931 lines
146 KiB
XML
931 lines
146 KiB
XML
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<mods:title id="11AEEF3BB147EBF9FF5495836F4DEAB2">Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<mods:title id="2E0F12A956F6DBCD2A6F462A096CDC26">Zootaxa</mods:title>
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<mods:part id="18E7E3B461495CF7B8CE4F7C0ADCECAE">
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<mods:date id="18B05C4463ACF7DB1C91FE60115E3A7A">2020</mods:date>
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<mods:number id="5D7EC700B99965CE1342DC718CA84E97">2020-07-29</mods:number>
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<treatment id="03AF3A058D76FFA6FAA9FB74A7E9FC0D" ID-DOI="http://doi.org/10.5281/zenodo.4440978" ID-GBIF-Taxon="176573416" ID-Zenodo-Dep="4440978" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D" httpUri="http://treatment.plazi.org/id/03AF3A058D76FFA6FAA9FB74A7E9FC0D" lastPageId="13" lastPageNumber="414" pageId="4" pageNumber="405">
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<subSubSection id="C31CD8988D76FFAFFAA9FB74A11CFB37" box="[151,635,1240,1267]" pageId="4" pageNumber="405" type="nomenclature">
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<paragraph id="8BB98B138D76FFAFFAA9FB74A11CFB37" blockId="4.[151,635,1240,1303]" box="[151,635,1240,1267]" pageId="4" pageNumber="405">
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<heading id="D0F13C7F8D76FFAFFAA9FB74A11CFB37" bold="true" box="[151,635,1240,1267]" fontSize="11" level="1" pageId="4" pageNumber="405" reason="1">
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<emphasis id="B97257018D76FFAFFAA9FB74A11CFB37" bold="true" box="[151,635,1240,1267]" pageId="4" pageNumber="405">
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<taxonomicName id="4C06F0908D76FFAFFAA9FB74A155FB37" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[151,562,1240,1267]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
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||
<emphasis id="B97257018D76FFAFFAA9FB74A274FB37" bold="true" box="[151,275,1240,1267]" italics="true" pageId="4" pageNumber="405">Procloeon</emphasis>
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||
(
|
||
<emphasis id="B97257018D76FFAFFB1AFB74A2D0FB37" bold="true" box="[292,439,1240,1267]" italics="true" pageId="4" pageNumber="405">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D76FFAFFBF6FB74A155FB37" bold="true" box="[456,562,1240,1267]" italics="true" pageId="4" pageNumber="405">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241EA7A8D76FFAFF807FB75A11CFB37" box="[569,635,1241,1267]" pageId="4" pageNumber="405" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD8988D76FFAFFAA9FB50A27EFAD3" box="[151,281,1276,1303]" pageId="4" pageNumber="405" type="description">
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<paragraph id="8BB98B138D76FFAFFAA9FB50A27EFAD3" blockId="4.[151,635,1240,1303]" box="[151,281,1276,1303]" pageId="4" pageNumber="405">
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(
|
||
<figureCitation id="133D97968D76FFAFFAA1FB50A277FAD3" box="[159,272,1276,1303]" captionStart-0="FIGURES 1–7" captionStart-1="FIGURES 8–16" captionStart-2="FIGURES 17–18" captionStart-3="FIGURES 19–29" captionStart-4="FIGURES 30–33" captionStart-5="FIGURES 34–35" captionStart-6="FIGURES 36–42" captionStartId-0="5.[151,264,1918,1942]" captionStartId-1="6.[151,264,1953,1977]" captionStartId-2="7.[151,264,1525,1549]" captionStartId-3="9.[151,264,1552,1576]" captionStartId-4="10.[151,264,1718,1742]" captionStartId-5="11.[151,264,1567,1591]" captionStartId-6="12.[151,264,1918,1942]" captionTargetBox-0="[154,1436,181,1894]" captionTargetBox-1="[155,1426,181,1930]" captionTargetBox-2="[151,1436,561,1502]" captionTargetBox-3="[153,1428,187,1527]" captionTargetBox-4="[151,1436,181,1694]" captionTargetBox-5="[162,1424,316,1534]" captionTargetBox-6="[151,1436,181,1894]" captionTargetId-0="figure-16@5.[151,1436,181,1894]" captionTargetId-1="figure-27@6.[155,1433,181,1930]" captionTargetId-2="figure-198@7.[151,1436,561,1502]" captionTargetId-3="figure-16@9.[151,1436,181,1529]" captionTargetId-4="figure-27@10.[151,1436,181,1694]" captionTargetId-5="figure-85@11.[151,1436,309,1544]" captionTargetId-6="figure-27@12.[151,1436,181,1894]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="9" captionTargetPageId-4="10" captionTargetPageId-5="11" captionTargetPageId-6="12" captionText-0="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." captionText-1="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." captionText-2="FIGURES 17–18. Larva of Procloeon (Oculogaster) barnardi sp. n., middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line)." captionText-3="FIGURES 19–29. Larvae of Procloeon (Oculogaster) barnardi sp. n. 19–25, tergalii I–VII; 26–27, tergalii V of other specimens; 28, tergalius VII of another specimen; 29, claw (holotype)." captionText-4="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." captionText-5="FIGURES 34–35. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 34, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 35, their subimaginal exuviae." captionText-6="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi-0="http://doi.org/10.5281/zenodo.4397863" figureDoi-1="http://doi.org/10.5281/zenodo.4397867" figureDoi-2="http://doi.org/10.5281/zenodo.4397869" figureDoi-3="http://doi.org/10.5281/zenodo.4397871" figureDoi-4="http://doi.org/10.5281/zenodo.4397875" figureDoi-5="http://doi.org/10.5281/zenodo.4397877" figureDoi-6="http://doi.org/10.5281/zenodo.4397879" httpUri-0="https://zenodo.org/record/4397863/files/figure.png" httpUri-1="https://zenodo.org/record/4397867/files/figure.png" httpUri-2="https://zenodo.org/record/4397869/files/figure.png" httpUri-3="https://zenodo.org/record/4397871/files/figure.png" httpUri-4="https://zenodo.org/record/4397875/files/figure.png" httpUri-5="https://zenodo.org/record/4397877/files/figure.png" httpUri-6="https://zenodo.org/record/4397879/files/figure.png" pageId="4" pageNumber="405">Figs 1–42</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="8BB98B138D76FFAFFAA9FAEDA023FA36" blockId="4.[151,1436,1345,1522]" pageId="4" pageNumber="405">
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||
<subSubSection id="C31CD8988D76FFAFFAA9FAEDA6FBFABC" pageId="4" pageNumber="405" type="reference_group">
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||
<treatmentCitationGroup id="AB16AC3D8D76FFAFFAA9FAEDA6FBFA9D" box="[151,1436,1345,1369]" pageId="4" pageNumber="405">
|
||
<treatmentCitation id="0AA7AD028D76FFAFFAA9FAEDA13EFA9D" author="Barnard, K. H." box="[151,601,1345,1369]" page="217" pageId="4" pageNumber="405" year="1932">
|
||
<taxonomicName id="4C06F0908D76FFAFFAA9FAEDA13EFA9D" authority=": Barnard 1932: 217" authorityName=": Barnard" authorityPageNumber="217" authorityYear="1932" box="[151,601,1345,1369]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D76FFAFFAA9FAEDA2EBFA9D" box="[151,396,1345,1369]" italics="true" pageId="4" pageNumber="405">Austrocloeon africanum</emphasis>
|
||
:
|
||
<bibRefCitation id="EF97F6E28D76FFAFFBA4FAEDA13EFA9D" author="Barnard, K. H." box="[410,601,1345,1369]" pageId="4" pageNumber="405" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932: 217</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(imago, larva);
|
||
<treatmentCitation id="0AA7AD028D76FFAFF8C3FAEDA0D8FA9D" author="Barnard, K. H." box="[765,959,1345,1369]" page="623" pageId="4" pageNumber="405" year="1940">
|
||
<bibRefCitation id="EF97F6E28D76FFAFF8C3FAEDA0D8FA9D" author="Barnard, K. H." box="[765,959,1345,1369]" pageId="4" pageNumber="405" pagination="609 - 661" refId="ref17288" refString="Barnard, K. H. (1940) Additional records, and descriptions of new species of South-African alder-flies (Megaloptera), May-flies (Ephemeroptera), caddies-flies (Trichoptera), stone-flies (Plecoptera), and dragon-flies (Odonata). Annals of the South African Museum, 32, 609 - 661." type="journal article" year="1940">Barnard 1940: 623</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA7AD028D76FFAFF9F4FAEDA701FA9D" author="Crass, R. S." box="[970,1126,1345,1369]" page="57" pageId="4" pageNumber="405" year="1947">
|
||
<bibRefCitation id="EF97F6E28D76FFAFF9F4FAEDA701FA9D" author="Crass, R. S." box="[970,1126,1345,1369]" pageId="4" pageNumber="405" pagination="37 - 100" refId="ref17425" refString="Crass, R. S. (1947) The May-flies (Ephemeroptera) of Natal and the Eastern Cape. Annals of the Natal Museum, 11 (1), 37 - 100." type="journal article" year="1947">Crass 1947: 57</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AA7AD028D76FFAFFE4FFAEDA65EFA9D" author="Harrison, A. C." box="[1137,1337,1345,1369]" page="113" pageId="4" pageNumber="405" year="1950">
|
||
<bibRefCitation id="EF97F6E28D76FFAFFE4FFAEDA65EFA9D" author="Harrison, A. C." box="[1137,1337,1345,1369]" pageId="4" pageNumber="405" pagination="112 - 115" refId="ref17771" refString="Harrison, A. C. (1950) Cape May-flies. Part VIII. The family Baetidae (continued). Piscator, 4 (16), 112 - 115." type="journal article" year="1950">Harrison 1950: 113</bibRefCitation>
|
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</treatmentCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D76FFAFFF7DFAEDA6FBFA9D" author="Harrison, A. D. & Agnew, J. D." box="[1347,1436,1345,1369]" pageId="4" pageNumber="405" pagination="273 - 291" refId="ref17804" refString="Harrison, A. D. & Agnew, J. D. (1962) The distribution of invertebrates endemic to acid streams in the Western and Southern Cape Province. Annals of the Cape Provincial Museums, 2, 273 - 291." type="journal article" year="1962">Harrison</bibRefCitation>
|
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</treatmentCitationGroup>
|
||
& Agnew 1962: 278: Table 4, 289;
|
||
<bibRefCitation id="EF97F6E28D76FFAFF817FACCA1D6FAB3" author="Chutter, F. M." box="[553,689,1375,1400]" pageId="4" pageNumber="405" pagination="1 - 65" refId="ref17366" refString="Chutter, F. M. (1963) Hydrobiological studies on the Vaal River in the Vereeniging area. Part 1. Introduction, water chemistry and studies on the fauna of habitats other than muddy bottom sediments. Hydrobiologia, 21, 1 - 65. https: // doi. org / 10.1007 / BF 00161377" type="journal article" year="1963">Chutter 1963</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D76FFAFF884FACCA0CBFAB3" author="Oliff, W. D. & King, J. L." box="[698,940,1375,1400]" pageId="4" pageNumber="405" pagination="567 - 583" refId="ref18861" refString="Oliff, W. D. & King, J. L. (1964) Hydrobiological studies of the Tugela River System. Part IV. The Mooi River. Hydrobiologia, 24 (4), 567 - 583. https: // doi. org / 10.1007 / BF 00142003" type="journal article" year="1964">Oliff & King 1964: 576</bibRefCitation>
|
||
: Table 3a (non
|
||
<taxonomicName id="4C06F0908D76FFAFFE7DFACCA6FBFABC" authority="Esben-Petersen 1913" authorityName="Esben-Petersen" authorityYear="1913" box="[1091,1436,1376,1400]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D76FFAFFE7DFACCA79DFABC" box="[1091,1274,1376,1400]" italics="true" pageId="4" pageNumber="405">Cloeon africanum</emphasis>
|
||
<bibRefCitation id="EF97F6E28D76FFAFFF3EFACCA6FBFABC" author="Esben-Petersen, P." box="[1280,1436,1376,1400]" pageId="4" pageNumber="405" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen</bibRefCitation>
|
||
</taxonomicName>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD8988D76FFAFFAF9FAD2A023FA36" pageId="4" pageNumber="405" type="description">
|
||
1913).
|
||
<taxonomicName id="4C06F0908D76FFAFFAA9FA31A12EFA71" authority=": Kimmins 1956: 866" authorityName=": Kimmins" authorityPageNumber="866" authorityYear="1956" box="[151,585,1437,1461]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D76FFAFFAA9FA31A209FA71" box="[151,366,1437,1461]" italics="true" pageId="4" pageNumber="405">Procloeon africanum</emphasis>
|
||
:
|
||
<bibRefCitation id="EF97F6E28D76FFAFFB42FA31A12EFA71" author="Kimmins, D. E." box="[380,585,1437,1461]" pageId="4" pageNumber="405" pagination="859 - 880" refId="ref17911" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880. [1955] https: // doi. org / 10.1080 / 00222935508655706" type="journal article" year="1956">Kimmins 1956: 866</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D76FFAFF86AFA31A060FA71" author="Gillies, M. T." box="[596,775,1437,1461]" pageId="4" pageNumber="405" pagination="247 - 250" refId="ref17733" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies 1997: 247</bibRefCitation>
|
||
.
|
||
<taxonomicName id="4C06F0908D76FFAFFAA9FA10A0EBFA10" authority=": Palmer, O'Keeffe, Palmer, Dunne & Radloff 1993: 448" authorityName="Palmer, O'Keeffe, Palmer, Dunne & Radloff" authorityPageNumber="448" authorityYear="1993" box="[151,908,1468,1492]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D76FFAFFAA9FA10A228FA10" box="[151,335,1468,1492]" italics="true" pageId="4" pageNumber="405">Cloeon africanum</emphasis>
|
||
:
|
||
<bibRefCitation id="EF97F6E28D76FFAFFB62FA10A0EBFA10" author="Palmer, C. & O'Keeffe, J. & Palmer, A. & Dunne, T. & Radloff, S." box="[348,908,1468,1492]" pageId="4" pageNumber="405" pagination="441 - 453" refId="ref18914" refString="Palmer, C., O'Keeffe, J., Palmer, A., Dunne, T. & Radloff, S. (1993) Macroinvertebrate functional feeding groups in the middle and lower reaches of the Buffalo River, Eastern Cape, South Africa. I. Dietary variability. Freshwater Biology, 29, 441 - 453. https: // doi. org / 10.1111 / j. 1365 - 2427.1993. tb 00778. x" type="journal article" year="1993">Palmer, O’Keeffe, Palmer, Dunne & Radloff 1993: 448</bibRefCitation>
|
||
</taxonomicName>
|
||
(non
|
||
<bibRefCitation id="EF97F6E28D76FFAFF9F9FA10A7C7FA10" author="Esben-Petersen, P." box="[967,1184,1468,1492]" pageId="4" pageNumber="405" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C06F0908D76FFAFFAA9FA76A14BFA36" authorityName=": Kluge" authorityYear="2016" box="[151,556,1498,1522]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D76FFAFFAA9FA76A267FA36" box="[151,256,1498,1522]" italics="true" pageId="4" pageNumber="405">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D76FFAFFB31FA76A2EBFA36" box="[271,396,1498,1522]" italics="true" pageId="4" pageNumber="405">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D76FFAFFBA5FA76A14BFA36" box="[411,556,1498,1522]" italics="true" pageId="4" pageNumber="405">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
(partim):
|
||
<bibRefCitation id="EF97F6E28D76FFAFF8ADFA76A027FA36" author="Kluge, N. J." box="[659,832,1498,1522]" pageId="4" pageNumber="405" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016: 495</bibRefCitation>
|
||
.
|
||
</subSubSection>
|
||
</paragraph>
|
||
<subSubSection id="C31CD8988D76FFAFFAA9F98DA26DF99B" pageId="4" pageNumber="405" type="etymology">
|
||
<paragraph id="8BB98B138D76FFAFFAA9F98DA26DF99B" blockId="4.[151,1437,1568,1993]" pageId="4" pageNumber="405">
|
||
<emphasis id="B97257018D76FFAFFAA9F98DA279F9FF" bold="true" box="[151,286,1569,1595]" pageId="4" pageNumber="405">Etymology.</emphasis>
|
||
The species is named in honour of K.H. Barnard, who described it under the name «
|
||
<taxonomicName id="4C06F0908D76FFAFFEDFF98CA393F99A" authorityName=": Barnard" authorityYear="1932" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="405" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D76FFAFFEDFF98CA393F99A" italics="true" pageId="4" pageNumber="405">Austrocloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
».
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD8988D76FFA0FAF9F9C4A127FED9" lastPageId="11" lastPageNumber="412" pageId="4" pageNumber="405" type="materials_examined">
|
||
<paragraph id="8BB98B138D76FFAFFAF9F9C4A616F802" blockId="4.[151,1437,1568,1993]" pageId="4" pageNumber="405">
|
||
<emphasis id="B97257018D76FFAFFAF9F9C4A2D6F946" bold="true" box="[199,433,1640,1666]" pageId="4" pageNumber="405">Material examined.</emphasis>
|
||
<materialsCitation id="3B6E814E8D76FFAFFB86F9C5A0ADF90E" ID-GBIF-Occurrence="3014991315" collectingDate="2019-01-22" collectionCode="AM, SE" collectorName="N. Kluge & L. Sheyko." country="South Africa" county="Winelands District" latitude="-33.958336" location="Jonkershoek Valley" longLatPrecision="767" longitude="18.916666" municipality="Boland District" pageId="4" pageNumber="405" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape" typeStatus="holotype">
|
||
<emphasis id="B97257018D76FFAFFB86F9C5A142F947" bold="true" box="[440,549,1641,1667]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54BD35B18D76FFAFFB86F9C5A142F947" box="[440,549,1641,1667]" pageId="4" pageNumber="405" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED1713D68D76FFAFF80AF9C5A100F947" box="[564,615,1641,1667]" country="Australia" name="Australian Museum" pageId="4" pageNumber="405" type="Museum">AM</collectionCode>
|
||
): L-S-I
|
||
<specimenCount id="9D00409A8D76FFAFF884F9C5A1AAF947" box="[698,717,1641,1667]" pageId="4" pageNumber="405" type="male">♂</specimenCount>
|
||
{specimen [III](10)2019},
|
||
<collectingCountry id="F311CB838D76FFAFFE3DF9C5A7B0F947" box="[1027,1239,1641,1667]" name="South Africa" pageId="4" pageNumber="405">SOUTH AFRICA</collectingCountry>
|
||
,
|
||
<collectingRegion id="49C245F18D76FFAFFEDBF9C5A678F947" box="[1253,1311,1641,1667]" country="South Africa" name="Western Cape" pageId="4" pageNumber="405">Cape</collectingRegion>
|
||
<collectingCounty id="62D8F39F8D76FFAFFF18F9C4A38CF962" pageId="4" pageNumber="405">Winelands District</collectingCounty>
|
||
(=
|
||
<collectingMunicipality id="6BDD11698D76FFAFFB28F920A2AFF963" box="[278,456,1676,1703]" pageId="4" pageNumber="405">Boland District</collectingMunicipality>
|
||
),
|
||
<location id="8ED9DDC88D76FFAFFBE2F920A1DEF962" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFBE2F920A1DEF962" box="[476,697,1676,1702]" country="South Africa" county="Winelands District" latitude="-33.958336" longLatPrecision="767" longitude="18.916666" municipality="Boland District" name="Jonkershoek Valley" pageId="4" pageNumber="405" stateProvince="Western Cape">Jonkershoek Valley</location>
|
||
,
|
||
<location id="8ED9DDC88D76FFAFF8FBF921A035F962" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF8FBF921A035F962" box="[709,850,1676,1703]" country="South Africa" county="Winelands District" latitude="-33.958336" longLatPrecision="767" longitude="18.916666" municipality="Boland District" name="Eerste River" pageId="4" pageNumber="405" stateProvince="Western Cape">Eerste River</location>
|
||
upstream
|
||
<location id="8ED9DDC88D76FFAFF9F4F920A73BF962" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF9F4F920A73BF962" box="[970,1116,1676,1702]" country="South Africa" county="Winelands District" latitude="-33.958336" longLatPrecision="767" longitude="18.916666" municipality="Boland District" name="Stellenbosch" pageId="4" pageNumber="405" stateProvince="Western Cape">Stellenbosch</location>
|
||
,
|
||
<location id="8ED9DDC88D76FFAFFE57F920A78EF962" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFE57F920A78EF962" box="[1129,1257,1676,1702]" country="South Africa" county="Winelands District" latitude="-33.958336" longLatPrecision="767" longitude="18.916666" municipality="Boland District" name="Swiss Club" pageId="4" pageNumber="405" stateProvince="Western Cape">Swiss Club</location>
|
||
(
|
||
<quantity id="4CFE26F68D76FFAFFEC5F920A651F962" box="[1275,1334,1676,1703]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="405" unit="km" value="5.0">5 km</quantity>
|
||
<collectionCode id="ED1713D68D76FFAFFF00F921A638F963" box="[1342,1375,1677,1703]" pageId="4" pageNumber="405">SE</collectionCode>
|
||
<location id="8ED9DDC88D76FFAFFF59F920A398F90F" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFF59F920A398F90F" country="South Africa" county="Winelands District" latitude="-33.958336" longLatPrecision="767" longitude="18.916666" municipality="Boland District" name="Stellenbosch" pageId="4" pageNumber="405" stateProvince="Western Cape">Stel-lenbosch</location>
|
||
),
|
||
<geoCoordinate id="EE32EDD48D76FFAFFB2DF91CA2F1F90F" box="[275,406,1712,1741]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="4" pageNumber="405" precision="15" seconds="30" value="-33.958336">33°57′30′′S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE32EDD48D76FFAFFB9DF91CA167F90F" box="[419,512,1712,1741]" degrees="18" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="405" precision="925" value="18.916666">18°55′E</geoCoordinate>
|
||
,
|
||
<date id="FFB8ADD38D76FFAFF833F91DA11DF90F" box="[525,634,1713,1739]" pageId="4" pageNumber="405" value="2019-01-22">
|
||
<collectingDate id="EFFC543B8D76FFAFF833F91DA11DF90F" box="[525,634,1713,1739]" pageId="4" pageNumber="405" value="2019-01-22">22.I.2019</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26F3EEC58D76FFAFF8FEF91DA04FF90E" box="[704,808,1712,1739]" pageId="4" pageNumber="405">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26F3EEC58D76FFAFF970F91DA0ADF90E" box="[846,970,1712,1739]" pageId="4" pageNumber="405">L. Sheyko.</collectorName>
|
||
</materialsCitation>
|
||
<emphasis id="B97257018D76FFAFF9EFF91DA72EF90F" bold="true" box="[977,1097,1713,1739]" pageId="4" pageNumber="405">
|
||
<typeStatus id="54BD35B18D76FFAFF9EFF91DA72EF90F" box="[977,1097,1713,1739]" pageId="4" pageNumber="405" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<collectionCode id="ED1713D68D76FFAFFE64F91DA7E8F90F" box="[1114,1167,1713,1739]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="405" type="Museum">ZIN</collectionCode>
|
||
): same locality and col-lectors,
|
||
<date id="FFB8ADD38D76FFAFFACDF979A2EBF92B" box="[243,396,1749,1775]" pageId="4" pageNumber="405" value="2019-01-18" valueMax="2019-01-19" valueMin="2019-01-18">
|
||
<collectingDate id="EFFC543B8D76FFAFFACDF979A2EBF92B" box="[243,396,1749,1775]" pageId="4" pageNumber="405" value="2019-01-18" valueMax="2019-01-19" valueMin="2019-01-18">18–19.I.2019</collectingDate>
|
||
</date>
|
||
: 1 L/S
|
||
<specimenCount id="9D00409A8D76FFAFFBE3F979A296F92B" box="[477,497,1749,1775]" pageId="4" pageNumber="405" type="male">♂</specimenCount>
|
||
, 2L/S
|
||
<specimenCount id="9D00409A8D76FFAFF800F979A12AF92B" box="[574,589,1749,1775]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D00409A8D76FFAFF862F979A1DCF92A" box="[604,699,1748,1775]" pageId="4" pageNumber="405" type="larva">3 larvae</specimenCount>
|
||
;
|
||
<date id="FFB8ADD38D76FFAFF8F7F979A03AF92B" box="[713,861,1749,1775]" pageId="4" pageNumber="405" value="2019-02-08" valueMax="2019-02-11" valueMin="2019-02-08">
|
||
<collectingDate id="EFFC543B8D76FFAFF8F7F979A03AF92B" box="[713,861,1749,1775]" pageId="4" pageNumber="405" value="2019-02-08" valueMax="2019-02-11" valueMin="2019-02-08">8–11.II.2019</collectingDate>
|
||
</date>
|
||
: 2 L/S
|
||
<specimenCount id="9D00409A8D76FFAFF990F979A0A5F92B" box="[942,962,1749,1775]" pageId="4" pageNumber="405" type="male">♂</specimenCount>
|
||
, 2 L-S
|
||
<specimenCount id="9D00409A8D76FFAFFE24F979A74EF92B" box="[1050,1065,1749,1775]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D00409A8D76FFAFFE06F978A7FFF92A" box="[1080,1176,1748,1775]" pageId="4" pageNumber="405" type="larva">5 larvae</specimenCount>
|
||
;
|
||
<location id="8ED9DDC88D76FFAFFE9BF978A61EF92A" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFE9BF978A61EF92A" box="[1189,1401,1748,1774]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Winelands District" pageId="4" pageNumber="405">Winelands District</location>
|
||
(=
|
||
<location id="8ED9DDC88D76FFAFFAA9F954A220F8D7" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFAA9F954A220F8D7" box="[151,327,1784,1811]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Boland District" pageId="4" pageNumber="405">Boland District</location>
|
||
),
|
||
<location id="8ED9DDC88D76FFAFFB67F954A140F8D6" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFB67F954A140F8D6" box="[345,551,1784,1810]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Wolwekloof River" pageId="4" pageNumber="405">Wolwekloof River</location>
|
||
near fall to
|
||
<location id="8ED9DDC88D76FFAFF892F954A04AF8D6" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF892F954A04AF8D6" box="[684,813,1784,1810]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Witte River" pageId="4" pageNumber="405">Witte River</location>
|
||
(
|
||
<location id="8ED9DDC88D76FFAFF902F954A0B6F8D7" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF902F954A0B6F8D7" box="[828,977,1784,1811]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Bain's Kloof" pageId="4" pageNumber="405">Bain’s Kloof</location>
|
||
),
|
||
<location id="8ED9DDC88D76FFAFF9DDF954A703F8D6" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF9DDF954A703F8D6" box="[995,1124,1784,1810]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Tweede Tol" pageId="4" pageNumber="405">Tweede Tol</location>
|
||
campsite,
|
||
<geoCoordinate id="EE32EDD48D76FFAFFEE5F955A652F8D7" box="[1243,1333,1785,1813]" degrees="33" direction="south" minutes="34" orientation="latitude" pageId="4" pageNumber="405" precision="925" value="-33.566666">33°34′S</geoCoordinate>
|
||
<geoCoordinate id="EE32EDD48D76FFAFFF05F955A6F0F8D7" box="[1339,1431,1785,1813]" degrees="19" direction="east" minutes="08" orientation="longitude" pageId="4" pageNumber="405" precision="925" value="19.133333">19°08′E</geoCoordinate>
|
||
,
|
||
<date id="FFB8ADD38D76FFAFFAA9F8B1A257F8F3" box="[151,304,1821,1847]" pageId="4" pageNumber="405" value="2019-01-20" valueMax="2019-01-22" valueMin="2019-01-20">
|
||
<collectingDate id="EFFC543B8D76FFAFFAA9F8B1A257F8F3" box="[151,304,1821,1847]" pageId="4" pageNumber="405" value="2019-01-20" valueMax="2019-01-22" valueMin="2019-01-20">20–22.I.2019</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26F3EEC58D76FFAFFB4CF8B1A2BFF8F2" box="[370,472,1820,1847]" pageId="4" pageNumber="405">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26F3EEC58D76FFAFFBC2F8B1A114F8F2" box="[508,627,1820,1847]" pageId="4" pageNumber="405">L. Sheyko</collectorName>
|
||
: 1 L/S
|
||
<specimenCount id="9D00409A8D76FFAFF8FDF8B1A1B4F8F3" box="[707,723,1821,1847]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D00409A8D76FFAFF8E0F8B1A049F8F2" box="[734,814,1820,1847]" pageId="4" pageNumber="405" type="larva">1 larva</specimenCount>
|
||
;
|
||
<location id="8ED9DDC88D76FFAFF904F8B0A766F8F2" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF904F8B0A766F8F2" box="[826,1025,1820,1846]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Overberg District" pageId="4" pageNumber="405">Overberg District</location>
|
||
, Wolfkloof,
|
||
<location id="8ED9DDC88D76FFAFFEB2F8B0A634F8F2" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFEB2F8B0A634F8F2" box="[1164,1363,1820,1846]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Keurbooms River" pageId="4" pageNumber="405">Keurbooms River</location>
|
||
(
|
||
<quantity id="4CFE26F68D76FFAFFF5DF8B1A6FBF8F2" box="[1379,1436,1820,1847]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="405" unit="km" value="8.0">8 km</quantity>
|
||
<collectionCode id="ED1713D68D76FFAFFAA9F8EDA387F89F" box="[151,224,1857,1883]" pageId="4" pageNumber="405">WNW</collectionCode>
|
||
<location id="8ED9DDC88D76FFAFFAD8F8ECA212F89F" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFAD8F8ECA212F89F" box="[230,373,1856,1883]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Swellendam" pageId="4" pageNumber="405">Swellendam</location>
|
||
),
|
||
<geoCoordinate id="EE32EDD48D76FFAFFBB8F8ECA2B8F89F" box="[390,479,1856,1885]" degrees="33" direction="south" minutes="59" orientation="latitude" pageId="4" pageNumber="405" precision="925" value="-33.983334">33°59′S</geoCoordinate>
|
||
<geoCoordinate id="EE32EDD48D76FFAFFBDBF8EDA10DF89F" box="[485,618,1857,1885]" degrees="20" direction="east" minutes="22" orientation="longitude" pageId="4" pageNumber="405" precision="15" seconds="30" value="20.375">20°22′30”E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26F3EEC58D76FFAFF895F8EDA077F89E" box="[683,784,1856,1883]" pageId="4" pageNumber="405">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26F3EEC58D76FFAFF90CF8EDA0CFF89E" box="[818,936,1856,1883]" pageId="4" pageNumber="405">L. Sheyko</collectorName>
|
||
:
|
||
<date id="FFB8ADD38D76FFAFF98FF8ECA72CF89F" box="[945,1099,1856,1883]" pageId="4" pageNumber="405" value="2019-01-25" valueMax="2019-01-26" valueMin="2019-01-25">
|
||
<collectingDate id="EFFC543B8D76FFAFF98FF8ECA72CF89F" box="[945,1099,1856,1883]" pageId="4" pageNumber="405" value="2019-01-25" valueMax="2019-01-26" valueMin="2019-01-25">25–26.I.2019</collectingDate>
|
||
</date>
|
||
:
|
||
<specimenCount id="9D00409A8D76FFAFFE6AF8EDA7C4F89E" box="[1108,1187,1856,1883]" pageId="4" pageNumber="405" type="larva">1 larva</specimenCount>
|
||
;
|
||
<location id="8ED9DDC88D76FFAFFE90F8ECA6FBF89E" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFE90F8ECA6FBF89E" box="[1198,1436,1856,1883]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Garden Route district" pageId="4" pageNumber="405">Garden Route district</location>
|
||
(=
|
||
<location id="8ED9DDC88D76FFAFFA88F8C8A22DF8BB" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFA88F8C8A22DF8BB" box="[182,330,1892,1919]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Eden district" pageId="4" pageNumber="405">Eden district</location>
|
||
):
|
||
<location id="8ED9DDC88D76FFAFFB65F8C9A170F8BA" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFFB65F8C9A170F8BA" box="[347,535,1892,1919]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Cango Mountain" pageId="4" pageNumber="405">Cango Mountain</location>
|
||
resort (
|
||
<quantity id="4CFE26F68D76FFAFF855F8C9A1D5F8BA" box="[619,690,1892,1919]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="405" unit="km" value="20.0">20 km</quantity>
|
||
<collectionCode id="ED1713D68D76FFAFF886F8C9A195F8BB" box="[696,754,1893,1919]" pageId="4" pageNumber="405">NNE</collectionCode>
|
||
<location id="8ED9DDC88D76FFAFF8C6F8C8A01EF8BB" LSID="urn:lsid:plazi:treatment:03AF3A058D76FFA6FAA9FB74A7E9FC0D:8ED9DDC88D76FFAFF8C6F8C8A01EF8BB" box="[760,889,1892,1919]" county="Nature's Valley" latitude="-33.95" longLatPrecision="925" longitude="23.525" municipality="Bobbejaans River" name="Oudshoorn" pageId="4" pageNumber="405">Oudshoorn</location>
|
||
),
|
||
<geoCoordinate id="EE32EDD48D76FFAFF9B5F8C8A083F8BB" box="[907,996,1892,1921]" degrees="33" direction="south" minutes="25" orientation="latitude" pageId="4" pageNumber="405" precision="925" value="-33.416668">33°25′S</geoCoordinate>
|
||
<geoCoordinate id="EE32EDD48D76FFAFF9D4F8C8A721F8BB" box="[1002,1094,1892,1921]" degrees="22" direction="east" minutes="16" orientation="longitude" pageId="4" pageNumber="405" precision="925" value="22.266666">22°16′E</geoCoordinate>
|
||
,
|
||
<date id="FFB8ADD38D76FFAFFE6FF8C9A793F8BB" box="[1105,1268,1893,1919]" pageId="4" pageNumber="405" value="2019-01-28" valueMax="2019-02-02" valueMin="2019-01-28">
|
||
<collectingDate id="EFFC543B8D76FFAFFE6FF8C9A793F8BB" box="[1105,1268,1893,1919]" pageId="4" pageNumber="405" value="2019-01-28" valueMax="2019-02-02" valueMin="2019-01-28">28.I–2.II.2019</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26F3EEC58D76FFAFFF08F8C9A6FBF8BA" box="[1334,1436,1892,1919]" pageId="4" pageNumber="405">N. Kluge</collectorName>
|
||
&
|
||
<collectorName id="26F3EEC58D76FFAFFA8BF825A24AF866" box="[181,301,1928,1955]" pageId="4" pageNumber="405">L. Sheyko</collectorName>
|
||
: 1 L/S
|
||
<specimenCount id="9D00409A8D76FFAFFB44F825A2E9F867" box="[378,398,1929,1955]" pageId="4" pageNumber="405" type="male">♂</specimenCount>
|
||
, 1 L-S-I
|
||
<specimenCount id="9D00409A8D76FFAFFBC9F825A162F867" box="[503,517,1929,1955]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
, 1 L-S
|
||
<specimenCount id="9D00409A8D76FFAFF864F825A10EF867" box="[602,617,1929,1955]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
; lodge «Wild Spirit», tributary of
|
||
<collectingMunicipality id="6BDD11698D76FFAFF9D2F824A7D3F866" box="[1004,1204,1928,1954]" pageId="4" pageNumber="405">Bobbejaans River</collectingMunicipality>
|
||
(
|
||
<quantity id="4CFE26F68D76FFAFFEFBF824A667F866" box="[1221,1280,1928,1955]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="405" unit="km" value="5.0">5 km</quantity>
|
||
<collectionCode id="ED1713D68D76FFAFFF39F825A652F867" box="[1287,1333,1929,1955]" pageId="4" pageNumber="405">NW</collectionCode>
|
||
<collectingCounty id="62D8F39F8D76FFAFFF03F825A384F803" pageId="4" pageNumber="405">Nature’s Valley</collectingCounty>
|
||
),
|
||
<geoCoordinate id="EE32EDD48D76FFAFFAC8F800A228F803" box="[246,335,1964,1993]" degrees="33" direction="south" minutes="57" orientation="latitude" pageId="4" pageNumber="405" precision="925" value="-33.95">33°57′S</geoCoordinate>
|
||
<geoCoordinate id="EE32EDD48D76FFAFFB68F801A2BCF803" box="[342,475,1965,1993]" degrees="23" direction="east" minutes="31" orientation="longitude" pageId="4" pageNumber="405" precision="15" seconds="30" value="23.525">23°31′30”E</geoCoordinate>
|
||
, 3rd pool from waterfall,
|
||
<date id="FFB8ADD38D76FFAFF8C5F801A0E7F803" box="[763,896,1965,1991]" pageId="4" pageNumber="405" value="2019-02-03" valueMax="2019-02-07" valueMin="2019-02-03">
|
||
<collectingDate id="EFFC543B8D76FFAFF8C5F801A0E7F803" box="[763,896,1965,1991]" pageId="4" pageNumber="405" value="2019-02-03" valueMax="2019-02-07" valueMin="2019-02-03">3–7.II.2019</collectingDate>
|
||
</date>
|
||
, coll.
|
||
<collectorName id="26F3EEC58D76FFAFF9FDF801A748F802" box="[963,1071,1964,1991]" pageId="4" pageNumber="405">N. Kluge</collectorName>
|
||
: 3 L-S-I
|
||
<specimenCount id="9D00409A8D76FFAFFEAAF801A7C5F803" box="[1172,1186,1965,1991]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
, 1 L-S
|
||
<specimenCount id="9D00409A8D76FFAFFECAF801A664F803" box="[1268,1283,1965,1991]" pageId="4" pageNumber="405" type="female">♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D00409A8D76FFAFFF2EF801A60AF802" box="[1296,1389,1964,1991]" pageId="4" pageNumber="405" type="larva">3 larvae</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF79DB9B8D77FFAEFAA9F8D2A6F1F81B" ID-DOI="http://doi.org/10.5281/zenodo.4397863" ID-Zenodo-Dep="4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="5" pageNumber="406" startId="5.[151,264,1918,1942]" targetBox="[154,1436,181,1894]" targetPageId="5">
|
||
<paragraph id="8BB98B138D77FFAEFAA9F8D2A6F1F81B" blockId="5.[151,1436,1918,2015]" pageId="5" pageNumber="406">
|
||
<emphasis id="B97257018D77FFAEFAA9F8D2A25BF852" bold="true" box="[151,316,1918,1942]" pageId="5" pageNumber="406">FIGURES 1–7.</emphasis>
|
||
<taxonomicName id="4C06F0908D77FFAEFB7DF8D3A1C5F853" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[323,674,1919,1943]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D77FFAEFB7DF8D3A2CBF853" box="[323,428,1919,1943]" italics="true" pageId="5" pageNumber="406">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D77FFAEFB82F8D3A15EF853" box="[444,569,1919,1943]" italics="true" pageId="5" pageNumber="406">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D77FFAEF877F8D3A1C5F853" box="[585,674,1919,1943]" italics="true" pageId="5" pageNumber="406">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D77FFAEF894F8D3A183F853" bold="true" box="[682,740,1919,1943]" pageId="5" pageNumber="406">
|
||
<taxonomicNameLabel id="A241EA7A8D77FFAEF894F8D3A183F853" box="[682,740,1919,1943]" pageId="5" pageNumber="406" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF79DB9B8D74FFADFAA9F80DA013F819" ID-DOI="http://doi.org/10.5281/zenodo.4397867" ID-Zenodo-Dep="4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="6" pageNumber="407" startId="6.[151,264,1953,1977]" targetBox="[155,1426,181,1930]" targetPageId="6">
|
||
<paragraph id="8BB98B138D74FFADFAA9F80DA013F819" blockId="6.[151,1437,1953,2014]" pageId="6" pageNumber="407">
|
||
<emphasis id="B97257018D74FFADFAA9F80DA22FF87D" bold="true" box="[151,328,1953,1977]" pageId="6" pageNumber="407">FIGURES 8–16.</emphasis>
|
||
Larvae of
|
||
<taxonomicName id="4C06F0908D74FFADFB89F80DA072F87D" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[439,789,1953,1978]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D74FFADFB89F80DA147F87D" box="[439,544,1953,1977]" italics="true" pageId="6" pageNumber="407">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D74FFADF811F80DA1CBF87D" box="[559,684,1953,1977]" italics="true" pageId="6" pageNumber="407">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D74FFADF882F80DA072F87D" box="[700,789,1953,1977]" italics="true" pageId="6" pageNumber="407">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D74FFADF925F80EA032F87E" bold="true" box="[795,853,1954,1978]" pageId="6" pageNumber="407">
|
||
<taxonomicNameLabel id="A241EA7A8D74FFADF925F80EA032F87E" box="[795,853,1954,1978]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB98B138D75FFACFAF9FF3AA172FDDD" blockId="7.[151,1437,150,537]" pageId="7" pageNumber="408">
|
||
<emphasis id="B97257018D75FFACFAF9FF3AA2D0FF75" bold="true" box="[199,439,150,177]" pageId="7" pageNumber="408">
|
||
Descriptions.
|
||
<emphasis id="B97257018D75FFACFB52FF3BA2D7FF75" bold="true" box="[364,432,151,177]" italics="true" pageId="7" pageNumber="408">Larva</emphasis>
|
||
.
|
||
</emphasis>
|
||
CUTICULAR COLORATION. Head with brownish and colorless areas. Pronotum and mesonotum ocher or brownish with lighter and darker areas (
|
||
<figureCitation id="133D97968D75FFACF965FF17A0C2FF11" box="[859,933,187,213]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="7" pageNumber="408">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="133D97968D75FFACF98BFF17A73EFF12" box="[949,1113,186,214]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="7" pageNumber="408">11, 13, 15, 16</figureCitation>
|
||
); mesonotum with more-orless large blanks approximately corresponding to blanks on submedioscutum and posterior scutal protuberance of subimaginal cuticle (
|
||
<figureCitation id="133D97968D75FFACFBBBFEAFA2AEFED9" box="[389,457,259,285]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="7" pageNumber="408">Fig. 1</figureCitation>
|
||
); cuticle of fore protoptera either nearly unicolor, or with darker brown stripes along convex veins and stripes lighter than background along concave veins (
|
||
<figureCitation id="133D97968D75FFACF9F5FE8BA746FE85" box="[971,1057,295,321]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="7" pageNumber="408">Fig. 13</figureCitation>
|
||
). Metanotum and thoracic pleura with colorless and brownish areas, thoracic sterna colorless. Legs colorless or light ocher, with diffusive brown bands in distal part of femur, in proximal part of tibia and in proximal part of tarsus. Abdominal terga with variable maculation, different on different segments; most or all terga with submedian pair of blanks, either corresponding to sigilla, or larger than sigilla; terga IV and VII more-or-less lighter than others (
|
||
<figureCitation id="133D97968D75FFACFE23FE1BA702FE15" box="[1053,1125,439,465]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="7" pageNumber="408">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="133D97968D75FFACFE4FFE1BA66BFE16" box="[1137,1292,439,466]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="7" pageNumber="408">10, 12, 14, 16</figureCitation>
|
||
). Abdominal sterna lighter than terga, either unicolor, or with lighter and darker areas. Caudalii ocher with posterior margin of every 4th segment brown (
|
||
<figureCitation id="133D97968D75FFACFBFCFE53A160FDDD" box="[450,519,511,537]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="7" pageNumber="408">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF79DB9B8D75FFACFAA9FA59A0B8F991" ID-DOI="http://doi.org/10.5281/zenodo.4397869" ID-Zenodo-Dep="4397869" httpUri="https://zenodo.org/record/4397869/files/figure.png" pageId="7" pageNumber="408" startId="7.[151,264,1525,1549]" targetBox="[151,1436,561,1502]" targetPageId="7">
|
||
<paragraph id="8BB98B138D75FFACFAA9FA59A0B8F991" blockId="7.[151,1436,1525,1621]" pageId="7" pageNumber="408">
|
||
<emphasis id="B97257018D75FFACFAA9FA59A233F9C9" bold="true" box="[151,340,1525,1549]" pageId="7" pageNumber="408">FIGURES 17–18.</emphasis>
|
||
Larva of
|
||
<taxonomicName id="4C06F0908D75FFACFB87FA59A071F9C9" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[441,790,1525,1550]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D75FFACFB87FA59A145F9C9" box="[441,546,1525,1549]" italics="true" pageId="7" pageNumber="408">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D75FFACF80EFA59A1CAF9C9" box="[560,685,1525,1549]" italics="true" pageId="7" pageNumber="408">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D75FFACF883FA59A071F9C9" box="[701,790,1525,1549]" italics="true" pageId="7" pageNumber="408">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D75FFACF923FA5AA031F9CA" bold="true" box="[797,854,1526,1550]" pageId="7" pageNumber="408">
|
||
<taxonomicNameLabel id="A241EA7A8D75FFACF923FA5AA031F9CA" box="[797,854,1526,1550]" pageId="7" pageNumber="408" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB98B138D75FFACFAF9F928A632F922" blockId="7.[151,1437,1668,2018]" pageId="7" pageNumber="408">
|
||
HYPODERMAL COLORATION. Mature larva with markings characteristic for winged stages, similar in male and female: abdominal terga II, IV and VII with large W-shaped reddish-brown macula (
|
||
<figureCitation id="133D97968D75FFACFEB4F904A677F907" box="[1162,1296,1704,1731]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="7" pageNumber="408">Figs 4, 6–7</figureCitation>
|
||
); abdominal sterna with lateral longitudinal brown maculae (
|
||
<figureCitation id="133D97968D75FFACF88EF960A071F923" box="[688,790,1740,1767]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="7" pageNumber="408">Figs 6–7</figureCitation>
|
||
), whose size varies individually (as in
|
||
<figureCitation id="133D97968D75FFACFEF6F960A62EF922" box="[1224,1353,1740,1766]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="7" pageNumber="408">Figs 37, 38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D75FFA3FAF9F95CA702FD09" blockId="7.[151,1437,1668,2018]" lastBlockId="8.[151,1437,151,2013]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">
|
||
<collectionCode id="ED1713D68D75FFACFAF9F95CA296F8CE" box="[199,497,1776,1802]" pageId="7" pageNumber="408">SHAPE AND SETATION</collectionCode>
|
||
. Labrum as in other
|
||
<taxonomicName id="4C06F0908D75FFACF8E7F95CA004F8CE" authorityName="Kluge" authorityYear="2016" box="[729,867,1776,1802]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D75FFACF8E7F95CA004F8CE" box="[729,867,1776,1802]" italics="true" pageId="7" pageNumber="408">Oculogaster</emphasis>
|
||
</taxonomicName>
|
||
(as in
|
||
<figureCitation id="133D97968D75FFACF98EF95CA756F8CF" box="[944,1073,1776,1803]" captionStart="FIGURES 49–56" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 49–56. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 49–50, labrum with focus on ventral and dorsal sides; 51–52, left and right mandibles; 53–54, maxilla (ds1–ds3, dentisetae); 55–56, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="7" pageNumber="408">Figs 49–50</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D75FFACFE03F95CA7A2F8CE" author="Kluge, N. J." box="[1085,1221,1776,1802]" pageId="7" pageNumber="408" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
: figs 29, 43). Mandibles with incisor and kinetodontium fused more than to middle, with ventral denticle well-developed; incisor of right mandible with 2nd denticle shortest; right prostheca apically with 3–6 short parallel processes [as in
|
||
<taxonomicName id="4C06F0908D75FFACFF70F895A3B5F8B2" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="421" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D75FFACFF70F895A638F896" box="[1358,1375,1849,1874]" italics="true" pageId="7" pageNumber="408">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D75FFACFF46F894A6EBF896" box="[1400,1420,1848,1874]" italics="true" pageId="7" pageNumber="408">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D75FFACFAA9F8F1A3B5F8B2" box="[151,210,1885,1910]" italics="true" pageId="7" pageNumber="408">niger</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D75FFACFAE5F8F0A27BF8B2" bold="true" box="[219,284,1884,1910]" pageId="7" pageNumber="408">
|
||
<taxonomicNameLabel id="A241EA7A8D75FFACFAE5F8F0A27BF8B2" box="[219,284,1884,1910]" pageId="7" pageNumber="408" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="133D97968D75FFACFB13F8F0A2E7F8B3" box="[301,384,1884,1911]" captionStart="FIGURES 49–56" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 49–56. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 49–50, labrum with focus on ventral and dorsal sides; 51–52, left and right mandibles; 53–54, maxilla (ds1–ds3, dentisetae); 55–56, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="7" pageNumber="408">Fig. 52</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C06F0908D75FFACFBFDF8F1A1DCF8B2" authorityName=": Kluge" authorityYear="2016" box="[451,699,1884,1910]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="422" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D75FFACFBFDF8F1A2B3F8B2" box="[451,468,1885,1910]" italics="true" pageId="7" pageNumber="408">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D75FFACFBD3F8F0A166F8B2" box="[493,513,1884,1910]" italics="true" pageId="7" pageNumber="408">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D75FFACF824F8F0A1DCF8B2" box="[538,699,1884,1910]" italics="true" pageId="7" pageNumber="408">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF97F6E28D75FFACF8F2F8F0A030F8B2" author="Kluge, N. J." box="[716,855,1884,1910]" pageId="7" pageNumber="408" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
: figs 9–10)]. Maxillary palp 2-segmented (
|
||
<bibRefCitation id="EF97F6E28D75FFACFF7EF8F0A3B4F85E" author="Barnard, K. H." pageId="7" pageNumber="408" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9e; as in
|
||
<figureCitation id="133D97968D75FFACFB51F82CA2A5F85F" box="[367,450,1920,1947]" captionStart="FIGURES 49–56" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 49–56. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 49–50, labrum with focus on ventral and dorsal sides; 51–52, left and right mandibles; 53–54, maxilla (ds1–ds3, dentisetae); 55–56, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="7" pageNumber="408">Fig. 54</figureCitation>
|
||
). Labium with glossae and paraglossae of subequal size, palp with median angle moderately projected (
|
||
<bibRefCitation id="EF97F6E28D75FFACFB77F808A280F87A" author="Barnard, K. H." box="[329,487,1956,1982]" pageId="7" pageNumber="408" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9d; as in
|
||
<figureCitation id="133D97968D75FFACF8BEF808A066F87B" box="[640,769,1956,1983]" captionStart="FIGURES 49–56" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 49–56. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 49–50, labrum with focus on ventral and dorsal sides; 51–52, left and right mandibles; 53–54, maxilla (ds1–ds3, dentisetae); 55–56, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="7" pageNumber="408">Figs 55–56</figureCitation>
|
||
). Femora with very small spine-like setae on outer and inner sides, with 2 spine-like apical setae located in plane of leg; apical setae pointed or blunt, larger than other spine-like setae or equal to them (
|
||
<figureCitation id="133D97968D7AFFA3FB9CFF3BA290FF75" box="[418,503,151,177]" captionStart="FIGURES 17–18" captionStartId="7.[151,264,1525,1549]" captionTargetBox="[151,1436,561,1502]" captionTargetId="figure-198@7.[151,1436,561,1502]" captionTargetPageId="7" captionText="FIGURES 17–18. Larva of Procloeon (Oculogaster) barnardi sp. n., middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line)." figureDoi="http://doi.org/10.5281/zenodo.4397869" httpUri="https://zenodo.org/record/4397869/files/figure.png" pageId="8" pageNumber="409">Fig. 17</figureCitation>
|
||
). Claws moderately long, with two rows of denticles, minute in proximal part and larger more distally, reaching mid-length of claw (
|
||
<figureCitation id="133D97968D7AFFA3F8FCFF17A071FF11" box="[706,790,187,213]" captionStart="FIGURES 19–29" captionStartId="9.[151,264,1552,1576]" captionTargetBox="[153,1428,187,1527]" captionTargetId="figure-16@9.[151,1436,181,1529]" captionTargetPageId="9" captionText="FIGURES 19–29. Larvae of Procloeon (Oculogaster) barnardi sp. n. 19–25, tergalii I–VII; 26–27, tergalii V of other specimens; 28, tergalius VII of another specimen; 29, claw (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397871" httpUri="https://zenodo.org/record/4397871/files/figure.png" pageId="8" pageNumber="409">Fig. 29</figureCitation>
|
||
). Hind protoptera completely absent. Lateral spines on abdominal segments V–IX, spines sometimes present only on some of these segments; segment IX with 2–7 spines on one side, other segments with equal or less number of spines (
|
||
<figureCitation id="133D97968D7AFFA3F96EFEAFA0D1FED9" box="[848,950,259,285]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="8" pageNumber="409">Figs 2–3</figureCitation>
|
||
). Posterolateral spines present on segments II–IX, being minute on anterior segments. Posterior margin of abdominal tergum I with few small sparse spine-like pointed denticles; posterior margins of terga II–X with larger spine-like pointed denticles of unequal length (
|
||
<figureCitation id="133D97968D7AFFA3FF51FEE7A3C1FE4D" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
|
||
); on tergum IX row of denticles interrupted medially, behind pair of submedian setae (as in
|
||
<figureCitation id="133D97968D7AFFA3FE86FEC3A66BFE4D" box="[1208,1292,367,393]" captionStart="FIGURES 43–48" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 43–48. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 43–44, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IX–X; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (43–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="8" pageNumber="409">Fig. 47</figureCitation>
|
||
). Abdominal terga with short scales in wide semilunar or W-shaped non-opercula-bearing sockets, with sparse simple fine setae; one pair of bifurcate setae (as in
|
||
<figureCitation id="133D97968D7AFFA3FBC1FE1BA128FE15" box="[511,591,439,465]" captionStart="FIGURES 43–48" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 43–48. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 43–44, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IX–X; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (43–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="8" pageNumber="409">Fig. 48</figureCitation>
|
||
) near midlength of each tergum
|
||
<collectionCode id="ED1713D68D7AFFA3F988FE1BA775FE15" box="[950,1042,439,465]" pageId="8" pageNumber="409">VIII–IX</collectionCode>
|
||
and one pair of bifurcate setae near posterior margin of each tergum VII and IX (
|
||
<figureCitation id="133D97968D7AFFA3F8ABFE77A1BDFE31" box="[661,730,475,501]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="8" pageNumber="409">Fig. 3</figureCitation>
|
||
). Posterior margins of abdominal sterna I–V smooth, posterior margins of sterna VI–IX and paraprocts with spine-like pointed denticles of unequal length (as in
|
||
<figureCitation id="133D97968D7AFFA3FEEDFE53A641FDDD" box="[1235,1318,511,537]" captionStart="FIGURES 43–48" captionStartId="14.[151,264,1916,1940]" captionTargetBox="[151,1436,181,1892]" captionTargetId="figure-27@14.[151,1436,181,1892]" captionTargetPageId="14" captionText="FIGURES 43–48. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 43–44, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IX–X; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (43–46, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397885" httpUri="https://zenodo.org/record/4397885/files/figure.png" pageId="8" pageNumber="409">Fig. 47</figureCitation>
|
||
). Abdominal sterna with sparse scales and sparse fine setae; in lateral areas of sterna both simple and bifid fine setae vary from small to very long, located sparsely and irregularly, not forming regular rows. Tergalius I with costal rib very short and lacking denticles; costal rib on tergalii II–VII longer, either without denticles or with few small denticles on apex; dorsal lamella present either on tergalii I–VI, I–V, or I–IV (
|
||
<figureCitation id="133D97968D7AFFA3F9A1FD23A744FD6D" box="[927,1059,655,681]" captionStart="FIGURES 19–29" captionStartId="9.[151,264,1552,1576]" captionTargetBox="[153,1428,187,1527]" captionTargetId="figure-16@9.[151,1436,181,1529]" captionTargetPageId="9" captionText="FIGURES 19–29. Larvae of Procloeon (Oculogaster) barnardi sp. n. 19–25, tergalii I–VII; 26–27, tergalii V of other specimens; 28, tergalius VII of another specimen; 29, claw (holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397871" httpUri="https://zenodo.org/record/4397871/files/figure.png" pageId="8" pageNumber="409">Figs 19–28</figureCitation>
|
||
). In distal part of cercus spine on lateral side of each segment subequal to or 1.5 times longer than next segment (
|
||
<figureCitation id="133D97968D7AFFA3FE2CFD1FA730FD09" box="[1042,1111,691,717]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="8" pageNumber="409">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7AFFA3FAF9FD7AA142FC45" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="409">
|
||
<emphasis id="B97257018D7AFFA3FAF9FD7AA224FD35" bold="true" box="[199,323,726,753]" italics="true" pageId="8" pageNumber="409">Subimago.</emphasis>
|
||
CUTICULAR COLORATION. Head colorless. Pronotum with brown and colorless areas. Mesonotum brown with sutures darker brown, with large colorless areas occupying anterior part of submedioscutum and posterior scutal protuberance (
|
||
<figureCitation id="133D97968D7AFFA3FBD4FCB3A159FCFD" box="[490,574,799,825]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-27@10.[151,1436,181,1694]" captionTargetPageId="10" captionText="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="8" pageNumber="409">Fig. 32</figureCitation>
|
||
). Thoracic pleura mostly colorless, with some sclerites brown (
|
||
<figureCitation id="133D97968D7AFFA3FEC1FCB3A635FCFD" box="[1279,1362,799,825]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-27@10.[151,1436,181,1694]" captionTargetPageId="10" captionText="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="8" pageNumber="409">Fig. 33</figureCitation>
|
||
). Legs nearly colorless, with base of tibia brown, sometimes brownish stripes on femur. Abdomen very light brownish with sigilla colorless. Caudalii colorless.
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7AFFA3FAF9FC27A7C6FC62" blockId="8.[151,1437,151,2013]" box="[199,1185,907,934]" pageId="8" pageNumber="409">HYPODERMAL COLORATION. As in imago, similar in male and female (see below).</paragraph>
|
||
<paragraph id="8BB98B138D7AFFA3FAF9FC03A60AFC29" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="409">
|
||
TEXTURE. On fore leg of male and female 1st tarsal segment covered partly with microtrichia (as tibia), partly with pointed microlepides; all other tarsal segments entirely covered with pointed microlepides (as in
|
||
<figureCitation id="133D97968D7AFFA3FF33FC7FA607FC29" box="[1293,1376,979,1005]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 42</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7AFFA3FAF9FC5BA0BCFA21" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="409">
|
||
<emphasis id="B97257018D7AFFA3FAF9FC5BA23CFBD5" bold="true" box="[199,347,1014,1041]" italics="true" pageId="8" pageNumber="409">Imago, male</emphasis>
|
||
(
|
||
<figureCitation id="133D97968D7AFFA3FB50FC5BA292FBD5" box="[366,501,1015,1041]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-27@10.[151,1436,181,1694]" captionTargetPageId="10" captionText="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="8" pageNumber="409">Figs 30–31</figureCitation>
|
||
). Head ocher with brown. Antenna pale ocher, scapus and pedicellus with small brown spot at apex on lateral and median sides. Turbinate eyes widened apically; facetted surface yellow, stem orange. Pronotum ocher with brown maculae; propleura and prosternum brown, articulations pale ocher. Meso- and metathorax with sclerites brown, membranes pale ocher. Legs pale ocher or whitish. In
|
||
<typeStatus id="54BD35B18D7AFFA3FE56FBCFA7AAFBB9" box="[1128,1229,1123,1149]" pageId="8" pageNumber="409" type="holotype">holotype</typeStatus>
|
||
, ratio femur / tibia / tarsal segments (mm) on fore leg 0.96: 1.20: 0.05: 0.52: 0.35: 0.19: 0.14; on middle and hind leg 0.85: 0.73: 0.28: 0.09: 0.04: 0.14. Middle and hind legs without apical spine on 1st+2nd tarsomere, with single apical spine on primary 3rd tarsomere (as in
|
||
<figureCitation id="133D97968D7AFFA3F83AFB63A13FFB2D" box="[516,600,1231,1257]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 42</figureCitation>
|
||
). Wing with membrane colorless, veins ocher; subcostal vein proximad of costal brace colored with brown. Pterostigma with one oblique cross vein; besides it, costal field with very slender cross vein belonging to middle cord of cross veins (as in female –
|
||
<bibRefCitation id="EF97F6E28D7AFFA3F94CFABBA769FAF5" author="Barnard, K. H." box="[882,1038,1303,1329]" pageId="8" pageNumber="409" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 8). Marginal intercalaries present in most of interspaces, both behind convex and concave longitudinal veins. Hind wing absent. Abdominal terga light ocher with large darker W-shaped maculae, contrastingly reddish-brown on terga II, IV and VII, less contrasting on other terga. Abdominal sternum I light brownish, with pair of round lateral brown maculae; sterna II–IX light ocher, with pair of longitudinal lateral brown stripes. Gonostyli pale ocher, penis brown. Median projection of penis semicircular (
|
||
<figureCitation id="133D97968D7AFFA3FB0FFA67A2E2FA21" box="[305,389,1483,1509]" captionStart="FIGURES 34–35" captionStartId="11.[151,264,1567,1591]" captionTargetBox="[162,1424,316,1534]" captionTargetId="figure-85@11.[151,1436,309,1544]" captionTargetPageId="11" captionText="FIGURES 34–35. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 34, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 35, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397877" httpUri="https://zenodo.org/record/4397877/files/figure.png" pageId="8" pageNumber="409">Fig. 34</figureCitation>
|
||
). Cerci with alternating whitish and brown segments.
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7AFFA3FAF9FA43A645F819" blockId="8.[151,1437,151,2013]" pageId="8" pageNumber="409">
|
||
<emphasis id="B97257018D7AFFA3FAF9FA43A20AF9CD" bold="true" box="[199,365,1518,1545]" italics="true" pageId="8" pageNumber="409">Imago, female</emphasis>
|
||
(
|
||
<figureCitation id="133D97968D7AFFA3FB45FA43A299F9CE" box="[379,510,1519,1546]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Figs 36–39</figureCitation>
|
||
). Head relatively wide, distance between eyes exceeds eye length; eyes only slightly elevated above head surface. Head ocher, with or without brown markings. Pronotum ocher, with more or less extensive brown spots. Mesonotum either uniformly pale ocher (
|
||
<figureCitation id="133D97968D7AFFA3F948F99BA0ACF995" box="[886,971,1591,1617]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 37</figureCitation>
|
||
), or with medioscutum darker, scutellum white and parascutellum brown (
|
||
<figureCitation id="133D97968D7AFFA3F831F9F7A102F9B1" box="[527,613,1627,1653]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 38</figureCitation>
|
||
). Postnotum either uniformly pale ocher, or with brown markings. Thoracic pleura and sterna ocher or light brownish, with brown and white maculae. Fore leg without apical spine on 2nd tarsomere, with single apical spine on primary 3rd tarsomere (as in
|
||
<figureCitation id="133D97968D7AFFA3F980F90FA775F97A" box="[958,1042,1699,1726]" captionStart="FIGURES 73–77" captionStartId="20.[151,264,1907,1931]" captionTargetBox="[151,1436,273,1884]" captionTargetId="figure-95@20.[151,1436,273,1884]" captionTargetPageId="20" captionText="FIGURES 73–77. Female Procloeon (Oculogaster) niger sp. n. 73–74, imago; 75, imaginal abdomen; 76–77, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st–5th, initial tarsomeres)." figureDoi="http://doi.org/10.5281/zenodo.4397897" httpUri="https://zenodo.org/record/4397897/files/figure.png" pageId="8" pageNumber="409">Fig. 76</figureCitation>
|
||
); same on middle and hind legs (as in
|
||
<figureCitation id="133D97968D7AFFA3FA8AF96BA26FF925" box="[180,264,1735,1761]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 42</figureCitation>
|
||
). Wings as in male (see above); on one wing of one specimen, pterostigma, besides one cross vein, with additional incomplete cross vein (
|
||
<figureCitation id="133D97968D7AFFA3F829F947A10BF8C1" box="[535,620,1771,1797]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 41</figureCitation>
|
||
). Abdominal terga ocher with contrasting reddish-brown maculae: most extensive and more or less W-shaped maculae on terga II, IV and VII; median unpaired spot or stripe (either separate or incorporated into W-shaped macula) on terga II–VII; pair of submedian spots on all or some of terga II–VII; pair of zigzag ocher or brown bands along lateral margins of abdominal terga, connecting W-shape maculae of terga II, IV and VII. Abdominal sterna ocher, sterna I–IX with pair of contrasting reddish-brown lateral maculae: these maculae either round on sternum I and longitudinal wedge-like on sterna II–IX (
|
||
<figureCitation id="133D97968D7AFFA3FE20F833A716F87D" box="[1054,1137,1951,1977]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 36</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D7AFFA3FE43F833A67CF87D" author="Barnard, K. H." box="[1149,1307,1951,1977]" pageId="8" pageNumber="409" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9a), or wide and parallel-sided on all sterna I–IX (
|
||
<figureCitation id="133D97968D7AFFA3F849F86FA1ABF819" box="[631,716,1987,2013]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="8" pageNumber="409">Fig. 39</figureCitation>
|
||
). Cerci with alternating whitish and brown segments.
|
||
</paragraph>
|
||
<caption id="DF79DB9B8D7BFFA2FAA9F9BCA049F989" ID-DOI="http://doi.org/10.5281/zenodo.4397871" ID-Zenodo-Dep="4397871" httpUri="https://zenodo.org/record/4397871/files/figure.png" pageId="9" pageNumber="410" startId="9.[151,264,1552,1576]" targetBox="[153,1428,187,1527]" targetPageId="9">
|
||
<paragraph id="8BB98B138D7BFFA2FAA9F9BCA049F989" blockId="9.[151,1437,1552,1613]" pageId="9" pageNumber="410">
|
||
<emphasis id="B97257018D7BFFA2FAA9F9BCA232F9ED" bold="true" box="[151,341,1552,1577]" pageId="9" pageNumber="410">FIGURES 19–29.</emphasis>
|
||
Larvae of
|
||
<taxonomicName id="4C06F0908D7BFFA2FBF9F9BDA041F9ED" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[455,806,1553,1577]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2FBF9F9BDA157F9ED" box="[455,560,1553,1577]" italics="true" pageId="9" pageNumber="410">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D7BFFA2F87EF9BDA1DAF9ED" box="[576,701,1553,1577]" italics="true" pageId="9" pageNumber="410">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D7BFFA2F8F3F9BDA041F9ED" box="[717,806,1553,1577]" italics="true" pageId="9" pageNumber="410">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7BFFA2F910F9BDA00FF9ED" bold="true" box="[814,872,1553,1577]" pageId="9" pageNumber="410">
|
||
<taxonomicNameLabel id="A241EA7A8D7BFFA2F910F9BDA00FF9ED" box="[814,872,1553,1577]" pageId="9" pageNumber="410" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
19–25, tergalii I–VII; 26–27, tergalii V of other specimens; 28, tergalius VII of another specimen; 29, claw (holotype).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB98B138D7BFFA2FAF9F9D7A1E2F952" blockId="9.[151,1437,1659,1973]" box="[199,645,1659,1686]" pageId="9" pageNumber="410">
|
||
<emphasis id="B97257018D7BFFA2FAF9F9D7A39BF951" bold="true" box="[199,252,1659,1685]" italics="true" pageId="9" pageNumber="410">Egg.</emphasis>
|
||
Without rigid chorion (viviparity).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7BFFA2FAF9F933A657F97D" blockId="9.[151,1437,1659,1973]" box="[199,1328,1695,1722]" pageId="9" pageNumber="410">
|
||
<emphasis id="B97257018D7BFFA2FAF9F933A229F97D" bold="true" box="[199,334,1695,1721]" pageId="9" pageNumber="410">Dimension.</emphasis>
|
||
Fore wing length of male and female
|
||
<quantity id="4CFE26F68D7BFFA2F8C7F933A00CF97D" box="[761,875,1695,1722]" metricMagnitude="-3" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="9" pageNumber="410" unit="mm" value="4.75" valueMax="5.0" valueMin="4.5">4.5–5 mm</quantity>
|
||
(
|
||
<quantity id="4CFE26F68D7BFFA2F944F933A08AF97D" box="[890,1005,1695,1722]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="9" pageNumber="410" unit="mm" value="5.25" valueMax="5.5" valueMin="5.0">5–5.5 mm</quantity>
|
||
according to
|
||
<bibRefCitation id="EF97F6E28D7BFFA2FEB8F933A644F97D" author="Barnard, K. H." box="[1158,1315,1695,1721]" pageId="9" pageNumber="410" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7BFFA2FAF9F96FA1BDF871" blockId="9.[151,1437,1659,1973]" pageId="9" pageNumber="410">
|
||
<emphasis id="B97257018D7BFFA2FAF9F96FA205F919" bold="true" box="[199,354,1731,1757]" pageId="9" pageNumber="410">Comparison.</emphasis>
|
||
The larva of
|
||
<taxonomicName id="4C06F0908D7BFFA2F83FF968A1DAF919" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[513,701,1731,1757]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2F83FF968A175F919" box="[513,530,1732,1757]" italics="true" pageId="9" pageNumber="410">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D7BFFA2F812F96FA127F919" box="[556,576,1731,1757]" italics="true" pageId="9" pageNumber="410">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D7BFFA2F864F96FA1DAF919" box="[602,701,1731,1757]" italics="true" pageId="9" pageNumber="410">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7BFFA2F8F8F96FA060F919" bold="true" box="[710,775,1731,1757]" pageId="9" pageNumber="410">
|
||
<taxonomicNameLabel id="A241EA7A8D7BFFA2F8F8F96FA060F919" box="[710,775,1731,1757]" pageId="9" pageNumber="410" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has the same structure and cuticular coloration as
|
||
<taxonomicName id="4C06F0908D7BFFA2FF6FF968A25FF8C5" authorityName=": Kluge" authorityYear="2016" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2FF6FF968A601F919" box="[1361,1382,1732,1757]" italics="true" pageId="9" pageNumber="410">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D7BFFA2FF46F96FA6F4F919" box="[1400,1427,1731,1757]" italics="true" pageId="9" pageNumber="410">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D7BFFA2FAA9F94BA25FF8C5" box="[151,312,1767,1793]" italics="true" pageId="9" pageNumber="410">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
. However, the hypodermal pigmentation of larva, subimago and imago is clearly different in these two species. While in
|
||
<taxonomicName id="4C06F0908D7BFFA2FBA8F8A0A1ECF8E1" authorityName=": Kluge" authorityYear="2016" box="[406,651,1803,1829]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="410" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2FBA8F8A0A2CCF8E1" box="[406,427,1804,1829]" italics="true" pageId="9" pageNumber="410">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D7BFFA2FB83F8A7A2BFF8E1" box="[445,472,1803,1829]" italics="true" pageId="9" pageNumber="410">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D7BFFA2FBD4F8A7A1ECF8E1" box="[490,651,1803,1829]" italics="true" pageId="9" pageNumber="410">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
the abdominal hypodermal pigmentation sharply differs in male and female (
|
||
<bibRefCitation id="EF97F6E28D7BFFA2FACDF883A21AF88D" author="Kluge, N. J." box="[243,381,1839,1865]" pageId="9" pageNumber="410" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
: figs 35–36), in
|
||
<taxonomicName id="4C06F0908D7BFFA2F808F89CA188F88D" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[566,751,1839,1865]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2F808F89CA120F88D" box="[566,583,1840,1865]" italics="true" pageId="9" pageNumber="410">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D7BFFA2F85EF883A113F88D" box="[608,628,1839,1865]" italics="true" pageId="9" pageNumber="410">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D7BFFA2F8B2F883A188F88D" box="[652,751,1839,1865]" italics="true" pageId="9" pageNumber="410">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7BFFA2F8C8F883A050F88D" bold="true" box="[758,823,1839,1865]" pageId="9" pageNumber="410">
|
||
<taxonomicNameLabel id="A241EA7A8D7BFFA2F8C8F883A050F88D" box="[758,823,1839,1865]" pageId="9" pageNumber="410" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
it is similar in both sexes (
|
||
<figureCitation id="133D97968D7BFFA2FE54F883A7A6F88D" box="[1130,1217,1839,1865]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-27@10.[151,1436,181,1694]" captionTargetPageId="10" captionText="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="9" pageNumber="410">Figs 30</figureCitation>
|
||
,
|
||
<figureCitation id="133D97968D7BFFA2FEF0F883A67FF88D" box="[1230,1304,1839,1865]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="9" pageNumber="410">37–38</figureCitation>
|
||
), especially in the subimaginal stage (
|
||
<figureCitation id="133D97968D7BFFA2FB89F8FFA179F8AA" box="[439,542,1875,1902]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="9" pageNumber="410">Figs 6–7</figureCitation>
|
||
). I was able to examine only one male imago of
|
||
<taxonomicName id="4C06F0908D7BFFA2FE7CF8F8A79DF8A9" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1090,1274,1875,1901]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="9" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7BFFA2FE7CF8F8A734F8A9" box="[1090,1107,1876,1901]" italics="true" pageId="9" pageNumber="410">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D7BFFA2FE55F8FFA718F8A9" box="[1131,1151,1875,1901]" italics="true" pageId="9" pageNumber="410">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D7BFFA2FEA9F8FFA79DF8A9" box="[1175,1274,1875,1901]" italics="true" pageId="9" pageNumber="410">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7BFFA2FF3FF8FFA626F8A9" bold="true" box="[1281,1345,1875,1901]" pageId="9" pageNumber="410">
|
||
<taxonomicNameLabel id="A241EA7A8D7BFFA2FF3FF8FFA626F8A9" box="[1281,1345,1875,1901]" pageId="9" pageNumber="410" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but the characteristic hypodermal pigmentation of the male was examined also on 7 mature male larvae (
|
||
<figureCitation id="133D97968D7BFFA2FEE7F8DBA659F856" box="[1241,1342,1911,1938]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="9" pageNumber="410">Figs 4, 6</figureCitation>
|
||
), so this character is assumed to be reliably species-specific.
|
||
</paragraph>
|
||
<caption id="DF79DB9B8D78FFA1FAA9F91AA10EF936" ID-DOI="http://doi.org/10.5281/zenodo.4397875" ID-Zenodo-Dep="4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="10" pageNumber="411" startId="10.[151,264,1718,1742]" targetBox="[151,1436,181,1694]" targetPageId="10">
|
||
<paragraph id="8BB98B138D78FFA1FAA9F91AA10EF936" blockId="10.[151,1436,1718,1778]" pageId="10" pageNumber="411">
|
||
<emphasis id="B97257018D78FFA1FAA9F91AA231F90A" bold="true" box="[151,342,1718,1742]" pageId="10" pageNumber="411">FIGURES 30–33.</emphasis>
|
||
Male
|
||
<taxonomicName id="4C06F0908D78FFA1FBA7F91AA19FF90A" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[409,760,1718,1742]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1FBA7F91AA165F90A" box="[409,514,1718,1742]" italics="true" pageId="10" pageNumber="411">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D78FFA1F82CF91AA1E8F90A" box="[530,655,1718,1742]" italics="true" pageId="10" pageNumber="411">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D78FFA1F8A1F91AA19FF90A" box="[671,760,1718,1742]" italics="true" pageId="10" pageNumber="411">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D78FFA1F93EF91AA05CF90A" bold="true" box="[768,827,1718,1742]" pageId="10" pageNumber="411">
|
||
<taxonomicNameLabel id="A241EA7A8D78FFA1F93EF91AA05CF90A" box="[768,827,1718,1742]" pageId="10" pageNumber="411" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB98B138D78FFA1FAF9F88DA6E9F82B" blockId="10.[151,1437,1824,2031]" pageId="10" pageNumber="411">
|
||
In contrast to male
|
||
<taxonomicName id="4C06F0908D78FFA1FBA3F88DA1E9F8FE" authorityName=": Kluge" authorityYear="2016" box="[413,654,1824,1851]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1FBA3F88DA2D4F8FE" box="[413,435,1825,1850]" italics="true" pageId="10" pageNumber="411">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D78FFA1FBFCF88DA2BAF8FF" box="[450,477,1825,1851]" italics="true" pageId="10" pageNumber="411">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D78FFA1FBD3F88CA1E9F8FE" box="[493,654,1824,1850]" italics="true" pageId="10" pageNumber="411">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
, whose middle abdominal terga are either non-pigmented, or with one pair of small spots on tergum IV only (
|
||
<bibRefCitation id="EF97F6E28D78FFA1F876F8E8A1B6F89A" author="Kluge, N. J." box="[584,721,1860,1886]" pageId="10" pageNumber="411" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
: figs 33–34), in male
|
||
<taxonomicName id="4C06F0908D78FFA1F9FDF8E9A71EF89A" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[963,1145,1860,1887]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1F9FDF8E9A0B3F89A" box="[963,980,1861,1886]" italics="true" pageId="10" pageNumber="411">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D78FFA1F9D5F8E9A098F89B" box="[1003,1023,1861,1887]" italics="true" pageId="10" pageNumber="411">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D78FFA1FE28F8E8A71EF89A" box="[1046,1145,1860,1886]" italics="true" pageId="10" pageNumber="411">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D78FFA1FE41F8E9A7D8F89B" bold="true" box="[1151,1215,1861,1887]" pageId="10" pageNumber="411">
|
||
<taxonomicNameLabel id="A241EA7A8D78FFA1FE41F8E9A7D8F89B" box="[1151,1215,1861,1887]" pageId="10" pageNumber="411" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
all abdominal terga have large W-shaped maculae (
|
||
<figureCitation id="133D97968D78FFA1FBCCF8C4A112F847" box="[498,629,1896,1923]" captionStart="FIGURES 30–33" captionStartId="10.[151,264,1718,1742]" captionTargetBox="[151,1436,181,1694]" captionTargetId="figure-27@10.[151,1436,181,1694]" captionTargetPageId="10" captionText="FIGURES 30–33. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 30–31, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397875" httpUri="https://zenodo.org/record/4397875/files/figure.png" pageId="10" pageNumber="411">Figs 30–31</figureCitation>
|
||
). Male imaginal turbinate eyes of
|
||
<taxonomicName id="4C06F0908D78FFA1F9D1F8C5A7C3F846" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1007,1188,1896,1923]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1F9D1F8C5A767F846" box="[1007,1024,1897,1922]" italics="true" pageId="10" pageNumber="411">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D78FFA1FE29F8C5A74CF847" box="[1047,1067,1897,1923]" italics="true" pageId="10" pageNumber="411">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D78FFA1FE7FF8C4A7C3F846" box="[1089,1188,1896,1922]" italics="true" pageId="10" pageNumber="411">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D78FFA1FE94F8C5A78EF847" bold="true" box="[1194,1257,1897,1923]" pageId="10" pageNumber="411">
|
||
<taxonomicNameLabel id="A241EA7A8D78FFA1FE94F8C5A78EF847" box="[1194,1257,1897,1923]" pageId="10" pageNumber="411" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
are much wider that that of
|
||
<taxonomicName id="4C06F0908D78FFA1FB26F821A16EF862" authorityName=": Kluge" authorityYear="2016" box="[280,521,1932,1959]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1FB26F821A24AF862" box="[280,301,1933,1958]" italics="true" pageId="10" pageNumber="411">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D78FFA1FB03F821A23FF863" box="[317,344,1933,1959]" italics="true" pageId="10" pageNumber="411">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D78FFA1FB56F820A16EF862" box="[360,521,1932,1958]" italics="true" pageId="10" pageNumber="411">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
. Kimmins (1965) correctly reported that the male imago of
|
||
<taxonomicName id="4C06F0908D78FFA1FE95F821A6FAF862" authorityName=": Kluge" authorityYear="2016" box="[1195,1437,1932,1959]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="411" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1FE95F821A7A7F862" box="[1195,1216,1933,1958]" italics="true" pageId="10" pageNumber="411">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D78FFA1FEEEF821A78CF863" box="[1232,1259,1933,1959]" italics="true" pageId="10" pageNumber="411">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D78FFA1FEC2F820A6FAF862" box="[1276,1437,1932,1958]" italics="true" pageId="10" pageNumber="411">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
differs by «more slender claspers» (i.e. gonostyli) (Kimmins 1965: fig. 2;
|
||
<bibRefCitation id="EF97F6E28D78FFA1F9D5F81CA712F80E" author="Kluge, N. J." box="[1003,1141,1968,1994]" pageId="10" pageNumber="411" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
: fig. 30), while in
|
||
<taxonomicName id="4C06F0908D78FFA1FF73F81DA39DF82A" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D78FFA1FF73F81DA639F80E" box="[1357,1374,1969,1994]" italics="true" pageId="10" pageNumber="411">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D78FFA1FF46F81DA6EBF80F" box="[1400,1420,1969,1995]" italics="true" pageId="10" pageNumber="411">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D78FFA1FAA9F878A39DF82A" box="[151,250,2004,2030]" italics="true" pageId="10" pageNumber="411">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D78FFA1FB3FF879A227F82B" bold="true" box="[257,320,2005,2031]" pageId="10" pageNumber="411">
|
||
<taxonomicNameLabel id="A241EA7A8D78FFA1FB3FF879A227F82B" box="[257,320,2005,2031]" pageId="10" pageNumber="411" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
the second segment of gonostylus is relatively shorter and thicker (
|
||
<figureCitation id="133D97968D78FFA1FE0DF878A7E0F82B" box="[1075,1159,2004,2031]" captionStart="FIGURES 34–35" captionStartId="11.[151,264,1567,1591]" captionTargetBox="[162,1424,316,1534]" captionTargetId="figure-85@11.[151,1436,309,1544]" captionTargetPageId="11" captionText="FIGURES 34–35. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 34, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 35, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397877" httpUri="https://zenodo.org/record/4397877/files/figure.png" pageId="10" pageNumber="411">Fig. 34</figureCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D78FFA1FEADF878A657F82B" author="Barnard, K. H." box="[1171,1328,2004,2031]" pageId="10" pageNumber="411" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9i).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D79FFA0FAF9FF3BA127FED9" blockId="11.[151,1436,151,285]" pageId="11" pageNumber="412">
|
||
In contrast to female
|
||
<taxonomicName id="4C06F0908D79FFA0FB91FF3BA1F9FF75" authorityName=": Kluge" authorityYear="2016" box="[431,670,151,177]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
|
||
<emphasis id="B97257018D79FFA0FB91FF3BA2A2FF74" box="[431,453,151,176]" italics="true" pageId="11" pageNumber="412">P.</emphasis>
|
||
(
|
||
<emphasis id="B97257018D79FFA0FBEDFF3BA289FF75" box="[467,494,151,177]" italics="true" pageId="11" pageNumber="412">O.</emphasis>
|
||
)
|
||
<emphasis id="B97257018D79FFA0FBC3FF3BA1F9FF75" box="[509,670,151,177]" italics="true" pageId="11" pageNumber="412">cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
, whose abdominal terga have constant zigzag lateral bands expressed both in imago and subimago, and have no median maculae (
|
||
<bibRefCitation id="EF97F6E28D79FFA0F90CFF17A0DDFF11" author="Kluge, N. J." box="[818,954,187,213]" pageId="11" pageNumber="412" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
; fig. 36), female
|
||
<taxonomicName id="4C06F0908D79FFA0FE48FF17A64BFF11" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1142,1324,187,213]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D79FFA0FE48FF17A7E0FF10" box="[1142,1159,187,212]" italics="true" pageId="11" pageNumber="412">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D79FFA0FEA0FF17A7D5FF11" box="[1182,1202,187,213]" italics="true" pageId="11" pageNumber="412">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D79FFA0FEF7FF17A64BFF11" box="[1225,1324,187,213]" italics="true" pageId="11" pageNumber="412">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D79FFA0FF0CFF17A616FF11" bold="true" box="[1330,1393,187,213]" pageId="11" pageNumber="412">
|
||
<taxonomicNameLabel id="A241EA7A8D79FFA0FF0CFF17A616FF11" box="[1330,1393,187,213]" pageId="11" pageNumber="412" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has median maculae on all or some terga, while zigzag lateral bands are expressed only in imago (
|
||
<figureCitation id="133D97968D79FFA0FEF8FF73A62DFF3E" box="[1222,1354,223,250]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="11" pageNumber="412">Figs 36–39</figureCitation>
|
||
), being absent in subimago and larva (
|
||
<figureCitation id="133D97968D79FFA0FBD3FEAFA156FED9" box="[493,561,259,285]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="11" pageNumber="412">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF79DB9B8D79FFA0FAA9F9B3A75FF998" ID-DOI="http://doi.org/10.5281/zenodo.4397877" ID-Zenodo-Dep="4397877" httpUri="https://zenodo.org/record/4397877/files/figure.png" pageId="11" pageNumber="412" startId="11.[151,264,1567,1591]" targetBox="[162,1424,316,1534]" targetPageId="11">
|
||
<paragraph id="8BB98B138D79FFA0FAA9F9B3A75FF998" blockId="11.[151,1436,1567,1628]" pageId="11" pageNumber="412">
|
||
<emphasis id="B97257018D79FFA0FAA9F9B3A232F9FC" bold="true" box="[151,341,1567,1592]" pageId="11" pageNumber="412">FIGURES 34–35.</emphasis>
|
||
Male
|
||
<taxonomicName id="4C06F0908D79FFA0FBA9F98CA191F9FC" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[407,758,1568,1592]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D79FFA0FBA9F98CA167F9FC" box="[407,512,1568,1592]" italics="true" pageId="11" pageNumber="412">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D79FFA0F82EF98CA1EAF9FC" box="[528,653,1568,1592]" italics="true" pageId="11" pageNumber="412">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D79FFA0F8A3F98CA191F9FC" box="[669,758,1568,1592]" italics="true" pageId="11" pageNumber="412">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D79FFA0F8C3F98CA050F9FC" bold="true" box="[765,823,1568,1592]" pageId="11" pageNumber="412">
|
||
<taxonomicNameLabel id="A241EA7A8D79FFA0F8C3F98CA050F9FC" box="[765,823,1568,1592]" pageId="11" pageNumber="412" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(holotype). 34, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 35, their subimaginal exuviae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31CD8988D79FFA6FAF9F926A7E9FC0D" lastPageId="13" lastPageNumber="414" pageId="11" pageNumber="412" type="discussion">
|
||
<paragraph id="8BB98B138D79FFA6FAF9F926A655FEA2" blockId="11.[151,1437,1674,2025]" lastBlockId="13.[151,1437,151,969]" lastPageId="13" lastPageNumber="414" pageId="11" pageNumber="412">
|
||
<emphasis id="B97257018D79FFA0FAF9F926A228F960" bold="true" box="[199,335,1674,1700]" pageId="11" pageNumber="412">Comments.</emphasis>
|
||
This species was described by
|
||
<bibRefCitation id="EF97F6E28D79FFA0F88CF926A007F960" author="Barnard, K. H." box="[690,864,1674,1700]" pageId="11" pageNumber="412" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932)</bibRefCitation>
|
||
, who regarded it to be conspecific with
|
||
<taxonomicName id="4C06F0908D79FFA0FF1BF926A28EF90C" authority="Esben-Petersen 1913" authorityName="Esben-Petersen" authorityYear="1913" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0FF1BF926A393F90C" italics="true" pageId="11" pageNumber="412">Cloeon africanum</emphasis>
|
||
<bibRefCitation id="EF97F6E28D79FFA0FAC2F902A28EF90C" author="Esben-Petersen, P." box="[252,489,1710,1736]" pageId="11" pageNumber="412" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
and placed it in a newly established genus
|
||
<taxonomicName id="4C06F0908D79FFA0F9EAF902A66AF90C" authority="Barnard 1932" authorityName="Barnard" authorityYear="1932" box="[980,1293,1710,1736]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D79FFA0F9EAF902A70FF90C" box="[980,1128,1710,1736]" italics="true" pageId="11" pageNumber="412">Austrocloeon</emphasis>
|
||
<bibRefCitation id="EF97F6E28D79FFA0FE4EF902A66AF90C" author="Barnard, K. H." box="[1136,1293,1710,1736]" pageId="11" pageNumber="412" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
</taxonomicName>
|
||
. The species identification was based not on the
|
||
<typeStatus id="54BD35B18D79FFA0F818F97EA1ECF928" box="[550,651,1746,1772]" pageId="11" pageNumber="412" type="holotype">holotype</typeStatus>
|
||
, but on «topotypes from Zululand» (
|
||
<bibRefCitation id="EF97F6E28D79FFA0FE19F97EA798F928" author="Barnard, K. H." box="[1063,1279,1746,1772]" pageId="11" pageNumber="412" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932: 219</bibRefCitation>
|
||
). The original description of
|
||
<taxonomicName id="4C06F0908D79FFA0FB03F95AA2D7F8D4" authorityName="Esben-Petersen" authorityYear="1913" box="[317,432,1782,1808]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0FB03F95AA2D7F8D4" box="[317,432,1782,1808]" italics="true" pageId="11" pageNumber="412">africanum</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="4C06F0908D79FFA0FBFDF95AA173F8D4" authorityName="Leach" authorityYear="1815" box="[451,532,1782,1808]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D79FFA0FBFDF95AA173F8D4" box="[451,532,1782,1808]" italics="true" pageId="11" pageNumber="412">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
] was based on a single male imago. Its genitalia (
|
||
<bibRefCitation id="EF97F6E28D79FFA0FE6BF95AA62EF8D4" author="Esben-Petersen, P." box="[1109,1353,1782,1808]" pageId="11" pageNumber="412" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913</bibRefCitation>
|
||
: fig. 8) have nothing in common with the genitalia of the species described by
|
||
<bibRefCitation id="EF97F6E28D79FFA0F997F8B6A728F8F0" author="Barnard, K. H." box="[937,1103,1818,1844]" pageId="11" pageNumber="412" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932</bibRefCitation>
|
||
: fig. 9i) under the name «
|
||
<taxonomicName id="4C06F0908D79FFA0FF57F8B6A21AF89C" authorityName=": Barnard" authorityYear="1932" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0FF57F8B6A21AF89C" italics="true" pageId="11" pageNumber="412">Austrocloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
». Because of this, I concluded that the genus-group name
|
||
<taxonomicName id="4C06F0908D79FFA0FE22F892A7D7F89C" authorityName="Barnard" authorityYear="1932" box="[1052,1200,1854,1880]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D79FFA0FE22F892A7D7F89C" box="[1052,1200,1854,1880]" italics="true" pageId="11" pageNumber="412">Austrocloeon</emphasis>
|
||
</taxonomicName>
|
||
(whose type species is
|
||
<taxonomicName id="4C06F0908D79FFA0FA8FF8CEA243F8B8" authorityName=": Barnard" authorityYear="1932" box="[177,292,1890,1916]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0FA8FF8CEA243F8B8" box="[177,292,1890,1916]" italics="true" pageId="11" pageNumber="412">africanum</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="4C06F0908D79FFA0FB08F8CEA2E0F8B8" authorityName="Leach" authorityYear="1815" box="[310,391,1890,1916]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D79FFA0FB08F8CEA2E0F8B8" box="[310,391,1890,1916]" italics="true" pageId="11" pageNumber="412">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
]) and the species name
|
||
<taxonomicName id="4C06F0908D79FFA0F8A9F8CEA06DF8B8" authorityName=": Barnard" authorityYear="1932" box="[663,778,1890,1916]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0F8A9F8CEA06DF8B8" box="[663,778,1890,1916]" italics="true" pageId="11" pageNumber="412">africanum</emphasis>
|
||
</taxonomicName>
|
||
should not be applied to the species described by Barnard under these names (
|
||
<bibRefCitation id="EF97F6E28D79FFA0FB49F82AA165F865" author="Kluge, N. J." box="[375,514,1926,1953]" pageId="11" pageNumber="412" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
). Recent examination of a male imago belonging to the Barnards’s «
|
||
<taxonomicName id="4C06F0908D79FFA0FF36F82AA26DF800" authorityName=": Barnard" authorityYear="1932" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0FF36F82AA26DF800" italics="true" pageId="11" pageNumber="412">Austrocloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
» reveals that it sharply differs from
|
||
<taxonomicName id="4C06F0908D79FFA0F890F806A046F800" authorityName=": Barnard" authorityYear="1932" box="[686,801,1962,1988]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D79FFA0F890F806A046F800" box="[686,801,1962,1988]" italics="true" pageId="11" pageNumber="412">africanum</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="4C06F0908D79FFA0F90DF806A0E3F800" authorityName="Leach" authorityYear="1815" box="[819,900,1962,1988]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="412" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D79FFA0F90DF806A0E3F800" box="[819,900,1962,1988]" italics="true" pageId="11" pageNumber="412">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
] not only in genital structure, but in coloration as well: according to the original description, «2nd–6th segments of abdomen transparent, whitish and with a faint reddish tinge; the hind borders of the segments darker. The 7th–9th segments reddish brown on the dorsal surface, whitish on the ventral» (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FB90FF17A1BBFF11" author="Esben-Petersen, P." box="[430,732,187,213]" pageId="13" pageNumber="414" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913: 184</bibRefCitation>
|
||
); such coloration occurs in many species of
|
||
<taxonomicName id="4C06F0908D7FFFA6FEE4FF17A658FF11" authorityName="Leach" authorityYear="1815" box="[1242,1343,187,213]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
|
||
, but not in
|
||
<taxonomicName id="4C06F0908D7FFFA6FA8AFF73A20BFF3D" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[180,364,223,249]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="422" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7FFFA6FA8AFF73A3A2FF3C" box="[180,197,223,248]" italics="true" pageId="13" pageNumber="414">P</emphasis>
|
||
. (
|
||
<emphasis id="B97257018D7FFFA6FAE3FF73A396FF3D" box="[221,241,223,249]" italics="true" pageId="13" pageNumber="414">O</emphasis>
|
||
.)
|
||
<emphasis id="B97257018D7FFFA6FB37FF73A20BFF3D" box="[265,364,223,249]" italics="true" pageId="13" pageNumber="414">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7FFFA6FB4DFF73A2D4FF3D" bold="true" box="[371,435,223,249]" pageId="13" pageNumber="414">
|
||
<taxonomicNameLabel id="A241EA7A8D7FFFA6FB4DFF73A2D4FF3D" box="[371,435,223,249]" pageId="13" pageNumber="414" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
described above. Judging by the structure of gonostyli (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FE14FF73A67BFF3D" author="Esben-Petersen, P." box="[1066,1308,223,249]" pageId="13" pageNumber="414" pagination="177 - 187" refId="ref17520" refString="Esben-Petersen, P. (1913) Ephemeridae from South Africa. Annals of the South African Museum, 10 (6), 177 - 187." type="journal article" year="1913">Esben-Petersen 1913</bibRefCitation>
|
||
: fig. 8), the species
|
||
<taxonomicName id="4C06F0908D7FFFA6FAD0FEAFA206FED9" authorityName=": Barnard" authorityYear="1932" box="[238,353,259,285]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6FAD0FEAFA206FED9" box="[238,353,259,285]" italics="true" pageId="13" pageNumber="414">africanum</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="4C06F0908D7FFFA6FB4EFEAFA2A6FED9" authorityName="Leach" authorityYear="1815" box="[368,449,259,285]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D7FFFA6FB4EFEAFA2A6FED9" box="[368,449,259,285]" italics="true" pageId="13" pageNumber="414">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
] can be a member of Rhithrocloeoninae Kluge 2012, which belong to the plesiomorphon Protopatellata
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FB04FE8BA137FE85" author="Kluge, N. J. & Novikova, E. A." box="[314,592,295,321]" pageId="13" pageNumber="414" pagination="1 - 56" refId="ref18669" refString="Kluge, N. J. & Novikova, E. A. (2011) Systematics of the mayfly taxon Acentrella (Ephemeroptera, Baetidae), with description of new Asian and African species. Russian Entomological Journal, 20 (1), 1 - 56." type="journal article" year="2011">Kluge & Novikova 2011</bibRefCitation>
|
||
(
|
||
<bibRefCitation id="EF97F6E28D7FFFA6F861FE8BA18EFE86" author="Kluge, N. J." box="[607,745,295,322]" pageId="13" pageNumber="414" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
|
||
). Recently, larvae of an unknown two-winged protopatellatan species were found in
|
||
<collectingCountry id="F311CB838D7FFFA6FBACFEE7A144FEA1" box="[402,547,331,357]" name="South Africa" pageId="13" pageNumber="414">South Africa</collectingCountry>
|
||
; possibly, they belong to the true
|
||
<taxonomicName id="4C06F0908D7FFFA6F9A5FEE7A769FEA1" authorityName=": Barnard" authorityYear="1932" box="[923,1038,331,357]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6F9A5FEE7A769FEA1" box="[923,1038,331,357]" italics="true" pageId="13" pageNumber="414">africanum</emphasis>
|
||
</taxonomicName>
|
||
[
|
||
<taxonomicName id="4C06F0908D7FFFA6FE21FEE7A717FEA1" authorityName="Leach" authorityYear="1815" box="[1055,1136,331,357]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97257018D7FFFA6FE21FEE7A717FEA1" box="[1055,1136,331,357]" italics="true" pageId="13" pageNumber="414">Cloeon</emphasis>
|
||
</taxonomicName>
|
||
] (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FEB6FEE7A641FEA2" author="Kluge, N. J." box="[1160,1318,331,358]" pageId="13" pageNumber="414" pagination="438 - 464" refId="ref18534" refString="Kluge, N. J. (2020 b) Demoulinia Gillies 1990 and two unnamed genera of the plesiomorphon Protopatellata (Ephemeroptera, Baetidae) from South Africa. Zootaxa, 4820 (3), 438 - 464 https: // doi. org / 10.11646 / zootaxa. 4820.3.2" type="journal article" year="2020">Kluge, 2020b</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF79DB9B8D7EFFA7FAA9F8D2A239F87F" ID-DOI="http://doi.org/10.5281/zenodo.4397879" ID-Zenodo-Dep="4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="12" pageNumber="413" startId="12.[151,264,1918,1942]" targetBox="[151,1436,181,1894]" targetPageId="12">
|
||
<paragraph id="8BB98B138D7EFFA7FAA9F8D2A239F87F" blockId="12.[151,1436,1918,1979]" pageId="12" pageNumber="413">
|
||
<emphasis id="B97257018D7EFFA7FAA9F8D2A231F852" bold="true" box="[151,342,1918,1942]" pageId="12" pageNumber="413">FIGURES 36–42.</emphasis>
|
||
Female
|
||
<taxonomicName id="4C06F0908D7EFFA7FB8FF8D3A075F853" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[433,786,1919,1943]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="12" pageNumber="413" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
|
||
<emphasis id="B97257018D7EFFA7FB8FF8D3A17DF853" box="[433,538,1919,1943]" italics="true" pageId="12" pageNumber="413">Procloeon</emphasis>
|
||
(
|
||
<emphasis id="B97257018D7EFFA7F815F8D3A1CFF853" box="[555,680,1919,1943]" italics="true" pageId="12" pageNumber="413">Oculogaster</emphasis>
|
||
)
|
||
<emphasis id="B97257018D7EFFA7F887F8D3A075F853" box="[697,786,1919,1943]" italics="true" pageId="12" pageNumber="413">barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7EFFA7F924F8D3A032F853" bold="true" box="[794,853,1919,1943]" pageId="12" pageNumber="413">
|
||
<taxonomicNameLabel id="A241EA7A8D7EFFA7F924F8D3A032F853" box="[794,853,1919,1943]" pageId="12" pageNumber="413" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB98B138D7FFFA6FAF9FEC3A647FE31" blockId="13.[151,1437,151,969]" pageId="13" pageNumber="414">
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FAF9FEC3A213FE4D" author="Barnard, K. H." box="[199,372,367,393]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932)</bibRefCitation>
|
||
described the imago of «
|
||
<taxonomicName id="4C06F0908D7FFFA6F8ADFEC3A0C5FE4D" authorityName=": Barnard" authorityYear="1932" box="[659,930,367,393]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6F8ADFEC3A0C5FE4D" box="[659,930,367,393]" italics="true" pageId="13" pageNumber="414">Austrocloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
» not indicating its sex; actually this description belongs to the female only, more precisely to the light form of female (the same as in
|
||
<figureCitation id="133D97968D7FFFA6FE93FE3FA657FE6A" box="[1197,1328,403,430]" captionStart="FIGURES 36–42" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 36–42. Female Procloeon (Oculogaster) barnardi sp. n. 36–39, imagines; 40–41, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="13" pageNumber="414">Figs 36–37</figureCitation>
|
||
). Besides the drawings of the female imaginal abdomen (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6F894FE1BA02EFE15" author="Barnard, K. H." box="[682,841,439,465]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: figs 9a–b), Barnard gave also a drawing of the male imaginal gonostylus (ibid.,
|
||
<figureCitation id="133D97968D7FFFA6FBF4FE77A170FE32" box="[458,535,475,502]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="13" pageNumber="414">Fig. 9i</figureCitation>
|
||
). Other characters of male imago or subimago had not been reported.
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7FFFA6FAF9FE53A098FD42" blockId="13.[151,1437,151,969]" pageId="13" pageNumber="414">
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FAF9FE53A213FDDD" author="Barnard, K. H." box="[199,372,511,537]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932)</bibRefCitation>
|
||
also described larvae of this species and noted «two varieties» of their abdominal coloration; probably, the first of them, with all terga similarly ornamented (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6F95CFD8FA765FDF9" author="Barnard, K. H." box="[866,1026,547,573]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9c) represents a combination of cuticular and hypodermal coloration (
|
||
<figureCitation id="133D97968D7FFFA6F87DFDEBA1EFFDA5" box="[579,648,583,609]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="13" pageNumber="414">Fig. 4</figureCitation>
|
||
), and the second one, with terga IV and VII much lighter than others (
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FAA1FDC7A259FD41" author="Barnard, K. H." box="[159,318,619,645]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard 1932</bibRefCitation>
|
||
: fig. 9h), represents the cuticular coloration solely (
|
||
<figureCitation id="133D97968D7FFFA6F940FDC7A0A0FD41" box="[894,967,619,645]" captionStart="FIGURES 1–7" captionStartId="5.[151,264,1918,1942]" captionTargetBox="[154,1436,181,1894]" captionTargetId="figure-16@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1–7. Procloeon (Oculogaster) barnardi sp. n. 1–6, male larva ready to moult to subimago: 1–2, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IV–IX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." figureDoi="http://doi.org/10.5281/zenodo.4397863" httpUri="https://zenodo.org/record/4397863/files/figure.png" pageId="13" pageNumber="414">Figs 2</figureCitation>
|
||
,
|
||
<figureCitation id="133D97968D7FFFA6F9EDFDC7A095FD42" box="[979,1010,619,646]" captionStart="FIGURES 8–16" captionStartId="6.[151,264,1953,1977]" captionTargetBox="[155,1426,181,1930]" captionTargetId="figure-27@6.[155,1433,181,1930]" captionTargetPageId="6" captionText="FIGURES 8–16. Larvae of Procloeon (Oculogaster) barnardi sp. n. 8–9, caudalii; 10–15, exuviae of abdomen and pronotum with mesonotum of three specimens (14–15, holotype); 16, young larva." figureDoi="http://doi.org/10.5281/zenodo.4397867" httpUri="https://zenodo.org/record/4397867/files/figure.png" pageId="13" pageNumber="414">16</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7FFFA6FAF9FD23A769FD35" blockId="13.[151,1437,151,969]" pageId="13" pageNumber="414">
|
||
Since the paper by
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FBA5FD23A121FD6D" author="Barnard, K. H." box="[411,582,655,681]" pageId="13" pageNumber="414" pagination="201 - 259" refId="ref17247" refString="Barnard, K. H. (1932) South African May-flies (Ephemeroptera). Transactions of the Royal Society of South Africa, 20, 201 - 259. https: // doi. org / 10.1080 / 00359193209518858" type="journal article" year="1932">Barnard (1932)</bibRefCitation>
|
||
was published, a variety of authors have applied the name «
|
||
<taxonomicName id="4C06F0908D7FFFA6FEDDFD23A393FD08" authorityName=": Barnard" authorityYear="1932" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6FEDDFD23A393FD08" italics="true" pageId="13" pageNumber="414">Austrocloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
» to the species described in this paper, including some (e.g.,
|
||
<bibRefCitation id="EF97F6E28D7FFFA6F99FFD1FA7D0FD09" author="Harrison, A. D. & Agnew, J. D." box="[929,1207,691,717]" pageId="13" pageNumber="414" pagination="273 - 291" refId="ref17804" refString="Harrison, A. D. & Agnew, J. D. (1962) The distribution of invertebrates endemic to acid streams in the Western and Southern Cape Province. Annals of the Cape Provincial Museums, 2, 273 - 291." type="journal article" year="1962">Harrison & Agnew 1962</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF97F6E28D7FFFA6FEFDFD1FA6F3FD0A" author="Oliff, W. D. & King, J. L." box="[1219,1428,691,718]" pageId="13" pageNumber="414" pagination="567 - 583" refId="ref18861" refString="Oliff, W. D. & King, J. L. (1964) Hydrobiological studies of the Tugela River System. Part IV. The Mooi River. Hydrobiologia, 24 (4), 567 - 583. https: // doi. org / 10.1007 / BF 00142003" type="journal article" year="1964">Oliff & King 1964</bibRefCitation>
|
||
) who erroneously reported its authorship as «
|
||
<taxonomicName id="4C06F0908D7FFFA6F8B7FD7BA099FD35" authority="Barnard" authorityName="" sensu Barnard" authorityYear="1932" box="[649,1022,727,753]" class="Insecta" family="Baetidae" genus="Austrocloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6F8B7FD7BA0F0FD35" box="[649,919,727,753]" italics="true" pageId="13" pageNumber="414">Austrocloeon africanum</emphasis>
|
||
Barnard
|
||
</taxonomicName>
|
||
».
|
||
</paragraph>
|
||
<paragraph id="8BB98B138D7FFFA6FAF0FD57A7E9FC0D" blockId="13.[151,1437,151,969]" pageId="13" pageNumber="414">
|
||
The coloration of winged males of this species was not described.
|
||
<bibRefCitation id="EF97F6E28D7FFFA6F985FD57A710FCD1" author="Kimmins, D. E." box="[955,1143,762,789]" pageId="13" pageNumber="414" pagination="859 - 880" refId="ref17911" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880. [1955] https: // doi. org / 10.1080 / 00222935508655706" type="journal article" year="1956">Kimmins (1956)</bibRefCitation>
|
||
, comparing the newly described species
|
||
<taxonomicName id="4C06F0908D7FFFA6FB77FCB3A102FCFD" authorityName=": Kluge" authorityYear="2016" box="[329,613,799,825]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cylindroculum">
|
||
<emphasis id="B97257018D7FFFA6FB77FCB3A102FCFD" box="[329,613,799,825]" italics="true" pageId="13" pageNumber="414">Procloeon cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
with what he called «
|
||
<taxonomicName id="4C06F0908D7FFFA6F95EFCB3A729FCFD" authorityName=": Kimmins" authorityYear="1956" box="[864,1102,799,825]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6F95EFCB3A729FCFD" box="[864,1102,799,825]" italics="true" pageId="13" pageNumber="414">Procloeon africanum</emphasis>
|
||
</taxonomicName>
|
||
(E.-P.)» (actually
|
||
<taxonomicName id="4C06F0908D7FFFA6FF23FCB3A6FAFCFD" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1309,1437,799,825]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov.">
|
||
<emphasis id="B97257018D7FFFA6FF23FCB3A6FAFCFD" box="[1309,1437,799,825]" italics="true" pageId="13" pageNumber="414">P. barnardi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B97257018D7FFFA6FAA9FCEFA3B1FC99" bold="true" box="[151,214,835,861]" pageId="13" pageNumber="414">
|
||
<taxonomicNameLabel id="A241EA7A8D7FFFA6FAA9FCEFA3B1FC99" box="[151,214,835,861]" pageId="13" pageNumber="414" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
) wrote that «it differs from that species in the translucent middle segments of the abdomen», but he did not report the source of information about abdomen coloration of the male «
|
||
<taxonomicName id="4C06F0908D7FFFA6F997FCCBA750FC45" authorityName=": Kimmins" authorityYear="1956" box="[937,1079,871,897]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="africanum">
|
||
<emphasis id="B97257018D7FFFA6F997FCCBA750FC45" box="[937,1079,871,897]" italics="true" pageId="13" pageNumber="414">P. africanum</emphasis>
|
||
</taxonomicName>
|
||
». Recent examination of reared male imago and male subimagines extracted from mature larvae reveals that the male of this species has middle segments of the abdomen intensively pigmented, which is different from
|
||
<taxonomicName id="4C06F0908D7FFFA6F9F5FC03A7EFFC0D" authorityName=": Kluge" authorityYear="2016" box="[971,1160,943,969]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="13" pageNumber="414" phylum="Arthropoda" rank="species" species="cylindroculum">
|
||
<emphasis id="B97257018D7FFFA6F9F5FC03A7EFFC0D" box="[971,1160,943,969]" italics="true" pageId="13" pageNumber="414">P. cylindroculum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |