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<document id="38A204E73B2CB5367A4B27CC9E234553" ID-CLB-Dataset="8958" ID-DOI="10.11646/zootaxa.4820.3.1" ID-GBIF-Dataset="fd413580-2960-463f-a924-8c27eaeafc6e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4397861" ID-ZooBank="44BD1E07-C9F3-4488-936E-819C2FCA18C8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609210733456" checkinUser="plazi" docAuthor="Kluge, Nikita J." docDate="2020" docId="03AF3A058D53FF89FAA9F970A037FEA1" docLanguage="en" docName="zootaxa.4820.3.1.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procloeon (Oculogaster) Kluge 2020" docType="treatment" docVersion="7" lastPageNumber="434" masterDocId="FF96427D8D72FFABFA3EFFACA367FFC4" masterDocTitle="Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)" masterLastPageNumber="437" masterPageNumber="401" pageNumber="434" updateTime="1698848343517" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1AE3882E9D40E044DFFB3CDB47A387E3">Review of Oculogaster Kluge 2016 (Ephemeroptera, Baetidae, Procloeon Bengtsson 1915)</mods:title>
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<treatment id="03AF3A058D53FF89FAA9F970A037FEA1" ID-DOI="http://doi.org/10.5281/zenodo.4436265" ID-GBIF-Taxon="176573424" ID-Zenodo-Dep="4436265" LSID="urn:lsid:plazi:treatment:03AF3A058D53FF89FAA9F970A037FEA1" httpUri="http://treatment.plazi.org/id/03AF3A058D53FF89FAA9F970A037FEA1" lastPageId="34" lastPageNumber="434" pageId="33" pageNumber="434">
<subSubSection id="C31CD8988D53FF8AFAA9F970A1EBF932" box="[151,652,1756,1783]" pageId="33" pageNumber="434" type="nomenclature">
<paragraph id="8BB98B138D53FF8AFAA9F970A1EBF932" blockId="33.[151,652,1756,1818]" box="[151,652,1756,1783]" pageId="33" pageNumber="434">
<heading id="D0F13C7F8D53FF8AFAA9F970A1EBF932" bold="true" box="[151,652,1756,1783]" fontSize="11" level="1" pageId="33" pageNumber="434" reason="1">
<emphasis id="B97257018D53FF8AFAA9F970A1EBF932" bold="true" box="[151,652,1756,1783]" pageId="33" pageNumber="434">
<taxonomicName id="4C06F0908D53FF8AFAA9F970A2D0F933" ID-CoL="63PRK" ID-ENA="248186" authorityName="Kluge" authorityYear="2020" box="[151,439,1756,1783]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="402" phylum="Arthropoda" rank="genus">
<emphasis id="B97257018D53FF8AFAA9F970A274F933" bold="true" box="[151,275,1756,1783]" italics="true" pageId="33" pageNumber="434">Procloeon</emphasis>
(
<emphasis id="B97257018D53FF8AFB1AF970A2D0F933" bold="true" box="[292,439,1756,1783]" italics="true" pageId="33" pageNumber="434">Oculogaster</emphasis>
</taxonomicName>
?) sp. «
<collectingRegion id="49C245F18D53FF8AF831F970A119F932" box="[527,638,1756,1782]" country="Indonesia" name="Sulawesi Utara" pageId="33" pageNumber="434">Sulawesi</collectingRegion>
»
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D53FF8AFAA9F8ACA224F8DE" box="[151,323,1792,1818]" pageId="33" pageNumber="434" type="description">
<paragraph id="8BB98B138D53FF8AFAA9F8ACA224F8DE" blockId="33.[151,652,1756,1818]" box="[151,323,1792,1818]" pageId="33" pageNumber="434">
(
<figureCitation id="133D97968D53FF8AFAA1F8ACA25CF8DE" box="[159,315,1792,1818]" captionStart="FIGURES 138141" captionStartId="34.[151,264,1809,1833]" captionTargetBox="[151,1436,381,1785]" captionTargetId="figure-132@34.[151,1436,381,1785]" captionTargetPageId="34" captionText="FIGURES 138141. Larva of Procloeon (Oculogaster?) sp. «Sulawesi». 138, glossae and paraglossae; 139, maxilla (ds1ds3, dentisetae); 140, right mandible; 141, part of cercus." figureDoi="http://doi.org/10.5281/zenodo.4397927" httpUri="https://zenodo.org/record/4397927/files/figure.png" pageId="33" pageNumber="434">Figs 138141</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D53FF8AFAA9F8E4A387F842" pageId="33" pageNumber="434" type="materials_examined">
<paragraph id="8BB98B138D53FF8AFAA9F8E4A387F842" blockId="33.[151,1437,1864,2035]" pageId="33" pageNumber="434">
<emphasis id="B97257018D53FF8AFAA9F8E4A2E2F8A6" bold="true" box="[151,389,1864,1890]" pageId="33" pageNumber="434">Material examined.</emphasis>
<materialsCitation id="3B6E814E8D53FF8AFBB1F8E4A3BBF842" ID-GBIF-Occurrence="3014991319" collectingDate="2009-08-15" collectingDateMax="2009-08-27" collectingDateMin="2009-08-15" collectionCode="ZIN" collectorName="N. Kluge &amp; L. Sheyko" country="Indonesia" location="Mamasa" pageId="33" pageNumber="434" specimenCount="1" specimenCount-larva="1" stateProvince="Sulawesi Utara">
<collectingCountry id="F311CB838D53FF8AFBB1F8E4A14DF8A6" box="[399,554,1864,1890]" name="Indonesia" pageId="33" pageNumber="434">INDONESIA</collectingCountry>
,
<collectingRegion id="49C245F18D53FF8AF804F8E4A1C5F8A6" box="[570,674,1864,1890]" country="Indonesia" name="Sulawesi Utara" pageId="33" pageNumber="434">Sulawesi</collectingRegion>
,
<location id="8ED9DDC88D53FF8AF88FF8E5A074F8A7" LSID="urn:lsid:plazi:treatment:03AF3A058D53FF89FAA9F970A037FEA1:8ED9DDC88D53FF8AF88FF8E5A074F8A7" box="[689,787,1865,1891]" country="Indonesia" name="Mamasa" pageId="33" pageNumber="434" stateProvince="Sulawesi Utara">Mamasa</location>
,
<date id="FFB8ADD38D53FF8AF91FF8E4A087F8A7" box="[801,992,1864,1891]" pageId="33" pageNumber="434" value="2009-08-15" valueMax="2009-08-27" valueMin="2009-08-15">
<collectingDate id="EFFC543B8D53FF8AF91FF8E4A087F8A7" box="[801,992,1864,1891]" pageId="33" pageNumber="434" value="2009-08-15" valueMax="2009-08-27" valueMin="2009-08-15">1527.VIII.2009</collectingDate>
</date>
, coll.
<collectorName id="26F3EEC58D53FF8AFE15F8E5A7F1F8A6" box="[1067,1174,1864,1891]" pageId="33" pageNumber="434">N. Kluge</collectorName>
&amp;
<collectorName id="26F3EEC58D53FF8AFEFCF8E5A65AF8A6" box="[1218,1341,1864,1891]" pageId="33" pageNumber="434">L. Sheyko</collectorName>
:
<specimenCount id="9D00409A8D53FF8AFF75F8E4A6FBF8A6" box="[1355,1436,1864,1890]" pageId="33" pageNumber="434" type="larva">1 larva</specimenCount>
(
<collectionCode id="ED1713D68D53FF8AFAA1F8C0A3B4F842" box="[159,211,1900,1926]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="33" pageNumber="434" type="Museum">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31CD8988D53FF89FAF9F83CA037FEA1" lastPageId="34" lastPageNumber="435" pageId="33" pageNumber="434" type="description">
<paragraph id="8BB98B138D53FF89FAF9F83CA037FEA1" blockId="33.[151,1437,1864,2035]" lastBlockId="34.[151,1437,151,357]" lastPageId="34" lastPageNumber="435" pageId="33" pageNumber="434">
<emphasis id="B97257018D53FF8AFAF9F83CA144F86E" bold="true" box="[199,547,1936,1962]" pageId="33" pageNumber="434">Description and comparison.</emphasis>
Femur with 2 apical setae (as in other
<taxonomicName id="4C06F0908D53FF8AF9DAF83CA709F86E" authorityName="Kluge" authorityYear="2016" box="[996,1134,1936,1962]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D53FF8AF9DAF83CA709F86E" box="[996,1134,1936,1962]" italics="true" pageId="33" pageNumber="434">Oculogaster</emphasis>
</taxonomicName>
); other characters of
<taxonomicName id="4C06F0908D53FF8AFF5AF83DA394F80A" authorityName="Kluge" authorityYear="2016" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D53FF8AFF5AF83DA394F80A" italics="true" pageId="33" pageNumber="434">Oculogaster</emphasis>
</taxonomicName>
(eggs and venation of pterostigma) unknown. Mouthparts longer and narrower than in other species (
<figureCitation id="133D97968D53FF8AFF55F818A39DF836" captionStart="FIGURES 138141" captionStartId="34.[151,264,1809,1833]" captionTargetBox="[151,1436,381,1785]" captionTargetId="figure-132@34.[151,1436,381,1785]" captionTargetPageId="34" captionText="FIGURES 138141. Larva of Procloeon (Oculogaster?) sp. «Sulawesi». 138, glossae and paraglossae; 139, maxilla (ds1ds3, dentisetae); 140, right mandible; 141, part of cercus." figureDoi="http://doi.org/10.5281/zenodo.4397927" httpUri="https://zenodo.org/record/4397927/files/figure.png" pageId="33" pageNumber="434">Figs 138140</figureCitation>
) (labial palps lost). Maxilla with 1st dentiseta stout, canine-like and bent toward canines («
<taxonomicName id="4C06F0908D53FF8AFF3BF875A62CF836" authorityName="Leach" authorityYear="1815" box="[1285,1355,2009,2034]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="33" pageNumber="434" phylum="Arthropoda" rank="genus">
<emphasis id="B97257018D53FF8AFF3BF875A62CF836" box="[1285,1355,2009,2034]" italics="true" pageId="33" pageNumber="434">Baetis</emphasis>
</taxonomicName>
-
<typeStatus id="54BD35B18D53FF8AFF6DF875A6E0F837" box="[1363,1415,2009,2035]" pageId="33" pageNumber="434">type</typeStatus>
») (
<figureCitation id="133D97968D50FF89FA9EFF3BA266FF75" box="[160,257,151,177]" captionStart="FIGURES 138141" captionStartId="34.[151,264,1809,1833]" captionTargetBox="[151,1436,381,1785]" captionTargetId="figure-132@34.[151,1436,381,1785]" captionTargetPageId="34" captionText="FIGURES 138141. Larva of Procloeon (Oculogaster?) sp. «Sulawesi». 138, glossae and paraglossae; 139, maxilla (ds1ds3, dentisetae); 140, right mandible; 141, part of cercus." figureDoi="http://doi.org/10.5281/zenodo.4397927" httpUri="https://zenodo.org/record/4397927/files/figure.png" pageId="34" pageNumber="435">Fig. 139</figureCitation>
), in contrast to other
<taxonomicName id="4C06F0908D50FF89FBCEFF3BA121FF75" authorityName="Leach" authorityYear="1815" box="[496,582,151,177]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, whose maxillae are of «
<taxonomicName id="4C06F0908D50FF89F9B6FF3BA0BEFF75" authorityName="Leach" authorityYear="1815" box="[904,985,151,177]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">
<emphasis id="B97257018D50FF89F9B6FF3BA0BEFF75" box="[904,985,151,177]" italics="true" pageId="34" pageNumber="435">Cloeon</emphasis>
</taxonomicName>
-
<typeStatus id="54BD35B18D50FF89F9DEFF3BA774FF75" box="[992,1043,151,177]" pageId="34" pageNumber="435">type</typeStatus>
» (
<bibRefCitation id="EF97F6E28D50FF89FE11FF3BA784FF75" author="Kluge, N. J." box="[1071,1251,151,177]" pageId="34" pageNumber="435" pagination="91 - 127" refId="ref18427" refString="Kluge, N. J. (2017) Contribution to the knowledge of Cloeodes Traver 1938 (Ephemeroptera, Baetidae). Zootaxa, 4319 (1), 91 - 127. https: // doi. org / 10.11646 / zootaxa. 4319.1.5" type="journal article" year="2017">Kluge 2017: 94</bibRefCitation>
). Maxillary palp 3-segmented, with 3rd segment much shorter than 2nd segment (in contrast to subequal in other described Asian
<taxonomicName id="4C06F0908D50FF89FAA9FF73A246FF3D" authorityName="Kluge" authorityYear="2016" box="[151,289,223,249]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="402" phylum="Arthropoda" rank="subGenus" subGenus="Oculogaster">
<emphasis id="B97257018D50FF89FAA9FF73A246FF3D" box="[151,289,223,249]" italics="true" pageId="34" pageNumber="435">Oculogaster</emphasis>
</taxonomicName>
). Hind protoptera absent. Denticles on claw minute, located in proximal part of claw only (as in
<taxonomicName id="4C06F0908D50FF89FFB6FF73A3BAFED9" authorityName="Kluge" authorityYear="2016" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="species" species="album">
<emphasis id="B97257018D50FF89FFB6FF73A3BAFED9" italics="true" pageId="34" pageNumber="435">P. album</emphasis>
</taxonomicName>
). Lateral denticles absent on all abdominal segments (in contrast to most other
<taxonomicName id="4C06F0908D50FF89FE65FEAFA7D6FED9" authorityName="Leach" authorityYear="1815" box="[1115,1201,259,285]" class="Insecta" family="Baetidae" genus="Cloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">Cloeon</taxonomicName>
/fg1, but similar to
<emphasis id="B97257018D50FF89FFBAFEAFA6F2FED8" box="[1412,1429,259,284]" italics="true" pageId="34" pageNumber="435">P</emphasis>
. (
<emphasis id="B97257018D50FF89FA9FFE8BA3D2FE85" box="[161,181,295,321]" italics="true" pageId="34" pageNumber="435">O</emphasis>
.) sp. «
<collectingCountry id="F311CB838D50FF89FB3FFE8BA2E1FE85" box="[257,390,295,321]" name="Philippines" pageId="34" pageNumber="435">Philippines</collectingCountry>
», below). Cerci with long lateral spines (
<figureCitation id="133D97968D50FF89F96BFE8BA0D2FE85" box="[853,949,295,321]" captionStart="FIGURES 138141" captionStartId="34.[151,264,1809,1833]" captionTargetBox="[151,1436,381,1785]" captionTargetId="figure-132@34.[151,1436,381,1785]" captionTargetPageId="34" captionText="FIGURES 138141. Larva of Procloeon (Oculogaster?) sp. «Sulawesi». 138, glossae and paraglossae; 139, maxilla (ds1ds3, dentisetae); 140, right mandible; 141, part of cercus." figureDoi="http://doi.org/10.5281/zenodo.4397927" httpUri="https://zenodo.org/record/4397927/files/figure.png" pageId="34" pageNumber="435">Fig. 141</figureCitation>
) (as in other
<taxonomicName id="4C06F0908D50FF89FE72FE8BA7A0FE86" authorityName="Kluge" authorityYear="2020" box="[1100,1223,295,322]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1). Tergalii VII as in other
<taxonomicName id="4C06F0908D50FF89FACBFEE7A20BFEA1" authorityName="Kluge" authorityYear="2020" box="[245,364,331,357]" class="Insecta" family="Baetidae" genus="Procloeon" kingdom="Animalia" order="Ephemeroptera" pageId="34" pageNumber="435" phylum="Arthropoda" rank="genus">Procloeon</taxonomicName>
/g1 (as in
<figureCitation id="133D97968D50FF89FBEBFEE7A151FEA1" box="[469,566,331,357]" captionStart="FIGURES 118126" captionStartId="30.[151,264,1969,1993]" captionTargetBox="[151,1430,424,1938]" captionTargetId="figure-126@30.[151,1436,417,1946]" captionTargetPageId="30" captionText="FIGURES 118126. Procloeon (Oculogaster) malabarensis sp. n. 118124, tergalii IVII (holotype); 125, genitalia of male imago (muscles not shown); 126, their subimaginal exuviae." figureDoi="http://doi.org/10.5281/zenodo.4397919" httpUri="https://zenodo.org/record/4397919/files/figure.png" pageId="34" pageNumber="435">Fig. 124</figureCitation>
), other tergalii unknown.
</paragraph>
</subSubSection>
</treatment>
</document>