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<document ID-DOI="http://doi.org/10.5281/zenodo.6462060" ID-GBIF-Dataset="cc86b94f-f8d5-42e0-bd4e-38b28b985a4e" ID-ISSN="2073-2333" ID-Zenodo-Dep="6462060" approvalRequired="1" approvalRequired_for_textStreams="1" checkinTime="1649773158173" checkinUser="felipe" docAuthor="Korneyev, V. A." docDate="2015" docId="03CF8785F216FF97F9FFFA1947037978" docLanguage="en" docName="VestnikZoologii.49.6.489-496.pdf" docOrigin="Vestnik Zoologii 49 (6)" docSource="http://dx.doi.org/10.1515/vzoo-2015-0060" docStyle="DocumentStyle:9FE6554C034D0A4D8EA9CEAE255F4368.2:VestnikZoologii.2014-2019.journal_article" docStyleId="9FE6554C034D0A4D8EA9CEAE255F4368" docStyleName="VestnikZoologii.2014-2019.journal_article" docStyleVersion="2" docTitle="Ramuliseta madagascariensis Hennig 1960" docType="treatment" docVersion="5" lastPageId="5" lastPageNumber="493" masterDocId="FFF6FFFDF215FF92F951FFE443257831" masterDocTitle="A New Species Of The Genus Ramuliseta (Diptera, Ctenostylidae) From Madagascar, With A Key To Species" masterLastPageNumber="496" masterPageNumber="489" pageId="3" pageNumber="492" updateTime="1649965230080" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A New Species Of The Genus Ramuliseta (Diptera, Ctenostylidae) From Madagascar, With A Key To Species</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Korneyev, V. A.</mods:namePart>
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<mods:title>Vestnik Zoologii</mods:title>
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<mods:date>2015</mods:date>
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<mods:number>2015-12-01</mods:number>
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<mods:number>49</mods:number>
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<mods:number>6</mods:number>
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<mods:identifier type="DOI">10.1515/vzoo-2015-0060</mods:identifier>
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<paragraph blockId="3.[174,834,1533,1560]" box="[174,834,1533,1560]" pageId="3" pageNumber="492">
<heading box="[174,834,1533,1560]" fontSize="36" level="2" pageId="3" pageNumber="492" reason="3">
<taxonomicName authority="Hennig, 1960" authorityName="Hennig" authorityYear="1960" box="[174,695,1533,1560]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="492" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis bold="true" box="[174,531,1533,1560]" italics="true" pageId="3" pageNumber="492">Ramuliseta madagascariensis</emphasis>
<bibRefCitation author="Hennig, W." box="[537,695,1533,1559]" pageId="3" pageNumber="492" pagination="321 - 353" refId="ref5006" refString="Hennig, W. 1960. Pyrgotidae de Madagascar (Diptera). Memoires de l'Institut Scientifique de Madagascar. Ser. E, 11 (1959), 321 - 353." type="journal article" year="1960">Hennig, 1960</bibRefCitation>
</taxonomicName>
(
<figureCitation box="[710,769,1533,1559]" captionStart="Fig" captionStartId="1.[174,207,1795,1817]" captionTargetBox="[216,1185,1632,1756]" captionTargetId="figure-569@1.[212,1195,1626,1784]" captionTargetPageId="1" captionText="Fig. 1. Ramuliseta, wing: 1 — R. dolini sp.n.; 2 — R. madagascariensis; 3 — R. palpifera (arrows show key characters)." figureDoi="http://doi.org/10.5281/zenodo.6452667" httpUri="https://zenodo.org/record/6452667/files/figure.png" pageId="3" pageNumber="492">fig. 1</figureCitation>
,
<figureCitation box="[783,797,1533,1559]" captionStart="Fig" captionStartId="3.[175,208,822,844]" captionTargetBox="[260,1144,174,801]" captionTargetId="figure-371@3.[255,1144,174,810]" captionTargetPageId="3" captionText="Fig. 2. Ramuliseta dolini sp. n., holotype ♀: 1 — habitus, lateral view; 25 — head (2 — left, 3 — anterior, 4 — dorsal view); 6 — wing; 7 — mesonotum, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6452669" httpUri="https://zenodo.org/record/6452669/files/figure.png" pageId="3" pageNumber="492">
<emphasis box="[783,797,1533,1559]" italics="true" pageId="3" pageNumber="492">2</emphasis>
</figureCitation>
;
<figureCitation box="[810,825,1533,1560]" captionStart="Fig" captionStartId="4.[166,199,750,772]" captionTargetBox="[272,1051,199,697]" captionTargetId="figure-449@4.[228,1133,172,735]" captionTargetPageId="4" captionText="Fig. 3. Ramuliseta madagascariensis, paralectotype ♀: 1 — head dorsal, 2 — same, right; 3 — wing." figureDoi="http://doi.org/10.5281/zenodo.6452673" httpUri="https://zenodo.org/record/6452673/files/figure.png" pageId="3" pageNumber="492">3</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="492" type="reference_group">
<paragraph blockId="3.[174,1232,1590,1726]" pageId="3" pageNumber="492">
<bibRefCitation author="Hennig, W." box="[174,349,1590,1612]" pageId="3" pageNumber="492" pagination="321 - 353" refId="ref5006" refString="Hennig, W. 1960. Pyrgotidae de Madagascar (Diptera). Memoires de l'Institut Scientifique de Madagascar. Ser. E, 11 (1959), 321 - 353." type="journal article" year="1960">Hennig, 1960: 326</bibRefCitation>
;
<bibRefCitation author="Rohlfien, K. &amp; Ewald, B." box="[358,613,1590,1612]" pageId="3" pageNumber="492" pagination="407 - 469" refId="ref5489" refString="Rohlfien, K., Ewald, B. 1970. Katalog der in den Sammlungen des ehemaligen Deutschen Entomologischen Institutes aufbewahrten Typen - VIII. (Diptera: Cyclorrhapa: Schizophora: Acalyptratae). Beitrage zur Entomologie, 22, 407 - 469." type="journal article" year="1970">Rohlfien, Ewald, 1970: 422</bibRefCitation>
;
<bibRefCitation author="Steyskal, G. C." box="[621,802,1590,1612]" pageId="3" pageNumber="492" pagination="556 - 562" refId="ref5535" refString="Steyskal, G. C. 1980. 42. Family Pyrgotidae. In: Crosskey, R. W., ed. Catalogue of the Diptera of the Afrotropical Region. British Museum (Natural History), London, 556 - 562." type="book chapter" year="1980">Steyskal, 1980: 556</bibRefCitation>
;
<bibRefCitation author="Barraclough, D. A." box="[810,1008,1590,1612]" pageId="3" pageNumber="492" pagination="5 - 14" refId="ref4848" refString="Barraclough, D. A. 1994. A review of the Afrotropical Ctenostylidae (Diptera: Schizophora:? Tephritoidea), with redescription of Ramuliseta lindneri Keiser, 1952. Annals of the Natal Museum, 35, 5 - 14." type="journal article" year="1994">Barraclough, 1994: 9</bibRefCitation>
,
<bibRefCitation author="Barraclough, D. A." box="[1017,1109,1590,1612]" pageId="3" pageNumber="492" pagination="115 - 126" refId="ref4931" refString="Barraclough, D. A. 1998. The missing males of Ramuliseta Keiser (Diptera: Schizophora: Ctenostylidae). Annals of the Natal Museum, 39, 115 - 126." type="journal article" year="1998">1998: 116</bibRefCitation>
;
<bibRefCitation author="Rafael, J. A. &amp; Pimentel, T. de &amp; Godoi, F. S. P. &amp; Machado, R. J. P." pageId="3" pageNumber="492" pagination="63 - 68" refId="ref5419" refString="Rafael, J. A., Pimentel, T. de J., Godoi, F. S. P., Machado, R. J. P. 2009. The enigmatic genus Ctenostylum Macquart with the description of a new species from the Brazilian Amazon Basin and a checklist of world Ctenostylidae species (Diptera). Zootaxa, 2026, 63 - 68." type="journal article" year="2009">Rafael et al., 2009: 64</bibRefCitation>
;
<bibRefCitation author="Pereira-Colavite, A. &amp; Mello, R. L." box="[265,611,1617,1639]" pageId="3" pageNumber="492" pagination="215 - 223" refId="ref5382" refString="Pereira-Colavite, A., Mello, R. L. 2014. Catalogue of the Ctenostylidae (Diptera, Schizophora) of the World. Zootaxa, 3838 (2), 215 - 223." type="journal article" year="2014">Pereira-Colavite &amp; Mello, 2014: 219</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="494" pageId="3" pageNumber="492" type="description">
<paragraph blockId="3.[174,1232,1590,1726]" pageId="3" pageNumber="492">
<materialsCitation collectionCode="MHNP, SDEI" country="Madagascar" location="Centre-Est" municipality="Moramanga" pageId="3" pageNumber="492" specimenCount="2" specimenCount-female="2" typeStatus="lectotype">
M a t e r i a l.
<emphasis bold="true" box="[341,397,1651,1672]" pageId="3" pageNumber="492">Type.</emphasis>
<typeStatus box="[402,496,1651,1673]" pageId="3" pageNumber="492">Lectotype</typeStatus>
<specimenCount box="[502,517,1650,1673]" pageId="3" pageNumber="492" type="female"></specimenCount>
(here designated): [
<emphasis bold="true" box="[709,835,1651,1672]" pageId="3" pageNumber="492">
<collectingCountry box="[709,831,1651,1672]" name="Madagascar" pageId="3" pageNumber="492">Madagascar</collectingCountry>
:
</emphasis>
] “Sandrangato” [“
<location LSID="urn:lsid:plazi:treatment:03CF8785F216FF97F9FFFA1947037978:8EB96048F216FF91FAA0F997477B7EB8" box="[1009,1118,1651,1673]" country="Madagascar" municipality="Moramanga" name="Centre-Est" pageId="3" pageNumber="492">Centre-Est</location>
,
<collectingMunicipality pageId="3" pageNumber="492">Moramanga</collectingMunicipality>
”], “Institute / Scientifique / Madagascar”, “
<typeStatus box="[620,722,1678,1700]" pageId="3" pageNumber="492">Holotypus</typeStatus>
” [dark red printed label] (
<collectionCode box="[981,1057,1678,1700]" country="France" httpUri="http://grbio.org/cool/a5xg-x4h4" name="Museum d'Histoire Naturelle Perpignan" pageId="3" pageNumber="492">MHNP</collectionCode>
).
<typeStatus box="[1074,1205,1678,1700]" pageId="3" pageNumber="492">Paralectotype</typeStatus>
<specimenCount box="[1211,1226,1677,1700]" pageId="3" pageNumber="492" type="female"></specimenCount>
: same labels as in
<typeStatus box="[338,427,1704,1726]" pageId="3" pageNumber="492">lectotype</typeStatus>
, except “
<typeStatus box="[512,609,1704,1726]" pageId="3" pageNumber="492">Paratypus</typeStatus>
” [dark red printed label] (
<collectionCode box="[861,917,1704,1726]" pageId="3" pageNumber="492">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[174,1233,1759,1856]" lastBlockId="4.[166,1225,822,1841]" lastPageId="4" lastPageNumber="493" pageId="3" pageNumber="492">
Diagnosis. Moderately large fly with yellow body and its vestiture, and wing with yellow base, two hyaline incisions in the costal cell, two triangular hyaline incisions at anterior margin distal to vein R
<subScript attach="left" box="[548,556,1842,1856]" fontSize="6" pageId="3" pageNumber="492">1</subScript>
reaching R
<subScript attach="left" box="[691,718,1842,1856]" fontSize="6" pageId="3" pageNumber="492">2+3</subScript>
and R
<subScript attach="left" box="[796,822,1842,1856]" fontSize="6" pageId="3" pageNumber="492">4+5</subScript>
vein, correspondingly, and 3 large hyaline incision at posterior margin, separated by yellow bands: two from cell cua1 (basal reaching CuA
<subScript attach="left" box="[331,339,873,887]" fontSize="6" pageId="4" pageNumber="493">1</subScript>
vein and distal entering through cell dm into r
<subScript attach="left" box="[900,926,873,887]" fontSize="6" pageId="4" pageNumber="493">4+5</subScript>
) and the apical crescentlike area covering almost all cell m and widely entering into cell r
<subScript attach="left" box="[946,972,905,919]" fontSize="6" pageId="4" pageNumber="493">4+5</subScript>
anterior of crossvein dm-cu and along costa; wing apex hyaline, at most costal vein narrowly yellow. It is similar to
<taxonomicName authorityName="V. Korneyev" authorityYear="2015" box="[197,296,950,976]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="dolini">
<emphasis box="[197,296,950,976]" italics="true" pageId="4" pageNumber="493">R. dolini</emphasis>
</taxonomicName>
, in most features, differing by the wing pattern as noted in the key.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.6452673" ID-Zenodo-Dep="6452673" box="[166,1108,749,772]" httpUri="https://zenodo.org/record/6452673/files/figure.png" pageId="4" pageNumber="493" startId="4.[166,199,750,772]" targetBox="[272,1051,199,697]" targetPageId="4">
<paragraph blockId="4.[166,1108,749,772]" box="[166,1108,749,772]" pageId="4" pageNumber="493">
Fig. 3.
<taxonomicName authorityName="Hennig" authorityYear="1960" box="[230,509,750,771]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis box="[230,509,750,771]" italics="true" pageId="4" pageNumber="493">Ramuliseta madagascariensis</emphasis>
</taxonomicName>
, paralectotype ♀:
<emphasis box="[681,692,750,771]" italics="true" pageId="4" pageNumber="493">1</emphasis>
— head dorsal,
<emphasis box="[847,858,750,771]" italics="true" pageId="4" pageNumber="493">2</emphasis>
— same, right;
<emphasis box="[1009,1020,750,771]" italics="true" pageId="4" pageNumber="493">3</emphasis>
— wing.
</paragraph>
</caption>
<paragraph blockId="4.[166,1225,822,1841]" pageId="4" pageNumber="493">
Description. Head as described for
<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[672,772,982,1008]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="dolini" status="sp. n.">
<emphasis box="[672,772,982,1008]" italics="true" pageId="4" pageNumber="493">R. dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[780,845,983,1009]" pageId="4" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
, in
<typeStatus box="[890,1048,982,1008]" pageId="4" pageNumber="493">paralectotype</typeStatus>
conspicuously compressed.
</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" pageId="4" pageNumber="493">
Thorax as described for
<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[496,594,1046,1072]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="dolini" status="sp. n.">
<emphasis box="[496,594,1046,1072]" italics="true" pageId="4" pageNumber="493">R. dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[599,660,1047,1073]" pageId="4" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
Scutum bearing: 0 acrostichal, 23 dorsocentral, 2 posterior notopleural, 1 intraalar and 1 postalar seta. Postpronotal lobe without setae, but 13 setae and 46 smaller setulae. Scutellum with 2 pairs of setae (basal and apical) and 1520 irregular setulae on disk and margin. Prosternum transverse, sclerotised at posteroventral margin.
</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" box="[219,962,1206,1233]" pageId="4" pageNumber="493">
Legs yellow, with yellow setulae, as described for
<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[794,893,1206,1232]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="dolini" status="sp. n.">
<emphasis box="[794,893,1206,1232]" italics="true" pageId="4" pageNumber="493">R. dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[900,962,1207,1233]" pageId="4" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" pageId="4" pageNumber="493">
Wing as on
<figureCitation box="[371,462,1238,1264]" captionStart="Fig" captionStartId="4.[166,199,750,772]" captionTargetBox="[272,1051,199,697]" captionTargetId="figure-449@4.[228,1133,172,735]" captionTargetPageId="4" captionText="Fig. 3. Ramuliseta madagascariensis, paralectotype ♀: 1 — head dorsal, 2 — same, right; 3 — wing." figureDoi="http://doi.org/10.5281/zenodo.6452673" httpUri="https://zenodo.org/record/6452673/files/figure.png" pageId="4" pageNumber="493">
fig. 3,
<emphasis box="[448,462,1238,1264]" italics="true" pageId="4" pageNumber="493">3</emphasis>
</figureCitation>
. Vein M almost parallel to R
<subScript attach="left" box="[822,848,1257,1271]" fontSize="6" pageId="4" pageNumber="493">4+5</subScript>
; ratio of its three last sections: M2: M3: M4 = 1: 0.9: 1.2. Wing pattern, as in most ctenostylines, with yellow basal cells and well-developed pattern of connected yellow crossbands and rounded subapical yellowbrown area separated by yellow bands: two from cell cua
<subScript attach="left" box="[845,853,1353,1367]" fontSize="6" pageId="4" pageNumber="493">1</subScript>
(basal reaching CuA
<subScript attach="left" box="[1105,1113,1353,1367]" fontSize="6" pageId="4" pageNumber="493">1</subScript>
vein and distal entering through cell dm into r
<subScript attach="left" box="[606,633,1385,1399]" fontSize="6" pageId="4" pageNumber="493">4+5</subScript>
) and the apical crescent-like area covering almost all cell m and widely entering into cell r
<subScript attach="left" box="[651,677,1417,1431]" fontSize="6" pageId="4" pageNumber="493">4+5</subScript>
anterior of crossvein dm-cu and along costa; wing apex is hyaline, at most costal vein narrowly yellow. The rest as in
<taxonomicName authority="Korneyev, 2015" authorityName="V. Korneyev" authorityYear="2015" box="[1004,1104,1430,1456]" class="Insecta" family="Ctenostylidae" genus="Ramuliseta" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="493" phylum="Arthropoda" rank="species" species="dolini" status="sp. n.">
<emphasis box="[1004,1104,1430,1456]" italics="true" pageId="4" pageNumber="493">R. dolini</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1110,1172,1431,1457]" pageId="4" pageNumber="493" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" pageId="4" pageNumber="493">Abdomen yellow. Syntergite 1+2 yellow, 1.7 times as long as wide, slightly longer than tergites 36 combined; tergites 5 and 6 with large laterobasal black spots. Sternites yellow, black setulose. Sternite 1 longitudinal trapezoid, as well as sternite 2, with clear seam between them; sternites 36 subquadrate, poorly sclerotized. Membrane without any setulae, short microtrichose. Oviscape convex, yellow, yellow setulose, as long as tergites 36 combined. Oviscape simple, tubular, with microtrichose cercal unit, flexibly joined to its apico-dorsal surface</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" box="[219,788,1686,1713]" pageId="4" pageNumber="493">
Measurements. Body
<quantity box="[474,565,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="4" pageNumber="493" unit="mm" value="8.4">8.4 mm</quantity>
, wing
<quantity box="[641,784,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" metricValueMax="7.4" metricValueMin="7.2" pageId="4" pageNumber="493" unit="mm" value="7.3" valueMax="7.4" valueMin="7.2">7.27.4 mm</quantity>
.
</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" box="[219,405,1718,1744]" pageId="4" pageNumber="493">Male unknown.</paragraph>
<paragraph blockId="4.[166,1225,822,1841]" lastBlockId="5.[174,1233,175,330]" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
D i s c u s s i o n.
<bibRefCitation author="Hennig, W." box="[387,612,1750,1777]" pageId="4" pageNumber="493" pagination="321 - 353" refId="ref5006" refString="Hennig, W. 1960. Pyrgotidae de Madagascar (Diptera). Memoires de l'Institut Scientifique de Madagascar. Ser. E, 11 (1959), 321 - 353." type="journal article" year="1960">Hennig (1960: 327)</bibRefCitation>
described this species based on
<specimenCount box="[982,1122,1750,1777]" pageId="4" pageNumber="493" type="female">two females</specimenCount>
, without designation of
<typeStatus box="[347,453,1782,1808]" pageId="4" pageNumber="493">holotype</typeStatus>
, so both specimens were
<typeStatus box="[764,868,1783,1809]" pageId="4" pageNumber="493">syntypes</typeStatus>
; the subsequent statement by
<bibRefCitation author="Pereira-Colavite, A. &amp; Mello, R. L." box="[166,547,1814,1841]" pageId="4" pageNumber="493" pagination="215 - 223" refId="ref5382" refString="Pereira-Colavite, A., Mello, R. L. 2014. Catalogue of the Ctenostylidae (Diptera, Schizophora) of the World. Zootaxa, 3838 (2), 215 - 223." type="journal article" year="2014">Pereira-Colavite &amp; Mello (2014)</bibRefCitation>
that one of them is a
<typeStatus box="[807,910,1814,1840]" pageId="4" pageNumber="493">holotype</typeStatus>
and the other one is paratype, based on the labels they bear, is therefore erroneous. I herewith designate the specimen deposited in MHNP (good condition, both wings and most legs present) as
<typeStatus box="[1126,1232,207,233]" pageId="5" pageNumber="494">lectotype</typeStatus>
and the specimen deposited in SDEI (fair condition: head shriveled, all tarsi and tibiae and femora of 3 legs missing, left wing partly broken; one wing and abdomen detached and possibly are stored separately on slides, but not located by far) as
<typeStatus box="[896,1058,303,329]" pageId="5" pageNumber="494">paralectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>