310 lines
37 KiB
XML
310 lines
37 KiB
XML
<document id="0C26D3150FB8EC42730DA5CA37343EF7" ID-CLB-Dataset="29442" ID-DOI="10.11646/zootaxa.4472.3.9" ID-GBIF-Dataset="e7a395e6-38d8-4cbc-8365-6e883cd0d060" ID-ISSN="1175-5326" ID-Zenodo-Dep="1440349" ID-ZooBank="DE17EA66-E8B8-4B64-9DDF-40D262A88CFB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538372643219" checkinUser="plazi" docAuthor="Siewert, Ricardo Russo, Madruga, Janaína, Dolibaina, Diego Rodrigo, Mielke, Olaf Hermann Hendrik & Casagrande, Mirna Martins" docDate="2018" docId="03AF30658E608047A0B1A422FE463FFE" docLanguage="en" docName="zootaxa.4472.3.9.pdf" docOrigin="Zootaxa 4472 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drephalys (Drephalys) citrinus Siewert & Madruga & Dolibaina & Mielke & Casagrande 2018, sp. n." docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="FF96481D8E638042A026A20FFFC23971" masterDocTitle="Hidden in plain sight: a morphological study revealing three new species of the skipper genus Drephalys Watson, 1893 (Lepidoptera: Hesperiidae) from Brazil" masterLastPageNumber="580" masterPageNumber="573" pageNumber="576" updateTime="1698559144843" updateUser="ExternalLinkService">
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<mods:title id="CBD853D9BBD47C8AA600AF44F95D90EB">Hidden in plain sight: a morphological study revealing three new species of the skipper genus Drephalys Watson, 1893 (Lepidoptera: Hesperiidae) from Brazil</mods:title>
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<mods:namePart id="D74B9CBB93EA1A520EF76F44743D7EB1">Siewert, Ricardo Russo</mods:namePart>
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<mods:namePart id="DE8BF720567033D519AFDF5F518C665C">Madruga, Janaína</mods:namePart>
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<mods:namePart id="D92DEAC82A4C038A9C2B01363254A069">Dolibaina, Diego Rodrigo</mods:namePart>
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<mods:namePart id="3110B6F6820FCC8AAC302D39F1B7A045">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<mods:namePart id="61C66E98A272B8818B4F8EE6D9057F63">Casagrande, Mirna Martins</mods:namePart>
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<mods:title id="9504CD65EDCB96A15FEEEA0EE8D47948">Zootaxa</mods:title>
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<mods:date id="671B084C18FD0551133838190085A7E8">2018</mods:date>
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<mods:number id="BC044602748CFB923B542911B723D8A4">2018-09-11</mods:number>
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<mods:number id="DAE2CDBDA8E1219A45DC2D3AB7C6C63E">4472</mods:number>
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<treatment id="03AF30658E608047A0B1A422FE463FFE" ID-DOI="http://doi.org/10.5281/zenodo.6494470" ID-GBIF-Taxon="148421426" ID-Zenodo-Dep="6494470" LSID="urn:lsid:plazi:treatment:03AF30658E608047A0B1A422FE463FFE" httpUri="http://treatment.plazi.org/id/03AF30658E608047A0B1A422FE463FFE" lastPageId="5" lastPageNumber="578" pageId="3" pageNumber="576">
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<subSubSection id="C31CD2F88E608041A0B1A422FE933F19" pageId="3" pageNumber="576" type="nomenclature">
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<paragraph id="8BB981738E608041A0B1A422FBAB3F36" blockId="3.[151,1129,1581,1641]" box="[151,1129,1581,1607]" pageId="3" pageNumber="576">
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<heading id="D0F1361F8E608041A0B1A422FBAB3F36" bold="true" box="[151,1129,1581,1607]" fontSize="11" level="1" pageId="3" pageNumber="576" reason="1">
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<emphasis id="B9725D618E608041A0B1A422FBAB3F36" bold="true" box="[151,1129,1581,1607]" pageId="3" pageNumber="576">
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<taxonomicName id="4C06FAF08E608041A0B1A422FBD93F36" authority="Madruga, Siewert, Mielke & Casagrande" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[151,1051,1581,1607]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov." subGenus="Drephalys">
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<emphasis id="B9725D618E608041A0B1A422FDCF3F36" bold="true" box="[151,525,1581,1607]" italics="true" pageId="3" pageNumber="576">Drephalys (Drephalys) citrinus</emphasis>
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Madruga, Siewert, Mielke & Casagrande
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</taxonomicName>
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,
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<taxonomicNameLabel id="A241E01A8E608041A401A422FBAB3F36" box="[1063,1129,1581,1607]" pageId="3" pageNumber="576" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB981738E608041A0B1A45FFE933F19" blockId="3.[151,1129,1581,1641]" box="[151,337,1616,1641]" pageId="3" pageNumber="576">
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(
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<figureCitation id="133D9DF68E608041A0B9A45FFEC13F18" box="[159,259,1616,1641]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1765,1788]" captionTargetBox="[198,1387,221,1717]" captionTargetId="figure@2.[198,1388,198,1717]" captionTargetPageId="2" captionText="FIGURES 1–6. Species of Drephalys in dorsal and ventral views. 1–2, D. dracarys sp. n. 3–4, D. electrinus sp. n. 5–6, D. citrinus sp. n." httpUri="https://zenodo.org/record/1440351/files/figure.png" pageId="3" pageNumber="576">Figs 5–6</figureCitation>
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,
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<figureCitation id="133D9DF68E608041A129A45FFEDD3F19" box="[271,287,1616,1640]" captionStart="FIGURES 7–9" captionStartId="4.[151,264,1161,1184]" captionTargetBox="[151,1437,199,1143]" captionTargetId="figure@4.[151,1437,199,1143]" captionTargetPageId="4" captionText="FIGURES 7–9. Male genitalia of Drephalys species. Tegumen, uncus, gnathos and saccus in lateral view; right valva in left, right and dorsal views; tegumen and uncus in dorsal view; aedeagus in left lateral and dorsal views; detail of the everted cornuti. 7. D. dracarys sp. n., 8. D. electrinus sp. n., 9. D. citrinus sp. n." httpUri="https://zenodo.org/record/1440353/files/figure.png" pageId="3" pageNumber="576">9</figureCitation>
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,
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<figureCitation id="133D9DF68E608041A10DA45FFE8A3F19" box="[299,328,1616,1640]" captionStart="FIGURE 13" captionStartId="6.[151,250,1536,1558]" captionTargetBox="[150,1437,761,1525]" captionTargetId="figure@6.[150,1437,761,1525]" captionTargetPageId="6" captionText="FIGURE 13. Distribution map of Drephalys species." httpUri="https://zenodo.org/record/1440357/files/figure.png" pageId="3" pageNumber="576">13</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31CD2F88E608041A0B1A49CFB563E41" pageId="3" pageNumber="576" type="diagnosis">
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<paragraph id="8BB981738E608041A0B1A49CFB563E41" blockId="3.[151,1436,1683,1841]" pageId="3" pageNumber="576">
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<emphasis id="B9725D618E608041A0B1A49CFED33FDD" bold="true" box="[151,273,1683,1708]" pageId="3" pageNumber="576">Diagnosis.</emphasis>
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<taxonomicName id="4C06FAF08E608041A13EA49CFE2A3FDA" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[280,488,1683,1708]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov.">
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<emphasis id="B9725D618E608041A13EA49CFE2A3FDA" box="[280,488,1683,1708]" italics="true" pageId="3" pageNumber="576">Drephalys citrinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A241E01A8E608041A1C9A49CFDED3FDD" box="[495,559,1683,1708]" pageId="3" pageNumber="576" rank="species">
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<emphasis id="B9725D618E608041A1C9A49CFDED3FDD" bold="true" box="[495,559,1683,1708]" pageId="3" pageNumber="576">sp. n.</emphasis>
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</taxonomicNameLabel>
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is nearly identical to
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<taxonomicName id="4C06FAF08E608041A306A49BFBB13FDD" authority="(Hewitson, 1867)" authorityName="Hewitson" authorityYear="1867" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[800,1139,1683,1708]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="phoenice">
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<emphasis id="B9725D618E608041A306A49BFC6A3FDD" box="[800,936,1683,1708]" italics="true" pageId="3" pageNumber="576">D. phoenice</emphasis>
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(Hewitson, 1867)
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</taxonomicName>
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and
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<taxonomicName id="4C06FAF08E608041A48FA49BFA823FDD" authorityName="Bell" authorityYear="1942" box="[1193,1344,1683,1708]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="heraclides">
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<emphasis id="B9725D618E608041A48FA49BFA823FDD" box="[1193,1344,1683,1708]" italics="true" pageId="3" pageNumber="576">D. heraclides</emphasis>
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</taxonomicName>
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in wing markings and in the wider white transversal band on the ventral hind wings. It is distinguished from
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<taxonomicName id="4C06FAF08E608041A4D5A4BAFAB83FBC" baseAuthorityName="Hewitson" baseAuthorityYear="1867" box="[1267,1402,1716,1741]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="phoenice">
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<emphasis id="B9725D618E608041A4D5A4BAFAB83FBC" box="[1267,1402,1716,1741]" italics="true" pageId="3" pageNumber="576">D. phoenice</emphasis>
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</taxonomicName>
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by the reduced lilac reflections on the ventral wings, and from
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<taxonomicName id="4C06FAF08E608041A363A4D9FC1C3F9F" authorityName="Bell" authorityYear="1942" box="[837,990,1749,1774]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="heraclides">
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<emphasis id="B9725D618E608041A363A4D9FC1C3F9F" box="[837,990,1749,1774]" italics="true" pageId="3" pageNumber="576">D. heraclides</emphasis>
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</taxonomicName>
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by the white band almost reaching the costal margin. Furthermore,
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<taxonomicName id="4C06FAF08E608041A1FFA4F7FD903E7E" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[473,594,1784,1807]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov.">
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<emphasis id="B9725D618E608041A1FFA4F7FD903E7E" box="[473,594,1784,1807]" italics="true" pageId="3" pageNumber="576">D. citrinus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A241E01A8E608041A27CA4F9FD593E7E" box="[602,667,1782,1807]" pageId="3" pageNumber="576" rank="species">
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<emphasis id="B9725D618E608041A27CA4F9FD593E7E" bold="true" box="[602,667,1782,1807]" pageId="3" pageNumber="576">sp. n.</emphasis>
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</taxonomicNameLabel>
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has a short uncus, with truncate arms, and not deeply bifid as in
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<taxonomicName id="4C06FAF08E608041A5A7A4F7FF3E3E40" baseAuthorityName="Hewitson" baseAuthorityYear="1867" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="576" phylum="Arthropoda" rank="species" species="phoenice">
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<emphasis id="B9725D618E608041A5A7A4F7FF3E3E40" italics="true" pageId="3" pageNumber="576">D. phoenice</emphasis>
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</taxonomicName>
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(
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<figureCitation id="133D9DF68E608041A12DA517FE9E3E41" box="[267,348,1816,1841]" captionStart="FIGURES 10–12" captionStartId="5.[151,264,1268,1291]" captionTargetBox="[151,1436,193,1248]" captionTargetId="figure@5.[151,1436,193,1248]" captionTargetPageId="5" captionText="FIGURES 10–12. Male genitalia of Drephalys species. Structures as presented in figures 7–9. 10. D. heraclides. 11. D. phoenice. 12. D. phoenicoides." httpUri="https://zenodo.org/record/1440355/files/figure.png" pageId="3" pageNumber="576">Fig. 11</figureCitation>
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), and the costa of the valva has a short and narrow distal process (
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<figureCitation id="133D9DF68E608041A464A517FB443E41" box="[1090,1158,1816,1841]" captionStart="FIGURES 7–9" captionStartId="4.[151,264,1161,1184]" captionTargetBox="[151,1437,199,1143]" captionTargetId="figure@4.[151,1437,199,1143]" captionTargetPageId="4" captionText="FIGURES 7–9. Male genitalia of Drephalys species. Tegumen, uncus, gnathos and saccus in lateral view; right valva in left, right and dorsal views; tegumen and uncus in dorsal view; aedeagus in left lateral and dorsal views; detail of the everted cornuti. 7. D. dracarys sp. n., 8. D. electrinus sp. n., 9. D. citrinus sp. n." httpUri="https://zenodo.org/record/1440353/files/figure.png" pageId="3" pageNumber="576">Fig. 9</figureCitation>
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).
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF79D1FB8E678046A0B1A686FCB43DAD" httpUri="https://zenodo.org/record/1440353/files/figure.png" pageId="4" pageNumber="577" startId="4.[151,264,1161,1184]" targetBox="[151,1437,199,1143]" targetPageId="4">
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<paragraph id="8BB981738E678046A0B1A686FCB43DAD" blockId="4.[151,1436,1161,1245]" pageId="4" pageNumber="577">
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<emphasis id="B9725D618E678046A0B1A686FEF93DEF" bold="true" box="[151,315,1161,1184]" pageId="4" pageNumber="577">FIGURES 7–9.</emphasis>
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Male genitalia of
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<taxonomicName id="4C06FAF08E678046A1DCA686FDA13DEE" authorityName="Watson" authorityYear="1893" box="[506,611,1161,1183]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725D618E678046A1DCA686FDA13DEE" box="[506,611,1161,1183]" italics="true" pageId="4" pageNumber="577">Drephalys</emphasis>
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</taxonomicName>
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species. Tegumen, uncus, gnathos and saccus in lateral view; right valva in left, right and dorsal views; tegumen and uncus in dorsal view; aedeagus in left lateral and dorsal views; detail of the everted cornuti. 7.
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<taxonomicName id="4C06FAF08E678046A122A6C7FEBF3DAC" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[260,381,1223,1245]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="dracarys" status="sp. nov.">
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<emphasis id="B9725D618E678046A122A6C7FEBF3DAC" box="[260,381,1223,1245]" italics="true" pageId="4" pageNumber="577">D. dracarys</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A241E01A8E678046A1A2A6C8FE7E3DAD" box="[388,444,1223,1244]" pageId="4" pageNumber="577" rank="species">
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||
<emphasis id="B9725D618E678046A1A2A6C8FE7E3DAD" bold="true" box="[388,444,1223,1244]" pageId="4" pageNumber="577">sp. n.</emphasis>
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||
</taxonomicNameLabel>
|
||
, 8.
|
||
<taxonomicName id="4C06FAF08E678046A1C5A6C7FDA43DAC" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[483,614,1223,1245]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="electrinus" status="sp. nov.">
|
||
<emphasis id="B9725D618E678046A1C5A6C7FDA43DAC" box="[483,614,1223,1245]" italics="true" pageId="4" pageNumber="577">D. electrinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241E01A8E678046A24AA6C8FD663DAD" box="[620,676,1223,1244]" pageId="4" pageNumber="577" rank="species">
|
||
<emphasis id="B9725D618E678046A24AA6C8FD663DAD" bold="true" box="[620,676,1223,1244]" pageId="4" pageNumber="577">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, 9.
|
||
<taxonomicName id="4C06FAF08E678046A2EDA6C7FCF53DAC" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[715,823,1224,1245]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov.">
|
||
<emphasis id="B9725D618E678046A2EDA6C7FCF53DAC" box="[715,823,1224,1245]" italics="true" pageId="4" pageNumber="577">D. citrinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241E01A8E678046A31BA6C8FCB43DAD" box="[829,886,1223,1244]" pageId="4" pageNumber="577" rank="species">sp. n.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31CD2F88E678046A0E1A706FD293F8E" pageId="4" pageNumber="577" type="description">
|
||
<paragraph id="8BB981738E678046A0E1A706FD293F8E" blockId="4.[151,1436,1289,2007]" pageId="4" pageNumber="577">
|
||
<emphasis id="B9725D618E678046A0E1A706FE623C53" bold="true" box="[199,416,1289,1314]" pageId="4" pageNumber="577">Description, male.</emphasis>
|
||
<emphasis id="B9725D618E678046A180A706FE213C53" box="[422,483,1289,1314]" italics="true" pageId="4" pageNumber="577">Head</emphasis>
|
||
: nudum of 30 segments (n=1).
|
||
<emphasis id="B9725D618E678046A365A706FC563C53" box="[835,916,1289,1314]" italics="true" pageId="4" pageNumber="577">Thorax</emphasis>
|
||
: as in
|
||
<taxonomicName id="4C06FAF08E678046A3FBA704FB9B3C53" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[989,1113,1289,1314]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="dacarys" status="sp. nov.">
|
||
<emphasis id="B9725D618E678046A3FBA704FB9B3C53" box="[989,1113,1289,1314]" italics="true" pageId="4" pageNumber="577">D. dacarys</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241E01A8E678046A479A706FB5C3C53" box="[1119,1182,1289,1314]" pageId="4" pageNumber="577" rank="species">
|
||
<emphasis id="B9725D618E678046A479A706FB5C3C53" bold="true" box="[1119,1182,1289,1314]" pageId="4" pageNumber="577">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B9725D618E678046A483A704FA563C53" box="[1189,1428,1289,1314]" italics="true" pageId="4" pageNumber="577">Forewing, upper side</emphasis>
|
||
: length
|
||
<quantity id="4CFE2C968E678046A0C3A721FEF73C34" box="[229,309,1326,1350]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="577" unit="mm" value="20.0">20 mm</quantity>
|
||
(n=1); costal fold absent; distal hyaline spot in discal cell wider than in previous species; one large and circular discal yellow spot in CuA2-2A, reaches CuA2; three apical hyaline spots in R3-R4, R4-R5, and R5-M1 longer than in previous species.
|
||
<emphasis id="B9725D618E678046A22EA776FD303CE0" box="[520,754,1400,1425]" italics="true" pageId="4" pageNumber="577">Forewing, underside</emphasis>
|
||
: as upper side, except for the lighter colors and the whitish area from CuA1 to the anal margin.
|
||
<emphasis id="B9725D618E678046A204A793FCDB3CC4" box="[546,793,1436,1461]" italics="true" pageId="4" pageNumber="577">Hind wing, upper side</emphasis>
|
||
: post-discal rectangular yellow spot in Rs-M1 longer than in previous species; post-discal yellow spot extends to 2A-3A; fringes mixed with brown and yellowish.
|
||
<emphasis id="B9725D618E678046A542A7CCFE893F71" italics="true" pageId="4" pageNumber="577">Hind wing, underside</emphasis>
|
||
: large white transversal band, from costal to anal margin; white submarginal markings in CuA2- 2A; anal margin white except on tornus.
|
||
<emphasis id="B9725D618E678046A24BA401FD153F56" box="[621,727,1550,1575]" italics="true" pageId="4" pageNumber="577">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="133D9DF68E678046A2CEA401FCED3F56" box="[744,815,1550,1575]" captionStart="FIGURES 7–9" captionStartId="4.[151,264,1161,1184]" captionTargetBox="[151,1437,199,1143]" captionTargetId="figure@4.[151,1437,199,1143]" captionTargetPageId="4" captionText="FIGURES 7–9. Male genitalia of Drephalys species. Tegumen, uncus, gnathos and saccus in lateral view; right valva in left, right and dorsal views; tegumen and uncus in dorsal view; aedeagus in left lateral and dorsal views; detail of the everted cornuti. 7. D. dracarys sp. n., 8. D. electrinus sp. n., 9. D. citrinus sp. n." httpUri="https://zenodo.org/record/1440353/files/figure.png" pageId="4" pageNumber="577">Fig. 9</figureCitation>
|
||
): tegumen rectangular, narrow anteriorly. Uncus bifid, “U”-shaped; distal arms short, robust and distally truncated. Disto-lateral margin of gnathos ribbed with rounded tip. Costa of valva with a narrow and long process curved inwards with two pointed tips; harpe not projected; ampulla undifferentiated from harpe, producing a serrated distal process curved up and inwards. Aedeagus thick and shorther than in previous species; ejaculatory bulb opening ovoid and broader than in previous species; distal margin ventro-laterally excavated, with a dorsal and small spine in the right distal margin; vesica with six needlelike cornuti of two different sizes.
|
||
<emphasis id="B9725D618E678046A23FA4E9FDAF3F8E" box="[537,621,1766,1791]" italics="true" pageId="4" pageNumber="577">Female</emphasis>
|
||
: unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD2F88E678046A0E1A505FDDA3E1B" pageId="4" pageNumber="577" type="materials_examined">
|
||
<paragraph id="8BB981738E678046A0E1A505FDDA3E1B" blockId="4.[151,1436,1289,2007]" pageId="4" pageNumber="577">
|
||
<emphasis id="B9725D618E678046A0E1A505FEB53E52" bold="true" box="[199,375,1802,1827]" pageId="4" pageNumber="577">
|
||
<typeStatus id="54BD3FD18E678046A0E1A505FEC03E52" box="[199,258,1802,1827]" pageId="4" pageNumber="577">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54BD3FD18E678046A1A6A505FE2A3E52" box="[384,488,1802,1827]" pageId="4" pageNumber="577" type="holotype">Holotype</typeStatus>
|
||
male with the following labels: /
|
||
<typeStatus id="54BD3FD18E678046A354A505FBDA3E53" box="[882,1048,1802,1826]" pageId="4" pageNumber="577" type="holotype">HOLOTYPUS</typeStatus>
|
||
/ Teffé [
|
||
<collectingRegion id="49C24F918E678046A45BA505FB3B3E53" box="[1149,1273,1802,1826]" country="Brazil" name="Amazonas" pageId="4" pageNumber="577">Amazonas</collectingRegion>
|
||
,
|
||
<collectingCountry id="F311C1E38E678046A520A505FA8F3E52" box="[1286,1357,1802,1827]" name="Brazil" pageId="4" pageNumber="577">Brazil</collectingCountry>
|
||
] / OM 12.853 / Gen. Prep. Mielke 1996 /
|
||
<typeStatus id="54BD3FD18E678046A26CA521FD7D3E36" box="[586,703,1838,1863]" pageId="4" pageNumber="577" type="holotype">Holotypus</typeStatus>
|
||
<taxonomicName id="4C06FAF08E678046A2EBA521FED83E1A" authority="Madruga, Siewert, Mielke & Casagrande" authorityName="Madruga, Siewert, Mielke & Casagrande" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="577" phylum="Arthropoda" rank="species" species="citrinus" subGenus="Drephalys">
|
||
<emphasis id="B9725D618E678046A2EBA521FBF53E37" box="[717,1079,1838,1863]" italics="true" pageId="4" pageNumber="577">Drephalys (Drephalys) citrinus</emphasis>
|
||
Madruga, Siewert, Mielke & Casagrande
|
||
</taxonomicName>
|
||
/. Deposited at
|
||
<collectionCode id="ED1719B68E678046A1ECA55CFDD63E1B" box="[458,532,1875,1898]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="4" pageNumber="577">DZUP</collectionCode>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD2F88E678046A0E1A579FC563EC2" pageId="4" pageNumber="577" type="etymology">
|
||
<paragraph id="8BB981738E678046A0E1A579FC563EC2" blockId="4.[151,1436,1289,2007]" pageId="4" pageNumber="577">
|
||
<emphasis id="B9725D618E678046A0E1A579FE8C3EFE" bold="true" box="[199,334,1910,1935]" pageId="4" pageNumber="577">Etymology.</emphasis>
|
||
The name is a reference to the yellow-colored quartz, citrino. The epithet is from the Latin
|
||
<emphasis id="B9725D618E678046A573A578FA593EFF" box="[1365,1435,1911,1934]" italics="true" pageId="4" pageNumber="577">citrus,</emphasis>
|
||
pure yellow, in reference to the yellow patches on the ventral wings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD2F88E678046A0E1A5B1FC3B3EA6" box="[199,1017,1982,2007]" pageId="4" pageNumber="577" type="distribution">
|
||
<paragraph id="8BB981738E678046A0E1A5B1FC3B3EA6" blockId="4.[151,1436,1289,2007]" box="[199,1017,1982,2007]" pageId="4" pageNumber="577">
|
||
<emphasis id="B9725D618E678046A0E1A5B1FEA23EA6" bold="true" box="[199,352,1982,2007]" pageId="4" pageNumber="577">Distribution.</emphasis>
|
||
This species is currently known only from its
|
||
<typeStatus id="54BD3FD18E678046A34EA5B0FC5A3EA6" box="[872,920,1983,2007]" pageId="4" pageNumber="577">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF79D1FB8E668047A0B1A6FBFE133C5B" httpUri="https://zenodo.org/record/1440355/files/figure.png" pageId="5" pageNumber="578" startId="5.[151,264,1268,1291]" targetBox="[151,1436,193,1248]" targetPageId="5">
|
||
<paragraph id="8BB981738E668047A0B1A6FBFE133C5B" blockId="5.[151,1437,1268,1322]" pageId="5" pageNumber="578">
|
||
<emphasis id="B9725D618E668047A0B1A6FBFE9A3C7B" bold="true" box="[151,344,1268,1291]" pageId="5" pageNumber="578">FIGURES 10–12.</emphasis>
|
||
Male genitalia of
|
||
<taxonomicName id="4C06FAF08E668047A204A6FAFD493C7A" authorityName="Watson" authorityYear="1893" box="[546,651,1269,1291]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725D618E668047A204A6FAFD493C7A" box="[546,651,1269,1291]" italics="true" pageId="5" pageNumber="578">Drephalys</emphasis>
|
||
</taxonomicName>
|
||
species. Structures as presented in figures 7–9. 10.
|
||
<taxonomicName id="4C06FAF08E668047A49AA6F9FA8B3C7A" authorityName="Bell" authorityYear="1942" box="[1212,1353,1269,1291]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="heraclides">
|
||
<emphasis id="B9725D618E668047A49AA6F9FA8B3C7A" box="[1212,1353,1269,1291]" italics="true" pageId="5" pageNumber="578">D. heraclides</emphasis>
|
||
</taxonomicName>
|
||
. 11.
|
||
<taxonomicName id="4C06FAF08E668047A5A5A6F9FF313C5B" baseAuthorityName="Hewitson" baseAuthorityYear="1867" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="phoenice">
|
||
<emphasis id="B9725D618E668047A5A5A6F9FF313C5B" italics="true" pageId="5" pageNumber="578">D. phoenice</emphasis>
|
||
</taxonomicName>
|
||
. 12.
|
||
<taxonomicName id="4C06FAF08E668047A103A71AFE0F3C5B" baseAuthorityName="Mabille & Boullet" baseAuthorityYear="1919" box="[293,461,1300,1322]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="phoenicoides">D. phoenicoides</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31CD2F88E668047A0E1A759FE463FFE" pageId="5" pageNumber="578" type="discussion">
|
||
<paragraph id="8BB981738E668047A0E1A759FE463FFE" blockId="5.[151,1436,1366,1679]" pageId="5" pageNumber="578">
|
||
<emphasis id="B9725D618E668047A0E1A759FE8D3C1E" bold="true" box="[199,335,1366,1391]" pageId="5" pageNumber="578">Comments.</emphasis>
|
||
The arm of the gnathos ribbed at the tip of
|
||
<taxonomicName id="4C06FAF08E668047A352A759FB883C1E" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[884,1098,1366,1391]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov.">
|
||
<emphasis id="B9725D618E668047A352A759FB883C1E" box="[884,1098,1366,1391]" italics="true" pageId="5" pageNumber="578">Drephalys citrinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241E01A8E668047A470A759FB5E3C1E" box="[1110,1180,1366,1391]" pageId="5" pageNumber="578" rank="species">
|
||
<emphasis id="B9725D618E668047A470A759FB5E3C1E" bold="true" box="[1110,1180,1366,1391]" pageId="5" pageNumber="578">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is also present in
|
||
<taxonomicName id="4C06FAF08E668047A5A7A757FECE3CE2" authorityName="Bell" authorityYear="1942" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="heraclides">
|
||
<emphasis id="B9725D618E668047A5A7A757FECE3CE2" italics="true" pageId="5" pageNumber="578">D. heraclides</emphasis>
|
||
</taxonomicName>
|
||
, suggesting the latter is a close relative of the former.
|
||
<taxonomicName id="4C06FAF08E668047A34BA773FC273CE2" authority="Siewert & Madruga & Dolibaina & Mielke & Casagrande, 2018" authorityName="Siewert & Madruga & Dolibaina & Mielke & Casagrande" authorityYear="2018" box="[877,997,1404,1427]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="citrinus" status="sp. nov.">
|
||
<emphasis id="B9725D618E668047A34BA773FC273CE2" box="[877,997,1404,1427]" italics="true" pageId="5" pageNumber="578">D. citrinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A241E01A8E668047A3CAA775FBEE3CE2" box="[1004,1068,1402,1427]" pageId="5" pageNumber="578" rank="species">
|
||
<emphasis id="B9725D618E668047A3CAA775FBEE3CE2" bold="true" box="[1004,1068,1402,1427]" pageId="5" pageNumber="578">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, together with
|
||
<taxonomicName id="4C06FAF08E668047A4F3A773FAAF3CE2" authorityName="Bell" authorityYear="1942" box="[1237,1389,1402,1427]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="heraclides">
|
||
<emphasis id="B9725D618E668047A4F3A773FAAF3CE2" box="[1237,1389,1402,1427]" italics="true" pageId="5" pageNumber="578">D. heraclides</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06FAF08E668047A0B1A7AFFE523CC6" authority="Evans, 1952" authorityName="Evans" authorityYear="1952" box="[151,400,1438,1463]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="opifex">
|
||
<emphasis id="B9725D618E668047A0B1A7AFFF3F3CC6" box="[151,253,1439,1463]" italics="true" pageId="5" pageNumber="578">D. opifex</emphasis>
|
||
Evans, 1952
|
||
</taxonomicName>
|
||
, are the only yellow-checkered species belonging to the subgenus
|
||
<taxonomicName id="4C06FAF08E668047A454A791FB243CC6" authorityName="Watson" authorityYear="1893" box="[1138,1254,1438,1463]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725D618E668047A454A791FB243CC6" box="[1138,1254,1438,1463]" italics="true" pageId="5" pageNumber="578">Drephalys</emphasis>
|
||
</taxonomicName>
|
||
without a costal fold on the forewing (
|
||
<figureCitation id="133D9DF68E668047A1B5A7CDFE1B3CAA" box="[403,473,1474,1499]" captionStart="FIGURES 1–6" captionStartId="2.[151,264,1765,1788]" captionTargetBox="[198,1387,221,1717]" captionTargetId="figure@2.[198,1388,198,1717]" captionTargetPageId="2" captionText="FIGURES 1–6. Species of Drephalys in dorsal and ventral views. 1–2, D. dracarys sp. n. 3–4, D. electrinus sp. n. 5–6, D. citrinus sp. n." httpUri="https://zenodo.org/record/1440351/files/figure.png" pageId="5" pageNumber="578">Fig. 5</figureCitation>
|
||
). The absence of the forewing costal fold in this subgenus is also shared by
|
||
<taxonomicName id="4C06FAF08E668047A56EA7CBFEE03C8E" authority="Evans, 1952" authorityName="Evans" authorityYear="1952" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="olva">
|
||
<emphasis id="B9725D618E668047A56EA7CBFA5E3CAA" box="[1352,1436,1474,1499]" italics="true" pageId="5" pageNumber="578">D. olva</emphasis>
|
||
Evans, 1952
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06FAF08E668047A17CA7E7FD9A3C8E" authority="Evans, 1952" authorityName="Evans" authorityYear="1952" box="[346,600,1510,1535]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="olvina">
|
||
<emphasis id="B9725D618E668047A17CA7E7FE013C8E" box="[346,451,1510,1535]" italics="true" pageId="5" pageNumber="578">D. olvina</emphasis>
|
||
Evans, 1952
|
||
</taxonomicName>
|
||
, while
|
||
<taxonomicName id="4C06FAF08E668047A28CA7E7FC703C8E" authority="Burns, 2000" authorityName="Burns" authorityYear="2000" box="[682,946,1510,1535]" class="Insecta" family="Hesperiidae" genus="Drephalys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="578" phylum="Arthropoda" rank="species" species="kidonoi">
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<emphasis id="B9725D618E668047A28CA7E7FCE33C8E" box="[682,801,1510,1535]" italics="true" pageId="5" pageNumber="578">D. kidonoi</emphasis>
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<bibRefCitation id="EF97FC828E668047A30EA7E7FC703C8E" box="[808,946,1511,1535]" pageId="5" pageNumber="578" refId="ref4060" refString="Burns, J. M. & Janzen, D. H. (2000) Drephalys: division of this showy Neotropical genus, plus a new species and the immatures and food plants of two species from Costa Rican dry forest (Hesperiidae: Pyrginae). Journal of the Lepidopterists' Society, 53, 77 - 89." type="journal article">Burns, 2000</bibRefCitation>
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</taxonomicName>
|
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has a vestigial costal fold (
|
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<bibRefCitation id="EF97FC828E668047A4C0A7E8FF163F52" pageId="5" pageNumber="578" refId="ref4060" refString="Burns, J. M. & Janzen, D. H. (2000) Drephalys: division of this showy Neotropical genus, plus a new species and the immatures and food plants of two species from Costa Rican dry forest (Hesperiidae: Pyrginae). Journal of the Lepidopterists' Society, 53, 77 - 89." type="journal article">Burns & Janzen 2000</bibRefCitation>
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). The evaluation of this character is useful to differentiate the males of species with similar wing phenotypes, as is the case with species possessing yellow spots on the dorsal wings. However, studies of other features are required for species-level determinations, such as the development of the ventral hind wing white band and details of the male genitalia.
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</paragraph>
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</subSubSection>
|
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</treatment>
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</document> |