J. Kugler, Diptera: Tephritidae (Fauna Palaestina: Insecta, IV), Jerusalem: The Israel Academy of Sciences and Humanities, 1989, fig. 202 (by Walter Ferguson), p. 190 © The Israel Academy of Sciences and Humanities)." httpUri="https://zenodo.org/record/267232/files/figure.png" pageId="4" pageNumber="58">Figs. 3, 4</figureCitation>
)
</bibRefCitation>
</taxonomicName>
; glans strongly sclerotized, with large bubble-like structure in ventrobasal part of sclerotization (
<figureCitation id="13422A5B43169D3F5D80F8B4FCD6F822" box="[676,812,1917,1942]" captionStart="FIGURES 9 12" captionStartId="7.[151,264,1855,1878]" captionTargetBox="[484,1099,934,1832]" captionTargetId="figure@7.[473,1114,927,1834]" captionTargetPageId="7" captionText="FIGURES 9 12. Goniglossum spp., male terminalia. 9. G. liat, epandrium, posterior view. 10. G. liat, epandrium, lateral view. 11. G. liat, phallus. 12. G. wiedemanni, phallus (Solid bar indicates the length of the “ bubble-like structure ”; dashed bar indicates the length of the distal part of the sclerotization)." httpUri="https://zenodo.org/record/267234/files/figure.png" pageId="4" pageNumber="58">Figs. 1112</figureCitation>
, bars). Tergite 6 of female much shorter than tergite 5; apex of aculeus arrow-shaped and serrated (
<figureCitation id="13422A5B43169D3F5DA7F868FD29F80E" box="[643,723,1953,1978]" captionStart="FIGURES 13 15" captionStartId="8.[151,264,1121,1144]" captionTargetBox="[477,1107,196,1093]" captionTargetId="figure@8.[472,1114,193,1100]" captionTargetPageId="8" captionText="FIGURES 13 15. Goniglossum spp., female terminalia. 13. G. liat, aculeus: 13 a. whole, 13 b. apex enlarged. 14. G. wiedemanni, aculeus: 14 a. whole, 14 b. apex enlarged. 15. G. liat, spermatheca: 15 a. spherical to ovoid, 15 b. elongate." httpUri="https://zenodo.org/record/267235/files/figure.png" pageId="4" pageNumber="58">Fig. 13</figureCitation>
b, 14b).
</paragraph>
</subSubSection>
<subSubSection id="C363655543169D3E5FE3F80CFE9AFF60" lastPageId="5" lastPageNumber="59" pageId="4" pageNumber="58" type="biology_ecology">
<paragraph id="8BC636DE43169D3E5FE3F80CFE9AFF60" blockId="4.[151,1436,1249,2014]" lastBlockId="5.[151,1436,151,213]" lastPageId="5" lastPageNumber="59" pageId="4" pageNumber="58">
<emphasis id="B90DEACC43169D3F5FE3F80CFE10F86A" bold="true" box="[199,490,1989,2014]" pageId="4" pageNumber="58">Host plants and biology.</emphasis>
The biology of
<taxonomicName id="4C794D5D43169D3F5D86F80FFCB3F86A" box="[674,841,1989,2014]" class="Insecta" family="Lauxaniidae" genus="Trypeta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="58" phylum="Arthropoda" rank="species" species="wiedemanni">
<emphasis id="B90DEACC43169D3F5D86F80FFCB3F86A" box="[674,841,1989,2014]" italics="true" pageId="4" pageNumber="58">G. wiedemanni</emphasis>
</taxonomicName>
was studied in detail by
<bibRefCitation id="EFE84B2F43169D3F5B41F80CFAE2F86A" author="Silvestri" box="[1125,1304,1989,2014]" pageId="4" pageNumber="58" refString="Silvestri, F. (1920) La mosca della Brionia, Gonyglossum wiedemanni Meig. (Diptera: Trypaneidae). Bollettino del Laboratorio di Zoologia Generale e Agraria della Real Scuola Superiore d'Agricoltura in Portici, 14, 205 - 215." type="journal article" year="1920">Silvestri (1920)</bibRefCitation>
. The larvae of both species develop in berries of
<taxonomicName id="4C794D5D43179D3E5D61FF50FD5AFF04" box="[581,672,153,176]" class="Magnoliopsida" family="Cucurbitaceae" genus="Bryonia" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="59" phylum="Tracheophyta" rank="genus">
<emphasis id="B90DEACC43179D3E5D61FF50FD5AFF04" box="[581,672,153,176]" italics="true" pageId="5" pageNumber="59">Bryonia</emphasis>
</taxonomicName>
spp. (
<taxonomicName id="4C794D5D43179D3E5DCCFF5EFC75FF04" box="[744,911,151,176]" class="Magnoliopsida" family="Cucurbitaceae" kingdom="Plantae" order="Cucurbitales" pageId="5" pageNumber="59" phylum="Tracheophyta" rank="family">Cucurbitaceae</taxonomicName>
). A brief overview of the biology is provided here under
<emphasis id="B90DEACC43179D3E5E31FF74FEAFFF60" box="[277,341,188,213]" italics="true" pageId="5" pageNumber="59">G. l i a t</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>