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<mods:title id="C1F3C922F9775F7DCFE655CB59E995D9">Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<bibRefCitation id="EF97F6E28D73FFAAFB0FF90FA2D9F979" author="Kluge, N. J." box="[305,446,1699,1725]" pageId="1" pageNumber="402" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
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Procloeon (Oculogaster) barnardi sp. n. 16, male larva ready to moult to subimago: 12, larval cuticle detached from thorax and abdomen; 3, fragment of abdomen (arrows show bases of bifurcate setae similar to Fig. 48); 4, intact specimen (IVIX, abdominal segments); 5, extracted subimaginal thorax; 6, extracted subimaginal abdomen; 7, same, female." captionText-1="FIGURES 816. Larvae of Procloeon (Oculogaster) barnardi sp. n. 89, caudalii; 1015, exuviae of abdomen and pronotum with mesonotum of three specimens (1415, holotype); 16, young larva." captionText-10="FIGURES 6770. Male Procloeon (Oculogaster) niger sp. n. 6768, male imagines; 69, subimaginal exuviae of postsubalar sclerite; 70, subimaginal exuviae of mesonotum (67, 69, holotype)." captionText-11="FIGURES 7172. Male Procloeon (Oculogaster) niger sp. n. 71, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 72, their subimaginal exuviae." captionText-12="FIGURES 7377. Female Procloeon (Oculogaster) niger sp. n. 7374, imago; 75, imaginal abdomen; 7677, subimaginal exuviae of fore and middle tarsi (arrows show spines; 1st5th, initial tarsomeres)." captionText-13="FIGURES 7883. Procloeon (Oculogaster) sp. «Wolfkloof». 7879, larval exuviae of abdominal terga, pronotum and mesonotum; 80, subimaginal abdomen; 81, larval hind leg with subimaginal leg inside (arrows show spines on subimaginal tarsus); 82, claw; 83, apex of wing extracted from larva." captionText-14="FIGURES 8489. Larvae of Procloeon (Oculogaster) regularum. 8488, exuviae: 84, abdomen; 8586, its caudalii; 8788, fragment of middle part of cercus, with focus on lateral and median sides; 89, male larva." captionText-15="FIGURES 9098. Procloeon (Oculogaster) spp. 9097, P. (O.) regularum: 9096, tergalii IVII; 97, genitalia of male imago (penis dotted, muscles not shown). 98, tergalius IV of Procloeon (Oculogaster?) sp. «Philippines»." captionText-16="FIGURES 99102. Male imagines of Procloeon (Oculogaster) regularum. 99101, general view; 102, abdominal tergum IV." captionText-17="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." captionText-18="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." captionText-19="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." captionText-2="FIGURES 1718. Larva of Procloeon (Oculogaster) barnardi sp. n., middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line)." captionText-20="FIGURES 118126. Procloeon (Oculogaster) malabarensis sp. n. 118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae." captionText-21="FIGURES 127133. Procloeon (Oculogaster) malabarensis sp. n. 127, abdominal terga IIIV of male imago; 128129, male imagines; 130, subimaginal exuviae of mesothoracic pleuron; 131, female imago; 132133, abdomina of female imagines (133, holotype)." captionText-3="FIGURES 1929. Larvae of Procloeon (Oculogaster) barnardi sp. n. 1925, tergalii IVII; 2627, tergalii V of other specimens; 28, tergalius VII of another specimen; 29, claw (holotype)." captionText-4="FIGURES 3033. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 3031, imago; 32, exuviae of subimaginal mesonotum; 33, exuviae of subimaginal pleuron." captionText-5="FIGURES 3435. Male Procloeon (Oculogaster) barnardi sp. n. (holotype). 34, genitalia of imago (penis dotted; gonostylar muscle not shown at right, penial muscles not shown at left); 35, their subimaginal exuviae." captionText-6="FIGURES 3642. Female Procloeon (Oculogaster) barnardi sp. n. 3639, imagines; 4041, veins of pterostigma; 42, hind tarsus of subimago." captionText-7="FIGURES 4348. Larval exuviae of Procloeon (Oculogaster) niger sp. n. 4344, caudalii; 45, abdomen; 46, half of pronotum and mesonotum; 47, male uromeres IXX; 48, fragment of abdominal tergum VIII (arrow shows long bifurcate seta near midlength) (4346, holotype)." captionText-8="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." captionText-9="FIGURES 5766. Larvae of Procloeon (Oculogaster) niger sp. n. 5762, tergalii IIII and VVII; 6365, tergalii I, IV and VII of other specimens (6465, holotype); 66, claw." figureDoi-0="http://doi.org/10.5281/zenodo.4397863" figureDoi-1="http://doi.org/10.5281/zenodo.4397867" figureDoi-10="http://doi.org/10.5281/zenodo.4397893" figureDoi-11="http://doi.org/10.5281/zenodo.4397895" figureDoi-12="http://doi.org/10.5281/zenodo.4397897" figureDoi-13="http://doi.org/10.5281/zenodo.4397901" figureDoi-14="http://doi.org/10.5281/zenodo.4397903" figureDoi-15="http://doi.org/10.5281/zenodo.4397907" figureDoi-16="http://doi.org/10.5281/zenodo.4397909" figureDoi-17="http://doi.org/10.5281/zenodo.4397911" figureDoi-18="http://doi.org/10.5281/zenodo.4397915" figureDoi-19="http://doi.org/10.5281/zenodo.4397917" figureDoi-2="http://doi.org/10.5281/zenodo.4397869" figureDoi-20="http://doi.org/10.5281/zenodo.4397919" figureDoi-21="http://doi.org/10.5281/zenodo.4397921" figureDoi-3="http://doi.org/10.5281/zenodo.4397871" figureDoi-4="http://doi.org/10.5281/zenodo.4397875" figureDoi-5="http://doi.org/10.5281/zenodo.4397877" figureDoi-6="http://doi.org/10.5281/zenodo.4397879" figureDoi-7="http://doi.org/10.5281/zenodo.4397885" figureDoi-8="http://doi.org/10.5281/zenodo.4397887" figureDoi-9="http://doi.org/10.5281/zenodo.4397889" httpUri-0="https://zenodo.org/record/4397863/files/figure.png" httpUri-1="https://zenodo.org/record/4397867/files/figure.png" httpUri-10="https://zenodo.org/record/4397893/files/figure.png" httpUri-11="https://zenodo.org/record/4397895/files/figure.png" httpUri-12="https://zenodo.org/record/4397897/files/figure.png" httpUri-13="https://zenodo.org/record/4397901/files/figure.png" httpUri-14="https://zenodo.org/record/4397903/files/figure.png" httpUri-15="https://zenodo.org/record/4397907/files/figure.png" httpUri-16="https://zenodo.org/record/4397909/files/figure.png" httpUri-17="https://zenodo.org/record/4397911/files/figure.png" httpUri-18="https://zenodo.org/record/4397915/files/figure.png" httpUri-19="https://zenodo.org/record/4397917/files/figure.png" httpUri-2="https://zenodo.org/record/4397869/files/figure.png" httpUri-20="https://zenodo.org/record/4397919/files/figure.png" httpUri-21="https://zenodo.org/record/4397921/files/figure.png" httpUri-3="https://zenodo.org/record/4397871/files/figure.png" httpUri-4="https://zenodo.org/record/4397875/files/figure.png" httpUri-5="https://zenodo.org/record/4397877/files/figure.png" httpUri-6="https://zenodo.org/record/4397879/files/figure.png" httpUri-7="https://zenodo.org/record/4397885/files/figure.png" httpUri-8="https://zenodo.org/record/4397887/files/figure.png" httpUri-9="https://zenodo.org/record/4397889/files/figure.png" pageId="1" pageNumber="402">Figs 1133</figureCitation>
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Subgenus
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<subSubSection id="C31CD8988D73FFAAFAA9F8E5A1ACF8A5" box="[151,715,1865,1889]" pageId="1" pageNumber="402" type="materials_examined">
<paragraph id="8BB98B138D73FFAAFAA9F8E5A1ACF8A5" blockId="1.[151,715,1865,1889]" box="[151,715,1865,1889]" pageId="1" pageNumber="402">
<typeStatus id="54BD35B18D73FFAAFAA9F8E5A3AEF8A5" box="[151,201,1865,1889]" pageId="1" pageNumber="402">Type</typeStatus>
species:
<taxonomicName id="4C06F0908D73FFAAFB18F8E5A1AFF8A5" authority="Kimmins 1956" authorityName="Kimmins" authorityYear="1956" box="[294,712,1865,1889]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="species" species="cylindroculum">
<emphasis id="B97257018D73FFAAFB18F8E5A141F8A5" box="[294,550,1865,1889]" italics="true" pageId="1" pageNumber="402">Procloeon cylindroculum</emphasis>
<bibRefCitation id="EF97F6E28D73FFAAF813F8E5A1AFF8A5" author="Kimmins, D. E." box="[557,712,1865,1889]" pageId="1" pageNumber="402" pagination="859 - 880" refId="ref17911" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880. [1955] https: // doi. org / 10.1080 / 00222935508655706" type="journal article" year="1956">Kimmins 1956</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D73FFA9FAA9F825A700FE4E" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402" type="diagnosis">
<paragraph id="8BB98B138D73FFA9FAA9F825A27AFF11" blockId="1.[151,1437,1929,2028]" lastBlockId="2.[151,1437,151,825]" lastPageId="2" lastPageNumber="403" pageId="1" pageNumber="402">
<emphasis id="B97257018D73FFAAFAA9F825A25FF867" bold="true" box="[151,312,1929,1955]" pageId="1" pageNumber="402">Diagnosis. (1)</emphasis>
Viviparous (
<bibRefCitation id="EF97F6E28D73FFAAFBF8F826A128F860" author="Kluge, N. J." box="[454,591,1930,1956]" pageId="1" pageNumber="402" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
: figs 3842, 7679). The same in
<taxonomicName id="4C06F0908D73FFAAF9FBF826A771F860" authorityName="Leach" authorityYear="1815" box="[965,1046,1930,1956]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97257018D73FFAAF9FBF826A771F860" box="[965,1046,1930,1956]" italics="true" pageId="1" pageNumber="402">Cloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241EA7A8D73FFAAFE22F826A731F860" box="[1052,1110,1930,1956]" pageId="1" pageNumber="402" sensu="stricto">s. str.</taxonomicNameLabel>
and some other mayflies, but not in other
<taxonomicName id="4C06F0908D73FFAAFB1DF802A2FAF80C" authorityName="sensu Kluge" authorityYear="2016" box="[291,413,1966,1992]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="402" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1.
<emphasis id="B97257018D73FFAAFBF6F801A28EF803" bold="true" box="[456,489,1965,1991]" pageId="1" pageNumber="402">(2)</emphasis>
Pterostigma normally either with 1 cross vein (
<figureCitation id="133D97968D73FFAAFE39F802A73BF80C" box="[1031,1116,1966,1992]" captionStart="FIGURES 3642" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 3642. Female Procloeon (Oculogaster) barnardi sp. n. 3639, imagines; 4041, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="1" pageNumber="402">Fig. 40</figureCitation>
), or with 2 cross veins (
<figureCitation id="133D97968D73FFAAFF51F802A3A5F828" captionStart="FIGURES 103108" captionStartId="26.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1895]" captionTargetId="figure-27@26.[151,1436,181,1895]" captionTargetPageId="26" captionText="FIGURES 103108. Females of Procloeon (Oculogaster) regularum. 103104, imagines; 105108, subimaginal exuviae: 105106, middle and fore tarsi (arrows show spines; 1st5th, initial tarsomeres); 107, half of mesonotum; 108, mesothoracic pleuron." figureDoi="http://doi.org/10.5281/zenodo.4397911" httpUri="https://zenodo.org/record/4397911/files/figure.png" pageId="1" pageNumber="402">Fig. 104</figureCitation>
) (on individual wings this number varies from 1 to 3) (
<figureCitation id="133D97968D73FFAAF918F87EA0CEF828" box="[806,937,2002,2028]" captionStart="FIGURES 3642" captionStartId="12.[151,264,1918,1942]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@12.[151,1436,181,1894]" captionTargetPageId="12" captionText="FIGURES 3642. Female Procloeon (Oculogaster) barnardi sp. n. 3639, imagines; 4041, veins of pterostigma; 42, hind tarsus of subimago." figureDoi="http://doi.org/10.5281/zenodo.4397879" httpUri="https://zenodo.org/record/4397879/files/figure.png" pageId="1" pageNumber="402">Figs 4041</figureCitation>
).
<emphasis id="B97257018D73FFAAF985F87DA0BBF82F" bold="true" box="[955,988,2001,2027]" pageId="1" pageNumber="402">(3)</emphasis>
Larval femur with two apical spine-like setae (
<figureCitation id="133D97968D70FFA9FAE1FF3BA250FF75" box="[223,311,151,177]" captionStart="FIGURES 1718" captionStartId="7.[151,264,1525,1549]" captionTargetBox="[151,1436,561,1502]" captionTargetId="figure-198@7.[151,1436,561,1502]" captionTargetPageId="7" captionText="FIGURES 1718. Larva of Procloeon (Oculogaster) barnardi sp. n., middle leg. 17, apex of femur and base of tibia, anterior view (patella-tibial suture and bases of fine setae are visible as light markings on darker background); 18, more distal part of tibia, posterior view (with continuation of patella-tibial suture visible as light line)." figureDoi="http://doi.org/10.5281/zenodo.4397869" httpUri="https://zenodo.org/record/4397869/files/figure.png" pageId="2" pageNumber="403">Figs 17</figureCitation>
,
<figureCitation id="133D97968D70FFA9FB7AFF3BA205FF75" box="[324,354,151,177]" captionStart="FIGURES 7883" captionStartId="21.[151,264,1920,1944]" captionTargetBox="[151,1436,453,1897]" captionTargetId="figure-172@21.[151,1436,453,1897]" captionTargetPageId="21" captionText="FIGURES 7883. Procloeon (Oculogaster) sp. «Wolfkloof». 7879, larval exuviae of abdominal terga, pronotum and mesonotum; 80, subimaginal abdomen; 81, larval hind leg with subimaginal leg inside (arrows show spines on subimaginal tarsus); 82, claw; 83, apex of wing extracted from larva." figureDoi="http://doi.org/10.5281/zenodo.4397901" httpUri="https://zenodo.org/record/4397901/files/figure.png" pageId="2" pageNumber="403">81</figureCitation>
,
<figureCitation id="133D97968D70FFA9FB51FF3BA2FBFF75" box="[367,412,151,177]" captionStart="FIGURES 109112" captionStartId="28.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-27@28.[151,1436,181,1894]" captionTargetPageId="28" captionText="FIGURES 109112. Larvae of Procloeon (Oculogaster) malabarensis sp. n. 109, mature male larva (arrows show reddish hypodermal markings on abdominal terga VI and VII); 110, fragment of fore leg, focus on anterior side; 111, claw; 112, caudalii (110, 112, holotype)." figureDoi="http://doi.org/10.5281/zenodo.4397915" httpUri="https://zenodo.org/record/4397915/files/figure.png" pageId="2" pageNumber="403">110</figureCitation>
). The same in many other taxa of
<taxonomicName id="4C06F0908D70FFA9F91CFF3BA0E0FF75" authorityName="Leach" authorityYear="1815" box="[802,903,151,177]" class="Insecta" family="Baetidae" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
, but not in other
<taxonomicName id="4C06F0908D70FFA9FE75FF3BA7A5FF75" authorityName="sensu Kluge" authorityYear="2016" box="[1099,1218,151,177]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1 and not in other
<taxonomicName id="4C06F0908D70FFA9FAA9FF17A38AFF11" authorityName="Leach" authorityYear="1815" box="[151,237,187,213]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1.
</paragraph>
<paragraph id="8BB98B138D70FFA9FAF9FF73A700FE4E" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="403">
Larvae of all
<taxonomicName id="4C06F0908D70FFA9FB5CFF73A28BFF3D" authorityName="Kluge" authorityYear="2016" box="[354,492,223,249]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FB5CFF73A28BFF3D" box="[354,492,223,249]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
have a similar structure of the labium, with a pair of submedian setae distinguishable among other setae of dorsal surface (
<figureCitation id="133D97968D70FFA9F855FEAFA1A5FED9" box="[619,706,258,285]" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="2" pageNumber="403">Figs 50</figureCitation>
,
<figureCitation id="133D97968D70FFA9F8F1FEAFA19BFED9" box="[719,764,259,285]" captionStart="FIGURES 113117" captionStartId="29.[151,264,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-16@29.[151,1436,181,1501]" captionTargetPageId="29" captionText="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.4397917" httpUri="https://zenodo.org/record/4397917/files/figure.png" pageId="2" pageNumber="403">113</figureCitation>
). Mandibles of the «
<taxonomicName id="4C06F0908D70FFA9F9DDFEAFA74EFED8" authorityName="Leach" authorityYear="1815" box="[995,1065,259,284]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus">
<emphasis id="B97257018D70FFA9F9DDFEAFA74EFED8" box="[995,1065,259,284]" italics="true" pageId="2" pageNumber="403">Baetis</emphasis>
</taxonomicName>
-
<typeStatus id="54BD35B18D70FFA9FE0FFEAFA703FED9" box="[1073,1124,259,285]" pageId="2" pageNumber="403">type</typeStatus>
» (
<bibRefCitation id="EF97F6E28D70FFA9FEBEFEAFA671FED9" author="Kluge, N. J." box="[1152,1302,259,285]" pageId="2" pageNumber="403" pagination="573 - 587" refId="ref18473" refString="Kluge, N. J. (2020 a) New subgenus Monilistylus subgen. n. and a new species Procloeon (Monilistylus) ornatipennis sp. n. (Ephemeroptera: Baetidae: Procloeon). Zootaxa, 4742 (3), 573 - 587. https: // doi. org / 10.11646 / zootaxa. 4742.3.11" type="journal article" year="2020">Kluge 2020a</bibRefCitation>
: table 1 and p. 578), i.e. with incisor and kinetodontium fused, left prostheca wide, and all denticles of incisor, kinetodontium, prostheca and mola visible in one view (
<figureCitation id="133D97968D70FFA9F854FEE7A189FEA1" box="[618,750,330,357]" captionStart="FIGURES 4956" captionStartId="15.[151,264,1887,1911]" captionTargetBox="[151,1436,181,1863]" captionTargetId="figure-16@15.[151,1436,181,1863]" captionTargetPageId="15" captionText="FIGURES 4956. Larva of Procloeon (Oculogaster) niger sp. n. (holotype): 4950, labrum with focus on ventral and dorsal sides; 5152, left and right mandibles; 5354, maxilla (ds1ds3, dentisetae); 5556, labium with focus on dorsal and ventral sides." figureDoi="http://doi.org/10.5281/zenodo.4397887" httpUri="https://zenodo.org/record/4397887/files/figure.png" pageId="2" pageNumber="403">Figs 5152</figureCitation>
,
<figureCitation id="133D97968D70FFA9F8C5FEE6A007FEA1" box="[763,864,330,357]" captionStart="FIGURES 113117" captionStartId="29.[151,264,1524,1548]" captionTargetBox="[151,1436,181,1501]" captionTargetId="figure-16@29.[151,1436,181,1501]" captionTargetPageId="29" captionText="FIGURES 113117. Larva of Procloeon (Oculogaster) malabarensis sp. n. (holotype). 113, labrum (focus on dorsal side); 114115, left and right mandibles; 116, maxilla; 117, labium (at left focus on dorsal side, at right focus on ventral side)." figureDoi="http://doi.org/10.5281/zenodo.4397917" httpUri="https://zenodo.org/record/4397917/files/figure.png" pageId="2" pageNumber="403">114115</figureCitation>
). All species of
<taxonomicName id="4C06F0908D70FFA9FE24FEE7A7C3FEA1" authorityName="Kluge" authorityYear="2016" box="[1050,1188,331,357]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FE24FEE7A7C3FEA1" box="[1050,1188,331,357]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
lack hind wings, and larvae have no vestiges of hind protoptera (
<bibRefCitation id="EF97F6E28D70FFA9F845FEC3A764FE4D" author="Muller-Liebenau, I. &amp; Hubbard, M. D." box="[635,1027,366,393]" pageId="2" pageNumber="403" pagination="537 - 561" refId="ref18778" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental Region (Insecta: Ephemeroptera). Florida Entomologist, 68 (4), 537 - 561. https: // doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
: fig. 9h).
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D70FFA9FAF9FE3EA0FBFE6A" box="[199,924,402,430]" pageId="2" pageNumber="403" type="distribution">
<paragraph id="8BB98B138D70FFA9FAF9FE3EA0FBFE6A" blockId="2.[151,1437,151,825]" box="[199,924,402,430]" pageId="2" pageNumber="403">
<emphasis id="B97257018D70FFA9FAF9FE3EA206FE68" bold="true" box="[199,353,402,428]" pageId="2" pageNumber="403">Distribution.</emphasis>
Paleotropical (Afrotropical and Oriental Regions).
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D70FFA9FAF9FE1AA66AFCFD" pageId="2" pageNumber="403" type="discussion">
<paragraph id="8BB98B138D70FFA9FAF9FE1AA658FD41" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="403">
<emphasis id="B97257018D70FFA9FAF9FE1AA22BFE14" bold="true" box="[199,332,438,464]" pageId="2" pageNumber="403">Discussion.</emphasis>
In the previous diagnosis, the number of cross veins in pterostigma was reported as one, with individual variability; this diagnosis was based on
<taxonomicName id="4C06F0908D70FFA9F8DAFE77A0BBFE31" authorityName="Kimmins" authorityYear="1956" box="[740,988,475,501]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F8DAFE77A19DFE30" box="[740,762,475,500]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9F932FE77A040FE31" box="[780,807,475,501]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9F905FE77A0BBFE31" box="[827,988,475,501]" italics="true" pageId="2" pageNumber="403">cylindroculum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C06F0908D70FFA9FE26FE77A7D2FE31" authorityName="Kluge" authorityYear="2016" box="[1048,1205,475,501]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="album" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FE26FE77A749FE30" box="[1048,1070,475,500]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FE7EFE77A73BFE31" box="[1088,1116,475,501]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9FE51FE77A7D2FE31" box="[1135,1205,475,501]" italics="true" pageId="2" pageNumber="403">album</emphasis>
</taxonomicName>
, whose pterostigma normally has one cross vein. Examination of
<taxonomicName id="4C06F0908D70FFA9F8AFFE53A046FDDD" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[657,801,511,537]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="regularum">
<emphasis id="B97257018D70FFA9F8AFFE53A046FDDD" box="[657,801,511,537]" italics="true" pageId="2" pageNumber="403">P. regularum</emphasis>
</taxonomicName>
reveals that it should be placed in
<taxonomicName id="4C06F0908D70FFA9FE9BFE53A648FDDD" authorityName="Kluge" authorityYear="2016" box="[1189,1327,511,537]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FE9BFE53A648FDDD" box="[1189,1327,511,537]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
, while its pterostigma normally has 2 cross veins; this fact expands the diagnosis of
<taxonomicName id="4C06F0908D70FFA9F9F1FD8FA73EFDF9" authorityName="Kluge" authorityYear="2016" box="[975,1113,547,573]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F9F1FD8FA73EFDF9" box="[975,1113,547,573]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
. While pterostigma with one cross vein is not found in any taxa other than
<taxonomicName id="4C06F0908D70FFA9F8A1FDEBA04EFDA5" authorityName="Kluge" authorityYear="2016" box="[671,809,583,609]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F8A1FDEBA04EFDA5" box="[671,809,583,609]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
, pterostigma with two veins occurs in some other taxa, particularly in some species of
<taxonomicName id="4C06F0908D70FFA9FBCAFDC7A122FD41" authorityName="Leach" authorityYear="1815" box="[500,581,619,645]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B97257018D70FFA9FBCAFDC7A122FD41" box="[500,581,619,645]" italics="true" pageId="2" pageNumber="403">Cloeon</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A241EA7A8D70FFA9F872FDC7A1EEFD41" box="[588,649,619,645]" pageId="2" pageNumber="403" sensu="stricto">s. str.</taxonomicNameLabel>
, which were confused with species belonging to
<taxonomicName id="4C06F0908D70FFA9FE90FDC7A65FFD41" authorityName="Kluge" authorityYear="2016" box="[1198,1336,619,645]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FE90FDC7A65FFD41" box="[1198,1336,619,645]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB98B138D70FFA9FAF9FD22A66AFCFD" blockId="2.[151,1437,151,825]" pageId="2" pageNumber="403">
<emphasis id="B97257018D70FFA9FAF9FD22A1F9FD6C" bold="true" box="[199,670,654,681]" pageId="2" pageNumber="403">Species composition and classification.</emphasis>
<taxonomicName id="4C06F0908D70FFA9F897FD23A054FD6D" authorityName="Kluge" authorityYear="2016" box="[681,819,655,681]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F897FD23A054FD6D" box="[681,819,655,681]" italics="true" pageId="2" pageNumber="403">Oculogaster</emphasis>
</taxonomicName>
can be divided into two species groups: the African group, which includes
<taxonomicName id="4C06F0908D70FFA9FBA3FD1FA02FFD09" authority="Kimmins 1956" authorityName="Kimmins" authorityYear="1956" box="[413,840,690,717]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="cylindroculum" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FBA3FD1FA2D5FD08" box="[413,434,691,716]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FBFAFD1FA2B8FD09" box="[452,479,691,717]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9FBCCFD1FA1F4FD09" box="[498,659,691,717]" italics="true" pageId="2" pageNumber="403">cylindroculum</emphasis>
<bibRefCitation id="EF97F6E28D70FFA9F8A2FD1FA02FFD09" author="Kimmins, D. E." box="[668,840,690,717]" pageId="2" pageNumber="403" pagination="859 - 880" refId="ref17911" refString="Kimmins, D. E. (1956) Ephemeroptera from Nyasaland, with description of three new species and some interesting nymphal forms. Annals and Magazine of Natural History, Series 12, 8 (95), 859 - 880. [1955] https: // doi. org / 10.1080 / 00222935508655706" type="journal article" year="1956">Kimmins 1956</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C06F0908D70FFA9F96BFD1FA76AFD09" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[853,1037,691,717]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="barnardi" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F96BFD1FA00CFD08" box="[853,875,691,716]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9F942FD1FA0FFFD09" box="[892,920,691,717]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9F994FD1FA76AFD09" box="[938,1037,691,717]" italics="true" pageId="2" pageNumber="403">barnardi</emphasis>
</taxonomicName>
<emphasis id="B97257018D70FFA9FE28FD1FA73FFD09" bold="true" box="[1046,1112,691,717]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A241EA7A8D70FFA9FE28FD1FA73FFD09" box="[1046,1112,691,717]" pageId="2" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C06F0908D70FFA9FE56FD1FA791FD09" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[1128,1270,691,717]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="421" phylum="Arthropoda" rank="species" species="niger" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FE56FD1FA71AFD08" box="[1128,1149,691,716]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FEB1FD1FA7CDFD09" box="[1167,1194,691,717]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
) niger
</taxonomicName>
<emphasis id="B97257018D70FFA9FEC1FD1FA626FD09" bold="true" box="[1279,1345,691,717]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A241EA7A8D70FFA9FEC1FD1FA626FD09" box="[1279,1345,691,717]" pageId="2" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B97257018D70FFA9FF6FFD1FA601FD08" box="[1361,1382,691,716]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FF46FD1FA6F4FD09" box="[1400,1427,691,717]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
) sp. «Wolfkloof» and presumably
<taxonomicName id="4C06F0908D70FFA9F82EFD7BA031FD35" authority="Gillies 1997" authorityName="Gillies" authorityYear="1997" box="[528,854,727,753]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="silvicola" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F82EFD7BA142FD34" box="[528,549,727,752]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9F80BFD7BA137FD35" box="[565,592,727,753]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9F85FFD7BA1A7FD35" box="[609,704,727,753]" italics="true" pageId="2" pageNumber="403">silvicola</emphasis>
<bibRefCitation id="EF97F6E28D70FFA9F8F6FD7BA031FD35" author="Gillies, M. T." box="[712,854,727,753]" pageId="2" pageNumber="403" pagination="247 - 250" refId="ref17733" refString="Gillies, M. T. (1997) A new species of Procloeon Bengtsson from the forest zone of West Africa (Ephemeroptera, Baetidae). Entomologist's Monthly Magazine, 133, 247 - 250." type="journal article" year="1997">Gillies 1997</bibRefCitation>
</taxonomicName>
, and the Asian group, which includes
<taxonomicName id="4C06F0908D70FFA9FF3AFD7BA279FCD1" authority="Kluge 2016" authorityName="Kluge" authorityYear="2016" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="album" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FF3AFD7BA67DFD34" box="[1284,1306,727,752]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FF14FD7BA622FD35" box="[1322,1349,727,753]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9FF68FD7BA6FBFD35" box="[1366,1436,727,753]" italics="true" pageId="2" pageNumber="403">album</emphasis>
<bibRefCitation id="EF97F6E28D70FFA9FAA9FD57A279FCD1" author="Kluge, N. J." box="[151,286,763,789]" pageId="2" pageNumber="403" pagination="491 - 516" refId="ref18350" refString="Kluge, N. J. (2016) A new subgenus Oculogaster subgen. n. for viviparous representatives of Procloeon s. l., with discussion about status of the generic name Austrocloeon Barnard 1932 and the species name africanum Esben-Petersen 1913 [Cloeon] (Ephemeroptera, Baetidae). Zootaxa, 4107 (4), 491 - 516. https: // doi. org / 10.11646 / zootaxa. 4107.4.2" type="journal article" year="2016">Kluge 2016</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C06F0908D70FFA9FB17FD57A01BFCD1" authority="Muller-Liebenau &amp; Hubbard 1985" authorityName="Muller-Liebenau &amp; Hubbard" authorityYear="1985" box="[297,892,762,789]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="regularum" subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9FB17FD57A259FCD0" box="[297,318,763,788]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9FB70FD57A20EFCD1" box="[334,361,763,789]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9FB44FD57A288FCD1" box="[378,495,763,789]" italics="true" pageId="2" pageNumber="403">regularum</emphasis>
<bibRefCitation id="EF97F6E28D70FFA9FBC9FD57A01BFCD1" author="Muller-Liebenau, I. &amp; Hubbard, M. D." box="[503,892,762,789]" pageId="2" pageNumber="403" pagination="537 - 561" refId="ref18778" refString="Muller-Liebenau, I. &amp; Hubbard, M. D. (1985) Baetidae from Sri Lanka with some general remarks on the Baetidae of the Oriental Region (Insecta: Ephemeroptera). Florida Entomologist, 68 (4), 537 - 561. https: // doi. org / 10.2307 / 3494855" type="journal article" year="1985">Müller-Liebenau &amp; Hubbard 1985</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C06F0908D70FFA9F9B6FD57A709FCD1" authority="Kluge, 2020" authorityName="Kluge" authorityYear="2020" box="[904,1134,763,789]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="malabarensis" status="sp. nov." subGenus="Oculogaster">
<emphasis id="B97257018D70FFA9F9B6FD57A0FAFCD0" box="[904,925,763,788]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
(
<emphasis id="B97257018D70FFA9F993FD57A0AFFCD1" box="[941,968,763,789]" italics="true" pageId="2" pageNumber="403">O.</emphasis>
)
<emphasis id="B97257018D70FFA9F9E6FD57A709FCD1" box="[984,1134,763,789]" italics="true" pageId="2" pageNumber="403">malabarensis</emphasis>
</taxonomicName>
<emphasis id="B97257018D70FFA9FE48FD57A7D2FCD1" bold="true" box="[1142,1205,763,789]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A241EA7A8D70FFA9FE48FD57A7D2FCD1" box="[1142,1205,763,789]" pageId="2" pageNumber="403" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, presumably
<taxonomicName id="4C06F0908D70FFA9FF73FD57A253FCFD" authority="(Gillies 1949)" authorityName="Kluge" authorityYear="2020" baseAuthorityName="Gillies" baseAuthorityYear="1949" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="403" phylum="Arthropoda" rank="species" species="julia" status="comb. nov.">
<emphasis id="B97257018D70FFA9FF73FD57A6FBFCD1" box="[1357,1436,763,789]" italics="true" pageId="2" pageNumber="403">P. julia</emphasis>
(
<bibRefCitation id="EF97F6E28D70FFA9FAA1FCB3A24CFCFD" author="Gillies, M. T." box="[159,299,799,825]" pageId="2" pageNumber="403" pagination="161 - 177" refId="ref17548" refString="Gillies, M. T. (1949) Notes on some Ephemeroptera Baetidae from India and South-East Asia. Transactions of the Royal Entomological Society London, 100, 161 - 177. https: // doi. org / 10.1111 / j. 1365 - 2311.1949. tb 01429. x" type="journal article" year="1949">Gillies 1949</bibRefCitation>
)
</taxonomicName>
<emphasis id="B97257018D70FFA9FB04FCB3A2C6FCFD" bold="true" box="[314,417,799,825]" pageId="2" pageNumber="403">
<taxonomicNameLabel id="A241EA7A8D70FFA9FB04FCB3A2C6FCFD" box="[314,417,799,825]" pageId="2" pageNumber="403" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
and unnamed
<emphasis id="B97257018D70FFA9F87AFCB3A13DFCFC" box="[580,602,799,824]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
sp. «Thailand-3»,
<emphasis id="B97257018D70FFA9F912FCB3A026FCFC" box="[812,833,799,824]" italics="true" pageId="2" pageNumber="403">P.</emphasis>
sp. «Sulawesi» and
<emphasis id="B97257018D70FFA9FE18FCB3A750FCFC" box="[1062,1079,799,824]" italics="true" pageId="2" pageNumber="403">P</emphasis>
. sp. «
<collectingCountry id="F311CB838D70FFA9FE46FCB3A79AFCFD" box="[1144,1277,799,825]" name="Philippines" pageId="2" pageNumber="403">Philippines</collectingCountry>
».
</paragraph>
</subSubSection>
</treatment>
</document>