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<document id="A0F9921E5CA57D74025A319C3A6069C1" ID-DOI="10.5852/ejt.2021.742.1291" ID-GBIF-Dataset="62f9dc57-bade-4fbc-8af7-f92ff107593c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4655404" ID-ZooBank="23424EC3-8440-4C72-AD9E-C1EC3087F7CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617262825393" checkinUser="plazi" docAuthor="De Oliveira, Luiz Fernando M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="03CF87ADFF9B661BFD0A9756FD60FC7E" docLanguage="en" docName="ejt-742_1-77.pdf" docOrigin="European Journal of Taxonomy 742" docSource="http://dx.doi.org/10.5852/ejt.2021.742.1291" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tafana oliviae De Oliveira &amp; Brescovit 2021, sp. nov." docType="treatment" docUuid="A7E716C9-A0F8-4F4E-A451-E6D94A3E82CD" docUuidSource="ZooBank" docVersion="11" lastPageNumber="54" masterDocId="FFF6FFD5FFA8662EFF99965EFF99FFCF" masterDocTitle="Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="52" updateTime="1701999878271" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="01A954D0E267E30C92FBC24BA2F189D1">Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species</mods:title>
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<mods:namePart id="4F4CB55171FCD9686315D6FF6C688900">De Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:affiliation id="E1B3B54B257E5230D6EA621890B6ABEC">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. &amp; Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A43D066DD762CA22454B98E950C7E52A" type="email">luiz.moura.lfm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5939D4AB7BBB2D80686F7E9BAF4EC87E">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="71325CCF24EE5B2EB90F31FAE254EBC6">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="B496E569C484D565D78278CD3F007B44" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:title id="4D75619D281DCBE0861C93065A958303">European Journal of Taxonomy</mods:title>
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<mods:date id="1A5755DEAB076F16E2E99ED9FC384288">2021</mods:date>
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<treatment id="03CF87ADFF9B661BFD0A9756FD60FC7E" ID-DOI="http://doi.org/10.5281/zenodo.4658632" ID-GBIF-Taxon="180784544" ID-Zenodo-Dep="4658632" LSID="urn:lsid:zoobank.org:act:A7E716C9-A0F8-4F4E-A451-E6D94A3E82CD" httpUri="http://treatment.plazi.org/id/03CF87ADFF9B661BFD0A9756FD60FC7E" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52">
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<paragraph id="8BD936BBFF9B661DFD0A9756FC38FEEC" blockId="51.[360,1227,264,362]" box="[659,929,264,291]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFD0A9756FC38FEEC" box="[659,929,264,291]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="52" reason="2">
<taxonomicName id="4C664D38FF9B661DFD0A9756FCA6FEEC" ID-CoL="98Z35" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[659,831,264,291]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oliviae" status="sp. nov.">
<emphasis id="B912EAA9FF9B661DFD0A9756FCA6FEEC" bold="true" box="[659,831,264,291]" italics="true" pageId="51" pageNumber="52">Tafana oliviae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF9B661DFCDE9757FC38FEEC" box="[839,929,265,291]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C37C6530FF9B661DFEF19772FCE5FEA5" pageId="51" pageNumber="52" type="description">
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BD936BBFF9B661DFD21970EFCE5FEA5" blockId="51.[360,1227,264,362]" box="[696,892,335,362]" pageId="51" pageNumber="52">
<figureCitation id="135D2A3EFF9B661DFD21970EFC88FEA5" box="[696,785,336,362]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Figs 29</figureCitation>
,
<figureCitation id="135D2A3EFF9B661DFC84970EFCCAFEA5" box="[797,851,336,362]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="51" pageNumber="52">33G</figureCitation>
,
<figureCitation id="135D2A3EFF9B661DFCC69711FCE5FEA5" box="[863,892,335,362]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="51" pageNumber="52">35</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9B661DFF2497CFFBA5FDA7" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8BD936BBFF9B661DFF2497CFFEAFFE64" blockId="51.[189,310,401,427]" box="[189,310,401,427]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFF2497CFFEAFFE64" bold="true" box="[189,310,401,427]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B912EAA9FF9B661DFF2497CFFEAFFE64" bold="true" box="[189,310,401,427]" pageId="51" pageNumber="52">Diagnosis</emphasis>
</heading>
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<paragraph id="8BD936BBFF9B661DFF24979EFBA5FDA7" blockId="51.[188,1399,448,616]" pageId="51" pageNumber="52">
Males of
<taxonomicName id="4C664D38FF9B661DFEB1979EFE56FE15" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[296,463,448,474]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oliviae" status="sp. nov.">
<emphasis id="B912EAA9FF9B661DFEB1979EFE56FE15" box="[296,463,448,474]" italics="true" pageId="51" pageNumber="52">Tafana oliviae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF9B661DFE4F979FFDA9FE14" box="[470,560,449,475]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
differ from those of the other species in the
<taxonomicName id="4C664D38FF9B661DFBB5979EFB1EFE15" authorityName="MNHN" baseAuthorityName="Caporiacco" baseAuthorityYear="1955" box="[1068,1159,448,474]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="silhavyi">
<emphasis id="B912EAA9FF9B661DFBB5979EFB1EFE15" box="[1068,1159,448,474]" italics="true" pageId="51" pageNumber="52">silhavyi</emphasis>
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group by having the apophysis at the base of the embolus finger-shaped, retrolateral tibial apophysis bifid with anterior branch short and flattened, posterior branch triangular and median tibial apophysis bifid in dorsal position on the tibia (
<figureCitation id="135D2A3EFF9B661DFE989474FE14FD8B" box="[257,397,554,580]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Fig. 29CE</figureCitation>
). Females differ by having the epigynal plate with lateral lobes extremely projected to the sides and by having the small primary spermathecae (
<figureCitation id="135D2A3EFF9B661DFCC49413FC74FDA7" box="[861,1005,589,616]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Figs 29FG</figureCitation>
,
<figureCitation id="135D2A3EFF9B661DFC619410FBB6FDA7" box="[1016,1071,590,616]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="51" pageNumber="52">33G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9B661DFF2494D1FAC7FD17" box="[189,1374,655,728]" pageId="51" pageNumber="52" type="etymology">
<paragraph id="8BD936BBFF9B661DFF2494D1FEDDFD66" blockId="51.[189,324,655,681]" box="[189,324,655,681]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFF2494D1FEDDFD66" bold="true" box="[189,324,655,681]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B912EAA9FF9B661DFF2494D1FEDDFD66" bold="true" box="[189,324,655,681]" pageId="51" pageNumber="52">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF9B661DFF2494E0FAC7FD17" blockId="51.[189,1374,702,728]" box="[189,1374,702,728]" pageId="51" pageNumber="52">The specific name is a matronymic in memory of Olivia de Moura Oliveira, the first authors mother.</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9B661DFF2494A1FDF6FBD8" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8BD936BBFF9B661DFF2494A1FE33FCD6" blockId="51.[189,426,767,793]" box="[189,426,767,793]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFF2494A1FE33FCD6" bold="true" box="[189,426,767,793]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B912EAA9FF9B661DFF2494A1FE33FCD6" bold="true" box="[189,426,767,793]" pageId="51" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF9B661DFF739570FD97FC5F" blockId="51.[189,1398,814,912]" pageId="51" pageNumber="52">
<materialsCitation id="3B0E3CE6FF9B661DFF739570FD97FC5F" ID-GBIF-Occurrence="3066856323" collectingDate="1913-11" collectingDateMax="1913-12-31" collectingDateMin="1913-11" collectionCode="MACN-Ar" collectorName="M. J. Viana" country="Argentina" latitude="-27.934444" location="Santa Maria" longLatPrecision="20" longitude="-55.406944" municipality="Santa Maria" pageId="51" pageNumber="52" specimenCode="MACN-Ar 3565" specimenCount="1" specimenCount-male="1" stateProvince="Misiones" typeStatus="holotype">
<emphasis id="B912EAA9FF9B661DFF739570FEC5FC87" bold="true" box="[234,348,814,840]" pageId="51" pageNumber="52">
<typeStatus id="54DD8819FF9B661DFF739570FEC5FC87" box="[234,348,814,840]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F371762BFF9B661DFF24950CFEF1FCA3" box="[189,360,850,876]" name="Argentina" pageId="51" pageNumber="52">ARGENTINA</collectingCountry>
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;
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,
<collectingMunicipality id="6BBDACC1FF9B661DFDCA950CFD72FCA3" box="[595,747,850,876]" pageId="51" pageNumber="52">Santa María</collectingMunicipality>
; [
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,
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];
<date id="FFD8107BFF9B661DFBF4950CFAA9FCA3" box="[1133,1328,850,876]" pageId="51" pageNumber="52" value="1913-11" valueMax="1913-12-31" valueMin="1913-11-00">
<collectingDate id="EF9CE993FF9B661DFBF4950CFAA9FCA3" box="[1133,1328,850,876]" pageId="51" pageNumber="52" value="1913-11" valueMax="1913-12-31" valueMin="1913-11-00">NovDec. 1913</collectingDate>
</date>
;
<collectorName id="2693536DFF9B661DFADB950CFE98FC40" pageId="51" pageNumber="52">M.J. Viana</collectorName>
leg.;
<specimenCode id="DBC09EC0FF9B661DFEDA952BFD90FC5F" box="[323,521,885,912]" collectionCode="MACN-Ar" pageId="51" pageNumber="52">MACN-Ar 3565</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD936BBFF9B661DFF7395E9FDF6FBD8" blockId="51.[189,1398,951,1048]" pageId="51" pageNumber="52">
<materialsCitation id="3B0E3CE6FF9B661DFF7395E9FDF3FBD8" ID-GBIF-Occurrence="3066856383" collectingDate="1970-11" collectionCode="MCZ" collectorName="M. E. Galiano" country="Argentina" county="Puerto Iguazu" latitude="-25.695276" location="Cataratas del Iguazu" longLatPrecision="21" longitude="-54.43667" municipality="Cataratas del Iguazu" pageId="51" pageNumber="52" specimenCount="1" specimenCount-female="1" stateProvince="Misiones" typeStatus="paratype">
<emphasis id="B912EAA9FF9B661DFF7395E9FEC5FC1E" bold="true" box="[234,348,951,977]" pageId="51" pageNumber="52">
<typeStatus id="54DD8819FF9B661DFF7395E9FEC5FC1E" box="[234,348,951,977]" pageId="51" pageNumber="52" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F371762BFF9B661DFF249584FEF1FC3B" box="[189,360,986,1012]" name="Argentina" pageId="51" pageNumber="52">ARGENTINA</collectingCountry>
<specimenCount id="9D60FD32FF9B661DFEE39584FE3CFC3B" box="[378,421,986,1012]" pageId="51" pageNumber="52" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49A2F859FF9B661DFE289584FDB9FC3B" box="[433,544,986,1012]" pageId="51" pageNumber="52">Misiones</collectingRegion>
,
<location id="8EB96060FF9B661DFDB19584FC8FFC3B" LSID="urn:lsid:plazi:treatment:03CF87ADFF9B661BFD0A9756FD60FC7E:8EB96060FF9B661DFDB19584FC8FFC3B" box="[552,790,986,1012]" country="Argentina" county="Puerto Iguazu" latitude="-25.695276" longLatPrecision="21" longitude="-54.43667" municipality="Cataratas del Iguazu" name="Cataratas del Iguazu" pageId="51" pageNumber="52" stateProvince="Misiones">Cataratas del Iguazú</location>
,
<locationDeviation id="9A6B2F99FF9B661DFC869584FBBAFC3B" box="[799,1059,986,1012]" pageId="51" pageNumber="52">near the Puerto Iguazú</locationDeviation>
; [
<geoCoordinate id="EE52507CFF9B661DFBAD9586FB5DFC3B" box="[1076,1220,984,1012]" degrees="25" direction="south" minutes="41" orientation="latitude" pageId="51" pageNumber="52" precision="15" seconds="43" value="-25.695276">25°4143″ S</geoCoordinate>
,
<geoCoordinate id="EE52507CFF9B661DFB579586FAF3FC3B" box="[1230,1386,984,1012]" degrees="54" direction="west" minutes="26" orientation="longitude" pageId="51" pageNumber="52" precision="15" seconds="12" value="-54.43667">54°2612″ W</geoCoordinate>
];
<date id="FFD8107BFF9B661DFF2495A0FEA3FBD8" box="[189,314,1021,1048]" pageId="51" pageNumber="52" value="1970-11">
<collectingDate id="EF9CE993FF9B661DFF2495A0FEA3FBD8" box="[189,314,1021,1048]" pageId="51" pageNumber="52" value="1970-11">Nov. 1970</collectingDate>
</date>
;
<collectorName id="2693536DFF9B661DFEDF95A0FE7CFBD8" box="[326,485,1021,1048]" pageId="51" pageNumber="52">M.E. Galiano</collectorName>
leg.;
<collectionCode id="ED77AE7EFF9B661DFDBE95A3FDF3FBD8" box="[551,618,1021,1047]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="51" pageNumber="52" type="Museum">MCZ</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9B661DFF249261FE7CFB00" pageId="51" pageNumber="52" type="discussion">
<paragraph id="8BD936BBFF9B661DFF249261FEBDFB96" blockId="51.[189,292,1087,1113]" box="[189,292,1087,1113]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFF249261FEBDFB96" bold="true" box="[189,292,1087,1113]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B912EAA9FF9B661DFF249261FEBDFB96" bold="true" box="[189,292,1087,1113]" pageId="51" pageNumber="52">Remark</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF9B661DFF249233FE7CFB00" blockId="51.[188,1398,1133,1231]" pageId="51" pageNumber="52">
The male and female were matched by their close localities in Misiones,
<collectingCountry id="F371762BFF9B661DFB9F9233FBE6FB48" box="[1030,1151,1133,1159]" name="Argentina" pageId="51" pageNumber="52">Argentina</collectingCountry>
. Despite the different colorations of the male and female, the morphology of the genitalia suggests that both exemplars belong to
<taxonomicName id="4C664D38FF9B661DFF4292EAFE1AFB01" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[219,387,1204,1230]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="oliviae" status="sp. nov.">
<emphasis id="B912EAA9FF9B661DFF4292EAFE1AFB01" box="[219,387,1204,1230]" italics="true" pageId="51" pageNumber="52">Tafana oliviae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF9B661DFE1392EBFE7CFB00" box="[394,485,1205,1231]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9B661BFF2492A8FAEEFC8E" lastPageId="53" lastPageNumber="54" pageId="51" pageNumber="52" type="description">
<paragraph id="8BD936BBFF9B661DFF2492A8FED7FADF" blockId="51.[189,334,1270,1296]" box="[189,334,1270,1296]" pageId="51" pageNumber="52">
<heading id="D09181D7FF9B661DFF2492A8FED7FADF" bold="true" box="[189,334,1270,1296]" fontSize="11" level="3" pageId="51" pageNumber="52" reason="3">
<emphasis id="B912EAA9FF9B661DFF2492A8FED7FADF" bold="true" box="[189,334,1270,1296]" pageId="51" pageNumber="52">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF9B661DFF73937BFDE4FAF0" blockId="51.[189,1398,1316,1449]" box="[234,637,1316,1343]" pageId="51" pageNumber="52">
<emphasis id="B912EAA9FF9B661DFF73937BFEB3FAF0" bold="true" box="[234,298,1317,1343]" pageId="51" pageNumber="52">Male</emphasis>
(
<typeStatus id="54DD8819FF9B661DFEA0937AFE3DFAF1" box="[313,420,1316,1342]" pageId="51" pageNumber="52" type="holotype">holotype</typeStatus>
, MACN-Ar 3565)
</paragraph>
<paragraph id="8BD936BBFF9B661DFF249316FD64FA66" blockId="51.[189,1398,1316,1449]" pageId="51" pageNumber="52">
COLOR. Exemplar probably discolored. Carapace orange, darker in cephalic region (
<figureCitation id="135D2A3EFF9B661DFB0B9316FA99FAAD" box="[1170,1280,1352,1378]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Fig. 29A</figureCitation>
). Labium, sternum and endites yellow. Chelicerae reddish. Legs IIV yellow. Abdomen yellow with lateral dark spots and a yellow spot in the middle (
<figureCitation id="135D2A3EFF9B661DFD1A93D1FD68FA66" box="[643,753,1423,1449]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Fig. 29A</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF9B661DFF24938FFE64F99B" blockId="51.[189,1398,1488,1620]" pageId="51" pageNumber="52">MEASUREMENTS. Total length 3.9, carapace length 1.7, width 1.4. Clypeus height 0.04. Eye diameters and interdistances: AME 0.06, ALE 0.08, PME 0.08, PLE 0.06; AMEAME 0.02, AMEALE 0.04, PMEPME 0.1, PMEPLE 0.06, ALEPLE 0.02. Chelicerae 1.0 long, with four promarginal teeth and six retromarginal denticles.</paragraph>
<paragraph id="8BD936BBFF9B661DFF249023FB4EF976" blockId="51.[189,1398,1660,1722]" pageId="51" pageNumber="52">LEG MEASUREMENTS. I—femur 2.1 /patella 0.7/ tibia 2.5/ metatarsus 2.2/ tarsus 1.1 /total 8.6; II— 1.6/ 0.5/ 1.7 /1.5 / 0.7/ 6.0; III—1.3 / 0.6/ 0.9/ 1.2/ 0.4/ 4.4; IV—2.0 / 0.6/ 1.3/ 2.1 /0.6 / 6.6.</paragraph>
<paragraph id="8BD936BBFF9B661DFF2490BCFCE3F8D1" blockId="51.[189,1399,1761,1823]" pageId="51" pageNumber="52">LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-1; II—tibia v1-2-0; III—tibia v1-1-0, p1-1-0, r1-1-0, metatarsus v2-1-2, r1-0-1; IV—tibia v0-1-1, p1-1-1, r1-1-0.</paragraph>
<paragraph id="8BD936BBFF9B661DFF249119FD73F868" blockId="51.[189,1399,1862,1959]" pageId="51" pageNumber="52">
PALP. Retrolateral tibial apophysis bifid; cymbium without retrolateral projection; embolus short and bifurcate at the apex; median apophysis long and curved apically; sperm duct with secondary looping reaching the apex of the tegulum (
<figureCitation id="135D2A3EFF9B661DFDC991D3FD44F868" box="[592,733,1933,1959]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="51" pageNumber="52">Fig. 29CE</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF9B661DFF249191FAEEF827" blockId="51.[189,1399,1998,2025]" box="[189,1399,1998,2025]" pageId="51" pageNumber="52">ABDOMEN. Length 2.1, epigastric furrow 0.6 from tracheal spiracle, spiracle 1.0 from base of spinnerets.</paragraph>
<paragraph id="8BD936BBFF9C661AFF739757FD8DFEEC" blockId="52.[189,1398,265,361]" box="[234,532,265,291]" pageId="52" pageNumber="53">
<emphasis id="B912EAA9FF9C661AFF739757FEDCFEEC" bold="true" box="[234,325,265,291]" pageId="52" pageNumber="53">Female</emphasis>
(
<typeStatus id="54DD8819FF9C661AFECC9757FE26FEEC" box="[341,447,265,291]" pageId="52" pageNumber="53" type="paratype">paratype</typeStatus>
, MCZ)
</paragraph>
<paragraph id="8BD936BBFF9C661AFF249772FD26FEA6" blockId="52.[189,1398,265,361]" pageId="52" pageNumber="53">
COLOR. Exemplar probably discolored. Coloration as in males of this species except for legs yellow, darker at tarsi. Abdomen yellow (
<figureCitation id="135D2A3EFF9C661AFDDF9711FD2BFEA6" box="[582,690,335,361]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="52" pageNumber="53">Fig. 29B</figureCitation>
).
</paragraph>
<caption id="DF196633FF9C661AFF249089FB2FF827" ID-DOI="http://doi.org/10.5281/zenodo.4655495" ID-Zenodo-Dep="4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="52" pageNumber="53" startId="52.[189,232,1751,1777]" targetBox="[189,1398,421,1706]" targetPageId="52">
<paragraph id="8BD936BBFF9C661AFF249089FB2FF827" blockId="52.[185,1399,1751,2024]" pageId="52" pageNumber="53">
<emphasis id="B912EAA9FF9C661AFF249089FE83F93E" bold="true" box="[189,282,1751,1777]" pageId="52" pageNumber="53">Fig. 29.</emphasis>
<taxonomicName id="4C664D38FF9C661AFEBB9089FE53F93E" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[290,458,1751,1777]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="oliviae" status="sp. nov.">
<emphasis id="B912EAA9FF9C661AFEBB9089FE53F93E" box="[290,458,1751,1777]" italics="true" pageId="52" pageNumber="53">Tafana oliviae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF9C661AFE4B9086FDB4F93D" box="[466,557,1752,1778]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
(A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ).
<emphasis id="B912EAA9FF9C661AFF2490A4FF4BF8DB" bold="true" box="[189,210,1786,1812]" pageId="52" pageNumber="53">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B912EAA9FF9C661AFE7190A4FE63F8DB" bold="true" box="[488,506,1786,1812]" pageId="52" pageNumber="53">B</emphasis>
. Habitus, dorsal view.
<emphasis id="B912EAA9FF9C661AFC8990A4FCBAF8DB" bold="true" box="[784,803,1786,1812]" pageId="52" pageNumber="53">C</emphasis>
. Left palp, ventral view.
<emphasis id="B912EAA9FF9C661AFBCC90A4FBF1F8DB" bold="true" box="[1109,1128,1786,1812]" pageId="52" pageNumber="53">D</emphasis>
. Left palp, retrolateral view.
<emphasis id="B912EAA9FF9C661AFE9A9140FE8EF8F7" bold="true" box="[259,279,1822,1848]" pageId="52" pageNumber="53">E</emphasis>
. Left palp, retrolateral view.
<emphasis id="B912EAA9FF9C661AFDE99140FD18F8F7" bold="true" box="[624,641,1822,1848]" pageId="52" pageNumber="53">F</emphasis>
. Epigynum, ventral view.
<emphasis id="B912EAA9FF9C661AFC239143FC49F8F8" bold="true" box="[954,976,1821,1847]" pageId="52" pageNumber="53">G</emphasis>
. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm.
</paragraph>
</caption>
<paragraph id="8BD936BBFF9D661BFF249757FE4DFE43" blockId="53.[188,1399,265,396]" pageId="53" pageNumber="54">MEASUREMENTS. Total length 5.7, carapace length 2.7, width 1.8. Clypeus height 0.04. Eye diameters and interdistances: AME 0.08, ALE 0.08, PME 0.1, PLE 0.1; AMEAME 0.08, AMEALE 0.08, PME PME 0.18, PMEPLE 0.16, ALEPLE 0.04. Chelicerae 1.1 long, with four promarginal teeth and four retromarginal denticles.</paragraph>
<paragraph id="8BD936BBFF9D661BFF2497EAFAD5FE3E" blockId="53.[189,1398,435,497]" pageId="53" pageNumber="54">LEG MEASUREMENTS. I—femur 2.0 /patella 0.9/ tibia 1.8/ metatarsus 1.4/ tarsus 0.8 /total 6.9; II— 1.9/ 0.9/ 1.1 /1.3 / 0.63 /5.83; III—1.5/ 0.72 / 1.1/ 1.23 / 0.5/ 5.05; IV—2.25/ 0.8/ 1.72 / 2.1/ 0.7/ 7.57.</paragraph>
<paragraph id="8BD936BBFF9D661BFF249446FD53FD9A" blockId="53.[189,1398,536,597]" pageId="53" pageNumber="54">LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-0; II—tibia v2-2-0; III—tibia v1-1-2, p0-0-1, r0-0-1, metatarsus v2-1-2; IV—tibia v1-1-2, p1-1-1.</paragraph>
<paragraph id="8BD936BBFF9D661BFF249423FCE6FD30" blockId="53.[187,1400,636,767]" pageId="53" pageNumber="54">
EPIGYNUM. Hood semicircular and thick; lateral lobes curved in anterior region; wide atrium; epigynal plate covered by cuticle in the posterior region. Internally, copulatory ducts long and narrow; secondary spermathecae in the anterior region of the copulatory ducts; primary spermathecae small and fertilization ducts longer than primary spermathecae (
<figureCitation id="135D2A3EFF9D661BFD3994BBFCA9FD30" box="[672,816,741,767]" captionStart="Fig" captionStartId="52.[189,232,1751,1777]" captionTargetBox="[189,1398,421,1706]" captionTargetId="figure-190@52.[189,1398,421,1706]" captionTargetPageId="52" captionText="Fig. 29. Tafana oliviae sp. nov. (A, CE =holotype, ♂, MACN-Ar 3565; B, FG=paratype, ♀, MCZ). A. Habitus, dorsal view. B. Habitus, dorsal view. C. Left palp, ventral view. D. Left palp, retrolateral view. E. Left palp, retrolateral view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A=atrium; ABE =apophysis of base of the embolus; aRTA= anterior branch of the retrolateral tibial apophysis; CD=copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL =lateral lobes; MA=median apophysis; MTA= median tibial apophysis; pRTA=posterior branch of the retrolateral tibial apophysis; RTA =retrolateral tibial apophysis; S1 =primary spermathecae; S2 =secondary spermathecae; T =tegulum. Scale bars: AB= 3.5 mm; CG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655495" httpUri="https://zenodo.org/record/4655495/files/figure.png" pageId="53" pageNumber="54">Figs 29FG</figureCitation>
,
<figureCitation id="135D2A3EFF9D661BFCA594BBFCEAFD30" box="[828,883,741,767]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="53" pageNumber="54">33G</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF9D661BFF249579FAEEFC8E" blockId="53.[189,1399,807,833]" box="[189,1399,807,833]" pageId="53" pageNumber="54">ABDOMEN. Length 2.9, epigastric furrow 0.7 from tracheal spiracle, spiracle 1.4 from base of spinnerets.</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF9D661BFF249536FD60FC7E" box="[189,761,872,945]" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8BD936BBFF9D661BFF249536FECFFC4D" blockId="53.[189,342,872,898]" box="[189,342,872,898]" pageId="53" pageNumber="54">
<heading id="D09181D7FF9D661BFF249536FECFFC4D" bold="true" box="[189,342,872,898]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="B912EAA9FF9D661BFF249536FECFFC4D" bold="true" box="[189,342,872,898]" pageId="53" pageNumber="54">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF9D661BFF2495C9FD60FC7E" blockId="53.[189,761,918,945]" box="[189,761,918,945]" pageId="53" pageNumber="54">
Known only from Misiones,
<collectingCountry id="F371762BFF9D661BFD9695C8FD1DFC7F" box="[527,644,918,944]" name="Argentina" pageId="53" pageNumber="54">Argentina</collectingCountry>
(
<figureCitation id="135D2A3EFF9D661BFD0D95C8FD75FC7E" box="[660,748,918,945]" captionStart="Fig" captionStartId="59.[189,232,1997,2023]" captionTargetBox="[222,1366,265,1955]" captionTargetId="figure-20@59.[222,1366,265,1955]" captionTargetPageId="59" captionText="Fig. 35. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655519" httpUri="https://zenodo.org/record/4655519/files/figure.png" pageId="53" pageNumber="54">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>