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<document id="3B54DBD150A8017BE8D37CA3B2D34C81" ID-DOI="10.5281/zenodo.175790" ID-GBIF-Dataset="2a6512b7-5bd9-4682-b5db-79c72373b68c" ID-ISSN="1175-5326" ID-Zenodo-Dep="175790" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459820893039" checkinUser="plazi" docAuthor="Freitag, Hendrik &amp; Jäch, Manfred A." docDate="2007" docId="03B9F24A7C04FF95FF3054A2618625C4" docLanguage="en" docName="zt01431p044.pdf" docOrigin="Zootaxa 1431" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Hydraena" docType="key" docVersion="9" lastPageNumber="6" masterDocId="FF808A327C00FF90FFA75435641C2645" masterDocTitle="Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925" masterLastPageNumber="44" masterPageNumber="1" pageNumber="5" updateTime="1698221773297" updateUser="plazi">
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<mods:title id="9648EF5F2880EA94E28A8D93E3F7426F">Revision of the species of Hydraena Kugelann (Coleoptera: Hydraenidae) from Palawan and Busuanga, with descriptions of eleven new species, and redescription of Hydraena (Hydraenopsis) scabra d’Orchymont, 1925</mods:title>
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<mods:namePart id="2529756C572470CD4B9468093D14DC38">Freitag, Hendrik</mods:namePart>
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<mods:namePart id="DBC1F91A9DB6B6563CAF0EDDEA2F235B">Jäch, Manfred A.</mods:namePart>
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<mods:title id="754D96B56C68A99C45DE71AFC526C0BB">Zootaxa</mods:title>
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<mods:date id="BAB02477C292AE52D3B5C90F4F9EB346">2007</mods:date>
<mods:detail id="87E69A9E18A80C6B250C59DFAE75367D" type="volume">
<mods:number id="B658B6AC773470CD11175A2333E55ED0">1431</mods:number>
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<mods:classification id="FE11CCE9D7C716756F378E4E3EF33E69">journal article</mods:classification>
<mods:identifier id="15965CA2F6B69E46A40D988E287A4E4D" type="DOI">10.5281/zenodo.175790</mods:identifier>
<mods:identifier id="5780D598DC1BE68EEB4AD99477E11B58" type="GBIF-Dataset">2a6512b7-5bd9-4682-b5db-79c72373b68c</mods:identifier>
<mods:identifier id="D80C3B25FED9C934E6E5035EE94DA739" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B9F24A7C04FF95FF3054A2618625C4" ID-DOI="http://doi.org/10.5281/zenodo.6248065" ID-GBIF-Taxon="119355651" ID-Zenodo-Dep="6248065" LSID="urn:lsid:plazi:treatment:03B9F24A7C04FF95FF3054A2618625C4" httpUri="http://treatment.plazi.org/id/03B9F24A7C04FF95FF3054A2618625C4" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection id="C30A10D77C04FF94FF3054A267FE26F4" box="[151,994,151,177]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8BAF435C7C04FF94FF3054A267FE26F4" blockId="4.[151,994,151,177]" box="[151,994,151,177]" pageId="4" pageNumber="5">
<heading id="D0E7F4307C04FF94FF3054A267FE26F4" bold="true" box="[151,994,151,177]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B9649F4E7C04FF94FF3054A267FE26F4" bold="true" box="[151,994,151,177]" pageId="4" pageNumber="5">
Key to the known species of
<taxonomicName id="4C1038DF7C04FF94FE5F54A2666D26F4" ID-CoL="8VRR8" box="[504,625,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9649F4E7C04FF94FE5F54A2666D26F4" bold="true" box="[504,625,151,177]" italics="true" pageId="4" pageNumber="5">Hydraena</emphasis>
</taxonomicName>
from Palawan and
<taxonomicName id="4C1038DF7C04FF94FCC154A267FE26F4" box="[870,994,151,177]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="43" phylum="Arthropoda" rank="species" species="busuanga" status="sp. nov.">Busuanga</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A10D77C04FF95FF3054D7618625C4" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="key">
<keyStep id="30E45BF97C04FF94FF3054D761862761" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF3054D7618626B9" blockId="4.[151,1437,226,2012]" box="[151,1434,226,252]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3054D7618626B9" box="[151,1434,226,252]" pageId="4" pageNumber="5">
1 Total length usually&gt;
<quantity id="4CE8EEB97C04FF94FE6A54D7663B26B9" box="[461,551,226,252]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="1.3">1.3 mm</quantity>
................................................................................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF30553F61862761" blockId="4.[151,1437,226,2012]" box="[151,1434,266,292]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30553F61862761" box="[151,1434,266,292]" pageId="4" pageNumber="5">
- Total length usually &lt;
<quantity id="4CE8EEB97C04FF94FE6A553F663B2761" box="[461,551,266,292]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="1.3">1.3 mm</quantity>
................................................................................................................... 9
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF305507618527A9" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF30550761862781" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30550761862781" pageId="4" pageNumber="5">
2 Entire body and pronotum about twice as wide as long; general habitus rather robust; elytra and pronotum distinctly punctate, punctures deeply impressed (Fig. 13). Male: aedeagus with long apical flagellum (
<figureCitation id="132B5FD97C04FF94FACB556F64FB27D9" captionStart="FIGURE 27" captionStartId="36.[151,255,1627,1651]" captionTargetBox="[264,1321,395,1600]" captionTargetId="figure@36.[264,1323,394,1604]" captionTargetPageId="36" captionText="FIGURE 27. Hydraena busuanga sp. n. a – c) aedeagus (paratype from Mabintangen, Busuanga) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175805/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
a). Female: gonocoxite subcrescentic, setae of subapical tufts furcate (
<figureCitation id="132B5FD97C04FF94FBBE55B7607227D9" box="[1049,1134,386,412]" captionStart="FIGURE 27" captionStartId="36.[151,255,1627,1651]" captionTargetBox="[264,1321,395,1600]" captionTargetId="figure@36.[264,1323,394,1604]" captionTargetPageId="36" captionText="FIGURE 27. Hydraena busuanga sp. n. a – c) aedeagus (paratype from Mabintangen, Busuanga) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175805/files/figure.png" pageId="4" pageNumber="5">Fig. 27</figureCitation>
e) .................................... ..............................................................................................................
<taxonomicName id="4C1038DF7C04FF94FC58559F61502781" ID-CoL="3N375" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1023,1356,426,452]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="busuanga" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FC58559F60892786" box="[1023,1173,426,451]" italics="true" pageId="4" pageNumber="5">H. busuanga</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C04FF94FAFF559F61862781" bold="true" box="[1368,1434,426,452]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF559F61862781" box="[1368,1434,426,452]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF3055E7618527A9" blockId="4.[151,1437,226,2012]" box="[151,1433,466,492]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3055E7618527A9" box="[151,1433,466,492]" pageId="4" pageNumber="5">- Pronotum wider than long, but never twice as wide as long (Figs. 3, 6–9, 11, 12) ................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF3055CF61852479" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF3055CF61862451" blockId="4.[151,1437,226,2012]" box="[151,1434,506,532]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3055CF61862451" box="[151,1434,506,532]" pageId="4" pageNumber="5">3 Pronotal disc with distinct dark, rectangular or irregular spot (Figs. 3, 7, 8, 11, 12) ................................ 4</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF30561761852479" blockId="4.[151,1437,226,2012]" box="[151,1433,546,572]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30561761852479" box="[151,1433,546,572]" pageId="4" pageNumber="5">- Pronotum evenly reddish or golden brown, without distinct dark rectangular spot (Figs. 6, 9) ............... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF30567F61862499" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF30567F618624C9" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30567F618624C9" pageId="4" pageNumber="5">4 Lateral elytral gutter distinctly explanate (Figs. 7, 8), pseudepipleuron broad, reaching beyond posterior 0.2 .............................................................................................................................................................. 5</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF3056AF61862499" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3056AF61862499" pageId="4" pageNumber="5">- Lateral elytral gutter only slightly explanate anteriorly, pseudepipleuron not reaching posterior 0.2 (Figs. 3, 11, 12) .................................................................................................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF3056DF61862539" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF3056DF61862569" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3056DF61862569" pageId="4" pageNumber="5">
<specimenCount id="9D1688D57C04FF94FF3056DF651B2541" box="[151,263,746,772]" pageId="4" pageNumber="5" type="male">5 Male</specimenCount>
: aedeagus about half as broad as long, ejaculatory duct ending in a strongly (more than 90°) curved, flagellum-like structure (
<figureCitation id="132B5FD97C04FF94FE4D572766232569" box="[490,575,786,812]" captionStart="FIGURE 20" captionStartId="22.[151,255,1412,1436]" captionTargetBox="[267,1320,200,1378]" captionTargetId="figure@22.[264,1323,194,1388]" captionTargetPageId="22" captionText="FIGURE 20. Hydraena claudia sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of parameres omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175798/files/figure.png" pageId="4" pageNumber="5">Fig. 20</figureCitation>
a) ...........................................................
<taxonomicName id="4C1038DF7C04FF94FBBD572761502569" ID-CoL="3N37N" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1050,1356,786,812]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="claudia" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FBBD5727608A256E" box="[1050,1174,786,811]" italics="true" pageId="4" pageNumber="5">H. claudia</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C04FF94FAFF572761862569" bold="true" box="[1368,1434,786,812]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF572761862569" box="[1368,1434,786,812]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF30570F61862539" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30570F61862539" pageId="4" pageNumber="5">
- Male: aedeagus about one third as broad as long, ejaculatory duct ending in an obtusely (less than 45°) bent tube-like structure (
<figureCitation id="132B5FD97C04FF94FE41575766252539" box="[486,569,866,892]" captionStart="FIGURE 19" captionStartId="19.[151,255,1921,1945]" captionTargetBox="[267,1323,495,1888]" captionTargetId="figure@19.[263,1323,474,1898]" captionTargetPageId="19" captionText="FIGURE 19. Hydraena castanescens sp. n. a – c) aedeagus (holotype) in ventral, dorsal (ventral setae of left paramere omitted) and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175797/files/figure.png" pageId="4" pageNumber="5">Fig. 19</figureCitation>
a) ..................................................
<taxonomicName id="4C1038DF7C04FF94FC7F575761502539" ID-CoL="3N37D" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[984,1356,866,892]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="castanescens" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FC7F57576089253E" box="[984,1173,866,891]" italics="true" pageId="4" pageNumber="5">H. castanescens</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C04FF94FAFF575761862539" bold="true" box="[1368,1434,866,892]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF575761862539" box="[1368,1434,866,892]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF3057BF61862229" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF3057BF61862259" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3057BF61862259" pageId="4" pageNumber="5">
6 Elytral punctures small and only moderately impressed (Fig. 12). Male: inner side of hind tibia unilaterally dilated (Fig. 12); parameres inserted at basal 0.2 of aedeagus, longer than median lobe (
<figureCitation id="132B5FD97C04FF94FA82578761632589" box="[1317,1407,946,972]" captionStart="FIGURE 26" captionStartId="33.[151,255,1629,1653]" captionTargetBox="[269,1322,194,1586]" captionTargetId="figure@33.[264,1323,184,1605]" captionTargetPageId="33" captionText="FIGURE 26. Hydraena hosiwergi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere, mesal setae cluster of left paramere and weakly sclerotized distal lobe omitted), dorsal (margin setae and mesal setae cluster of left paramere omitted) and lateral view (margin setae of left and right paramere and weakly sclerotized distal lobe omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175804/files/figure.png" pageId="4" pageNumber="5">Fig. 26</figureCitation>
b). Female: gonocoxite crescentic, with two distinctly separated cavities, short apical tufts of setae not distinctly longer than setae on disc (
<figureCitation id="132B5FD97C04FF94FDED503766BD2259" box="[586,673,1026,1052]" captionStart="FIGURE 26" captionStartId="33.[151,255,1629,1653]" captionTargetBox="[269,1322,194,1586]" captionTargetId="figure@33.[264,1323,184,1605]" captionTargetPageId="33" captionText="FIGURE 26. Hydraena hosiwergi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere, mesal setae cluster of left paramere and weakly sclerotized distal lobe omitted), dorsal (margin setae and mesal setae cluster of left paramere omitted) and lateral view (margin setae of left and right paramere and weakly sclerotized distal lobe omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175804/files/figure.png" pageId="4" pageNumber="5">Fig. 26</figureCitation>
e)
<emphasis id="B9649F4E7C04FF94FD1050376089225E" box="[695,1173,1026,1051]" italics="true" pageId="4" pageNumber="5">
............................................
<taxonomicName id="4C1038DF7C04FF94FC5850376089225E" ID-CoL="6MJBC" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1023,1173,1026,1051]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hosiwergi" status="sp. nov.">H. hosiwergi</taxonomicName>
</emphasis>
Freitag &amp; Jäch,
<emphasis id="B9649F4E7C04FF94FAFF503761862259" bold="true" box="[1368,1434,1026,1052]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF503761862259" box="[1368,1434,1026,1052]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF30501F61862229" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30501F61862229" pageId="4" pageNumber="5">
- Elytral punctures large and distinctly impressed (Fig. 3, 11). Male hind tibia not unilaterally dilated (Figs. 3, 11); gonocoxite and aedeagus different (
<figureCitation id="132B5FD97C04FF94FD1C506767052229" box="[699,793,1106,1132]" captionStart="FIGURE 15" captionStartId="11.[151,255,1670,1694]" captionTargetBox="[290,1312,200,1641]" captionTargetId="figure@11.[264,1323,194,1646]" captionTargetPageId="11" captionText="FIGURE 15. Hydraena scabra d’Orchymont. a – c) aedeagus (specimen from Taytay, Palawan) in ventral, dorsal (parameral setae omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175793/files/figure.png" pageId="4" pageNumber="5">Figs. 15</figureCitation>
a–c, e; 25a–c, e) ........................................................ 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF30504F61852349" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF30504F618622F9" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30504F618622F9" pageId="4" pageNumber="5">
<specimenCount id="9D1688D57C04FF94FF30504F651B22D1" box="[151,263,1146,1172]" pageId="4" pageNumber="5" type="male">7 Male</specimenCount>
: aedeagus Y-shaped, with medioapical gap (
<figureCitation id="132B5FD97C04FF94FCA8504F677A22D1" box="[783,870,1146,1172]" captionStart="FIGURE 25" captionStartId="32.[151,255,1659,1683]" captionTargetBox="[266,1318,183,1635]" captionTargetId="figure@32.[264,1323,183,1635]" captionTargetPageId="32" captionText="FIGURE 25. Hydraena manguao sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175803/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
a). Female: gonocoxite subcrescentic to subrectangular, condyles inconspicuous (
<figureCitation id="132B5FD97C04FF94FDF5509766BB22F9" box="[594,679,1186,1212]" captionStart="FIGURE 25" captionStartId="32.[151,255,1659,1683]" captionTargetBox="[266,1318,183,1635]" captionTargetId="figure@32.[264,1323,183,1635]" captionTargetPageId="32" captionText="FIGURE 25. Hydraena manguao sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175803/files/figure.png" pageId="4" pageNumber="5">Fig. 25</figureCitation>
e) ..........................................
<taxonomicName id="4C1038DF7C04FF94FBA35096615022F9" ID-CoL="6MJB3" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1028,1356,1186,1212]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FBA35096608922F9" box="[1028,1173,1187,1212]" italics="true" pageId="4" pageNumber="5">H. manguao</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C04FF94FAFF5097618622F9" bold="true" box="[1368,1434,1186,1212]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF5097618622F9" box="[1368,1434,1186,1212]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF3050FF61852349" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
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- Male: aedeagus compact and intricately shaped (
<figureCitation id="132B5FD97C04FF94FCAD50FF677D22A1" box="[778,865,1226,1252]" captionStart="FIGURE 15" captionStartId="11.[151,255,1670,1694]" captionTargetBox="[290,1312,200,1641]" captionTargetId="figure@11.[264,1323,194,1646]" captionTargetPageId="11" captionText="FIGURE 15. Hydraena scabra d’Orchymont. a – c) aedeagus (specimen from Taytay, Palawan) in ventral, dorsal (parameral setae omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175793/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
a). Female: gonocoxite subtrapezoidal, condyles prominent (
<figureCitation id="132B5FD97C04FF94FEE850C765BF2349" box="[335,419,1266,1292]" captionStart="FIGURE 15" captionStartId="11.[151,255,1670,1694]" captionTargetBox="[290,1312,200,1641]" captionTargetId="figure@11.[264,1323,194,1646]" captionTargetPageId="11" captionText="FIGURE 15. Hydraena scabra d’Orchymont. a – c) aedeagus (specimen from Taytay, Palawan) in ventral, dorsal (parameral setae omitted) and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175793/files/figure.png" pageId="4" pageNumber="5">Fig. 15</figureCitation>
e) ....................................................................................
<taxonomicName id="4C1038DF7C04FF94FB9F50C661852349" ID-CoL="3N3HB" authority="d'Orchymont, 1925" authorityName="d'Orchymont" authorityYear="1925" box="[1080,1433,1266,1292]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="scabra">
<emphasis id="B9649F4E7C04FF94FB9F50C660B02349" box="[1080,1196,1267,1292]" italics="true" pageId="4" pageNumber="5">H. scabra</emphasis>
d’Orchymont, 1925
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</paragraph>
</keyStep>
<keyStep id="30E45BF97C04FF94FF30512F618623B9" pageId="4" pageNumber="5">
<paragraph id="8BAF435C7C04FF94FF30512F618623C1" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30512F618623C1" pageId="4" pageNumber="5">
8 Elytron with&gt; 13 rows of punctures (Fig. 6). Male: distal lobe of aedeagus distinctly asymmetric, tapered and hood-like; parameres short and stout (
<figureCitation id="132B5FD97C04FF94FD1D5177670D2319" box="[698,785,1346,1372]" captionStart="FIGURE 18" captionStartId="17.[151,255,1492,1516]" captionTargetBox="[270,1316,195,1454]" captionTargetId="figure@17.[264,1323,194,1469]" captionTargetPageId="17" captionText="FIGURE 18. Hydraena zetteli sp. n. a – c) aedeagus (paratype from Cabayugan, Palawan) in ventral (parameral setae omitted), dorsal (parameral tufts omitted) and lateral view (tuft of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175796/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
a). Female: disc of tergite X with squamose setae, with an apical notch (
<figureCitation id="132B5FD97C04FF94FE20515F65C623C1" box="[391,474,1386,1412]" captionStart="FIGURE 18" captionStartId="17.[151,255,1492,1516]" captionTargetBox="[270,1316,195,1454]" captionTargetId="figure@17.[264,1323,194,1469]" captionTargetPageId="17" captionText="FIGURE 18. Hydraena zetteli sp. n. a – c) aedeagus (paratype from Cabayugan, Palawan) in ventral (parameral setae omitted), dorsal (parameral tufts omitted) and lateral view (tuft of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175796/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
h) ..........................................................................
<taxonomicName id="4C1038DF7C04FF94FB8B515E615023C1" ID-CoL="3N3L4" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1068,1356,1386,1412]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="zetteli" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FB8B515E608923C1" box="[1068,1173,1387,1412]" italics="true" pageId="4" pageNumber="5">H. zetteli</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C04FF94FAFF515F618623C1" bold="true" box="[1368,1434,1386,1412]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF515F618623C1" box="[1368,1434,1386,1412]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF3051A7618623B9" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
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- Elytron with &lt;13 rows of punctures (Fig. 9). Male: distal lobe of aedeagus broad, subsymmetric; parameres long and slender (
<figureCitation id="132B5FD97C04FF94FDBC518F666D2391" box="[539,625,1466,1492]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
a). Female: disc of tergite X with trichoid setae, without apical notch (
<figureCitation id="132B5FD97C04FF94FF6851D7653F23B9" box="[207,291,1506,1532]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
h) ................................................................................................
<taxonomicName id="4C1038DF7C04FF94FBB451D6615023B9" ID-CoL="3N3CD" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1043,1356,1506,1532]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
<emphasis id="B9649F4E7C04FF94FBB451D6608A23B9" box="[1043,1174,1507,1532]" italics="true" pageId="4" pageNumber="5">H. kodadai</emphasis>
Freitag &amp; Jäch
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,
<emphasis id="B9649F4E7C04FF94FAFF51D7618623B9" bold="true" box="[1368,1434,1506,1532]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A25722357C04FF94FAFF51D7618623B9" box="[1368,1434,1506,1532]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="8BAF435C7C04FF94FF30523F61852031" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30523F61852031" pageId="4" pageNumber="5">9 Pronotum distinctly sinuously or arcuately convergent posteriorly; posterior margin only 0.75–0.80 times as broad as pronotal width across middle, dark rectangular spot usually indistinct or lacking (Figs. 4–6, 9) .................................................................................................................................................................. 10</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF3052B761862081" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3052B761862081" pageId="4" pageNumber="5">- Pronotum only slightly and evenly convergent posteriorly; posterior margin more than 0.80 times as broad as pronotal width across middle, dark rectangular spot distinct (Figs. 10, 11) ....................................... 12</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BAF435C7C04FF94FF3052E761862121" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF3052E761862121" pageId="4" pageNumber="5">
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10 Total length ca.
<quantity id="4CE8EEB97C04FF94FE2152E765FF20A9" box="[390,483,1746,1772]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="1.3">1.3 mm</quantity>
. Lateral elytral gutter widely explanate, rather straight and subparallel at anterior 0.2–0.7; body usually reddish to golden brown (Figs. 6, 9). Male: aedeagal parameres either asymmetric, stout, with tufts of fused setae (
<figureCitation id="132B5FD97C04FF94FDEA531766B92179" box="[589,677,1826,1852]" captionStart="FIGURE 18" captionStartId="17.[151,255,1492,1516]" captionTargetBox="[270,1316,195,1454]" captionTargetId="figure@17.[264,1323,194,1469]" captionTargetPageId="17" captionText="FIGURE 18. Hydraena zetteli sp. n. a – c) aedeagus (paratype from Cabayugan, Palawan) in ventral (parameral setae omitted), dorsal (parameral tufts omitted) and lateral view (tuft of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175796/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
a) or subsymmetric and elongate (
<figureCitation id="132B5FD97C04FF94FBE1531760BC2179" box="[1094,1184,1826,1852]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
a). Female: tergite X either with squamose setae on disc (
<figureCitation id="132B5FD97C04FF94FDCB537F66DC2121" box="[620,704,1866,1892]" captionStart="FIGURE 18" captionStartId="17.[151,255,1492,1516]" captionTargetBox="[270,1316,195,1454]" captionTargetId="figure@17.[264,1323,194,1469]" captionTargetPageId="17" captionText="FIGURE 18. Hydraena zetteli sp. n. a – c) aedeagus (paratype from Cabayugan, Palawan) in ventral (parameral setae omitted), dorsal (parameral tufts omitted) and lateral view (tuft of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175796/files/figure.png" pageId="4" pageNumber="5">Fig. 18</figureCitation>
h) or without apical notch (
<figureCitation id="132B5FD97C04FF94FC59537F604E2121" box="[1022,1106,1866,1892]" captionStart="FIGURE 21" captionStartId="24.[151,255,1670,1694]" captionTargetBox="[274,1322,194,1644]" captionTargetId="figure@24.[264,1323,194,1646]" captionTargetPageId="24" captionText="FIGURE 21. Hydraena kodadai sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view (setae of left paramere omitted); d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175799/files/figure.png" pageId="4" pageNumber="5">Fig. 21</figureCitation>
h) .................................... 8
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C04FF94FF30534761802199" blockId="4.[151,1437,226,2012]" pageId="4" pageNumber="5">
<keyLead id="30E1E0697C04FF94FF30534761802199" pageId="4" pageNumber="5">
- Total length &lt;
<quantity id="4CE8EEB97C04FF94FED2534765C821C9" box="[373,468,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="4" pageNumber="5" unit="mm" value="1.3">1.3 mm</quantity>
. Elytra rather ovoid, lateral margins moderately explanate; usually greyish to yellowish brown (Figs. 4, 5). Male: aedeagal parameres always asymmetrical, stout, without fused tufts of setae, but with separate setae. Female: tergite X with one apical notch, without squamose setae on disc
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C05FF95FF6054AD61862744" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
(
<figureCitation id="132B5FD97C05FF95FF6854AD653326F7" box="[207,303,152,178]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="5" pageNumber="6">Figs. 16</figureCitation>
h, 17h) ........................................................................................................................................ 11
<keyLead id="30E1E0697C05FF95FF3054F561862744" pageId="5" pageNumber="6">
<keyStep id="30E45BF97C05FF95FF3054F561862744" pageId="5" pageNumber="6">
11 Total length&gt;
<quantity id="4CE8EEB97C05FF95FEC854F565D1269F" box="[367,461,192,218]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="mm" value="1.2">1.2 mm</quantity>
. Male: distal lobe of aedeagus ovoidly rounded (
<figureCitation id="132B5FD97C05FF95FBA554F5604B269F" box="[1026,1111,192,218]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
b). Female: subapical fringe of tergite X with vermiform setae (
<figureCitation id="132B5FD97C05FF95FDF854DD66AE2747" box="[607,690,232,258]" captionStart="FIGURE 16" captionStartId="12.[151,255,1659,1683]" captionTargetBox="[270,1322,190,1625]" captionTargetId="figure@12.[264,1323,182,1634]" captionTargetPageId="12" captionText="FIGURE 16. Hydraena jojoorculloi sp. n. a – c) aedeagus (holotype) in ventral (setae of right paramere omitted), dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175794/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
h) ....................................
<taxonomicName id="4C1038DF7C05FF95FC4F54DD61502747" ID-CoL="3N3C7" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1000,1356,232,258]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="jojoorculloi" status="sp. nov.">
<emphasis id="B9649F4E7C05FF95FC4F54DD60892744" box="[1000,1173,232,257]" italics="true" pageId="5" pageNumber="6">H. jojoorculloi</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C05FF95FAFF54D261862744" bold="true" box="[1368,1434,231,257]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A25722357C05FF95FAFF54D261862744" box="[1368,1434,231,257]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C05FF95FF30552561862714" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
<keyLead id="30E1E0697C05FF95FF30552561862714" pageId="5" pageNumber="6">
- Total length &lt;
<quantity id="4CE8EEB97C05FF95FEDE552565C5276F" box="[377,473,272,298]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="6" unit="mm" value="1.2">1.2 mm</quantity>
. Male: distal lobe of aedeagus not ovoid, with few short appendages (
<figureCitation id="132B5FD97C05FF95FA805525619C276F" box="[1319,1408,272,298]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
a). Female: tergite X without vermiform setae (
<figureCitation id="132B5FD97C05FF95FD6D550D67022717" box="[714,798,312,338]" captionStart="FIGURE 17" captionStartId="15.[151,255,1670,1694]" captionTargetBox="[271,1314,196,1635]" captionTargetId="figure@15.[264,1323,194,1646]" captionTargetPageId="15" captionText="FIGURE 17. Hydraena nielshaggei sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175795/files/figure.png" pageId="5" pageNumber="6">Fig. 17</figureCitation>
h) .....................
<taxonomicName id="4C1038DF7C05FF95FC4D550D61502717" ID-CoL="6MJCK" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1002,1356,312,338]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nielshaggei" status="sp. nov.">
<emphasis id="B9649F4E7C05FF95FC4D550D60892714" box="[1002,1173,312,337]" italics="true" pageId="5" pageNumber="6">H. nielshaggei</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
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<paragraph id="8BAF435C7C05FF95FF305555618627B4" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
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12 Mesoventral intercoxal process narrower than pseudepipleuron, not reaching posterior margin of mesocoxa, not firmly connected with metaventrite. Male: aedeagus Y-shaped, with medioapical gap (
<figureCitation id="132B5FD97C05FF95FA8E55BD619C27E7" box="[1321,1408,392,418]" captionStart="FIGURE 25" captionStartId="32.[151,255,1659,1683]" captionTargetBox="[266,1318,183,1635]" captionTargetId="figure@32.[264,1323,183,1635]" captionTargetPageId="32" captionText="FIGURE 25. Hydraena manguao sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175803/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
a). Female: gonocoxite subcrescentic to subrectangular (
<figureCitation id="132B5FD97C05FF95FC9F55856793278F" box="[824,911,432,458]" captionStart="FIGURE 25" captionStartId="32.[151,255,1659,1683]" captionTargetBox="[266,1318,183,1635]" captionTargetId="figure@32.[264,1323,183,1635]" captionTargetPageId="32" captionText="FIGURE 25. Hydraena manguao sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175803/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
e); tergite X with only one apical notch, subapical fringe with vermiform setae (
<figureCitation id="132B5FD97C05FF95FDCC55ED66DC27B7" box="[619,704,472,498]" captionStart="FIGURE 25" captionStartId="32.[151,255,1659,1683]" captionTargetBox="[266,1318,183,1635]" captionTargetId="figure@32.[264,1323,183,1635]" captionTargetPageId="32" captionText="FIGURE 25. Hydraena manguao sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175803/files/figure.png" pageId="5" pageNumber="6">Fig. 25</figureCitation>
h) ......................................
<taxonomicName id="4C1038DF7C05FF95FBA355ED615027B7" ID-CoL="6MJB3" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[1028,1356,472,498]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="manguao" status="sp. nov.">
<emphasis id="B9649F4E7C05FF95FBA355ED608927B4" box="[1028,1173,472,497]" italics="true" pageId="5" pageNumber="6">H. manguao</emphasis>
Freitag &amp; Jäch
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,
<emphasis id="B9649F4E7C05FF95FAFF55E2618627B4" bold="true" box="[1368,1434,471,497]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A25722357C05FF95FAFF55E2618627B4" box="[1368,1434,471,497]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C05FF95FF305635618524D7" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
<keyLead id="30E1E0697C05FF95FF305635618524D7" pageId="5" pageNumber="6">
- Mesoventral intercoxal process very wide and long, broader than pseudepipleuron, reaching beyond posterior margin of mesocoxa, firmly connected with metaventrite. Male: aedeagus compact and intricately shaped (
<figureCitation id="132B5FD97C05FF95FE8C56656592242F" box="[299,398,592,618]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="5" pageNumber="6">Figs. 22</figureCitation>
a–c, 23a–c). Female: gonocoxite subtriangular (
<figureCitation id="132B5FD97C05FF95FC6E5665603F242F" box="[969,1059,592,618]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="5" pageNumber="6">Fig. 22</figureCitation>
e); tergite X without vermiform setae, apical margin with three notches (
<figureCitation id="132B5FD97C05FF95FD3A564D66EE24D7" box="[669,754,632,658]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="5" pageNumber="6">Fig. 22</figureCitation>
h) .................................................................................. 13
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<paragraph id="8BAF435C7C05FF95FF305695618524FF" blockId="5.[151,1437,152,898]" box="[151,1433,672,698]" pageId="5" pageNumber="6">
<keyLead id="30E1E0697C05FF95FF305695618524FF" box="[151,1433,672,698]" pageId="5" pageNumber="6">
<specimenCount id="9D1688D57C05FF95FF305695651124FF" box="[151,269,672,698]" pageId="5" pageNumber="6" type="male">13 Males</specimenCount>
........................................................................................................................................................ 14
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C05FF95FF3056FD618524A7" blockId="5.[151,1437,152,898]" box="[151,1433,712,738]" pageId="5" pageNumber="6">
<keyLead id="30E1E0697C05FF95FF3056FD618524A7" box="[151,1433,712,738]" pageId="5" pageNumber="6">
- Females (not clearly distinguishable) ........
<emphasis id="B9649F4E7C05FF95FD6B56FD608424A4" box="[716,1176,712,737]" italics="true" pageId="5" pageNumber="6">
<taxonomicName id="4C1038DF7C05FF95FD6B56FD679324A4" ID-CoL="6MJBR" box="[716,911,712,737]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="palawanensis">H. palawanensis</taxonomicName>
/
<taxonomicName id="4C1038DF7C05FF95FC0156FD608424A4" ID-CoL="3N3G3" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[934,1176,712,737]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pseudopalawanensis">pseudopalawanensis</taxonomicName>
</emphasis>
Freitag &amp; Jäch spp.n.
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</paragraph>
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<paragraph id="8BAF435C7C05FF95FF3056C561862574" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
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14 Main piece of aedeagus emarginate near insertion of right paramere; left paramere not longer than width of phallobase (
<figureCitation id="132B5FD97C05FF95FED4572D65D52577" box="[371,457,792,818]" captionStart="FIGURE 22" captionStartId="27.[151,255,1189,1213]" captionTargetBox="[269,1321,197,1154]" captionTargetId="figure@27.[264,1323,194,1166]" captionTargetPageId="27" captionText="FIGURE 22. Hydraena palawanensis sp. n. a – c) aedeagus (holotype) in ventral, dorsal and lateral view; d) male sternite X and spiculum; e) gonocoxite; f – g) spermatheca; h) female tergite X." httpUri="https://zenodo.org/record/175800/files/figure.png" pageId="5" pageNumber="6">Fig. 22</figureCitation>
a) .................................................................
<taxonomicName id="4C1038DF7C05FF95FC75572D61502577" ID-CoL="6MJBR" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[978,1356,792,818]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="palawanensis" status="sp. nov.">
<emphasis id="B9649F4E7C05FF95FC75572D60892574" box="[978,1173,792,817]" italics="true" pageId="5" pageNumber="6">H. palawanensis</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
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<taxonomicNameLabel id="A25722357C05FF95FAFF572261862574" box="[1368,1434,791,817]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAF435C7C05FF95FF305775618625C4" blockId="5.[151,1437,152,898]" pageId="5" pageNumber="6">
<keyLead id="30E1E0697C05FF95FF305775618625C4" pageId="5" pageNumber="6">
- Main piece of aedeagus not emarginate near right paramere insertion; left paramere bent, distinctly longer than width of phallobase (
<figureCitation id="132B5FD97C05FF95FE50575D665025C7" box="[503,588,872,898]" captionStart="FIGURE 23" captionStartId="29.[151,255,1223,1247]" captionTargetBox="[267,1322,197,1187]" captionTargetId="figure@29.[264,1323,194,1199]" captionTargetPageId="29" captionText="FIGURE 23. Hydraena pseudopalawanensis sp. n. a – c) aedeagus (paratype from Cabayugan, Palawan) in ventral, dorsal and lateral view; d) male sternite X and spiculum." httpUri="https://zenodo.org/record/175801/files/figure.png" pageId="5" pageNumber="6">Fig. 23</figureCitation>
a) ....................................
<taxonomicName id="4C1038DF7C05FF95FCD8575D615025C7" ID-CoL="3N3G3" authority="Freitag &amp; Jach" authorityName="Freitag &amp; Jach" box="[895,1356,872,898]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="pseudopalawanensis" status="sp. nov.">
<emphasis id="B9649F4E7C05FF95FCD8575D608925C4" box="[895,1173,872,897]" italics="true" pageId="5" pageNumber="6">H. pseudopalawanensis</emphasis>
Freitag &amp; Jäch
</taxonomicName>
,
<emphasis id="B9649F4E7C05FF95FAFF5752618625C4" bold="true" box="[1368,1434,871,897]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A25722357C05FF95FAFF5752618625C4" box="[1368,1434,871,897]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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</treatment>
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