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<document id="348B6FA6D236FA16509017C9E799C972" ID-CLB-Dataset="29479" ID-DOI="10.11646/zootaxa.4471.2.3" ID-GBIF-Dataset="694f85ad-7913-43ea-89b9-8eb4e20f0821" ID-ISSN="1175-5326" ID-Zenodo-Dep="1439674" ID-ZooBank="CE4C6E73-09CB-4E74-91C5-771B3696DC52" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538364447371" checkinUser="plazi" docAuthor="Smith, Andrew B. T. &amp; Evans, Arthur V." docDate="2018" docId="03AF87E48E4CFFDBFF49FDE1FD2685EF" docLanguage="en" docName="zootaxa.4471.2.3.pdf" docOrigin="Zootaxa 4471 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Apteroathlia nox Smith &amp; Evans 2018, new species" docType="treatment" docUuid="70ABBE95-B0D0-4C43-A2DF-4038525D507C" docUuidSource="ZooBank" docVersion="8" lastPageNumber="282" masterDocId="FF96FF9C8E4FFFD8FFDEFFC7FFE38255" masterDocTitle="Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America" masterLastPageNumber="308" masterPageNumber="279" pageNumber="282" updateTime="1698558211898" updateUser="ExternalLinkService">
<mods:mods id="8A2467CE522D4549991DA0941BC23D79" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6CD97AF1319D5D99B69919BA74051CDF">
<mods:title id="5B852C8F0558F1112FBD1DFD525729B2">Taxonomic review of Athliini (Coleoptera: Scarabaeidae: Melolonthinae), a new tribe of scarab beetles endemic to South America</mods:title>
</mods:titleInfo>
<mods:name id="7DA627B7BAAD96C662A3EE7EB50357DD" type="personal">
<mods:role id="BF84DF85A29B997A9E746096B4BD5827">
<mods:roleTerm id="75830D458F4B76077B19ACB510FF09E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="740A5E6D884C118B88E3306DDCF4E4A6">Smith, Andrew B. T.</mods:namePart>
</mods:name>
<mods:name id="0CA306C9EF4C7791E9827C41C819BAB3" type="personal">
<mods:role id="0C4B5573AC3D049782CFD3E7CF5A4EB0">
<mods:roleTerm id="AA09DD861D313DB143359E135C68156E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="504C3DAC135A3A335EC41EF8FA72268B">Evans, Arthur V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4EA613DD1BEC709983393F16D092CDE6">text</mods:typeOfResource>
<mods:relatedItem id="CCDE34FFB0DFEC1DC07C0E05038B689E" type="host">
<mods:titleInfo id="CB74D243B2C16CAF4A6B2749F9B7385D">
<mods:title id="990FD060BA20B73647F11D6F3D33A00A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E7793715643768649AC095F461B1CD29">
<mods:date id="960C4982BBE4F074EA24CEB8E47D298B">2018</mods:date>
<mods:detail id="5ACA8A58451CFA9142D0DC0476A01113" type="pubDate">
<mods:number id="AE78A8DEFE5566E04C9B0C5582222FA1">2018-09-05</mods:number>
</mods:detail>
<mods:detail id="C3781653E0A22A4D6E87858B04DC9B83" type="volume">
<mods:number id="DA38CD280481DB9838915D1D0C6D38F1">4471</mods:number>
</mods:detail>
<mods:detail id="F977A616968971561215236838243368" type="issue">
<mods:number id="EBB3711F347AB8CF7ADA185FA70561F9">2</mods:number>
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<mods:start id="6B8390CEF4F6D0D4C57C0B9F1D839AFD">279</mods:start>
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</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="07A0CF21CD9B2E2367F942F275E08339">journal article</mods:classification>
<mods:identifier id="F7BDD003307C07AD683859CDB840FAB3" type="CLB-Dataset">29479</mods:identifier>
<mods:identifier id="5AF7BCEA56C83D6FF354E50037B2BA60" type="DOI">10.11646/zootaxa.4471.2.3</mods:identifier>
<mods:identifier id="BA0956DA015E3653D757FEAA7338E733" type="GBIF-Dataset">694f85ad-7913-43ea-89b9-8eb4e20f0821</mods:identifier>
<mods:identifier id="127C22470605DAB61EC153AB4548C153" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8ACE0AB6A79566011D3AED42F765C861" type="Zenodo-Dep">1439674</mods:identifier>
<mods:identifier id="B4AFAB6923FB0A5C820D0258638BC324" type="ZooBank">CE4C6E73-09CB-4E74-91C5-771B3696DC52</mods:identifier>
</mods:mods>
<treatment id="03AF87E48E4CFFDBFF49FDE1FD2685EF" ID-DOI="http://doi.org/10.5281/zenodo.6485491" ID-GBIF-Taxon="148415680" ID-Zenodo-Dep="6485491" LSID="urn:lsid:zoobank.org:act:70ABBE95-B0D0-4C43-A2DF-4038525D507C" httpUri="http://treatment.plazi.org/id/03AF87E48E4CFFDBFF49FDE1FD2685EF" lastPageNumber="282" pageId="3" pageNumber="282">
<subSubSection id="C31C65798E4CFFDBFF49FDE1FED38037" pageId="3" pageNumber="282" type="nomenclature">
<paragraph id="8BB936F28E4CFFDBFF49FDE1FD248015" blockId="3.[151,1190,550,646]" box="[151,711,550,576]" pageId="3" pageNumber="282">
<heading id="D0F1819E8E4CFFDBFF49FDE1FD248015" bold="true" box="[151,711,550,576]" fontSize="11" level="1" pageId="3" pageNumber="282" reason="1">
<emphasis id="B972EAE08E4CFFDBFF49FDE1FD248015" bold="true" box="[151,711,550,576]" pageId="3" pageNumber="282">
<taxonomicName id="4C064D718E4CFFDBFF49FDE1FDC98015" ID-CoL="FYHY" authority="Smith &amp; Evans" authorityName="Smith &amp; Evans" authorityYear="2018" box="[151,554,550,576]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="nox" status="sp. nov.">
<emphasis id="B972EAE08E4CFFDBFF49FDE1FE808015" bold="true" box="[151,355,550,576]" italics="true" pageId="3" pageNumber="282">Apteroathlia nox</emphasis>
Smith &amp; Evans
</taxonomicName>
,
<taxonomicNameLabel id="A241579B8E4CFFDBFDE6FDE1FD248015" box="[568,711,550,576]" pageId="3" pageNumber="282" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB936F28E4CFFDBFF49FD8EFED38037" blockId="3.[151,1190,550,646]" box="[151,304,585,610]" pageId="3" pageNumber="282">
<figureCitation id="133D2A778E4CFFDBFF49FD8EFEE28037" box="[151,257,585,610]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 1–5. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Figs. 1–5</figureCitation>
,
<figureCitation id="133D2A778E4CFFDBFED3FD8EFECF8037" box="[269,300,585,610]" captionStart="FIGURE 66" captionStartId="21.[151,250,1961,1983]" captionTargetBox="[153,1433,499,1939]" captionTargetId="figure@21.[153,1433,499,1939]" captionTargetPageId="21" captionText="FIGURE 66. Distribution map of Apteroathlia nox Smith &amp; Evans, new species (black circles) and Apteroathlia translucida Smith &amp; Evans, new species (red circles)." httpUri="https://zenodo.org/record/1439706/files/figure.png" pageId="3" pageNumber="282">66</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF49FDAAFB4580D3" box="[151,1190,621,646]" pageId="3" pageNumber="282" type="description">
<paragraph id="8BB936F28E4CFFDBFF49FDAAFB4580D3" blockId="3.[151,1190,550,646]" box="[151,1190,621,646]" pageId="3" pageNumber="282">http://zoobank.org/urn:lsid:zoobank.org:act:70ABBE95-B0D0-4C43-A2DF-4038525D507C</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF49FD72FB0E860F" pageId="3" pageNumber="282" type="materials_examined">
<paragraph id="8BB936F28E4CFFDBFF49FD72FC59809B" blockId="3.[151,1437,693,1978]" box="[151,954,693,718]" pageId="3" pageNumber="282">
<materialsCitation id="3B6E3CAF8E4CFFDBFF49FD72FC59809B" ID-GBIF-Occurrence="1913888843" box="[151,954,693,718]" country="Argentina" location="Playas de Juancho" municipality="Madariaga" pageId="3" pageNumber="282" specimenCount="1" stateProvince="Buenos Aires" typeStatus="holotype">
<emphasis id="B972EAE08E4CFFDBFF49FD72FED6809B" bold="true" box="[151,309,693,718]" pageId="3" pageNumber="282">
<typeStatus id="54BD88508E4CFFDBFF49FD72FF31809B" box="[151,210,693,718]" pageId="3" pageNumber="282">Type</typeStatus>
locality.
</emphasis>
<location id="8ED960298E4CFFDBFEE2FD72FDEF809B" LSID="urn:lsid:plazi:treatment:03AF87E48E4CFFDBFF49FDE1FD2685EF:8ED960298E4CFFDBFEE2FD72FDEF809B" box="[316,524,693,718]" country="Argentina" municipality="Madariaga" name="Playas de Juancho" pageId="3" pageNumber="282" stateProvince="Buenos Aires">Playas de Juancho</location>
,
<collectingMunicipality id="6BDDAC888E4CFFDBFDC9FD72FD71809B" box="[535,658,693,718]" pageId="3" pageNumber="282">Madariaga</collectingMunicipality>
,
<collectingRegion id="49C2F8108E4CFFDBFD43FD71FCD4809B" box="[669,823,693,718]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">Buenos Aires</collectingRegion>
,
<collectingCountry id="F31176628E4CFFDBFC9DFD72FC55809B" box="[835,950,693,718]" name="Argentina" pageId="3" pageNumber="282">Argentina</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB936F28E4CFFDBFF19FD1EFB0E860F" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<materialsCitation id="3B6E3CAF8E4CFFDBFF19FD1EFF28816F" ID-GBIF-Occurrence="1913888836" collectionCode="CMNC, BUENOS, AIRES" collectorName="Coll. Martinez" country="Argentina" county="Do" location="Nov" municipality="Playas de Juancho" pageId="3" pageNumber="282" specimenCount="9" specimenCount-female="2" specimenCount-male="4" stateProvince="BUENOS AIRES" typeStatus="holotype">
<emphasis id="B972EAE08E4CFFDBFF19FD1EFEB580A7" bold="true" box="[199,342,729,754]" pageId="3" pageNumber="282">
<typeStatus id="54BD88508E4CFFDBFF19FD1EFEE180A7" box="[199,258,729,754]" pageId="3" pageNumber="282">Type</typeStatus>
series.
</emphasis>
One male
<typeStatus id="54BD88508E4CFFDBFE02FD1EFDA280A7" box="[476,577,729,754]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D00FD7B8E4CFFDBFD8EFD1DFD7C80A7" box="[592,671,729,754]" pageId="3" pageNumber="282" type="male">4 male</specimenCount>
<typeStatus id="54BD88508E4CFFDBFD75FD1DFCFA80A7" box="[683,793,730,754]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
, and
<specimenCount id="9D00FD7B8E4CFFDBFC85FD1DFC2380A7" box="[859,960,729,754]" pageId="3" pageNumber="282" type="female">2 female</specimenCount>
<typeStatus id="54BD88508E4CFFDBFC12FD1DFBD880A7" box="[972,1083,730,754]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
.
<typeStatus id="54BD88508E4CFFDBFB97FD1EFB5280A7" box="[1097,1201,729,754]" pageId="3" pageNumber="282" type="holotype">Holotype</typeStatus>
male (
<figureCitation id="133D2A778E4CFFDBFAD8FD1EFA9080A7" box="[1286,1395,729,754]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 1–5. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Figs. 1–5</figureCitation>
) at
<collectionCode id="ED17AE378E4CFFDBFF49FD39FF098143" box="[151,234,766,790]" pageId="3" pageNumber="282">CMNC</collectionCode>
labeled a) “
<collectingCountry id="F31176628E4CFFDBFEACFD39FDFB8143" box="[370,536,766,790]" name="Argentina" pageId="3" pageNumber="282">ARGENTINA</collectingCountry>
/
<collectingRegion id="49C2F8108E4CFFDBFDF1FD39FD1A8143" box="[559,761,766,790]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">
<collectionCode id="ED17AE378E4CFFDBFDF1FD39FD1A8143" box="[559,761,766,790]" pageId="3" pageNumber="282">BUENOS AIRES</collectionCode>
</collectingRegion>
/
<collectingCounty id="62D84E7E8E4CFFDBFCCEFD39FCD18140" box="[784,818,766,789]" pageId="3" pageNumber="282">Do</collectingCounty>
MADARIAGA /
<collectingMunicipality id="6BDDAC888E4CFFDBFBDFFD3AFB338143" box="[1025,1232,765,790]" pageId="3" pageNumber="282">Playas de Juancho</collectingMunicipality>
/
<collectorName id="26F353248E4CFFDBFB36FD3AFA6F8143" box="[1256,1420,765,790]" pageId="3" pageNumber="282">Coll. Martínez</collectorName>
/
<location id="8ED960298E4CFFDBFF49FCE5FF28816F" LSID="urn:lsid:plazi:treatment:03AF87E48E4CFFDBFF49FDE1FD2685EF:8ED960298E4CFFDBFF49FCE5FF28816F" box="[151,203,802,826]" country="Argentina" county="Do" municipality="Playas de Juancho" name="Nov" pageId="3" pageNumber="282" stateProvince="BUENOS AIRES">Nov</location>
</materialsCitation>
.
<materialsCitation id="3B6E3CAF8E4CFFDBFF11FCE5FD6081D7" ID-GBIF-Occurrence="1913888837" collectionCode="CMNC, BUENOS, AIRES" collectorName="A. Martinez &amp; Coll. Martinez" country="Argentina" county="Pdo." location="Nov" municipality="Playas de Juancho" pageId="3" pageNumber="282" specimenCount="1" stateProvince="BUENOS AIRES" typeStatus="paratype">
-941” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4CFFDBFC1CFCE5FC38816C" box="[962,987,802,825]" italics="true" pageId="3" pageNumber="282">ex</emphasis>
.
<collectorName id="26F353248E4CFFDBFC32FCE5FB8E816F" box="[1004,1133,802,826]" pageId="3" pageNumber="282">A.Martínez</collectorName>
coll.” (typeset with black border). Two male
<typeStatus id="54BD88508E4CFFDBFEA7FC81FE07810B" box="[377,484,838,862]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
at
<collectionCode id="ED17AE378E4CFFDBFDD0FC81FD83810B" box="[526,608,838,862]" pageId="3" pageNumber="282">CMNC</collectionCode>
labeled a) “
<collectingCountry id="F31176628E4CFFDBFD2FFC81FC74810B" box="[753,919,838,862]" name="Argentina" pageId="3" pageNumber="282">ARGENTINA</collectingCountry>
/
<collectingRegion id="49C2F8108E4CFFDBFC6BFC81FB62810B" box="[949,1153,838,862]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">
<collectionCode id="ED17AE378E4CFFDBFC6BFC81FB62810B" box="[949,1153,838,862]" pageId="3" pageNumber="282">BUENOS AIRES</collectionCode>
</collectingRegion>
/
<collectingCounty id="62D84E7E8E4CFFDBFB40FC82FB32810B" box="[1182,1233,837,862]" pageId="3" pageNumber="282">Pdo.</collectingCounty>
MADARIAGA /
<collectingMunicipality id="6BDDAC888E4CFFDBFF49FCAEFE8881D7" box="[151,363,873,898]" pageId="3" pageNumber="282">Playas de Juancho</collectingMunicipality>
/
<collectorName id="26F353248E4CFFDBFE55FCAEFDD381D7" box="[395,560,873,898]" pageId="3" pageNumber="282">Coll. Martínez</collectorName>
/
<location id="8ED960298E4CFFDBFD8EFCADFD6081D7" LSID="urn:lsid:plazi:treatment:03AF87E48E4CFFDBFF49FDE1FD2685EF:8ED960298E4CFFDBFD8EFCADFD6081D7" box="[592,643,874,898]" country="Argentina" county="Pdo." municipality="Playas de Juancho" name="Nov" pageId="3" pageNumber="282" stateProvince="BUENOS AIRES">Nov</location>
</materialsCitation>
.
<materialsCitation id="3B6E3CAF8E4CFFDBFD58FCADFB5B8647" ID-GBIF-Occurrence="1913888844" collectingDate="1961-08-11" collectionCode="CMNC" collectorName="A. Martinez &amp; Two &amp; S. Ventana &amp; Gallardo-leg. &amp; Coll. Martinez" country="Argentina" county="Villa Gesell" location="Ene" municipality="Horacio-leg." pageId="3" pageNumber="282" specimenCount="3" stateProvince="BUENOS AIRES" typeStatus="paratype">
-941” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4CFFDBFAA3FCADFA7681D4" box="[1405,1429,874,897]" italics="true" pageId="3" pageNumber="282">ex</emphasis>
.
<collectorName id="26F353248E4CFFDBFF49FC49FEF781F3" box="[151,276,910,934]" pageId="3" pageNumber="282">A.Martínez</collectorName>
coll.” (typeset with black border).
<collectorName id="26F353248E4CFFDBFD79FC49FD3481F0" box="[679,727,910,933]" pageId="3" pageNumber="282">Two</collectorName>
male
<typeStatus id="54BD88508E4CFFDBFCFFFC49FC6981F3" box="[801,906,910,934]" pageId="3" pageNumber="282" type="paratype">paratypes</typeStatus>
and one female
<typeStatus id="54BD88508E4CFFDBFB92FC49FB4981F3" box="[1100,1194,910,934]" pageId="3" pageNumber="282" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4CFFDBFB0DFC49FAC781F3" box="[1235,1316,910,934]" pageId="3" pageNumber="282">CMNC</collectionCode>
labeled a) “
<collectingCountry id="F31176628E4CFFDBFF7EFC75FEA7819F" box="[160,324,946,970]" name="Argentina" pageId="3" pageNumber="282">ARGENTINA</collectingCountry>
/
<collectingRegion id="49C2F8108E4CFFDBFE86FC75FDFF819F" box="[344,540,946,970]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">BUENOS AIRES</collectingRegion>
/
<collectorName id="26F353248E4CFFDBFDEEFC75FD46819C" box="[560,677,946,970]" pageId="3" pageNumber="282">S. Ventana</collectorName>
/
<collectorName id="26F353248E4CFFDBFD67FC76FCA8819F" box="[697,843,945,970]" pageId="3" pageNumber="282">Gallardo-leg.</collectorName>
/
<collectorName id="26F353248E4CFFDBFC80FC76FC1E819F" box="[862,1021,945,970]" pageId="3" pageNumber="282">Coll. Martínez</collectorName>
/
<collectingDate id="EFFCE9DA8E4CFFDBFBCFFC76FB69819F" box="[1041,1162,945,970]" pageId="3" pageNumber="282" value="1961-08-11">
<date id="FFB810328E4CFFDBFBCFFC76FB69819F" box="[1041,1162,945,970]" pageId="3" pageNumber="282" value="1961-08-11">11-VIII-61</date>
</collectingDate>
” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4CFFDBFDCCFC11FDC981B8" box="[530,554,982,1005]" italics="true" pageId="3" pageNumber="282">ex</emphasis>
.
<collectorName id="26F353248E4CFFDBFDE7FC11FD5581BB" box="[569,694,982,1006]" pageId="3" pageNumber="282">A.Martínez</collectorName>
coll.” (typeset with black border). Ono female
<typeStatus id="54BD88508E4CFFDBFB16FC11FAC581BB" box="[1224,1318,982,1006]" pageId="3" pageNumber="282" type="paratype">paratype</typeStatus>
at
<collectionCode id="ED17AE378E4CFFDBFA95FC11FA7F81BB" box="[1355,1436,982,1006]" pageId="3" pageNumber="282">CMNC</collectionCode>
labeled a) “
<collectingCountry id="F31176628E4CFFDBFECCFC3DFE568647" box="[274,437,1018,1042]" name="Argentina" pageId="3" pageNumber="282">ARGENTINA</collectingCountry>
/
<collectingRegion id="49C2F8108E4CFFDBFE14FC3DFD6D8647" box="[458,654,1018,1042]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">BUENOS AIRES</collectingRegion>
/
<collectingCounty id="62D84E7E8E4CFFDBFD7DFC3EFCC78647" box="[675,804,1017,1042]" pageId="3" pageNumber="282">Villa Gesell</collectingCounty>
/
<collectingMunicipality id="6BDDAC888E4CFFDBFCE6FC3EFC208647" box="[824,963,1017,1042]" pageId="3" pageNumber="282">Horacio-leg.</collectingMunicipality>
/
<collectorName id="26F353248E4CFFDBFC09FC3EFB958647" box="[983,1142,1017,1042]" pageId="3" pageNumber="282">Coll. Martínez</collectorName>
/
<location id="8ED960298E4CFFDBFB54FC3EFB5B8647" LSID="urn:lsid:plazi:treatment:03AF87E48E4CFFDBFF49FDE1FD2685EF:8ED960298E4CFFDBFB54FC3EFB5B8647" box="[1162,1208,1017,1042]" country="Argentina" county="Villa Gesell" municipality="Horacio-leg." name="Ene" pageId="3" pageNumber="282" stateProvince="BUENOS AIRES">Ene</location>
</materialsCitation>
.-976” (handwritten), b) “H. &amp; A. HOWDEN / COLLECTION /
<emphasis id="B972EAE08E4CFFDBFD54FBD9FD418660" box="[650,674,1054,1077]" italics="true" pageId="3" pageNumber="282">ex</emphasis>
. A.Martínez coll.” (typeset with black border). All paratypes listed above also bear a yellow paratype label and a database label with the database number and a barcode.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF19FBA2FA9684EB" pageId="3" pageNumber="282" type="description">
<paragraph id="8BB936F28E4CFFDBFF19FBA2FD648423" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19FBA2FE07862B" bold="true" box="[199,484,1125,1150]" pageId="3" pageNumber="282">
Description of
<typeStatus id="54BD88508E4CFFDBFEA3FBA2FE07862B" box="[381,484,1125,1150]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
</emphasis>
(
<figureCitation id="133D2A778E4CFFDBFE29FBA2FD85862B" box="[503,614,1125,1150]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 1–5. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Figs. 1–5</figureCitation>
). Male. Length 7.0 mm, width
<quantity id="4CFE9B178E4CFFDBFC06FBA1FBD68628" box="[984,1077,1126,1150]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="282" unit="mm" value="3.1">3.1 mm</quantity>
. Head (at base), elytra, ventral surface black; head (apex), pronotum, legs dark tan. Dorsal surface glabrous (except sparse row of setae adjacent to apex of clypeus).
<emphasis id="B972EAE08E4CFFDBFEB5FB6AFE448690" box="[363,423,1197,1221]" italics="true" pageId="3" pageNumber="282">Head</emphasis>
: clypeus densely punctate, frons moderately punctate. Labrum with apex reflexed and concave medially; underside concave medially. Clypeus with apex straight, abutted to labrum. Frontoclypeal suture visible. Eyes round, weakly protruding laterally. Antennae with 8 antennomeres; antennal club with 3 antennomeres, club approximately half the length of antennomeres 1–5 combined, club surface with some setae. Maxillae robust, triangular, basal width slightly less than width of mentum; maxillary palpus with 4 palpomeres, length approximately approximately half that of antennae. Mentum prominent, slightly longer than wide, apex straight, surface slightly convex; labial palpus reduced.
<emphasis id="B972EAE08E4CFFDBFCD2FA41FC9887C8" box="[780,891,1414,1437]" italics="true" pageId="3" pageNumber="282">Pronotum</emphasis>
: surface moderately to sparsely punctate; apical angle slightly acute, basal angle rounded. Pronotum widest medially with smooth margins.
<emphasis id="B972EAE08E4CFFDBFB4AFA6EFB398794" box="[1172,1242,1449,1473]" italics="true" pageId="3" pageNumber="282">Elytra</emphasis>
: surface sparsely punctate, striae punctate but somewhat obscured by other surface punctures.
<emphasis id="B972EAE08E4CFFDBFC2AFA09FBD487B0" box="[1012,1079,1486,1509]" italics="true" pageId="3" pageNumber="282">Wings</emphasis>
: apterous.
<emphasis id="B972EAE08E4CFFDBFB6BFA09FB1A87B0" box="[1205,1273,1486,1509]" italics="true" pageId="3" pageNumber="282">Venter</emphasis>
: metasternum with dense punctures. Pygidium flat; surface sparsely punctate with microsculpturing.
<emphasis id="B972EAE08E4CFFDBFBB9FA35FB7F845C" box="[1127,1180,1522,1545]" italics="true" pageId="3" pageNumber="282">Legs</emphasis>
: protibia with 3 apical teeth on outer margin of apical half. Claws symmetrical, each with a minute medial tooth (metatarsal claws appear simple). Protibial spur present, metatibial spurs approximately equal in length.
<emphasis id="B972EAE08E4CFFDBFBCCF9FEFB9F8404" box="[1042,1148,1593,1617]" italics="true" pageId="3" pageNumber="282">Genitalia</emphasis>
: parameres long, curving and tapering into needle-like apices (
<figureCitation id="133D2A778E4CFFDBFDEAF99AFD9A8423" box="[564,633,1629,1654]" captionStart="FIGURES 1–5" captionStartId="4.[151,264,1267,1289]" captionTargetBox="[318,1274,195,1255]" captionTargetId="figure@4.[318,1274,195,1255]" captionTargetPageId="4" captionText="FIGURES 1–5. Apteroathlia nox Smith &amp; Evans, new species, holotype. 1, dorsal habitus; 2, oblique habitus; 3, ventral habitus; 4, head and pronotum; 5, parameres." httpUri="https://zenodo.org/record/1439676/files/figure.png" pageId="3" pageNumber="282">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BB936F28E4CFFDBFF19F946FA9684EB" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19F946FEDF84CF" bold="true" box="[199,316,1665,1690]" pageId="3" pageNumber="282">Variation.</emphasis>
Length 7.0–
<quantity id="4CFE9B178E4CFFDBFE13F945FDC684CC" box="[461,549,1666,1690]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="3" pageNumber="282" unit="mm" value="9.4">9.4 mm</quantity>
. Pronotum black, dark tan, or black lateral and dark tan/brown medially. Claws all appear simple on the
<typeStatus id="54BD88508E4CFFDBFE77F961FDEA84EB" box="[425,521,1702,1726]" pageId="3" pageNumber="282" type="paratype">paratype</typeStatus>
specimens. Females with slightly more inflated, convex abdominal segments.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF19F90EFBFC8553" pageId="3" pageNumber="282" type="etymology">
<paragraph id="8BB936F28E4CFFDBFF19F90EFBFC8553" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19F90EFEAD84B7" bold="true" box="[199,334,1737,1762]" pageId="3" pageNumber="282">Etymology.</emphasis>
The Latin word
<taxonomicName id="4C064D718E4CFFDBFDCDF90DFDD884B4" authorityName="Smith &amp; Evans" authorityYear="2018" box="[531,571,1738,1761]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="nox">
<emphasis id="B972EAE08E4CFFDBFDCDF90DFDD884B4" box="[531,571,1738,1761]" italics="true" pageId="3" pageNumber="282">nox</emphasis>
</taxonomicName>
means night and is used here in reference to the night-like darkness of the dorsal and ventral surfaces. This name is an adjective in the nominative singular.
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF19F8D6FC81851B" pageId="3" pageNumber="282" type="distribution">
<paragraph id="8BB936F28E4CFFDBFF19F8D6FC81851B" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19F8D6FEBA857F" bold="true" box="[199,345,1809,1834]" pageId="3" pageNumber="282">Distribution</emphasis>
(
<figureCitation id="133D2A778E4CFFDBFEB5F8D6FE21857F" box="[363,450,1809,1834]" captionStart="FIGURE 66" captionStartId="21.[151,250,1961,1983]" captionTargetBox="[153,1433,499,1939]" captionTargetId="figure@21.[153,1433,499,1939]" captionTargetPageId="21" captionText="FIGURE 66. Distribution map of Apteroathlia nox Smith &amp; Evans, new species (black circles) and Apteroathlia translucida Smith &amp; Evans, new species (red circles)." httpUri="https://zenodo.org/record/1439706/files/figure.png" pageId="3" pageNumber="282">Fig. 66</figureCitation>
).
<collectingCountry id="F31176628E4CFFDBFE09F8D5FD98857F" box="[471,635,1810,1834]" name="Argentina" pageId="3" pageNumber="282">ARGENTINA</collectingCountry>
(7):
<collectingRegion id="49C2F8108E4CFFDBFD68F8D5FCB3857F" box="[694,848,1809,1834]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="282">Buenos Aires</collectingRegion>
(7): Playas de Juancho, Sierra de la Ventana, Villa Gesell.
<emphasis id="B972EAE08E4CFFDBFF32F8F2FE40851B" bold="true" box="[236,419,1845,1870]" pageId="3" pageNumber="282">Temporal data.</emphasis>
January (1), August (3), November (3).
</paragraph>
</subSubSection>
<subSubSection id="C31C65798E4CFFDBFF19F89EFD2685EF" pageId="3" pageNumber="282" type="discussion">
<paragraph id="8BB936F28E4CFFDBFF19F89EFD2685EF" blockId="3.[151,1437,693,1978]" pageId="3" pageNumber="282">
<emphasis id="B972EAE08E4CFFDBFF19F89EFED78527" bold="true" box="[199,308,1881,1906]" pageId="3" pageNumber="282">Remarks</emphasis>
.
<taxonomicName id="4C064D718E4CFFDBFE9CF89EFE1C8524" authority="Smith &amp; Evans, 2018" authorityName="Smith &amp; Evans" authorityYear="2018" box="[322,511,1881,1905]" class="Insecta" family="Scarabaeidae" genus="Apteroathlia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="282" phylum="Arthropoda" rank="species" species="nox" status="sp. nov.">
<emphasis id="B972EAE08E4CFFDBFE9CF89EFE1C8524" box="[322,511,1881,1905]" italics="true" pageId="3" pageNumber="282">Apteroathlia nox</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241579B8E4CFFDBFDD8F89DFD6E8527" box="[518,653,1881,1906]" pageId="3" pageNumber="282" rank="species">new species</taxonomicNameLabel>
is endemic to the southern part of
<collectingRegion id="49C2F8108E4CFFDBFBCAF89DFAF88527" box="[1044,1307,1881,1906]" country="Argentina" name="Buenos Aires" pageId="3" pageNumber="282">Buenos Aires Province</collectingRegion>
,
<collectingCountry id="F31176628E4CFFDBFAF8F89EFA7A8527" box="[1318,1433,1881,1906]" name="Argentina" pageId="3" pageNumber="282">Argentina</collectingCountry>
. The
<typeStatus id="54BD88508E4CFFDBFF14F8BAFECF85C3" box="[202,300,1917,1942]" pageId="3" pageNumber="282" type="holotype">holotype</typeStatus>
specimen has claws with a minute, medial tooth but the tooth was not seen in other specimens. This perhaps indicates that the tooth is easily worn off.
</paragraph>
</subSubSection>
</treatment>
</document>
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