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<document id="57F8FCF1BC0E6D396B5674C604878CE5" ID-DOI="10.5281/zenodo.4665621" ID-ISSN="1638-9395" ID-Zenodo-Dep="4665621" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617712844033" checkinUser="felipe" docAuthor="Venczel, Márton &amp; Stiuc, Emanoil" docDate="2008" docId="03D08790FFC2FFE0548AA2A4FCCF5370" docLanguage="en" docName="Geodiversitas.30.4.731-763.pdf" docOrigin="Geodiversitas 30 (4)" docStyle="DocumentStyle:9DCE1246056655C2F9CBE9822E831E4E.3:Geodiversitas.2000-2008.journal_article.type3" docStyleId="9DCE1246056655C2F9CBE9822E831E4E" docStyleName="Geodiversitas.2000-2008.journal_article.type3" docStyleVersion="3" docTitle="Lissotriton Bell 1839" docType="treatment" docVersion="3" lastPageNumber="739" masterDocId="FFE9FFE8FFC5FFE85703A736FFA55105" masterDocTitle="Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania" masterLastPageNumber="763" masterPageNumber="731" pageNumber="738" updateTime="1698901057390" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods id="BB8A47118EFF782A0C113BF66E822AC3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="91B5A8C17609912846665DBE578094E6">
<mods:title id="D377ACE38FFED69533EEC5E574B851C5">Late middle Miocene amphibians and squamate reptiles from Taut ,, Romania</mods:title>
</mods:titleInfo>
<mods:name id="A2F2848B4F82E2414507C493DA87DA5B" type="personal">
<mods:role id="0D12192F123816D62AB2C7B897710B90">
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<mods:namePart id="7E9276A17EC7C3026CE107F96AEC8945">Venczel, Márton</mods:namePart>
<mods:affiliation id="89BA4A120938151FD4C0EF2B65617254">Tării ‚ Crişurilor Museum, B-dul Dacia 1 - 3, RO- 410464 Oradea (Romania) mvenczel @ rdslink. ro</mods:affiliation>
<mods:nameIdentifier id="14ED61F3869C66A4CBE9AB84A62DC46F" type="email">mvenczel@rdslink.ro</mods:nameIdentifier>
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<mods:name id="39DA26CCA10FFC23E349A7B3D7C3DFD1" type="personal">
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</mods:role>
<mods:namePart id="821D1742DD036209DE9C2000A2966BAA">Stiuc, Emanoil</mods:namePart>
<mods:affiliation id="6AC395B6B52D54B196E0F6BB8C76DE9B">Speleological Institute “ Emil Racovit, ă ”, Calea 13 Septembrie nr. 13, RO- 050711 Bucureşti (Romania) stiucaemil @ yahoo. com.</mods:affiliation>
<mods:nameIdentifier id="C0913BFB5104F4FA50426819446102A1" type="email">stiucaemil@yahoo.com</mods:nameIdentifier>
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<mods:title id="F2CEA907A91289DD3AE44B448E3C18BE">Geodiversitas</mods:title>
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<mods:part id="2EFBD3CEFA4702285ECB0E7A5AF70077">
<mods:date id="DD3461C690A49E5A4766E40D738D8F95">2008</mods:date>
<mods:detail id="2D2A82E6C9A158BD55822185B0786735" type="volume">
<mods:number id="B5913E6523EFB2FDC28D0B790F376E2F">30</mods:number>
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<mods:detail id="58ECC503F5D2830D3EEC7452C8A8D52E" type="issue">
<mods:number id="3DB28AFC8B068D148E5DE33D786C936B">4</mods:number>
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<mods:classification id="3C45D6C4B16FEAA6A165BCA0A28B51CB">journal article</mods:classification>
<mods:identifier id="457455B1CE139202EB456AA76BFB20A4" type="DOI">10.5281/zenodo.4665621</mods:identifier>
<mods:identifier id="B2A3C30B7FA9A379B3A8EA62B2299B0B" type="ISSN">1638-9395</mods:identifier>
<mods:identifier id="DAB2587E884FD6A9DF5709F2E7A01603" type="Zenodo-Dep">4665621</mods:identifier>
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<treatment id="03D08790FFC2FFE0548AA2A4FCCF5370" LSID="urn:lsid:plazi:treatment:03D08790FFC2FFE0548AA2A4FCCF5370" httpUri="http://treatment.plazi.org/id/03D08790FFC2FFE0548AA2A4FCCF5370" lastPageId="8" lastPageNumber="739" pageId="7" pageNumber="738">
<subSubSection id="C363650DFFC2FFEF548AA2A4FC5954C9" pageId="7" pageNumber="738" type="nomenclature">
<paragraph id="8BC63686FFC2FFEF548AA2A4FC5954C9" blockId="7.[905,1057,1426,1484]" pageId="7" pageNumber="738">
<heading id="D08E81EAFFC2FFEF548AA2A4FC5954C9" centered="true" fontSize="11" level="2" pageId="7" pageNumber="738" reason="2">
<taxonomicName id="4C794D05FFC2FFEF548AA2A4FC5E54A9" ID-CoL="5FHR" authorityName="Bell" authorityYear="1839" box="[905,1019,1426,1452]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="7" pageNumber="738" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFC2FFEF548AA2A4FC5E54A9" bold="true" box="[905,1019,1426,1452]" italics="true" pageId="7" pageNumber="738">Lissotriton</emphasis>
</taxonomicName>
sp. (
<figureCitation id="13422A03FFC2FFEF54B6A287FC5054C9" box="[949,1013,1457,1484]" captionStart="FIG" captionStartId="7.[151,161,647,664]" captionTargetBox="[158,675,255,597]" captionTargetId="figure-15@7.[153,683,238,612]" captionTargetPageId="7" captionText="FIG. 4.— Lissotriton sp.:A, frontal (ISER Tt-0350);B, atlas (ISER Tt- 0351);C-C’’, presacral vertebra (ISER Tt-0352/1);A, C, dorsal views; B, C’, ventral views; C’’, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665634" httpUri="https://zenodo.org/record/4665634/files/figure.png" pageId="7" pageNumber="738">Fig. 4</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFC2FFEF55C9A2DDFC3F5732" pageId="7" pageNumber="738" type="materials_examined">
<paragraph id="8BC63686FFC2FFEF55C9A2DDFC3F5732" blockId="7.[714,1247,1515,1591]" pageId="7" pageNumber="738">
MATERIAL EXAMINED. — One frontal (
<collectionCode id="ED68AE43FFC2FFEF5340A2DDFBDC5706" box="[1091,1145,1515,1539]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="7" pageNumber="738">ISER</collectionCode>
Tt-0350), one atlas (
<collectionCode id="ED68AE43FFC2FFEF5436A133FCC85718" box="[821,877,1541,1565]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="7" pageNumber="738">ISER</collectionCode>
Tt-0351), eight presacral vertebrae (
<collectionCode id="ED68AE43FFC2FFEF55D0A129FCAC5732" box="[723,777,1567,1591]" country="Romania" httpUri="http://grbio.org/cool/911h-bwmh" name="Institutul Speologie Emil G. Racovita" pageId="7" pageNumber="738">ISER</collectionCode>
Tt-0352/1-8).
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFC2FFE055C9A166FEDA5514" lastPageId="8" lastPageNumber="739" pageId="7" pageNumber="738" type="description">
<paragraph id="8BC63686FFC2FFEF55C9A166FCFE576D" blockId="7.[714,1247,1616,1706]" box="[714,859,1616,1642]" pageId="7" pageNumber="738">DESCRIPTION</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A159FCB3578C" blockId="7.[714,1247,1616,1706]" box="[714,790,1647,1673]" pageId="7" pageNumber="738">
<emphasis id="B90DEA94FFC2FFEF55C9A159FCB3578C" box="[714,790,1647,1673]" italics="true" pageId="7" pageNumber="738">Frontal</emphasis>
</paragraph>
<paragraph id="8BC63686FFC2FFEF55C9A1B9FB7A57AF" blockId="7.[714,1247,1616,1706]" box="[714,1247,1679,1706]" pageId="7" pageNumber="738">The dorsal surface bears a fine sculpture formed by</paragraph>
<paragraph id="8BC63686FFC2FFE055C9A199FEE650D2" blockId="7.[714,1247,1711,1738]" lastBlockId="8.[151,685,254,471]" lastPageId="8" lastPageNumber="739" pageId="7" pageNumber="738">
shallow ridges and pits (
<figureCitation id="13422A03FFC2FFEF54CAA199FBBB57CF" box="[969,1054,1711,1738]" captionStart="FIG" captionStartId="7.[151,161,647,664]" captionTargetBox="[158,675,255,597]" captionTargetId="figure-15@7.[153,683,238,612]" captionTargetPageId="7" captionText="FIG. 4.— Lissotriton sp.:A, frontal (ISER Tt-0350);B, atlas (ISER Tt- 0351);C-C’’, presacral vertebra (ISER Tt-0352/1);A, C, dorsal views; B, C’, ventral views; C’’, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665634" httpUri="https://zenodo.org/record/4665634/files/figure.png" pageId="7" pageNumber="738">Fig. 4A</figureCitation>
). The supraorbital margin is low; parallel to the latter, a deep and widening furrow is directed anteriorly. The relatively short posterolateral process tapers distally, and its posterior edge only slightly exceeds the posterior margin of the frontal. This morphology is suggestive of an incomplete or a somewhat weak frontosquamosal arch.
</paragraph>
<paragraph id="8BC63686FFCDFFE05794A6CDFF6F5310" blockId="8.[149,686,507,788]" box="[151,202,507,533]" pageId="8" pageNumber="739">
<emphasis id="B90DEA94FFCDFFE05794A6CDFF6F5310" box="[151,202,507,533]" italics="true" pageId="8" pageNumber="739">Atlas</emphasis>
</paragraph>
<paragraph id="8BC63686FFCDFFE05794A52DFDEA5211" blockId="8.[149,686,507,788]" pageId="8" pageNumber="739">
In the single specimen only the centrum is preserved. The subcentral surface is slightly concave, triangleshaped and provided with two relatively large subcentral foramina (
<figureCitation id="13422A03FFCDFFE05656A54CFE025390" box="[341,423,634,661]" captionStart="FIG" captionStartId="7.[151,161,647,664]" captionTargetBox="[158,675,255,597]" captionTargetId="figure-15@7.[153,683,238,612]" captionTargetPageId="7" captionText="FIG. 4.— Lissotriton sp.:A, frontal (ISER Tt-0350);B, atlas (ISER Tt- 0351);C-C’’, presacral vertebra (ISER Tt-0352/1);A, C, dorsal views; B, C’, ventral views; C’’, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665634" httpUri="https://zenodo.org/record/4665634/files/figure.png" pageId="8" pageNumber="739">Fig. 4B</figureCitation>
). The articular surface of the paired atlantal condyles is of circular shape. The odontoid process is partially divided exhibiting two shallowly convex ventrolateral articular surfaces of oval shape. The atlantal cotyle is circular.
</paragraph>
<paragraph id="8BC63686FFCDFFE05794A40EFEF35257" blockId="8.[149,686,824,1041]" box="[151,342,824,850]" pageId="8" pageNumber="739">
<emphasis id="B90DEA94FFCDFFE05794A40EFEF35257" box="[151,342,824,850]" italics="true" pageId="8" pageNumber="739">Presacral vertebrae</emphasis>
</paragraph>
<paragraph id="8BC63686FFCDFFE05794A46EFEDA5514" blockId="8.[149,686,824,1041]" pageId="8" pageNumber="739">
All the specimens are small sized with relatively short and opisthocoelous centrum (
<figureCitation id="13422A03FFCDFFE05522A441FDC65297" box="[545,611,887,914]" captionStart="FIG" captionStartId="7.[151,161,647,664]" captionTargetBox="[158,675,255,597]" captionTargetId="figure-15@7.[153,683,238,612]" captionTargetPageId="7" captionText="FIG. 4.— Lissotriton sp.:A, frontal (ISER Tt-0350);B, atlas (ISER Tt- 0351);C-C’’, presacral vertebra (ISER Tt-0352/1);A, C, dorsal views; B, C’, ventral views; C’’, right lateral view. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4665634" httpUri="https://zenodo.org/record/4665634/files/figure.png" pageId="8" pageNumber="739">Fig. 4</figureCitation>
C-C’’). The neural arch is moderately vaulted and provided with a relatively high neural spine forking posteriorly. The dorsal margin of the latter structure is not widened laterally.
</paragraph>
</subSubSection>
<subSubSection id="C363650DFFCDFFE05794A300FCCF5370" pageId="8" pageNumber="739" type="discussion">
<paragraph id="8BC63686FFCDFFE05794A300FF5E554B" blockId="8.[149,686,1078,1706]" box="[151,251,1078,1104]" pageId="8" pageNumber="739">REMARKS</paragraph>
<paragraph id="8BC63686FFCDFFE05794A363FD0857AF" blockId="8.[149,686,1078,1706]" pageId="8" pageNumber="739">
Based on the available material, the above taxon may be securely placed in the genus
<taxonomicName id="4C794D05FFCDFFE056E0A343FDEB558B" authorityName="Bell" authorityYear="1839" box="[483,590,1141,1166]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE056E0A343FDEB558B" box="[483,590,1141,1166]" italics="true" pageId="8" pageNumber="739">Lissotriton</emphasis>
</taxonomicName>
. Among members of
<taxonomicName id="4C794D05FFCDFFE05620A3A3FE3555AB" authorityName="Bell" authorityYear="1839" box="[291,400,1173,1198]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE05620A3A3FE3555AB" box="[291,400,1173,1198]" italics="true" pageId="8" pageNumber="739">Lissotriton</emphasis>
</taxonomicName>
the postfrontal process is rather long in
<taxonomicName id="4C794D05FFCDFFE05635A383FD0955CA" authority="(Razoumovsky, 1789)" baseAuthorityName="Razoumovsky" baseAuthorityYear="1789" box="[310,684,1204,1231]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="helveticus">
<emphasis id="B90DEA94FFCDFFE05635A383FE1255CB" box="[310,439,1204,1230]" italics="true" pageId="8" pageNumber="739">L. helveticus</emphasis>
(Razoumovsky, 1789)
</taxonomicName>
and
<taxonomicName id="4C794D05FFCDFFE057C6A3E2FE6D55EA" authority="(Lataste, 1879)" baseAuthorityName="Lataste" baseAuthorityYear="1879" box="[197,456,1236,1263]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="boscai">
<emphasis id="B90DEA94FFCDFFE057C6A3E2FEBA55EB" box="[197,287,1236,1262]" italics="true" pageId="8" pageNumber="739">L. boscai</emphasis>
(Lataste, 1879)
</taxonomicName>
but is relatively short in
<taxonomicName id="4C794D05FFCDFFE057B1A3C2FDB5540B" authority="(Boulenger, 1880)" baseAuthorityName="Boulenger" baseAuthorityYear="1880" box="[178,528,1267,1294]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="montandoni">
<emphasis id="B90DEA94FFCDFFE057B1A3C2FEEF5408" box="[178,330,1267,1293]" italics="true" pageId="8" pageNumber="739">L. montandoni</emphasis>
(Boulenger, 1880)
</taxonomicName>
and
<taxonomicName id="4C794D05FFCDFFE05541A3C2FEF7542B" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="vulgaris">
<emphasis id="B90DEA94FFCDFFE05541A3C2FD085408" box="[578,685,1267,1293]" italics="true" pageId="8" pageNumber="739">L. vulgaris</emphasis>
(Linnaeus, 1758)
</taxonomicName>
, the later displaying a morphology fairly similar with that of ISER Tt-0350. The size and morphology of the trunk vertebrae, with relatively high neural arch and well-developed neural spine forking posteriorly, support also the taxonomic assignment of the above fossils.
<taxonomicName id="4C794D05FFCDFFE05571A284FE1E54E9" authority="(Herre, 1955)" baseAuthorityName="Herre" baseAuthorityYear="1955" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="roehrsi">
<emphasis id="B90DEA94FFCDFFE05571A284FEB954EE" italics="true" pageId="8" pageNumber="739">Lissotriton roehrsi</emphasis>
(Herre, 1955)
</taxonomicName>
, an extinct member of the genus, known from the early to late Miocene of Central Europe (
<bibRefCitation id="EFE84B77FFCDFFE0566FA127FDAB5729" author="SANCHIZ B." box="[364,526,1553,1580]" pageId="8" pageNumber="739" pagination="13 - 29" refId="ref17825" refString="SANCHIZ B. 1998 a. - Vertebrates from the early Miocene lignite deposits of the opencast mine Oberdorf (Western Styrian Basin, Austria). 2. Amphibia. Annalen des Naturhistorischen Museums in Wien 99 A: 13 - 29." type="journal article" year="1998">Sanchiz 1998a</bibRefCitation>
;
<bibRefCitation id="EFE84B77FFCDFFE05519A127FF76574E" author="BOHME M. &amp; ILG A." pageId="8" pageNumber="739" refId="ref16033" refString="BOHME M. &amp; ILG A. 2003. - FosFARbase, www. wahrestaerke. com (accessed on April, 2008)." type="book" year="2003">Böhme &amp; Ilg 2003</bibRefCitation>
), differs from the other ingroup taxa by having a relatively high neural spine provided with a distinct dorsal enlargement of the dorsal margin. Furthermore,
<bibRefCitation id="EFE84B77FFCDFFE0562DA1B9FE7E57AF" author="SANCHIZ B." box="[302,475,1679,1706]" pageId="8" pageNumber="739" pagination="13 - 29" refId="ref17825" refString="SANCHIZ B. 1998 a. - Vertebrates from the early Miocene lignite deposits of the opencast mine Oberdorf (Western Styrian Basin, Austria). 2. Amphibia. Annalen des Naturhistorischen Museums in Wien 99 A: 13 - 29." type="journal article" year="1998">Sanchiz (1998a)</bibRefCitation>
reported numerous
</paragraph>
<paragraph id="8BC63686FFCDFFE05796A199FD0957CF" blockId="8.[149,684,1711,1738]" box="[149,684,1711,1738]" pageId="8" pageNumber="739">
skeletal remains assigned to
<taxonomicName id="4C794D05FFCDFFE056C3A186FD8E57CC" authorityName="Bell" authorityYear="1839" box="[448,555,1712,1737]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE056C3A186FD8E57CC" box="[448,555,1712,1737]" italics="true" pageId="8" pageNumber="739">Lissotriton</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C794D05FFCDFFE0554DA186FD0657CC" authorityName="Rafinesque" authorityYear="1815" box="[590,675,1712,1737]" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE0554DA186FD0657CC" box="[590,675,1712,1737]" italics="true" pageId="8" pageNumber="739">Triturus</emphasis>
</taxonomicName>
)
</paragraph>
<paragraph id="8BC63686FFCDFFE055E4A742FB6D5180" blockId="8.[743,1224,115,136]" box="[743,1224,115,136]" pageId="8" pageNumber="739">
Late middle Miocene amphibians and reptiles from Taut,,
<collectingCountry id="F36E7616FFCDFFE05384A745FB6D5180" box="[1159,1224,115,133]" name="Romania" pageId="8" pageNumber="739">Romania</collectingCountry>
</paragraph>
<paragraph id="8BC63686FFCDFFE055C9A7C8FC715092" blockId="8.[713,1248,254,629]" pageId="8" pageNumber="739">
<taxonomicName id="4C794D05FFCDFFE055C9A7C8FCB7501D" baseAuthorityName="Herre" baseAuthorityYear="1955" box="[714,786,254,280]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="roehrsi">
<emphasis id="B90DEA94FFCDFFE055C9A7C8FCB7501D" box="[714,786,254,280]" italics="true" pageId="8" pageNumber="739">roehrsi</emphasis>
</taxonomicName>
from the late early Miocene (MN 4) of Obersdorf,
<collectingCountry id="F36E7616FFCDFFE0544BA62BFC3C503D" box="[840,921,285,312]" name="Austria" pageId="8" pageNumber="739">Austria</collectingCountry>
. The frontals are provided with a relatively long postfrontal process and with an extensive sculptured dorsal surface differing clearly from the Tauţ specimen.
</paragraph>
<paragraph id="8BC63686FFCDFFE055E2A6AAFCCF5370" blockId="8.[713,1248,254,629]" pageId="8" pageNumber="739">
The earliest fossil record of the genus
<taxonomicName id="4C794D05FFCDFFE05377A6ABFB7A50B3" authorityName="Bell" authorityYear="1839" box="[1140,1247,413,438]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE05377A6ABFB7A50B3" box="[1140,1247,413,438]" italics="true" pageId="8" pageNumber="739">Lissotriton</emphasis>
</taxonomicName>
might be from the late early Miocene (MN 4) of Béon 1,
<collectingCountry id="F36E7616FFCDFFE05424A6EDFCD750F3" box="[807,882,475,502]" name="France" pageId="8" pageNumber="739">France</collectingCountry>
, described under the name
<taxonomicName id="4C794D05FFCDFFE053A3A6EAFD4C5310" authorityName="Rafinesque" authorityYear="1815" class="Amphibia" family="Salamandridae" genus="Triturus" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="genus">
<emphasis id="B90DEA94FFCDFFE053A3A6EAFD4C5310" italics="true" pageId="8" pageNumber="739">Triturus</emphasis>
</taxonomicName>
aff.
<taxonomicName id="4C794D05FFCDFFE0541AA6CAFC325310" baseAuthorityName="Razoumovsky" baseAuthorityYear="1789" box="[793,919,507,533]" class="Amphibia" family="Salamandridae" genus="Lissotriton" kingdom="Animalia" order="Caudata" pageId="8" pageNumber="739" phylum="Chordata" rank="species" species="helveticus">
<emphasis id="B90DEA94FFCDFFE0541AA6CAFC325310" box="[793,919,507,533]" italics="true" pageId="8" pageNumber="739">T. helveticus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE84B77FFCDFFE054BCA6CDFB155313" author="RAGE J. - C. &amp; BAILON S." box="[959,1200,507,534]" pageId="8" pageNumber="739" pagination="413 - 441" refId="ref17513" refString="RAGE J. - C. &amp; BAILON S. 2005. - Amphibians and squamate reptiles from the late early Miocene (MN 4) of Beon 1 (Montreal-du-Gers, southwestern France). Geodiversitas 27 (3): 413 - 441." type="journal article" year="2005">Rage &amp; Bailon (2005)</bibRefCitation>
and Obersdorf,
<collectingCountry id="F36E7616FFCDFFE05446A52DFC375333" box="[837,914,539,566]" name="Austria" pageId="8" pageNumber="739">Austria</collectingCountry>
(Sanchiz 1988a), suggesting an already wide distribution of this group during late early Miocene.
</paragraph>
</subSubSection>
</treatment>
</document>
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