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<document id="EE1C60C8BB6380B6DD26C203289A3DF2" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFEEE5025098825B610EFDD5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) densihirtus Assing 2017, spec. nov." docType="treatment" docUuid="E47C7587-9B06-4CD1-821D-6074284CA5BB" docUuidSource="ZooBank" docVersion="12" lastPageNumber="165" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="164" updateTime="1699471407760" updateUser="ExternalLinkService">
<mods:mods id="D9FF04AB79ADE091AA9491222313FF0F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D40A4EC3ABDA26BA1A4CE577FF0E9E80">
<mods:title id="DB2AEDF300936D8CF1E0D59FE84A9858">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
<mods:name id="0A33C2C3855C74A024559DF80D185121" type="personal">
<mods:role id="5A9B488ADC9EF4E5BA3E012CBF6B9A23">
<mods:roleTerm id="8E9FAAEC066BA591DDB88A9370540C15">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA5DA5F5846FE8C767EC1C5C415E73B2">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="11A7008499844A38A00DB8BC2379B309">text</mods:typeOfResource>
<mods:relatedItem id="968309C225CF550A35D862A5FF694993" type="host">
<mods:titleInfo id="82AD03DA286CC9C0E40F47AD2FE52C4D">
<mods:title id="6C8FCF373FFFE80E9180222FA9DEB664">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="76757AF69BC153BB398AE7DB8BA2424C">
<mods:date id="ED029853AECC16FF5F2E662C6FC06A33">2017</mods:date>
<mods:detail id="182250329FCE680850286381CCF40631" type="pubDate">
<mods:number id="38064329DF62228F079638D3E077BDAA">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="95D92D9087A659A612269D401C7068C3" type="volume">
<mods:number id="0648228727EA01296FDE89E6D8808DDB">67</mods:number>
</mods:detail>
<mods:detail id="3A07F1F6677C717E11E9B1CD0E88FE6E" type="issue">
<mods:number id="BEECF6F77AE973903D0606CC4228F5CC">1</mods:number>
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<mods:start id="3BAC6511EF5DC8FB03B67142B348BAB2">117</mods:start>
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<mods:classification id="0C15E621D6AA179D456D419D30FE7FE2">journal article</mods:classification>
<mods:identifier id="B6863BFFEB09AD72888AC46702FFD814" type="CLB-Dataset">2472</mods:identifier>
<mods:identifier id="48EF70245941EEA45C18A1A8A27A1776" type="DOI">10.21248/contrib.entomol.67.1.117-192</mods:identifier>
<mods:identifier id="5EE7A3A971195084CC19F8A1921574F3" type="GBIF-Dataset">39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43</mods:identifier>
<mods:identifier id="48BF5215D311C6AFB50A0D06BBBE84A3" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="918542372A70235627D1D740B7DC761E" type="Zenodo-Dep">5742363</mods:identifier>
<mods:identifier id="C67A019068B6B5B3FC5A243B696C84C9" type="ZooBank">FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE</mods:identifier>
</mods:mods>
<treatment id="03A787BAFFEEE5025098825B610EFDD5" ID-DOI="http://doi.org/10.5281/zenodo.5888535" ID-GBIF-Taxon="192498194" ID-Zenodo-Dep="5888535" LSID="urn:lsid:zoobank.org:act:E47C7587-9B06-4CD1-821D-6074284CA5BB" httpUri="http://treatment.plazi.org/id/03A787BAFFEEE5025098825B610EFDD5" lastPageId="48" lastPageNumber="165" pageId="47" pageNumber="164">
<subSubSection id="C3146527FFEEE51D5098825B61CCFA3B" pageId="47" pageNumber="164" type="nomenclature">
<paragraph id="8BB136ACFFEEE51D5098825B612DFADA" blockId="47.[189,711,1363,1455]" box="[189,550,1363,1390]" pageId="47" pageNumber="164">
<heading id="D0F981C0FFEEE51D5098825B612DFADA" box="[189,550,1363,1390]" fontSize="12" level="2" pageId="47" pageNumber="164" reason="2">
<taxonomicName id="4C0E4D2FFFEEE51D5098825B62BDFADA" ID-CoL="5DDBR" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,438,1363,1390]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="densihirtus" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFEEE51D5098825B63F8FADA" box="[189,243,1363,1390]" italics="true" pageId="47" pageNumber="164">Zyras</emphasis>
(
<emphasis id="B97AEABEFFEEE51D5124825B623CFADA" box="[257,311,1363,1390]" italics="true" pageId="47" pageNumber="164">Zyras</emphasis>
)
<emphasis id="B97AEABEFFEEE51D5160825B62BDFADA" box="[325,438,1363,1390]" italics="true" pageId="47" pageNumber="164">densihirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEEE51D5199825B612DFADA" box="[444,550,1363,1390]" pageId="47" pageNumber="164" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BB136ACFFEEE51D5098827361CCFA3B" blockId="47.[189,711,1363,1455]" box="[189,711,1403,1423]" pageId="47" pageNumber="164">
<uri id="FF9F3AAEFFEEE51D5098827361CCFA3B" box="[189,711,1403,1423]" pageId="47" pageNumber="164">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFEEE51D514A827361CCFA3B" box="[367,711,1403,1423]" pageId="47" pageNumber="164">E47C7587-9B06-4CD1-821D-6074284CA5BB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEEE51D5098829362C6FA1B" box="[189,461,1435,1455]" pageId="47" pageNumber="164" type="description">
<paragraph id="8BB136ACFFEEE51D5098829362C6FA1B" blockId="47.[189,711,1363,1455]" box="[189,461,1435,1455]" pageId="47" pageNumber="164">
(
<figureCitation id="13352A29FFEEE51D50E4829363F1FA1B" box="[193,250,1435,1455]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="47" pageNumber="164">Figs 37</figureCitation>
,
<figureCitation id="13352A29FFEEE51D51248293621DFA1B" box="[257,278,1435,1455]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="47" pageNumber="164">77</figureCitation>
,
<figureCitation id="13352A29FFEEE51D513882936230FA1B" box="[285,315,1435,1455]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="47" pageNumber="164">115</figureCitation>
,
<figureCitation id="13352A29FFEEE51D516782936283FA1B" box="[322,392,1435,1455]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">250–254</figureCitation>
,
<figureCitation id="13352A29FFEEE51D51AB829362C2FA1B" box="[398,457,1435,1455]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="47" pageNumber="164">Map 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEEE51D50B282C16246F975" pageId="47" pageNumber="164" type="materials_examined">
<paragraph id="8BB136ACFFEEE51D50B282C16246F975" blockId="47.[151,757,1481,1729]" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D50B282C16224FA55" bold="true" box="[151,303,1481,1505]" pageId="47" pageNumber="164">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFEEE51D511882C161DDF9B5" ID-GBIF-Occurrence="3450162319" collectingDate="1985-11" country="Indonesia" location="Plot" municipality="Dumoga-Bone N. P." pageId="47" pageNumber="164" specimenCount="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<typeStatus id="54B5880EFFEEE51D511882C162ABFA55" box="[317,416,1481,1505]" pageId="47" pageNumber="164" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFEEE51D518D82C262B6FA55" box="[424,445,1482,1505]" pageId="47" pageNumber="164">♂</specimenCount>
: “
<collectingCountry id="F319763CFFEEE51D51F682C16168FA55" box="[467,611,1481,1505]" name="Indonesia" pageId="47" pageNumber="164">INDONESIA</collectingCountry>
:
<collectingRegion id="49CAF84EFFEEE51D525282C163FAF9B5" country="Indonesia" name="Sulawesi Utara" pageId="47" pageNumber="164">SULAWESI UTARA</collectingRegion>
,
<collectingMunicipality id="6BD5ACD6FFEEE51D50DB82E262D9F9B5" box="[254,466,1514,1538]" pageId="47" pageNumber="164">Dumoga-Bone N.P.</collectingMunicipality>
,
<date id="FFB0106CFFEEE51D51FB82E16186F9B5" box="[478,653,1513,1537]" pageId="47" pageNumber="164" value="1985-11">
<collectingDate id="EFF4E984FFEEE51D51FB82E16186F9B5" box="[478,653,1513,1537]" pageId="47" pageNumber="164" value="1985-11">November 1985</collectingDate>
</date>
. /
<location id="8ED16077FFEEE51D528E82E161DDF9B5" LSID="urn:lsid:plazi:treatment:03A787BAFFEEE5025098825B610EFDD5:8ED16077FFEEE51D528E82E161DDF9B5" box="[683,726,1513,1537]" country="Indonesia" municipality="Dumoga-Bone N. P." name="Plot" pageId="47" pageNumber="164" stateProvince="SULAWESI UTARA">Plot</location>
</materialsCitation>
<materialsCitation id="3B663CF1FFEEE51D52FA82E263F6F935" ID-GBIF-Occurrence="3450162311" collectingMethod="Malaise trap" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="300" location="Ent. Soc. Lond." pageId="47" pageNumber="164" specimenCount="1" stateProvince="B" typeStatus="holotype">
<collectingRegion id="49CAF84EFFEEE51D52FA82E261FBF9B5" box="[735,752,1514,1537]" country="Indonesia" name="B" pageId="47" pageNumber="164">B</collectingRegion>
, ca
<quantity id="4CF69B49FFEEE51D509C810163F5F995" box="[185,254,1545,1569]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="164" unit="m" value="300.0">
<elevation id="0023D19FFFEEE51D509C810163F5F995" box="[185,254,1545,1569]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="47" pageNumber="164" unit="m" value="300.0">300 m</elevation>
</quantity>
, lowland forest /
<collectingMethod id="524F4EBBFFEEE51D51E281016144F995" box="[455,591,1545,1569]" pageId="47" pageNumber="164">Malaise trap</collectingMethod>
2 / R.
<location id="8ED16077FFEEE51D52BE810163D3F9F5" LSID="urn:lsid:plazi:treatment:03A787BAFFEEE5025098825B610EFDD5:8ED16077FFEEE51D52BE810163D3F9F5" country="Indonesia" name="Ent. Soc. Lond." pageId="47" pageNumber="164" stateProvince="B">Ent.Soc. Lond.</location>
, Project Wallace, B.M. 1985-10 / 29.123 /
<typeStatus id="54B5880EFFEEE51D5290812163DAF9D6" pageId="47" pageNumber="164" type="holotype">Holotypus</typeStatus>
♂
<taxonomicName id="4C0E4D2FFFEEE51D50DB814262B3F9D5" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[254,440,1609,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="densihirtus" status="sp. nov.">
<emphasis id="B97AEABEFFEEE51D50DB814262B3F9D5" box="[254,440,1609,1633]" italics="true" pageId="47" pageNumber="164">Zyras densihirtus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEEE51D51E181426109F9D5" box="[452,514,1610,1633]" pageId="47" pageNumber="164" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E6DDA68FFEEE51D5262814261A2F9D5" box="[583,681,1609,1633]" pageId="47" pageNumber="164">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFEEE51D50BA816163FEF935" box="[159,245,1641,1665]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="47" pageNumber="164">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B663CF1FFEEE51D512D81626242F975" ID-GBIF-Occurrence="3455147302" collectingDate="1985-05" collectingMethod="Flight interception trap" collectionCode="BMNH" country="Indonesia" determinerName="V. Assing" elevation="300" location="Ent. Soc. Lond." pageId="47" pageNumber="164" specimenCount="1" stateProvince="B" typeStatus="paratype">
<typeStatus id="54B5880EFFEEE51D512D8162626EF936" box="[264,357,1642,1666]" pageId="47" pageNumber="164" type="paratype">Paratype</typeStatus>
♂: same data as holotype, but “
<date id="FFB0106CFFEEE51D52E1816163C6F915" pageId="47" pageNumber="164" value="1985-05">
<collectingDate id="EFF4E984FFEEE51D52E1816163C6F915" pageId="47" pageNumber="164" value="1985-05">May 1985</collectingDate>
</date>
. / Plot
<collectingRegion id="49CAF84EFFEEE51D513081816223F915" box="[277,296,1673,1697]" country="Indonesia" name="C" pageId="47" pageNumber="164">C</collectingRegion>
, ca
<quantity id="4CF69B49FFEEE51D516A8181629FF915" box="[335,404,1673,1697]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="47" pageNumber="164" unit="m" value="400.0">
<elevation id="0023D19FFFEEE51D516A8181629FF915" box="[335,404,1673,1697]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="47" pageNumber="164" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<location id="8ED16077FFEEE51D51B881816137F915" LSID="urn:lsid:plazi:treatment:03A787BAFFEEE5025098825B610EFDD5:8ED16077FFEEE51D51B881816137F915" box="[413,572,1673,1697]" country="Indonesia" name="Lowland forest" pageId="47" pageNumber="164" stateProvince="C">Lowland forest</location>
/
<collectingMethod id="524F4EBBFFEEE51D5275818163FFF975" pageId="47" pageNumber="164">Flight interception trap</collectingMethod>
” (cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEEE51D50B281E16186F8D5" pageId="47" pageNumber="164" type="etymology">
<paragraph id="8BB136ACFFEEE51D50B281E16186F8D5" blockId="47.[151,756,1769,1889]" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D50B281E16205F8B5" bold="true" box="[151,270,1769,1793]" pageId="47" pageNumber="164">Etymology</emphasis>
: The specific epithet (adjective) is composed of the Latin adjectives densus (dense) and hirtus (pubescent). It alludes to the dense punctation of the elytra and the hypothesized close relationship to
<taxonomicName id="4C0E4D2FFFEEE51D52088042618CF8D5" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[557,647,1865,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFEEE51D52088042618CF8D5" box="[557,647,1865,1889]" italics="true" pageId="47" pageNumber="164">Z. hirtus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEEE50250B280816290FE15" lastPageId="48" lastPageNumber="165" pageId="47" pageNumber="164" type="description">
<paragraph id="8BB136ACFFEEE51D50B280816725FED5" blockId="47.[151,756,1929,2081]" lastBlockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D50B280816212F815" bold="true" box="[151,281,1929,1953]" pageId="47" pageNumber="164">Description</emphasis>
: Body length
<quantity id="4CF69B49FFEEE51D519A8081614BF815" box="[447,576,1929,1953]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" metricValueMax="7.5" metricValueMin="7.3" pageId="47" pageNumber="164" unit="mm" value="7.4" valueMax="7.5" valueMin="7.3">7.3–7.5 mm</quantity>
; length of forebody
<quantity id="4CF69B49FFEEE51D50F780A16258F875" box="[210,339,1961,1985]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="47" pageNumber="164" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.1–3.3 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFEEE51D51F980A16120F875" box="[476,555,1961,1985]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="47" pageNumber="164">Figs 37</figureCitation>
,
<figureCitation id="13352A29FFEEE51D521080A2615AF875" box="[565,593,1962,1985]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="47" pageNumber="164">77</figureCitation>
,
<figureCitation id="13352A29FFEEE51D527F80A1618FF875" box="[602,644,1961,1985]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="47" pageNumber="164">115</figureCitation>
): forebody black; abdomen blackish with the posterior margins of the segments and the paratergites partly paler brown; legs pale-yellowish with the profemora slightly darker and the apices of the meso- and metafemora narrowly infuscate; antennae black with the apical two antennomeres paleyellow; maxillary palpi brown to dark-brown, with the terminal palpomere paler.
</paragraph>
<paragraph id="8BB136ACFFEEE51D533F8661671AFD15" blockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
Head (
<figureCitation id="13352A29FFEEE51D5350866160CBFE35" box="[885,960,361,385]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="47" pageNumber="164">Fig. 77</figureCitation>
) moderately transverse, extensively impunctate along middle; lateral dorsal portions with rather dense and coarse punctation. Eyes large and bulging, much longer than postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFEEE51D53AE86E160DDFDB5" box="[907,982,489,513]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="47" pageNumber="164">Fig. 37</figureCitation>
) approximately
<quantity id="4CF69B49FFEEE51D54B586E167E9FDB5" box="[1168,1250,489,513]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="47" pageNumber="164" unit="mm" value="2.2">2.2 mm</quantity>
long; antennomeres IV weakly oblong, V approximately as long as broad, VI–X of gradually increasing width and increasingly transverse, X more than 1.5 times as broad as long, and XI of conical shape, shorter than the combined length of VIII–X, or nearly so.
</paragraph>
<paragraph id="8BB136ACFFEEE51D533F85A26084FC55" blockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
Pronotum (
<figureCitation id="13352A29FFEEE51D53BE85A160EDFD75" box="[923,998,681,705]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="47" pageNumber="164">Fig. 77</figureCitation>
) weakly transverse, 1.03–1.05 times as broad as long and approximately 1.3 times as broad as head, broadest near anterior angles; lateral margins straight in posterior two-thirds (dorsal view); punctation coarse, dense, and regularly or slightly irregularly distributed, laterally with or without impunctate patch on either side; midline with or without narrowly impunctate band; lateral margins each with four long and erect black setae; anterior margin with one long and erect black seta on either side.
</paragraph>
<paragraph id="8BB136ACFFEEE51D533F84E167BDFB35" blockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
Elytra (
<figureCitation id="13352A29FFEEE51D534284E160BAFBB5" box="[871,945,1001,1025]" captionStart="Figs 68–81" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 68–81: Forebody (68–77), head and pronotum (78), and abdomen (79–81) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="47" pageNumber="164">Fig. 77</figureCitation>
) approximately 0.8 times as long as pronotum; punctation rather coarse and very dense, denser anteriorly than posteriorly. Hind wings fully developed. Legs very slender; metatarsomere I as long as the combined length of II–IV, or nearly so.
</paragraph>
<paragraph id="8BB136ACFFEEE51D533F838160D2F975" blockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
Abdomen (
<figureCitation id="13352A29FFEEE51D53BF838160FFFB15" box="[922,1012,1161,1185]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="47" pageNumber="164">Fig. 115</figureCitation>
) narrower than elytra, with deep anterior impressions on tergites III–V; tergites III–V each with a transverse row of non-setiferous punctures in anterior impressions, those of tergite V dense, those of tergite IV less dense, and those of tergite III sparse; tergites III–V with a transverse row of four setiferous setae across middle and with 6–8 (tergites III–IV) or 8 setiferous setae (tergite V) at posterior margins; tergite VI with a transverse band of non-setiferous punctures anteriorly, with a transverse row of setiferous punctures across middle, and with approximately ten setiferous punctures at posterior margin; tergite VII with a broad transverse band of coarse non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFEEE51D53DC81616758F935" box="[1017,1107,1641,1665]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 253</figureCitation>
) with numerous long and erect black setae in posterior third, posterior margin distinctly convex.
</paragraph>
<paragraph id="8BB136ACFFEEE51D533F81C26603F835" blockId="47.[794,1399,233,1921]" pageId="47" pageNumber="164">
♂: sternite VIII (
<figureCitation id="13352A29FFEEE51D53C481C16733F955" box="[993,1080,1737,1761]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 254</figureCitation>
) with numerous long black setae in posterior half, posterior margin strongly convex; median lobe of aedeagus (
<figureCitation id="13352A29FFEEE51D5474800167EAF895" box="[1105,1249,1801,1825]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Figs 250–251</figureCitation>
)
<quantity id="4CF69B49FFEEE51D54D28001667CF895" box="[1271,1399,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.0" metricValueMin="8.0" pageId="47" pageNumber="164" unit="mm" value="0.85" valueMax="0.9" valueMin="0.8">0.8–0.9 mm</quantity>
long; ventral process subapically abruptly angled; paramere (
<figureCitation id="13352A29FFEEE51D5347804160B1F8D5" box="[866,954,1865,1889]" captionStart="Figs 245–259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245–259: Zyras titan (245–249), Z. densihirtus (250–254), and Z. nigrihirtus (255–259): median lobe of aedeagus in lateral and in ventral view (245–246, 250–251, 255–256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 252</figureCitation>
) as long as median lobe and with very small and flattened apical lobe with long setae.
</paragraph>
<paragraph id="8BB136ACFFEEE502533F80A16290FE15" blockId="47.[794,1398,1961,2081]" lastBlockId="48.[189,794,233,417]" lastPageId="48" lastPageNumber="165" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D533F80A160E0F875" bold="true" box="[794,1003,1961,1985]" pageId="47" pageNumber="164">Comparative notes</emphasis>
: As can be inferred from the conical shape of antennomere XI, the shape of the ventral process of the aedeagus, and particularly by the short and flattened apical lobe of the paramere,
<taxonomicName id="4C0E4D2FFFEEE51D54A68F026618F795" authorityName="Assing" authorityYear="2017" box="[1155,1299,2057,2081]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="densihirtus">
<emphasis id="B97AEABEFFEEE51D54A68F026618F795" box="[1155,1299,2057,2081]" italics="true" pageId="47" pageNumber="164">Z. densihirtus</emphasis>
</taxonomicName>
is related to
<taxonomicName id="4C0E4D2FFFF1E50250FF87E2623FFEB5" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[218,308,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFF1E50250FF87E2623FFEB5" box="[218,308,233,257]" italics="true" pageId="48" pageNumber="165">Z. hirtus</emphasis>
</taxonomicName>
and allied species. Among the species of this group, it is characterized particularly by the coloration, the dense and coarse punctation of the elytra, the punctation pattern of the abdomen, the rather sparse long setation of the forebody, and by the shape of the median lobe of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFF1E502509886C1610EFDD5" pageId="48" pageNumber="165" type="distribution">
<paragraph id="8BB136ACFFF1E502509886C1610EFDD5" blockId="48.[189,794,457,609]" pageId="48" pageNumber="165">
<emphasis id="B97AEABEFFF1E502509886C1613AFE55" bold="true" box="[189,561,457,481]" pageId="48" pageNumber="165">Distribution and natural history</emphasis>
: The
<typeStatus id="54B5880EFFF1E502525386C261AEFE56" box="[630,677,458,482]" pageId="48" pageNumber="165">type</typeStatus>
locality is identical to that of
<taxonomicName id="4C0E4D2FFFF1E50251AF86E2612FFDB5" authorityName="Assing" authorityYear="2017" box="[394,548,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="densissimus">
<emphasis id="B97AEABEFFF1E50251AF86E2612FFDB5" box="[394,548,489,513]" italics="true" pageId="48" pageNumber="165">Z. densissimus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0E4D2FFFF1E502527C86E261A2FDB5" authorityName="Assing" authorityYear="2017" box="[601,681,490,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="165" phylum="Arthropoda" rank="species" species="titan">
<emphasis id="B97AEABEFFF1E502527C86E261A2FDB5" box="[601,681,490,513]" italics="true" pageId="48" pageNumber="165">Z. titan</emphasis>
</taxonomicName>
(
<figureCitation id="13352A29FFF1E502529386E16006FDB5" box="[694,781,489,513]" captionStart="Map 10" captionStartId="70.[189,234,1271,1292]" captionTargetBox="[189,1436,232,1262]" captionTargetId="figure-89@70.[189,1436,232,1262]" captionTargetPageId="70" captionText="Map 10: Revised distributions of Zyras preangeranus (black circles), Z. brevilobatus (white star), Z. parvicollis (white star), Z. ambulans (white circle), Z. densissimus (black star), Z. titan (black star), Z. densihirtus (black star), and Z. nigrihirtus (black star) in the southern East Palaearctic and Oriental regions." figureDoi="http://doi.org/10.5281/zenodo.5742424" httpUri="https://zenodo.org/record/5742424/files/figure.png" pageId="48" pageNumber="165">Map 10</figureCitation>
). The specimens were collected with a Malaise trap and a flight interception trap in lowland forest at altitudes of approximately 300 and
<specimenCount id="9D08FD25FFF1E502519E8541610EFDD5" box="[443,517,585,609]" pageId="48" pageNumber="165" type="generic">
<quantity id="4CF69B49FFF1E502519E85416109FDD5" box="[443,514,585,609]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="48" pageNumber="165" unit="m" value="400.0">400 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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