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<document id="FCCB03E8E681AF893D592D94F1D8D892" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C15FF87FF30553A67A4276F" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena (Hydraenopsis) claudia Freitag &amp; Jach, sp. n." docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="22" updateTime="1698221773297" updateUser="plazi">
<mods:mods id="73A7E95B7BD105DF622E0B7B1EAC7F9E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C29E11A5B2A3ADC9503D5464E41B280A">
<mods:title id="AEB4F1557204587BE09C22A6C166A67C">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
</mods:titleInfo>
<mods:name id="D0FDCDE7D67E33DAD017CF01663B87B4" type="personal">
<mods:role id="CFC5EB9F985DF29F7B4CBC3F31ADCC45">
<mods:roleTerm id="E33602796253CEBB9CE46C39FD27153E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B81641C2BD9894E07030CCD4C1391CC0">Freitag, Hendrik</mods:namePart>
</mods:name>
<mods:name id="BC7CCA21F3FC0252E7961006D132F3BF" type="personal">
<mods:role id="C38EEE392B8500645B9C0DFF41E5A481">
<mods:roleTerm id="B6A9BA1E0A99933B347424EF8F1C8786">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BB8D3A3B66C8A3C69555C627BA77CEEC">Jäch, Manfred A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D05DBA7CDBAF26FEFE13646BEDDB8A8F">text</mods:typeOfResource>
<mods:relatedItem id="5F99BEC6AA9686DD2CAF43C7229EAD5B" type="host">
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<mods:title id="A0CE01EE0562F66870C7F5375B28986B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8C970B850436B7C79F486F0F283FA20A">
<mods:date id="ED3149753B7BB09A31CB45F7D765FC6D">2007</mods:date>
<mods:detail id="5DB927870D1A13CF825D798CBE96FBFE" type="volume">
<mods:number id="27CD7BA2D4A693912857AA84ACC161FA">1431</mods:number>
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<mods:extent id="931C728437C55DD7F32B2F23C79CF5A2" unit="page">
<mods:start id="2AAB275DF9A50EA92FE883ADFD9503A9">1</mods:start>
<mods:end id="CD43999D576758CDCA906D5B466AACB5">44</mods:end>
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<mods:classification id="EF970C9006B3D7BB4162C26CF42881EE">journal article</mods:classification>
<mods:identifier id="EFF9FC8B2AD037F82DA8AE5405AD0CDC" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="BCC1BDCBF7F8A5395AA97540C2E58618" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="57B9894C47C437F200D52E46EA552C94" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F22B342F14EDEE33C04EA8165EC46576" type="Zenodo-Dep">175790</mods:identifier>
</mods:mods>
<treatment id="03B9F24A7C15FF87FF30553A67A4276F" ID-DOI="http://doi.org/10.5281/zenodo.6248079" ID-GBIF-Taxon="119355658" ID-Zenodo-Dep="6248079" LSID="urn:lsid:plazi:treatment:03B9F24A7C15FF87FF30553A67A4276F" httpUri="http://treatment.plazi.org/id/03B9F24A7C15FF87FF30553A67A4276F" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
<subSubSection id="C30A10D77C15FF85FF30553A654C2709" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="8BAF435C7C15FF85FF30553A6759276C" blockId="21.[151,837,271,332]" box="[151,837,271,297]" pageId="21" pageNumber="22">
<heading id="D0E7F4307C15FF85FF30553A6759276C" bold="true" box="[151,837,271,297]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B9649F4E7C15FF85FF30553A6759276C" bold="true" box="[151,837,271,297]" pageId="21" pageNumber="22">
<taxonomicName id="4C1038DF7C15FF85FF30553A66EA276C" ID-CoL="3N37N" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[151,758,271,297]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="claudia" status="sp. nov." subGenus="Hydraenopsis">
<emphasis id="B9649F4E7C15FF85FF30553A650C276C" bold="true" box="[151,272,271,297]" italics="true" pageId="21" pageNumber="22">Hydraena</emphasis>
(
<emphasis id="B9649F4E7C15FF85FE87553A65DA276C" bold="true" box="[288,454,271,297]" italics="true" pageId="21" pageNumber="22">Hydraenopsis</emphasis>
)
<emphasis id="B9649F4E7C15FF85FE70553A662D276C" bold="true" box="[471,561,271,297]" italics="true" pageId="21" pageNumber="22">claudia</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<taxonomicNameLabel id="A25722357C15FF85FCA4553A6759276C" box="[771,837,271,297]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF435C7C15FF85FF305507654C2709" blockId="21.[151,837,271,332]" box="[151,336,306,332]" pageId="21" pageNumber="22">(Figs. 8, 20a–h)</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C15FF85FF3055B7677324F1" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="8BAF435C7C15FF85FF3055B764C02781" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">
<emphasis id="B9649F4E7C15FF85FF3055B7652127D9" bold="true" box="[151,317,386,412]" pageId="21" pageNumber="22">
<typeStatus id="54ABFDFE7C15FF85FF3055B764C827D9" box="[151,212,386,412]" pageId="21" pageNumber="22">Type</typeStatus>
locality.
</emphasis>
Alipuran Stream (Lake
<taxonomicName id="4C1038DF7C15FF85FDFB55B766D727D9" box="[604,715,386,412]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
tributary), Municipality of Taytay, northern Palawan,
<collectingCountry id="F30703CC7C15FF85FAEB55B764C52781" name="Philippines" pageId="21" pageNumber="22">Philippines</collectingCountry>
.
</paragraph>
<paragraph id="8BAF435C7C15FF85FF6055E7677324F1" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">
<emphasis id="B9649F4E7C15FF85FF6055E7661827A9" bold="true" box="[199,516,466,492]" pageId="21" pageNumber="22">
<typeStatus id="54ABFDFE7C15FF85FF6055E7651827A9" box="[199,260,466,492]" pageId="21" pageNumber="22">Type</typeStatus>
material.
<typeStatus id="54ABFDFE7C15FF85FE3555E7661827A9" box="[402,516,466,492]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
</emphasis>
ɗ (
<collectionCode id="ED01DB997C15FF85FD9A55E7668C27A9" box="[573,656,466,492]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C15FF85FD1255E7671C27A9" box="[693,768,466,492]" pageId="21" pageNumber="22">PHIL</collectionCode>
.: Palawan, Taytay; Lake
<taxonomicName id="4C1038DF7C15FF85FB9955E760B127A9" box="[1086,1197,466,492]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
tributary, Alipuran Stream, semi-prim. forest c.
<quantity id="4CE8EEB97C15FF85FE7055CF660C2451" box="[471,528,506,532]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="22" unit="m" value="40.0">40m</quantity>
asl,
<geoCoordinate id="EE24259B7C15FF85FDE255CF66BA2451" box="[581,678,506,532]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="10.75">10°45'N</geoCoordinate>
<geoCoordinate id="EE24259B7C15FF85FD0B55CF670B2451" box="[684,791,506,532]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="119.53333">
119°32'
<geoCoordinate id="EE24259B7C15FF85FCA355CF670B2451" box="[772,791,506,532]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="5555" value="29.4">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C15FF85FCB955CF678A2451" box="[798,918,506,532]" pageId="21" pageNumber="22" value="1995-04-29">29.4.1995</date>
, leg. Freitag (15)M”, terminal parts of abdomen, aedeagus and one mesotarsus glued separately, ultimate and penultimate segments of maxillary palpi and one protarsus lacking.
<emphasis id="B9649F4E7C15FF85FE38567F663A2421" bold="true" box="[415,550,586,612]" pageId="21" pageNumber="22">
<typeStatus id="54ABFDFE7C15FF85FE38567F663D2421" box="[415,545,586,612]" pageId="21" pageNumber="22" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 ɗ (
<collectionCode id="ED01DB997C15FF85FDC2567F66AA2421" box="[613,694,586,612]" httpUri="http://grbio.org/cool/134h-pm2f" name="Museum of Natural History, University of the Philippines" pageId="21" pageNumber="22">UPLB</collectionCode>
), 1 Ψ (
<collectionCode id="ED01DB997C15FF85FCA5567F67482421" box="[770,852,586,612]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22">NMW</collectionCode>
): same locality data as
<typeStatus id="54ABFDFE7C15FF85FBC7567F60D62421" box="[1120,1226,586,612]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
; 1 ɗ (
<collectionCode id="ED01DB997C15FF85FAA9567F61432421" box="[1294,1375,586,612]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22">NMW</collectionCode>
), 1 Ψ (
<collectionCode id="ED01DB997C15FF85FF38564764EE24C9" box="[159,242,626,652]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="21" pageNumber="22">NMW</collectionCode>
): “
<collectionCode id="ED01DB997C15FF85FEB55647654124C9" box="[274,349,626,652]" pageId="21" pageNumber="22">PHIL</collectionCode>
.: Palawan, Taytay; 1/3way high-way to Lake
<taxonomicName id="4C1038DF7C15FF85FC23564767E924C9" box="[900,1013,626,652]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
,
<taxonomicName id="4C1038DF7C15FF85FBA35647606F24C9" box="[1028,1139,626,652]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
Stream trib., prim.forest c.
<quantity id="4CE8EEB97C15FF85FF0F56AF64FC24F1" box="[168,224,666,692]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="21" pageNumber="22" unit="m" value="35.0">35m</quantity>
asl,
<geoCoordinate id="EE24259B7C15FF85FEB156AF656A24F1" box="[278,374,666,692]" direction="north" orientation="latitude" pageId="21" pageNumber="22" precision="925" value="10.783334">10°47'N</geoCoordinate>
<geoCoordinate id="EE24259B7C15FF85FED956AF65F524F1" box="[382,489,666,692]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="925" value="119.51667">
119°31'
<geoCoordinate id="EE24259B7C15FF85FE7256AF65F524F1" box="[469,489,666,692]" direction="east" orientation="longitude" pageId="21" pageNumber="22" precision="5555" value="29.4">E</geoCoordinate>
</geoCoordinate>
<date id="FFAE659C7C15FF85FE5756AF667424F1" box="[496,616,666,692]" pageId="21" pageNumber="22" value="1995-04-29">29.4.1995</date>
, leg. Freitag (13b)M”.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C15FF86FF6056F7611421D2" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
<paragraph id="8BAF435C7C15FF85FF6056F760ED2569" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">
<emphasis id="B9649F4E7C15FF85FF6056F765432499" bold="true" box="[199,351,706,732]" pageId="21" pageNumber="22">Description.</emphasis>
Body
<quantity id="4CE8EEB97C15FF85FE0A56F766442499" box="[429,600,706,732]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="21" pageNumber="22" unit="mm" value="1.5" valueMax="1.55" valueMin="1.45">1.45–1.55 mm</quantity>
long. Habitus as in Fig. 8. Elytra brown; rectangular spot in the middle of pronotal disc dark brown; head medially and anteriorly brown turning into almost black posteriorly and laterally (ocular region); legs, maxillary palpi and usually pronotal margins distinctly pale brown.
</paragraph>
<paragraph id="8BAF435C7C15FF85FF60570F66E82201" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">Pronotum wider than long, broadest at about anterior 0.35–0.50; entire pronotum moderately densely punctate, punctures moderately large and moderately deeply impressed, interstices glabrous; foveae slightly impressed or hardly perceptible; lateral margins anteriorly and posteriorly slightly convergent; anterior angles rounded, posterior angles more or less rectangular; pronoto-elytral angle obtuse; lateral rim denticulate; anterior margin slightly concave, posterior margin straight to slightly convex; lateral portions of pronotum slightly deflexed; lateral hypomeron approximately as broad as or slightly broader than profemur, inner margin convex; mesal hypomeron very narrow, inconspicuous.</paragraph>
<paragraph id="8BAF435C7C15FF85FF605067600F2371" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">Elytra elongately oval, apically almost conjointly rounded, with ca. 15 longitudinal, more or less regular, not impressed rows of punctures (eight between suture and shoulder); punctures moderately large and moderately impressed; interstices and intervals convex, glabrous; lateral margins of elytra widely explanate, almost reaching elytral apex, not or imperceptibly denticulate; inflexed lateral portion (pseudepipleuron) approximately as broad as metafemur, almost reaching elytral apex (at about posterior 0.15), with indistinct foveae and one inconspicuous row of punctures; epipleuron narrow, inconspicuous.</paragraph>
<paragraph id="8BAF435C7C15FF85FF605177668C23B9" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">Mesoventral longitudinal ridges narrow; meso- and metaventrite distinctly pubescent; mesoventral intercoxal process narrow (less than half as broad as pseudepipleuron), truncate, moderately long, margins exserted, medially flat, distinctly pubescent; metaventral disc moderately impressed; metaventral plaques inconspicuous, pubescent; intercoxal sternite broader than long, slightly narrower than metafemur; metaventrite and ventrites 1–4 distinctly pubescent.</paragraph>
<paragraph id="8BAF435C7C15FF85FF60523F65362009" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">Femora with inner margin straight to concave, outer margin conspicuously convex. Metatibiae more or less straight.</paragraph>
<paragraph id="8BAF435C7C15FF85FF60526F66422121" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">
Aedeagus (
<figureCitation id="132B5FD97C15FF85FEED526F65B02031" box="[330,428,1626,1652]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="21" pageNumber="22">Figs. 20</figureCitation>
a–c). Main piece moderately slender and dorso-ventrally bent, with one inconspicuous dorsal seta and several ventro-lateral micropores; phallobase subsymmetric. Distal lobe not clearly delimited from main piece; ejaculatory duct distally ending in a moderately long, bent, distinctly sclerotized, flagellumlike process, which is embedded in a less sclerotized flat structure. Parameres apically attenuately rounded; left paramere moderately long, articulately connected, inserted at about basal 0.35 of aedeagus, with a fringe of long setae (ca. seven ventral subapical, ca. four dorsal subapical); right paramere short, inserted at ca. basal 0.45 of aedeagus, with nine long setae.
</paragraph>
<paragraph id="8BAF435C7C15FF85FF605347679121F1" blockId="21.[151,1437,386,2012]" pageId="21" pageNumber="22">
Gonocoxite (
<figureCitation id="132B5FD97C15FF85FEC1534765DF21C9" box="[358,451,1906,1932]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="21" pageNumber="22">Fig. 20</figureCitation>
e) subsemicircular, almost as in female of
<taxonomicName id="4C1038DF7C15FF85FC78534660E721C9" box="[991,1275,1907,1932]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="castanescens">
<emphasis id="B9649F4E7C15FF85FC78534660E721C9" box="[991,1275,1907,1932]" italics="true" pageId="21" pageNumber="22">Hydraena castanescens</emphasis>
</taxonomicName>
, but slightly smaller and only with one subsemicircular cavity of dorsal plate.
</paragraph>
<paragraph id="8BAF435C7C15FF85FF6053F7662C2199" blockId="21.[151,1437,386,2012]" box="[199,560,1986,2012]" pageId="21" pageNumber="22">
Spermatheca as in
<figureCitation id="132B5FD97C15FF85FE0453F7661E2199" box="[419,514,1986,2012]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="21" pageNumber="22">Figs. 20</figureCitation>
f–g.
</paragraph>
<caption id="DF6F13D47C16FF86FF3051B1655723A4" httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="22" pageNumber="23" targetBox="[267,1320,200,1378]" targetPageId="22">
<paragraph id="8BAF435C7C16FF86FF3051B1655723A4" blockId="22.[151,1436,1412,1505]" pageId="22" pageNumber="23">
<emphasis id="B9649F4E7C16FF86FF3051B1653423D9" bold="true" box="[151,296,1412,1436]" pageId="22" pageNumber="23">FIGURE 20.</emphasis>
<taxonomicName id="4C1038DF7C16FF86FE9751B065EE23D9" box="[304,498,1413,1436]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="claudia" status="sp. nov.">
<emphasis id="B9649F4E7C16FF86FE9751B065EE23D9" box="[304,498,1413,1436]" italics="true" pageId="22" pageNumber="23">Hydraena claudia</emphasis>
</taxonomicName>
<emphasis id="B9649F4E7C16FF86FE5C51B1662423D9" bold="true" box="[507,568,1412,1436]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A25722357C16FF86FE5C51B1662423D9" box="[507,568,1412,1436]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a–c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f–g) spermatheca; h) female tergite X.
</paragraph>
</caption>
<paragraph id="8BAF435C7C16FF86FF60522065A4215A" blockId="22.[151,1437,1557,2023]" pageId="22" pageNumber="23">
Secondary sexual characters. Male profemur with inconspicuous transverse ledge (denticle) near base of ventral face. Mesoventral intercoxal process of male narrower. Male terminal sternite similar to that of
<taxonomicName id="4C1038DF7C16FF86FF30525065B2203B" box="[151,430,1637,1662]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="castanescens">
<emphasis id="B9649F4E7C16FF86FF30525065B2203B" box="[151,430,1637,1662]" italics="true" pageId="22" pageNumber="23">Hydraena castanescens</emphasis>
</taxonomicName>
, subovoid, apical margin convex, base moderately broad, basal corners less strongly produced; spiculum gastrale moderately long, entirely curved (
<figureCitation id="132B5FD97C16FF86FCD552B867D520E2" box="[882,969,1677,1703]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="22" pageNumber="23">Fig. 20</figureCitation>
d). Male tergite X distinctly excised apically. Female tergite X (
<figureCitation id="132B5FD97C16FF86FE1752806618208A" box="[432,516,1717,1743]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="22" pageNumber="23">Fig. 20</figureCitation>
h) almost as in female of
<taxonomicName id="4C1038DF7C16FF86FC8C528067F4208B" box="[811,1000,1717,1742]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="castanescens">
<emphasis id="B9649F4E7C16FF86FC8C528067F4208B" box="[811,1000,1717,1742]" italics="true" pageId="22" pageNumber="23">H. castanescens</emphasis>
</taxonomicName>
, but slightly smaller and with shorter discal and lateral setae; subapical fringe of moderately long trichoid setae medially interrupted; hyaline apical margin notched apically.
</paragraph>
<paragraph id="8BAF435C7C16FF86FF605319611421D2" blockId="22.[151,1437,1557,2023]" pageId="22" pageNumber="23">
<emphasis id="B9649F4E7C16FF86FF60531965C62103" bold="true" box="[199,474,1836,1862]" pageId="22" pageNumber="23">Differential diagnosis.</emphasis>
Except for its slightly bigger size and straight metatibia,
<taxonomicName id="4C1038DF7C16FF86FBDC5318614C2103" box="[1147,1360,1837,1862]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="claudia">
<emphasis id="B9649F4E7C16FF86FBDC5318614C2103" box="[1147,1360,1837,1862]" italics="true" pageId="22" pageNumber="23">Hydraena claudia</emphasis>
</taxonomicName>
externally resembles
<taxonomicName id="4C1038DF7C16FF86FEF25360660D212B" box="[341,529,1877,1902]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="castanescens">
<emphasis id="B9649F4E7C16FF86FEF25360660D212B" box="[341,529,1877,1902]" italics="true" pageId="22" pageNumber="23">H. castanescens</emphasis>
</taxonomicName>
, from which it can easily be distinguished by its distally broad aedeagus with a bent flagellum-like process and unequal parameres (left one longer and articulately connected).
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C16FF87FF60539166422747" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="distribution">
<paragraph id="8BAF435C7C16FF86FF605391657021A2" blockId="22.[151,1437,1557,2023]" pageId="22" pageNumber="23">
<emphasis id="B9649F4E7C16FF86FF605391657C21FB" bold="true" box="[199,352,1956,1982]" pageId="22" pageNumber="23">Distribution</emphasis>
(
<figureCitation id="132B5FD97C16FF86FED6539065D621FA" box="[369,458,1957,1983]" captionStart="FIGURE 30" captionStartId="41.[151,255,1142,1166]" captionTargetBox="[267,1314,183,1127]" captionTargetId="figure@41.[264,1323,183,1135]" captionTargetPageId="41" captionText="FIGURE 30. Collecting sites and geographical distribution of Hydraena species in Palawan and Busuanga, including enlarged map of St. Paul National Park: Hydraena scabra d’Orchymont, H. jojoorculloi sp. n., H. nielshaggei sp. n., H. zetteli sp. n., H. castanescens sp. n., H. claudia sp. n., H. kodadai sp. n." httpUri="https://zenodo.org/record/175808/files/figure.png" pageId="22" pageNumber="23">Fig. 30</figureCitation>
). This species is only known from the headwaters of tributaries of Lake
<taxonomicName id="4C1038DF7C16FF86FA815390618421FA" box="[1318,1432,1957,1983]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
, northern Palawan.
</paragraph>
<paragraph id="8BAF435C7C17FF87FF6054A266422747" blockId="23.[151,1437,151,298]" pageId="23" pageNumber="24">
<emphasis id="B9649F4E7C17FF87FF6054A2652C26F4" bold="true" box="[199,304,151,177]" pageId="23" pageNumber="24">Ecology.</emphasis>
<taxonomicName id="4C1038DF7C17FF87FE9054AD661726F4" box="[311,523,152,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="claudia">
<emphasis id="B9649F4E7C17FF87FE9054AD661726F4" box="[311,523,152,177]" italics="true" pageId="23" pageNumber="24">Hydraena claudia</emphasis>
</taxonomicName>
was collected together with
<taxonomicName id="4C1038DF7C17FF87FCFD54AD600B26F4" box="[858,1047,152,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="castanescens">
<emphasis id="B9649F4E7C17FF87FCFD54AD600B26F4" box="[858,1047,152,177]" italics="true" pageId="23" pageNumber="24">H. castanescens</emphasis>
</taxonomicName>
and additionally in another similar headwater site of the same catchment (Lake
<taxonomicName id="4C1038DF7C17FF87FD7354F56754269F" box="[724,840,192,218]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="43" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">Manguao</taxonomicName>
). The species likely has a strong affinity to undisturbed habitats in forest-covered areas.
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C17FF87FF60553A67A4276F" box="[199,952,271,298]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BAF435C7C17FF87FF60553A67A4276F" blockId="23.[151,1437,151,298]" box="[199,952,271,298]" pageId="23" pageNumber="24">
<emphasis id="B9649F4E7C17FF87FF60553A654F276C" bold="true" box="[199,339,271,297]" pageId="23" pageNumber="24">Etymology.</emphasis>
Named for
<taxonomicName id="4C1038DF7C17FF87FE4755256627276F" box="[480,571,272,298]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="42" phylum="Arthropoda" rank="species" species="claudia" status="sp. nov.">Claudia</taxonomicName>
Zlabinger (Vienna), pedagogue.
</paragraph>
</subSubSection>
</treatment>
</document>
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