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<document id="5F1A9A8C45A92F7AA53177C9B909E584" ID-CLB-Dataset="30873" ID-DOI="10.5852/ejt.2018.399" ID-GBIF-Dataset="b389e1f3-82be-4258-9b09-b7ef9d839f99" ID-Zenodo-Dep="1163422" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1517404992146" checkinUser="plazi" docAuthor="Zonstein, Sergei, Kunt, Kadir B. &amp; Yağmur, Ersen A." docDate="2018" docId="03B9B44C56700C2E514A87CFFB1E4DBF" docLanguage="en" docName="ejt-399_zonstein_kunt.pdf.imf" docOrigin="European Journal of Taxonomy 399" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Raveniola adjarica Zonstein &amp; Kunt &amp; Yağmur 2018, sp. nov." docType="treatment" docUuid="BCF871CB-E0C5-4A75-B376-EA83C72B04A5" docUuidSource="ZooBank" docVersion="14" lastPageNumber="10" masterDocId="FF80CC3456770C27533C8513FFDA4805" masterDocTitle="A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia" masterLastPageNumber="93" masterPageNumber="1" pageNumber="8" updateTime="1698518020641" updateUser="ExternalLinkService">
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<mods:title id="EBC5773D39A5E726C7F210BBA0CAFCEB">A revision of the spider genus Raveniola (Araneae, Nemesiidae). I. Species from Western Asia</mods:title>
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<mods:namePart id="4ECF9C3CE8077B2DB489473582CE61BB">Zonstein, Sergei</mods:namePart>
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<mods:namePart id="27B0F834B58AC422F7209EB34A786268">Kunt, Kadir B.</mods:namePart>
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<mods:title id="553447E7938992C0C4AE1177A29506C4">European Journal of Taxonomy</mods:title>
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<mods:date id="20F7D50218F7EB29C8A2DEC099B08673">2018</mods:date>
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<treatment id="03B9B44C56700C2E514A87CFFB1E4DBF" ID-DOI="http://doi.org/10.5281/zenodo.5980277" ID-GBIF-Taxon="140561340" ID-Zenodo-Dep="5980277" LSID="urn:lsid:zoobank.org:act:BCF871CB-E0C5-4A75-B376-EA83C72B04A5" httpUri="http://treatment.plazi.org/id/03B9B44C56700C2E514A87CFFB1E4DBF" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8">
<subSubSection id="C30A56D156700C20514A87CFFB164B1E" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BAF055A56700C20514A87CFFC674AF2" blockId="7.[359,1228,732,829]" box="[630,957,732,759]" pageId="7" pageNumber="8">
<heading id="D0E7B23656700C20514A87CFFC674AF2" box="[630,957,732,759]" centered="true" fontSize="11" level="2" pageId="7" pageNumber="8" reason="2">
<taxonomicName id="4C107ED956700C20514A87CFFC814AF2" ID-CoL="79596" authority="Zonstein &amp; Kunt &amp; Yağmur, 2018" authorityName="Zonstein &amp; Kunt &amp; Yağmur" authorityYear="2018" box="[630,859,732,759]" class="Arachnida" family="Nemesiidae" genus="Raveniola" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="adjarica" status="sp. nov.">
<emphasis id="B964D94856700C20514A87CFFC814AF2" bold="true" box="[630,859,732,759]" italics="true" pageId="7" pageNumber="8">Raveniola adjarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257643356700C20505F87CEFC674AF2" box="[867,957,733,759]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BAF055A56700C20525B8613FB164B1E" blockId="7.[359,1228,732,829]" box="[359,1228,768,795]" pageId="7" pageNumber="8">urn:lsid:zoobank.org:act:BCF871CB-E0C5-4A75-B376-EA83C72B04A5</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C2052E48630FB814B38" box="[472,1115,803,829]" pageId="7" pageNumber="8" type="description">
<paragraph id="8BAF055A56700C2052E48630FB814B38" blockId="7.[359,1228,732,829]" box="[472,1115,803,829]" pageId="7" pageNumber="8">
<figureCitation id="132B19DF56700C2052E48630FD844B38" box="[472,606,803,829]" captionStart="Figs4351" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs4351. Raveniolaspp.,habitusof ♂♂(43,4546,48,5051)and ♀♀(44,47,49),dorsalviews.— 4344. R. adjarica sp. nov. 45. R. dunini sp. nov. 4647. R. hyrcanica Dunin, 1988. 4849. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="7" pageNumber="8">Figs 4344</figureCitation>
,
<figureCitation id="132B19DF56700C2051568630FD504B38" box="[618,650,803,829]" captionStart="Figs 8489" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 8489. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="7" pageNumber="8">84</figureCitation>
,
<figureCitation id="132B19DF56700C2051AA8630FD1F4B38" box="[662,709,803,829]" captionStart="Figs 102116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="7" pageNumber="8">102</figureCitation>
,
<figureCitation id="132B19DF56700C2051ED8630FD254B38" box="[721,767,803,829]" captionStart="Figs 117128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="7" pageNumber="8">117</figureCitation>
,
<figureCitation id="132B19DF56700C2050368630FCE34B38" box="[778,825,803,829]" captionStart="Figs129132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="7" pageNumber="8">129</figureCitation>
,
<figureCitation id="132B19DF56700C2050798630FCAE4B38" box="[837,884,803,829]" captionStart="Figs 149152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="7" pageNumber="8">149</figureCitation>
,
<figureCitation id="132B19DF56700C2050438630FC334B38" box="[895,1001,803,829]" captionStart="Figs 171179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171179. Raveniola hyrcanica species group, holotypic (171177) and conspecific (178179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171173. R. adjarica sp. nov. 174175. R. dunini sp. nov. 176 177. R. hyrcanica Dunin, 1988. 178179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="7" pageNumber="8">171173</figureCitation>
,
<figureCitation id="132B19DF56700C2050C98630FBF94B38" box="[1013,1059,803,829]" captionStart="Figs 198203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="7" pageNumber="8">198</figureCitation>
,
<figureCitation id="132B19DF56700C2057138630FB814B38" box="[1071,1115,803,829]" captionStart="Figure 219" captionStartId="85.[189,272,1534,1560]" captionTargetBox="[189,1398,265,1471]" captionTargetId="figure@85.[189,1398,265,1471]" captionText="Figure 219. Localities of Raveniola spp. in the western part of the Caucasus: R. adjarica sp. nov. () and R. pontica (Spassky, 1937) ()." httpUri="https://zenodo.org/record/1163494/files/figure.png" pageId="7" pageNumber="8">219</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C2053818676FC684B7A" box="[189,946,868,895]" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="8BAF055A56700C2053818676FC684B7A" blockId="7.[189,946,868,895]" box="[189,946,868,895]" pageId="7" pageNumber="8">
<taxonomicName id="4C107ED956700C2053818676FD4E4B7B" authority="Spassky, 1937: 368" authorityName="Spassky" authorityPageNumber="368" authorityYear="1937" box="[189,660,868,895]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856700C2053818676FE704B7B" box="[189,426,869,895]" italics="true" pageId="7" pageNumber="8">Brachythele pontica</emphasis>
<bibRefCitation id="EF8178AB56700C20528D8677FD874B7A" author="Spassky S. A." box="[433,605,868,895]" journalOrPublisher="Festschrift Embrik Strand" pageId="7" pageNumber="8" pagination="361 - 368" part="3" refString="Spassky S. A. 1937. Araneae palaearcticae novae. Mygalomorphae. I. Festschrift Embrik Strand 3: 361 - 368." title="Araneae palaearcticae novae. Mygalomorphae. I" type="journal article" year="1937">Spassky, 1937</bibRefCitation>
: 368
</taxonomicName>
(♀, part; misidentified).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C20538186B5FBBC4C30" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BAF055A56700C20538186B5FEEC4BC5" blockId="7.[189,1398,934,1077]" box="[189,310,934,960]" pageId="7" pageNumber="8">
<heading id="D0E7B23656700C20538186B5FEEC4BC5" bold="true" box="[189,310,934,960]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B964D94856700C20538186B5FEEC4BC5" bold="true" box="[189,310,934,960]" pageId="7" pageNumber="8">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56700C20538186C7FBBC4C30" blockId="7.[189,1398,934,1077]" pageId="7" pageNumber="8">
Different from other species in the same group by having a relatively short embolus with a rounded submedian keel (
<figureCitation id="132B19DF56700C2052B486EBFDF54C17" box="[392,559,1016,1042]" captionStart="Figs 171179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171179. Raveniola hyrcanica species group, holotypic (171177) and conspecific (178179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171173. R. adjarica sp. nov. 174175. R. dunini sp. nov. 176 177. R. hyrcanica Dunin, 1988. 178179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="7" pageNumber="8">Figs 171173</figureCitation>
; cf.
<figureCitation id="132B19DF56700C20515986EBFCD34C17" box="[613,777,1016,1042]" captionStart="Figs 171179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171179. Raveniola hyrcanica species group, holotypic (171177) and conspecific (178179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171173. R. adjarica sp. nov. 174175. R. dunini sp. nov. 176 177. R. hyrcanica Dunin, 1988. 178179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="7" pageNumber="8">Figs 174179</figureCitation>
) and long, asymmetrical spermathecae with median receptacles being considerably longer than laterals (
<figureCitation id="132B19DF56700C2050218108FC584C30" box="[797,898,1051,1077]" captionStart="Figs 198203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="7" pageNumber="8">Fig. 198</figureCitation>
, cf.
<figureCitation id="132B19DF56700C2050888108FB834C30" box="[948,1113,1051,1077]" captionStart="Figs 198203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="7" pageNumber="8">Figs 199203</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C205381814EFB754CCD" pageId="7" pageNumber="8" type="etymology">
<paragraph id="8BAF055A56700C205381814EFE9E4C72" blockId="7.[189,1399,1117,1225]" box="[189,324,1117,1143]" pageId="7" pageNumber="8">
<heading id="D0E7B23656700C205381814EFE9E4C72" bold="true" box="[189,324,1117,1143]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B964D94856700C205381814EFE9E4C72" bold="true" box="[189,324,1117,1143]" pageId="7" pageNumber="8">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56700C2053818198FB754CCD" blockId="7.[189,1399,1117,1225]" pageId="7" pageNumber="8">The specific epithet, a Latinized adjective, is derived from Adjaria (also Adjara or Adzharia), the historical region and administrative province of Georgia where the species was found.</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C20538181FCFD574EB7" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="8BAF055A56700C20538181FCFE704D0C" blockId="7.[189,426,1263,1289]" box="[189,426,1263,1289]" pageId="7" pageNumber="8">
<heading id="D0E7B23656700C20538181FCFE704D0C" bold="true" box="[189,426,1263,1289]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B964D94856700C20538181FCFE704D0C" bold="true" box="[189,426,1263,1289]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A56700C2053D6800DFE864D3D" blockId="7.[189,1399,1310,1407]" box="[234,348,1310,1336]" pageId="7" pageNumber="8">
<materialsCitation id="3B780F0756700C2053D6800DFE864D3D" ID-GBIF-Occurrence="1806244360" box="[234,348,1310,1336]" collectingDate="1989-09-07" collectionCode="TAU" collectorName="S. L. Zonstein" country="Georgia" elevation="925" latitude="41.73" location="Kintrishi Valley, 3 km NE of Khino" longLatPrecision="115" longitude="42.058334" municipality="South slope of Meskheti Mts" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54ABBBF856700C2053D6800DFE864D3D" box="[234,348,1310,1336]" pageId="7" pageNumber="8" type="holotype">
<emphasis id="B964D94856700C2053D6800DFE864D3D" bold="true" box="[234,348,1310,1336]" pageId="7" pageNumber="8">Holotype</emphasis>
</typeStatus>
</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56700C2053818052FCCA4D7A" blockId="7.[189,1399,1310,1407]" pageId="7" pageNumber="8">
<collectingCountry id="F30745CA56700C2053818052FE9C4D5E" box="[189,326,1345,1371]" name="Georgia" pageId="7" pageNumber="8">GEORGIA</collectingCountry>
:
<specimenCount id="9D16CED356700C2052728051FEBC4D59" box="[334,358,1346,1372]" pageId="7" pageNumber="8" type="male"></specimenCount>
,
<collectingMunicipality id="6BCB9F2056700C20524C8052FD614D59" box="[368,699,1345,1372]" pageId="7" pageNumber="8">South slope of Meskheti Mts</collectingMunicipality>
,
<location id="8ECF538156700C2051F98052FB804D5E" LSID="urn:lsid:plazi:treatment:03B9B44C56700C2E514A87CFFB1E4DBF:8ECF538156700C2051F98052FB804D5E" box="[709,1114,1345,1372]" country="Georgia" latitude="41.73" longLatPrecision="115" longitude="42.058334" municipality="South slope of Meskheti Mts" name="Kintrishi Valley, 3 km NE of Khino" pageId="7" pageNumber="8">
Kintrishi Valley,
<quantity id="4CE8A8BF56700C2050BB8052FC1A4D5E" box="[903,960,1345,1371]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="8" unit="km" value="3.0">3 km</quantity>
NE of Khino
</location>
,
<elevation id="003DE26956700C2057588052FB324D59" box="[1124,1256,1345,1372]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="7" pageNumber="8" unit="m" value="925.0" valueMax="950.0" valueMin="900.0">
<quantity id="4CE8A8BF56700C2057588052FB324D59" box="[1124,1256,1345,1372]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="9.5" metricValueMin="9.0" pageId="7" pageNumber="8" unit="m" value="925.0" valueMax="950.0" valueMin="900.0">900950 m</quantity>
</elevation>
,
<geoCoordinate id="EE24639D56700C2057CE802CFAA84D59" box="[1266,1394,1343,1372]" direction="north" orientation="latitude" pageId="7" pageNumber="8" precision="92" value="41.73">41°43.8 N</geoCoordinate>
,
<geoCoordinate id="EE24639D56700C2053818070FEE14D7A" box="[189,315,1379,1407]" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="92" value="42.058334">42°03.5 E</geoCoordinate>
,
<collectingDate id="EFEADA7256700C2052748076FE0F4D7A" box="[328,469,1381,1407]" pageId="7" pageNumber="8" value="1989-09-07">
<date id="FFAE239A56700C2052748076FE0F4D7A" box="[328,469,1381,1407]" pageId="7" pageNumber="8" value="1989-09-07">7 Sep. 1989</date>
</collectingDate>
,
<collectorName id="26E5608C56700C2052DE8076FD5B4D7B" box="[482,641,1380,1407]" pageId="7" pageNumber="8">S.L. Zonstein</collectorName>
leg. (
<collectionCode id="ED019D9F56700C2051FF8076FCDE4D7A" box="[707,772,1381,1407]" collectionName="Israel, Tel Aviv, Tel Aviv University" pageId="7" pageNumber="8">TAU</collectionCode>
).
</paragraph>
<paragraph id="8BAF055A56700C2053D680B5FDDA4DC4" blockId="7.[189,1399,1446,1543]" box="[234,512,1446,1473]" pageId="7" pageNumber="8">
<materialsCitation id="3B780F0756700C2053D680B5FDDA4DC4" ID-GBIF-Occurrence="1835183789" box="[234,512,1446,1473]" pageId="7" pageNumber="8">
<emphasis id="B964D94856700C2053D680B5FEBD4DC5" bold="true" box="[234,359,1446,1472]" pageId="7" pageNumber="8">Paratypes</emphasis>
(1 ♂, 3 ♀♀)
</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56700C20538180DAFAAD4E03" blockId="7.[189,1399,1446,1543]" pageId="7" pageNumber="8">
GEORGIA: 2 ♀♀, same collection data as for holotype, but 78 Sep. 1989 (TAU);
<materialsCitation id="3B780F0756700C2057AB80DAFAAD4E03" ID-GBIF-Occurrence="1806244409" pageId="7" pageNumber="8">1 ♂, 1 ♀, Kintrishi Valley, 0.5 km W of Zerabozeli, 450 m, 41°44.0 N, 41°59.2 E, 15 Sep. 1989, S.L. Zonstein leg. (TAU).</materialsCitation>
</paragraph>
<paragraph id="8BAF055A56700C2053D6833DFD544E4D" blockId="7.[189,1399,1582,1714]" box="[234,654,1582,1608]" pageId="7" pageNumber="8">
<emphasis id="B964D94856700C2053D6833DFE3B4E4D" bold="true" box="[234,481,1582,1608]" pageId="7" pageNumber="8">Additional material</emphasis>
(2 ♀♀, 1 juv.)
</paragraph>
<paragraph id="8BAF055A56700C2053818342FD574EB7" blockId="7.[189,1399,1582,1714]" pageId="7" pageNumber="8">
<materialsCitation id="3B780F0756700C2053818342FAE24E69" ID-GBIF-Occurrence="1806244394" box="[189,1336,1617,1644]" pageId="7" pageNumber="8">
GEORGIA: 1 ♀, Batumi, 1909, S.A. Spassky leg. (ZISP, paralectotype of
<taxonomicName id="4C107ED956700C2057048342FAFD4E6E" authorityName="Spassky" authorityYear="1937" box="[1080,1319,1617,1643]" class="Arachnida" family="Nemesiidae" genus="Brachythele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pontica">
<emphasis id="B964D94856700C2057048342FAFD4E6E" box="[1080,1319,1617,1643]" italics="true" pageId="7" pageNumber="8">Brachythele pontica</emphasis>
</taxonomicName>
);
</materialsCitation>
<materialsCitation id="3B780F0756700C20567D8342FCE54E8B" ID-GBIF-Occurrence="1806244389" pageId="7" pageNumber="8">1 ♀, same locality, no other collecting data (MNHN 10338);</materialsCitation>
<materialsCitation id="3B780F0756700C2050798367FD574EB7" ID-GBIF-Occurrence="1806244387" pageId="7" pageNumber="8">1 juv., surroundings of Batumi, Mt Feria, 250 m, 9 Feb. 1982, D.V. Logunov leg. (TAU).</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30A56D156700C2E538183CAFD174C7A" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="description">
<paragraph id="8BAF055A56700C20538183CAFE944EF6" blockId="7.[189,334,1753,1778]" box="[189,334,1753,1779]" pageId="7" pageNumber="8">
<emphasis id="B964D94856700C20538183CAFE944EF6" bold="true" box="[189,334,1753,1779]" pageId="7" pageNumber="8">Description</emphasis>
</paragraph>
<paragraph id="8BAF055A56700C2053D6821BFE6D4F40" blockId="7.[189,439,1799,1861]" pageId="7" pageNumber="8">
<emphasis id="B964D94856700C2053D6821BFEF04F27" bold="true" box="[234,298,1800,1826]" pageId="7" pageNumber="8">Male</emphasis>
(holotype) HABITUS. See
<figureCitation id="132B19DF56700C2052618238FE6D4F40" box="[349,439,1835,1861]" captionStart="Figs4351" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs4351. Raveniolaspp.,habitusof ♂♂(43,4546,48,5051)and ♀♀(44,47,49),dorsalviews.— 4344. R. adjarica sp. nov. 45. R. dunini sp. nov. 4647. R. hyrcanica Dunin, 1988. 4849. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="7" pageNumber="8">Fig. 43.</figureCitation>
</paragraph>
<paragraph id="8BAF055A56700C205381827EFB7D4F83" blockId="7.[189,1191,1900,1927]" box="[189,1191,1900,1927]" pageId="7" pageNumber="8">MEASUREMENTS. TBL 13.10, CL 5.43, CW 4.88, LL 0.47, LW 1.03, SL 2.52, SW 2.38.</paragraph>
<paragraph id="8BAF055A56700C2F538182BDFD2F49B2" blockId="7.[189,1399,1965,2027]" lastBlockId="8.[189,1399,267,439]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">COLOUR. Carapace brownish rufous with slightly lighter yellowish rufous pattern covering foveal area and protruding into few weak radial rays on darker background; eye tubercle with central and two symmetrical lateral brownish black spots surrounding AMEs and lateral eyes respectively; chelicerae and first pair of legs reddish brown; palps and legs IIIV brownish rufous; sternum, labium and maxillae light yellowish rufous; abdomen dorsally light greyish brown, with weak and darker brownish pattern represented by median lanceolate patch fused and crossed with few transverse fasciae; spinnerets and ventral surface of abdomen light yellowish grey.</paragraph>
<paragraph id="8BAF055A567F0C2F538184F3FB5B4A63" blockId="8.[189,1399,479,615]" pageId="8" pageNumber="9">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF567F0C2F503E84CCFC8649FC" box="[770,860,479,505]" captionStart="Figs 5967" captionStartId="59.[189,241,1662,1688]" captionTargetBox="[189,1398,265,1617]" captionTargetId="figure@59.[189,1398,265,1617]" captionText="Figs 5967. Raveniola spp., ♀♀ (59, 63, 66) and ♂♂ (6062, 6465, 67). Left chelicera (5961; white arrow indicates male intercheliceral organ), sternum, labium and maxillae (6264), and spinnerets (65 67); prolateral (5961,65, 67)and ventral (6264,66) views.— 5960. R.pontica (Spassky, 1937). 61, 63, 66. R. zaizevi (Charitonov, 1948). 62, 65. R. hyrcanica Dunin, 1988. 64, 67. R. vonwicki Zonstein, 2000. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163442/files/figure.png" pageId="8" pageNumber="9">Fig. 66</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF567F0C2F57EE84CCFAE049FC" box="[1234,1338,479,505]" captionStart="Figs 102116" captionStartId="66.[189,241,1755,1781]" captionTargetBox="[204,1382,265,1739]" captionTargetId="figure@66.[187,1396,250,1752]" captionText="Figs 102116. Raveniola spp., ♂♂. Eyes and clypeus, dorsal views. — 102. R. adjarica sp. nov. (holotype).103. R.duninisp.nov.(holotype).104. R.hyrcanicaDunin, 1988(holotype).105.R.pontica (Spassky, 1937) (Lazarevskoe). 106.R.birecikensis sp. nov.(holotype).107. Samespecies as preceding (dwarf conspecific male?). 108. R. anadolu sp. nov. (holotype). 109. R. arthuri Kunt &amp; Yağmur, 2010 (holotype). 110. R. turcica sp. nov. (holotype). 111. R. micropa (Ausserer, 1871) (Uludağ). 112. R. mazandaranica Marusik, Zamani &amp; Mirshamsi, 2014 (holotype). 113. R. zaitzevi (Charitonov, 1948) (Car). 114. R. marusiki sp. nov. (holotype). 115. R. vonwicki (Zonstein, 2000) (holotype). 116. R. niedermeyeri (Brignoli, 1972) (holotype). Scalebar: 0.5 mm." httpUri="https://zenodo.org/record/1163456/files/figure.png" pageId="8" pageNumber="9">Fig. 102</figureCitation>
. Eye diameters and interdistances: AME 0.15(0.20), ALE 0.23, PLE 0.15, PME 0.13, AMEAME 0.11(0.06), ALEAME 0.10(0.08), ALEPLE 0.08, PLEPME 0.04, PMEPME 0.40. Each cheliceral furrow with 10 promarginal teeth and 5 mesobasal denticles. Maxillae with 78 cuspules each.
</paragraph>
<paragraph id="8BAF055A567F0C2F5381879CFF214B3F" blockId="8.[189,1399,655,826]" pageId="8" pageNumber="9">
LEGS. Tibia and metatarsus I as in
<figureCitation id="132B19DF567F0C2F5161879CFD1F4AAC" box="[605,709,655,681]" captionStart="Figs129132" captionStartId="68.[189,241,1870,1896]" captionTargetBox="[220,1291,268,1789]" captionTargetId="figure@68.[189,1323,265,1791]" captionText="Figs129132.Raveniolaspp.,♂♂.TibiaandmetatarsusI,retrolateralviews.—129.R.adjaricasp.nov. (holotype).130.R.duninisp.nov.(holotype).131.R.hyrcanicaDunin,1988(holotype).132.R.pontica (Spassky, 1937) (Lazarevskoe). Scalebars: 1.0 mm." httpUri="https://zenodo.org/record/1163460/files/figure.png" pageId="8" pageNumber="9">Fig. 129</figureCitation>
. Scopula: in distal 1/3 of metatarsi III, entire on tarsi I II, vestigial and widely divided by setae on tarsus III, absent on tarsus IV. Trichobothria: 2 rows of 911 per row on tibiae, 1015 on metatarsi, 1012 on tarsi, 9 on cymbium. Paired claws of tarsi III: outer and inner margins with 78 teeth each; claws of tarsi IIIIV: outer and inner margins with 810 teeth each.
</paragraph>
<paragraph id="8BAF055A567F0C2F53818670FD2A4C52" blockId="8.[189,1399,867,1112]" pageId="8" pageNumber="9">SPINATION. Palp: femur d1111(2), pd1, rd1; patella p1; tibia d1, p111, r11, v222; tarsus d4. Leg I: femur d1111, pd111, rd1; tibia p11(0), pv11, rv11mm; metatarsus v003. Leg II: femur d1111, pd111; patella p1; tibia p111, v223; metatarsus p1, v123. Leg III: femur d1111, pd11(0)1, rd11(0) 1; patella p1, r1; tibia d11, p11, r111, v223; metatarsus d11, p111, r111, v233. Leg IV: femur d11111(0), pd1, rd1(0)1; patella r1; tibia d11, p11, r11, v2 23; metatarsus d11, p1111, r11(0)212, v122133. Patella I and tarsi IIV (except right tarsus III carrying 1 retrolateral spine) aspinose.</paragraph>
<paragraph id="8BAF055A567F0C2F53818193FA874CBB" blockId="8.[189,1398,1151,1214]" pageId="8" pageNumber="9">
PALP. Tibia, cymbium and palpal organ as shown in
<figureCitation id="132B19DF567F0C2F5018816CFC534C9C" box="[804,905,1151,1177]" captionStart="Figs 149152" captionStartId="73.[189,241,1930,1956]" captionTargetBox="[194,1398,266,1866]" captionTargetId="figure@73.[189,1398,265,1875]" captionText="Figs 149152. Raveniola spp., ♂♂. Palpal tibia, cymbium and palpal organ, retrolateral views. — 149. R. adjarica sp. nov. (holotype). 150. R. dunini sp. nov. (holotype). 151. R. hyrcanica Dunin, 1988 (holotype). 152. R. pontica (Spassky, 1937) (Lazarevskoe). Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/1163470/files/figure.png" pageId="8" pageNumber="9">Fig. 149</figureCitation>
. Wide and flattened basal part of embolus provided with subapical keel extends into twisted and narrow apical part of embolus (
<figureCitation id="132B19DF567F0C2F579681B7FA8B4CBB" box="[1194,1361,1188,1214]" captionStart="Figs 171179" captionStartId="78.[189,241,1846,1872]" captionTargetBox="[189,1399,265,1803]" captionTargetId="figure@78.[189,1399,265,1803]" captionText="Figs 171179. Raveniola hyrcanica species group, holotypic (171177) and conspecific (178179) ♂♂. Details of palpal organ; retroventral (171), ventral (172, 175, 177, 179), proventral (173) and retrolateral (174, 176, 178) views. — 171173. R. adjarica sp. nov. 174175. R. dunini sp. nov. 176 177. R. hyrcanica Dunin, 1988. 178179. R. pontica (Spassky, 1937). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163480/files/figure.png" pageId="8" pageNumber="9">Figs 171173</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A567F0C2F538181F5FB904D21" blockId="8.[189,1398,1254,1317]" pageId="8" pageNumber="9">SPINNERETS. PMS: length 0.75; diameter 0.28. PLS: maximal diameter 0.65; length of basal, medial and apical segments 1.22, 0.85, 1.23; total length 3.30; apical segment digitiform.</paragraph>
<paragraph id="8BAF055A567F0C2F5381805DFE354D62" blockId="8.[189,495,1357,1384]" box="[189,495,1357,1384]" pageId="8" pageNumber="9">LEG MEASUREMENTS. ♂(♀).</paragraph>
<paragraph id="8BAF055A567F0C2F5259809AFABC4E76" pageId="8" pageNumber="9">
<table id="F910F7FA567FF3D853F4809BFABC4E76" box="[200,1382,1416,1651]" colsContinueIn="22.[200,1341,1653,1889]" gridcols="7" gridrows="6" pageId="8" pageNumber="9">
<tr id="35200718567FF3D853F4809BFABC4DA6" box="[200,1382,1416,1443]" gridrow="0" pageId="8" pageNumber="9" rowspan-0="1">
<th id="76F16E64567FF3D85277809BFE164DA6" box="[331,460,1416,1443]" gridcol="1" gridrow="0" pageId="8" pageNumber="9">Femur</th>
<th id="76F16E64567FF3D8513C809BFD584DA6" box="[512,642,1416,1443]" gridcol="2" gridrow="0" pageId="8" pageNumber="9">Patella</th>
<th id="76F16E64567FF3D8518A809BFCED4DA6" box="[694,823,1416,1443]" gridcol="3" gridrow="0" pageId="8" pageNumber="9">Tibia</th>
<th id="76F16E64567FF3D85057809BFC364DA6" box="[875,1004,1416,1443]" gridcol="4" gridrow="0" pageId="8" pageNumber="9">Metatarsus</th>
<th id="76F16E64567FF3D8571C809BFB784DA6" box="[1056,1186,1416,1443]" gridcol="5" gridrow="0" pageId="8" pageNumber="9">Tarsus</th>
<th id="76F16E64567FF3D857FB809BFABC4DA6" box="[1223,1382,1416,1443]" gridcol="6" gridrow="0" pageId="8" pageNumber="9">Total</th>
</tr>
<tr id="35200718567FF3D853F480A2FABC4DC9" box="[200,1382,1457,1484]" gridrow="1" pageId="8" pageNumber="9">
<th id="76F16E64567FF3D853F480A2FEC64DC9" box="[200,284,1457,1484]" gridcol="0" gridrow="1" pageId="8" pageNumber="9">Palp</th>
<td id="76F16E64567FF3D8527780A2FE164DC9" box="[331,460,1457,1484]" gridcol="1" gridrow="1" pageId="8" pageNumber="9">2.95 (3.07)</td>
<td id="76F16E64567FF3D8513C80A2FD584DC9" box="[512,642,1457,1484]" gridcol="2" gridrow="1" pageId="8" pageNumber="9">1.67 (1.95)</td>
<td id="76F16E64567FF3D8518A80A2FCED4DC9" box="[694,823,1457,1484]" gridcol="3" gridrow="1" pageId="8" pageNumber="9">2.35 (2.20)</td>
<td id="76F16E64567FF3D8505780A2FC364DC9" box="[875,1004,1457,1484]" gridcol="4" gridrow="1" pageId="8" pageNumber="9"></td>
<td id="76F16E64567FF3D8571C80A2FB784DC9" box="[1056,1186,1457,1484]" gridcol="5" gridrow="1" pageId="8" pageNumber="9">1.02 (2.12)</td>
<td id="76F16E64567FF3D857FB80A2FABC4DC9" box="[1223,1382,1457,1484]" gridcol="6" gridrow="1" pageId="8" pageNumber="9">7.99 (9.34)</td>
</tr>
<tr id="35200718567FF3D853F480C8FABC4DF3" box="[200,1382,1499,1526]" gridrow="2" pageId="8" pageNumber="9">
<th id="76F16E64567FF3D853F480C8FEC64DF3" box="[200,284,1499,1526]" gridcol="0" gridrow="2" pageId="8" pageNumber="9">Leg I</th>
<td id="76F16E64567FF3D8527780C8FE164DF3" box="[331,460,1499,1526]" gridcol="1" gridrow="2" pageId="8" pageNumber="9">4.55 (4.07)</td>
<td id="76F16E64567FF3D8513C80C8FD584DF3" box="[512,642,1499,1526]" gridcol="2" gridrow="2" pageId="8" pageNumber="9">2.60 (2.57)</td>
<td id="76F16E64567FF3D8518A80C8FCED4DF3" box="[694,823,1499,1526]" gridcol="3" gridrow="2" pageId="8" pageNumber="9">3.57 (2.95)</td>
<td id="76F16E64567FF3D8505780C8FC364DF3" box="[875,1004,1499,1526]" gridcol="4" gridrow="2" pageId="8" pageNumber="9">3.28 (2.43)</td>
<td id="76F16E64567FF3D8571C80C8FB784DF3" box="[1056,1186,1499,1526]" gridcol="5" gridrow="2" pageId="8" pageNumber="9">2.01 (2.12)</td>
<td id="76F16E64567FF3D857FB80C8FABC4DF3" box="[1223,1382,1499,1526]" gridcol="6" gridrow="2" pageId="8" pageNumber="9">16.01 (13.77)</td>
</tr>
<tr id="35200718567FF3D853F48316FABC4E25" box="[200,1382,1541,1568]" gridrow="3" pageId="8" pageNumber="9">
<th id="76F16E64567FF3D853F48316FEC64E25" box="[200,284,1541,1568]" gridcol="0" gridrow="3" pageId="8" pageNumber="9">Leg II</th>
<td id="76F16E64567FF3D852778316FE164E25" box="[331,460,1541,1568]" gridcol="1" gridrow="3" pageId="8" pageNumber="9">4.02 (3.55)</td>
<td id="76F16E64567FF3D8513C8316FD584E25" box="[512,642,1541,1568]" gridcol="2" gridrow="3" pageId="8" pageNumber="9">2.25 (2.23)</td>
<td id="76F16E64567FF3D8518A8316FCED4E25" box="[694,823,1541,1568]" gridcol="3" gridrow="3" pageId="8" pageNumber="9">3.03 (2.72)</td>
<td id="76F16E64567FF3D850578316FC364E25" box="[875,1004,1541,1568]" gridcol="4" gridrow="3" pageId="8" pageNumber="9">2.85 (2.40)</td>
<td id="76F16E64567FF3D8571C8316FB784E25" box="[1056,1186,1541,1568]" gridcol="5" gridrow="3" pageId="8" pageNumber="9">1.78 (1.73)</td>
<td id="76F16E64567FF3D857FB8316FABC4E25" box="[1223,1382,1541,1568]" gridcol="6" gridrow="3" pageId="8" pageNumber="9">14.23 (12.63)</td>
</tr>
<tr id="35200718567FF3D853F4833CFABC4E4F" box="[200,1382,1583,1610]" gridrow="4" pageId="8" pageNumber="9">
<th id="76F16E64567FF3D853F4833CFEC64E4F" box="[200,284,1583,1610]" gridcol="0" gridrow="4" pageId="8" pageNumber="9">Leg III</th>
<td id="76F16E64567FF3D85277833CFE164E4F" box="[331,460,1583,1610]" gridcol="1" gridrow="4" pageId="8" pageNumber="9">3.17 (3.03)</td>
<td id="76F16E64567FF3D8513C833CFD584E4F" box="[512,642,1583,1610]" gridcol="2" gridrow="4" pageId="8" pageNumber="9">1.72 (1.90)</td>
<td id="76F16E64567FF3D8518A833CFCED4E4F" box="[694,823,1583,1610]" gridcol="3" gridrow="4" pageId="8" pageNumber="9">2.46 (2.17)</td>
<td id="76F16E64567FF3D85057833CFC364E4F" box="[875,1004,1583,1610]" gridcol="4" gridrow="4" pageId="8" pageNumber="9">3.22 (2.75)</td>
<td id="76F16E64567FF3D8571C833CFB784E4F" box="[1056,1186,1583,1610]" gridcol="5" gridrow="4" pageId="8" pageNumber="9">1.79 (1.73)</td>
<td id="76F16E64567FF3D857FB833CFABC4E4F" box="[1223,1382,1583,1610]" gridcol="6" gridrow="4" pageId="8" pageNumber="9">12.36 (11.58)</td>
</tr>
<tr id="35200718567FF3D853F4834BFABC4E76" box="[200,1382,1624,1651]" gridrow="5" pageId="8" pageNumber="9">
<th id="76F16E64567FF3D853F4834BFEC64E76" box="[200,284,1624,1651]" gridcol="0" gridrow="5" pageId="8" pageNumber="9">Leg IV</th>
<td id="76F16E64567FF3D85277834BFE164E76" box="[331,460,1624,1651]" gridcol="1" gridrow="5" pageId="8" pageNumber="9">4.47 (4.10)</td>
<td id="76F16E64567FF3D8513C834BFD584E76" box="[512,642,1624,1651]" gridcol="2" gridrow="5" pageId="8" pageNumber="9">2.20 (2.25)</td>
<td id="76F16E64567FF3D8518A834BFCED4E76" box="[694,823,1624,1651]" gridcol="3" gridrow="5" pageId="8" pageNumber="9">3.43 (3.23)</td>
<td id="76F16E64567FF3D85057834BFC364E76" box="[875,1004,1624,1651]" gridcol="4" gridrow="5" pageId="8" pageNumber="9">4.15 (3.88)</td>
<td id="76F16E64567FF3D8571C834BFB784E76" box="[1056,1186,1624,1651]" gridcol="5" gridrow="5" pageId="8" pageNumber="9">2.02 (2.07)</td>
<td id="76F16E64567FF3D857FB834BFABC4E76" box="[1223,1382,1624,1651]" gridcol="6" gridrow="5" pageId="8" pageNumber="9">16.27 (15.53)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAF055A567F0C2F53D6838FFD844EB2" blockId="8.[189,606,1692,1755]" box="[234,606,1692,1719]" pageId="8" pageNumber="9">
<emphasis id="B964D948567F0C2F53D6838FFE9F4EB3" bold="true" box="[234,325,1692,1718]" pageId="8" pageNumber="9">Female</emphasis>
(paratype, from Khino)
</paragraph>
<paragraph id="8BAF055A567F0C2F538183D2FE6D4EDF" blockId="8.[189,606,1692,1755]" box="[189,439,1728,1755]" pageId="8" pageNumber="9">
HABITUS. See
<figureCitation id="132B19DF567F0C2F526183D3FE6D4EDF" box="[349,439,1728,1754]" captionStart="Figs4351" captionStartId="57.[189,241,1838,1864]" captionTargetBox="[190,1398,265,1787]" captionTargetId="figure@57.[189,1398,265,1787]" captionText="Figs4351. Raveniolaspp.,habitusof ♂♂(43,4546,48,5051)and ♀♀(44,47,49),dorsalviews.— 4344. R. adjarica sp. nov. 45. R. dunini sp. nov. 4647. R. hyrcanica Dunin, 1988. 4849. R. pontica (Spassky, 1937). 50. R. birecikensis sp. nov. 51. R. anadolu sp. nov. Scale bars: 5.0 mm." httpUri="https://zenodo.org/record/1163438/files/figure.png" pageId="8" pageNumber="9">Fig. 44.</figureCitation>
</paragraph>
<paragraph id="8BAF055A567F0C2F53818210FB7D4F18" blockId="8.[189,1191,1795,1822]" box="[189,1191,1795,1822]" pageId="8" pageNumber="9">MEASUREMENTS. TBL 15.90, CL 6.17, CW 4.48, LL 0.62, LW 1.33, SL 2.85, SW 2.60.</paragraph>
<paragraph id="8BAF055A567F0C2F53818255FEA04F81" blockId="8.[189,1399,1861,1925]" pageId="8" pageNumber="9">COLOUR. As in male, but with lighter, quite indistinct radial pattern on carapace and with lighter (yellowish rufous) clypeus.</paragraph>
<paragraph id="8BAF055A567F0C2E538182BEFB444949" blockId="8.[189,1399,1964,2027]" lastBlockId="9.[189,1398,267,333]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
PROSOMA. Carapace and chelicerae as shown in
<figureCitation id="132B19DF567F0C2F503E82BFFC864FC3" box="[770,860,1964,1990]" captionStart="Figs 8489" captionStartId="63.[189,241,1892,1918]" captionTargetBox="[189,1398,266,1873]" captionTargetId="figure@63.[189,1398,265,1878]" captionText="Figs 8489. Raveniola spp., ♀♀. Carapace and chelicerae, dorsal views. — 84. R. adjarica sp. nov. (paratype). 85. R. dunini sp. nov. (paratype). 86. R. hyrcanica Dunin, 1988 (paratype). 87. R. pontica (Spassky, 1937) (Lazarevskoe). 88. R. sinani sp. nov. (holotype). 89. R. nana sp. nov. (holotype). Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1163450/files/figure.png" pageId="8" pageNumber="9">Fig. 84</figureCitation>
. Clypeus and eye group as in
<figureCitation id="132B19DF567F0C2F57EF82BFFAE04FC3" box="[1235,1338,1964,1990]" captionStart="Figs 117128" captionStartId="67.[189,241,1621,1647]" captionTargetBox="[193,1382,278,1581]" captionTargetId="figure@67.[189,1398,265,1581]" captionText="Figs 117128. Raveniola spp., ♀♀. Eyes and clypeus, dorsal views. — 117. R. adjarica sp. nov. (paratype).118. R.dunini sp. nov.(paratype).119.R.hyrcanica Dunin, 1988(paratype).120. R.pontica (Spassky, 1937) (Lazarevskoe). 121. R. sinani sp. nov. (holotype). 122. R. nana sp. nov. (holotype). 123. R. turcica sp. nov. (paratype). 124. R. micropa (Ausserer, 1871) (holotype). 125. Same species as preceding (Uludağ). 126. R. zaitzevi (Charitonov, 1948) (Lagodekhi). 127. R. vonwicki Zonstein, 2000 (paratype). 128. R. niedermeyeri (Brignoli, 1972) (paratype). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/1163458/files/figure.png" pageId="8" pageNumber="9">Fig. 117</figureCitation>
. Eye diameters and interdistances: AME 0.14(0.19), ALE 0.22, PLE 0.13, PME 0.13, AMEAME 0.14(0.08), ALEAME 0.15(0.12), ALEPLE 0.07, PLEPME 0.07, PMEPME 0.51. Each cheliceral furrow with 10 promarginal teeth and 45 mesobasal denticles. Maxillae with 57 cuspules each.
</paragraph>
<paragraph id="8BAF055A567E0C2E53818464FE604A00" blockId="9.[189,1399,375,517]" pageId="9" pageNumber="10">LEGS. Scopula: distal on metatarsi III, entire on tarsus I and on palpal tarsus, narrowly divided by setae on tarsus II, elsewhere absent. Trichobothria: 2 rows of 1114 each on tibiae, 1114 on metatarsi, 1012 on tarsi, 8 on palpal tarsus. Paired claws: outer and inner margins with 68 teeth each, palpal claw with 4 teeth on promargin.</paragraph>
<paragraph id="8BAF055A567E0C2E53818723FC9F4AE1" blockId="9.[189,1399,559,741]" pageId="9" pageNumber="10">SPINATION. Palp: femur d1+2b, pd1; tibia v223; tarsus v223. Leg I: femur d1+3b, pd1; tibia v2 22; metatarsus v223. Leg II: femur d1+3b, pd1; tibia p11, v223; metatarsus p1, v223. Leg III: femur d1+3(4)b, pd2b, rd11; patella r1; tibia p11, r11, v223; metatarsus p11, r11, v223. Leg IV: femur d1+3b, pd2b, rd1; patella r1; tibia p11, r11, v223; metatarsus p1(0)11, r111, v21(0)23. Patellae III, and palpal patella aspinose.</paragraph>
<paragraph id="8BAF055A567E0C2E5381861CFB554B2C" blockId="9.[189,1167,783,809]" box="[189,1167,783,809]" pageId="9" pageNumber="10">
SPERMATHECAE. Broad and asymmetrical, with widely spaced receptacles (
<figureCitation id="132B19DF567E0C2E5727861CFB584B2C" box="[1051,1154,783,809]" captionStart="Figs 198203" captionStartId="81.[189,241,1819,1845]" captionTargetBox="[222,1376,271,1766]" captionTargetId="figure@81.[189,1398,265,1770]" captionText="Figs 198203. Raveniola spp., ♀♀. Vulva, dorsal views. — 198. R. adjarica sp. nov. (paratype). 199.R.duninisp.nov.(paratype).200.R.hyrcanicaDunin, 1988(paratype,Avas).201. Samespeciesas preceding (Gaftoni). 202. R. pontica (Spassky, 1937) (Lazarevskoe). 203. R. sinani sp. nov. (holotype). Scale bars: 0.25 mm." httpUri="https://zenodo.org/record/1163486/files/figure.png" pageId="9" pageNumber="10">Fig. 198</figureCitation>
).
</paragraph>
<paragraph id="8BAF055A567E0C2E53818647FB794B91" blockId="9.[189,1398,851,917]" pageId="9" pageNumber="10">SPINNERETS. PMS: length 0.36; diameter 0.26. PLS: maximal diameter 0.55; length of basal, medial and apical segments 0.70, 0.37, 0.65; total length 1.72; apical segment shortly digitiform.</paragraph>
<paragraph id="8BAF055A567E0C2E538186ACFEE94BDC" blockId="9.[189,307,959,985]" box="[189,307,959,985]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E538186ACFEE94BDC" bold="true" box="[189,307,959,985]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B964D948567E0C2E538186ACFEE94BDC" bold="true" box="[189,307,959,985]" pageId="9" pageNumber="10">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567E0C2E538186E2FD174C7A" blockId="9.[189,1399,1009,1151]" pageId="9" pageNumber="10">Carapace length in males varies from 5.15 to 5.38, in females from 4.55 to 6.20. No variation in the structure of the male palpal organ has been observed. Female vulvas look surprisingly uniform and similar to the described type, even that in the conspecific female MNHN 10338 from Batumi, the known locality that is farthest from the type locality.</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567E0C2E538181B9FE8B4D46" pageId="9" pageNumber="10" type="biology_ecology">
<paragraph id="8BAF055A567E0C2E538181B9FEFA4CC1" blockId="9.[189,288,1194,1220]" box="[189,288,1194,1220]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E538181B9FEFA4CC1" bold="true" box="[189,288,1194,1220]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B964D948567E0C2E538181B9FEFA4CC1" bold="true" box="[189,288,1194,1220]" pageId="9" pageNumber="10">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567E0C2E538181CFFE8B4D46" blockId="9.[189,1399,1244,1347]" pageId="9" pageNumber="10">
The spiders collected from the Kintrishi area were found hiding in soil cavities under stones in chestnut and spruce mountain forests dominated by
<taxonomicName id="4C107ED9567E0C2E51E68010FC3E4D19" authority="Mill." authorityName="Mill." box="[730,996,1282,1309]" class="Magnoliopsida" family="Fagaceae" genus="Castanea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="sativa">
<emphasis id="B964D948567E0C2E51E68010FC464D19" box="[730,924,1283,1309]" italics="true" pageId="9" pageNumber="10">Castanea sativa</emphasis>
Mill.
</taxonomicName>
and
<taxonomicName id="4C107ED9567E0C2E57148010FB324D19" box="[1064,1256,1282,1308]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="10" phylum="Tracheophyta" rank="species" species="orientalis">
<emphasis id="B964D948567E0C2E57148010FB324D19" box="[1064,1256,1282,1308]" italics="true" pageId="9" pageNumber="10">Picea orientalis</emphasis>
</taxonomicName>
L. (Link.), respectively.
</paragraph>
</subSubSection>
<subSubSection id="C30A56D1567E0C2E5381807DFB1E4DBF" box="[189,1220,1390,1467]" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8BAF055A567E0C2E5381807DFE8C4D8D" blockId="9.[189,342,1390,1416]" box="[189,342,1390,1416]" pageId="9" pageNumber="10">
<heading id="D0E7B236567E0C2E5381807DFE8C4D8D" bold="true" box="[189,342,1390,1416]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B964D948567E0C2E5381807DFE8C4D8D" bold="true" box="[189,342,1390,1416]" pageId="9" pageNumber="10">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF055A567E0C2E538180B3FB1E4DBF" blockId="9.[189,1220,1440,1467]" box="[189,1220,1440,1467]" pageId="9" pageNumber="10">
Known from the western part of the Caucasus Minor (Georgia: Adzharia). See
<figureCitation id="132B19DF567E0C2E576780B3FB1A4DBF" box="[1115,1216,1440,1466]" captionStart="Figure 219" captionStartId="85.[189,272,1534,1560]" captionTargetBox="[189,1398,265,1471]" captionTargetId="figure@85.[189,1398,265,1471]" captionText="Figure 219. Localities of Raveniola spp. in the western part of the Caucasus: R. adjarica sp. nov. () and R. pontica (Spassky, 1937) ()." httpUri="https://zenodo.org/record/1163494/files/figure.png" pageId="9" pageNumber="10">Fig. 219</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>