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<document id="72B09CD8BB99C47217E05CCD21379F0F" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFEDE51D50C6807B62C6FAB5" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) titan Assing 2017, spec. nov." docType="treatment" docUuid="04E2A49E-E856-490E-AAB3-333A8F481D98" docUuidSource="ZooBank" docVersion="13" lastPageNumber="164" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="161" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="F7FA71895686D51FF5EABAB08D65DA61">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:date id="B5191ED9C10D422AFEE66CE956BC7CA9">2017</mods:date>
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<treatment id="03A787BAFFEDE51D50C6807B62C6FAB5" ID-DOI="http://doi.org/10.5281/zenodo.5888440" ID-GBIF-Taxon="192498200" ID-Zenodo-Dep="5888440" LSID="urn:lsid:zoobank.org:act:04E2A49E-E856-490E-AAB3-333A8F481D98" httpUri="http://treatment.plazi.org/id/03A787BAFFEDE51D50C6807B62C6FAB5" lastPageId="47" lastPageNumber="164" pageId="44" pageNumber="161">
<subSubSection id="C3146527FFEDE51E50C6807B6102F83A" box="[227,521,1907,1934]" pageId="44" pageNumber="161" type="nomenclature">
<paragraph id="8BB136ACFFEDE51E50C6807B6102F83A" blockId="44.[227,746,1907,1999]" box="[227,521,1907,1934]" pageId="44" pageNumber="161">
<heading id="D0F981C0FFEDE51E50C6807B6102F83A" box="[227,521,1907,1934]" fontSize="12" level="2" pageId="44" pageNumber="161" reason="2">
<taxonomicName id="4C0E4D2FFFEDE51E50C6807B6292F83A" ID-CoL="7GHY2" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[227,409,1907,1934]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="161" phylum="Arthropoda" rank="species" species="titan" status="sp. nov." subGenus="Zyras">
<emphasis id="B97AEABEFFEDE51E50C6807B6212F83A" box="[227,281,1907,1934]" italics="true" pageId="44" pageNumber="161">Zyras</emphasis>
(
<emphasis id="B97AEABEFFEDE51E5102807B6256F83A" box="[295,349,1907,1934]" italics="true" pageId="44" pageNumber="161">Zyras</emphasis>
)
<emphasis id="B97AEABEFFEDE51E514F807B6292F83A" box="[362,409,1907,1934]" italics="true" pageId="44" pageNumber="161">titan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEDE51E5185807B6102F83A" box="[416,521,1907,1934]" pageId="44" pageNumber="161" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C3146527FFEDE51E50C6809361E1F81B" box="[227,746,1947,1967]" pageId="44" pageNumber="161" type="description">
<paragraph id="8BB136ACFFEDE51E50C6809361E1F81B" blockId="44.[227,746,1907,1999]" box="[227,746,1947,1967]" pageId="44" pageNumber="161">
<uri id="FF9F3AAEFFEDE51E50C6809361E1F81B" box="[227,746,1947,1967]" pageId="44" pageNumber="161">
urn:lsid:zoobank.org:act:
<uuid id="FFA80C79FFEDE51E51B6809361E1F81B" box="[403,746,1947,1967]" pageId="44" pageNumber="161">04E2A49E-E856-490E-AAB3-333A8F481D98</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEDE51E50C680B362A4F87B" box="[227,431,1979,1999]" pageId="44" pageNumber="161" type="description">
<paragraph id="8BB136ACFFEDE51E50C680B362A4F87B" blockId="44.[227,746,1907,1999]" box="[227,431,1979,1999]" pageId="44" pageNumber="161">
(
<figureCitation id="13352A29FFEDE51E50C280B3622BF87B" box="[231,288,1979,1999]" captionStart="Figs 141" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 141:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (35),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (1718), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (2829),novinversus (3031), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="44" pageNumber="161">Figs 41</figureCitation>
,
<figureCitation id="13352A29FFEDE51E510280B36237F87B" box="[295,316,1979,1999]" captionStart="Figs 6881" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 6881: Forebody (6877), head and pronotum (78), and abdomen (7981) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="44" pageNumber="161">76</figureCitation>
,
<figureCitation id="13352A29FFEDE51E516680B3626AF87B" box="[323,353,1979,1999]" captionStart="Figs 100116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="44" pageNumber="161">113</figureCitation>
,
<figureCitation id="13352A29FFEDE51E514D80B362A7F87B" box="[360,428,1979,1999]" captionStart="Figs 245259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245259: Zyras titan (245249), Z. densihirtus (250254), and Z. nigrihirtus (255259): median lobe of aedeagus in lateral and in ventral view (245246, 250251, 255256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="44" pageNumber="161">245249</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEDE51E509880E16757FE15" pageId="44" pageNumber="161" type="materials_examined">
<paragraph id="8BB136ACFFEDE51E509880E16757FE15" blockId="44.[189,794,2025,2082]" lastBlockId="44.[831,1436,233,417]" pageId="44" pageNumber="161">
<emphasis id="B97AEABEFFEDE51E509880E1625EF7B5" bold="true" box="[189,341,2025,2049]" pageId="44" pageNumber="161">Type material</emphasis>
:
<materialsCitation id="3B663CF1FFEDE51E514680E160A3FED5" ID-GBIF-Occurrence="3450162378" collectingDate="1985-02-26" collectingDateMax="1985-04" collectingDateMin="1985-02-24" collectingMethod="Flight interception trap" collectionCode="BMNH" collectorName="Dumoga-Bone N. P." country="Indonesia" determinerName="V. Assing" elevation="200" location="Dumoga-Bone N. P." pageId="44" pageNumber="161" specimenCount="1" stateProvince="SULAWESI UTARA" typeStatus="holotype">
<typeStatus id="54B5880EFFEDE51E514680E162CDF7B5" box="[355,454,2025,2049]" pageId="44" pageNumber="161" type="holotype">Holotype</typeStatus>
<specimenCount id="9D08FD25FFEDE51E51E880E262E9F7B5" box="[461,482,2026,2049]" pageId="44" pageNumber="161"></specimenCount>
: “
<collectingCountry id="F319763CFFEDE51E51DC80E16182F7B5" box="[505,649,2025,2049]" name="Indonesia" pageId="44" pageNumber="161">INDONESIA</collectingCountry>
:
<collectingRegion id="49CAF84EFFEDE51E52B880E1621CF795" country="Indonesia" name="Sulawesi Utara" pageId="44" pageNumber="161">SULAWESI UTARA</collectingRegion>
,
<location id="8ED16077FFEDE51E51048F0262F9F795" LSID="urn:lsid:plazi:treatment:03A787BAFFEDE51D50C6807B62C6FAB5:8ED16077FFEDE51E51048F0262F9F795" box="[289,498,2058,2082]" country="Indonesia" name="Dumoga-Bone N. P." pageId="44" pageNumber="161" stateProvince="SULAWESI UTARA">Dumoga-Bone N.P.</location>
,
<date id="FFB0106CFFEDE51E51D98F0161B3F795" box="[508,696,2057,2081]" pageId="44" pageNumber="161" value="1985-02-26">
<collectingDate id="EFF4E984FFEDE51E51D98F0161B3F795" box="[508,696,2057,2081]" pageId="44" pageNumber="161" value="1985-02-26">26 February 1985</collectingDate>
</date>
. / Plot A, ca
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<elevation id="0023D19FFFEDE51E537887E160AAFEB5" box="[861,929,233,257]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="44" pageNumber="161" unit="m" value="200.0">200 m</elevation>
</quantity>
, lowland forest /
<collectingMethod id="524F4EBBFFEDE51E544887E16664FEB5" box="[1133,1391,233,257]" pageId="44" pageNumber="161">Flight interception trap</collectingMethod>
2 / R.Ent.Soc. Lond., Project Wallace, B.M. 1985-10 / 29.16 / Holotypus ♂
<taxonomicName id="4C0E4D2FFFEDE51E53CB8622676EFEF5" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[1006,1125,298,321]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="161" phylum="Arthropoda" rank="species" species="titan" status="sp. nov.">
<emphasis id="B97AEABEFFEDE51E53CB8622676EFEF5" box="[1006,1125,298,321]" italics="true" pageId="44" pageNumber="161">Zyras titan</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957C5FFEDE51E5455862267A6FEF5" box="[1136,1197,298,321]" pageId="44" pageNumber="161" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E6DDA68FFEDE51E54D486226658FEF5" box="[1265,1363,297,321]" pageId="44" pageNumber="161">V. Assing</determinerName>
2016” (
<collectionCode id="ED1FAE69FFEDE51E536286416096FED5" box="[839,925,329,353]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="161">BMNH</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B663CF1FFEDE51E539086426753FE15" ID-GBIF-Occurrence="3450162437" collectingDate="1985-02-24" collectingDateMax="1985-04" collectingDateMin="1985-02-24" collectionCode="BMNH" collectorName="Dumoga-Bone N. P." country="Indonesia" determinerName="V. Assing" elevation="200" location="Plot" pageId="44" pageNumber="161" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="54B5880EFFEDE51E539086426714FED6" box="[949,1055,330,354]" pageId="44" pageNumber="161" type="paratype">Paratypes</typeStatus>
: 1 ♂: same data as holotype, but “
<date id="FFB0106CFFEDE51E536C8661670EFE35" box="[841,1029,361,385]" pageId="44" pageNumber="161" value="1985-02-24">
<collectingDate id="EFF4E984FFEDE51E536C8661670EFE35" box="[841,1029,361,385]" pageId="44" pageNumber="161" value="1985-02-24">24 February 1985</collectingDate>
</date>
.” (cAss); 2 ♀♀: same data as holotype, but “
<date id="FFB0106CFFEDE51E5354868160EBFE15" box="[881,992,393,417]" pageId="44" pageNumber="161" value="1985-04">
<collectingDate id="EFF4E984FFEDE51E5354868160EBFE15" box="[881,992,393,417]" pageId="44" pageNumber="161" value="1985-04">April 1985</collectingDate>
</date>
.” (
<collectionCode id="ED1FAE69FFEDE51E53DF8681675BFE15" box="[1018,1104,393,417]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="44" pageNumber="161">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEDE51E531A86C167DBFD95" pageId="44" pageNumber="161" type="etymology">
<paragraph id="8BB136ACFFEDE51E531A86C167DBFD95" blockId="44.[831,1437,457,545]" pageId="44" pageNumber="161">
<emphasis id="B97AEABEFFEDE51E531A86C160BDFE55" bold="true" box="[831,950,457,481]" pageId="44" pageNumber="161">Etymology</emphasis>
: The specific epithet (noun in apposition) alludes to the conspicuously large size of this species. Titans in Greek mythology are giants.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEDE51D531A85416221FBD5" lastPageId="47" lastPageNumber="164" pageId="44" pageNumber="161" type="description">
<paragraph id="8BB136ACFFEDE51E531A85416736FC15" blockId="44.[831,1437,585,2081]" pageId="44" pageNumber="161">
<emphasis id="B97AEABEFFEDE51E531A854160CAFDD5" bold="true" box="[831,961,585,609]" pageId="44" pageNumber="161">Description</emphasis>
: Very large species; body length 9.511.0 mm; length of forebody 4.0
<quantity id="4CF69B49FFEDE51E54C285616631FD35" box="[1255,1338,617,641]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="44" pageNumber="161" unit="mm" value="4.3">4.3 mm</quantity>
. Coloration (
<figureCitation id="13352A29FFEDE51E5359858160C0FD15" box="[892,971,649,673]" captionStart="Figs 141" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 141:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (35),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (1718), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (2829),novinversus (3031), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="44" pageNumber="161">Figs 41</figureCitation>
,
<figureCitation id="13352A29FFEDE51E53FF858160FDFD15" box="[986,1014,649,673]" captionStart="Figs 6881" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 6881: Forebody (6877), head and pronotum (78), and abdomen (7981) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="44" pageNumber="161">76</figureCitation>
,
<figureCitation id="13352A29FFEDE51E542085816724FD15" box="[1029,1071,649,673]" captionStart="Figs 100116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="44" pageNumber="161">113</figureCitation>
): head and pronotum reddish to reddish-brown, pronotum sometimes with paler lateral margins; elytra pale-reddish, occasionally with the lateral margins slightly darker; abdomen with tergites IIVI dark-reddish to brown with the posterior margin broadly pale-reddish, tergites VIIVIII reddish; antennae brown to dark-brown with the basal three antennomeres palereddish and the apical 23 antennomeres dark-yellowish; maxillary palpi reddish.
</paragraph>
<paragraph id="8BB136ACFFEDE51E531A84A16717FAB5" blockId="44.[831,1437,585,2081]" pageId="44" pageNumber="161">
Head (
<figureCitation id="13352A29FFEDE51E53AF84A160DFFC75" box="[906,980,937,961]" captionStart="Figs 6881" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 6881: Forebody (6877), head and pronotum (78), and abdomen (7981) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="44" pageNumber="161">Fig. 76</figureCitation>
) strongly transverse, extensively impunctate along middle; lateral dorsal portions with sparse and coarse punctation. Eyes large and bulging, more than twice as long as postocular region in dorsal view. Antenna (
<figureCitation id="13352A29FFEDE51E5397832160F6FBF5" box="[946,1021,1065,1089]" captionStart="Figs 141" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 141:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (35),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (1718), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (2829),novinversus (3031), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="44" pageNumber="161">Fig. 41</figureCitation>
)
<quantity id="4CF69B49FFEDE51E543683216799FBF5" box="[1043,1170,1065,1089]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" metricValueMax="3.3" metricValueMin="3.1" pageId="44" pageNumber="161" unit="mm" value="3.2" valueMax="3.3" valueMin="3.1">3.13.3 mm</quantity>
long; antennomeres IV weakly oblong, V weakly oblong or approximately as long as broad, VI approximately as broad as long or weakly transverse, VIIX of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI as long as the combined length of VIIIX, or nearly so.
</paragraph>
<paragraph id="8BB136ACFFEDE51E536582026719F9D5" blockId="44.[831,1437,585,2081]" pageId="44" pageNumber="161">
Pronotum (
<figureCitation id="13352A29FFEDE51E53E282016719FA95" box="[967,1042,1289,1313]" captionStart="Figs 6881" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 6881: Forebody (6877), head and pronotum (78), and abdomen (7981) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="44" pageNumber="161">Fig. 76</figureCitation>
) rather weakly transverse, 1.07 1.09 times as broad as long and approximately 1.27 times as broad as head, broadest in anterior half; lateral margins straight in posterior two-thirds (dorsal view); punctation coarse, dense, and somewhat irregularly distributed; laterally with impunctate patches; midline moderately broadly impunctate; lateral margins each with approximately eight stout, long, and erect black setae; anterior and posterior margins, too, with stout, long, and erect black setae; pubescence of disc pale, fine, moderately long, and sub-erect.
</paragraph>
<paragraph id="8BB136ACFFEDE51E536581616680F8B5" blockId="44.[831,1437,585,2081]" pageId="44" pageNumber="161">
Elytra (
<figureCitation id="13352A29FFEDE51E53A8816160DCF935" box="[909,983,1641,1665]" captionStart="Figs 6881" captionStartId="50.[189,229,2017,2039]" captionTargetBox="[189,1436,239,1959]" captionTargetId="figure-127@50.[189,1436,234,1973]" captionTargetPageId="50" captionText="Figs 6881: Forebody (6877), head and pronotum (78), and abdomen (7981) of Zyras spp.: brevilobatus (68), parvicollis (69), ambulans (70), nitens, holotype (71), gratellus (Malaysia) (72), matangensis (73), gratellus (Sulawesi) (74), densissimus (75), titan (76), densihirtus (77), nigrihirtus (78), kraatzi (79), asperatus (80), nigroaeneus (81). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742379" httpUri="https://zenodo.org/record/5742379/files/figure.png" pageId="44" pageNumber="161">Fig. 76</figureCitation>
) approximately 0.8 times as long as pronotum; punctation dense, near scutellum and anterior portion of suture very dense, asperate, and somewhat granulose. Hind wings fully developed. Metatarsomere I as long as, or longer than the combined length of IIIV.
</paragraph>
<paragraph id="8BB136ACFFEDE51D536580016154FE75" blockId="44.[831,1437,585,2081]" lastBlockId="47.[151,756,233,673]" lastPageId="47" lastPageNumber="164" pageId="44" pageNumber="161">
Abdomen (
<figureCitation id="13352A29FFEDE51E53E580016711F895" box="[960,1050,1801,1825]" captionStart="Figs 100116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="44" pageNumber="161">Fig. 113</figureCitation>
) narrower than elytra, with deep anterior impressions on tergites IIIV; tergites IIIV each with a transverse row of coarse non-setiferous punctures in anterior impressions, with a transverse row of usually four setiferous punctures across the middle, and with four to six setiferous punctures at posterior margins; tergite VI with a transverse band of coarse non-setiferous punctures anteriorly, with a transverse row of four setiferous punctures at posterior third, and with six setiferous punctures at posterior margin; tergite VII with a transverse band of coarse non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII (
<figureCitation id="13352A29FFEEE51D51CF8661614FFE35" box="[490,580,361,385]" captionStart="Figs 245259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245259: Zyras titan (245249), Z. densihirtus (250254), and Z. nigrihirtus (255259): median lobe of aedeagus in lateral and in ventral view (245246, 250251, 255256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 248</figureCitation>
) with approximately 20 long black setae posteriorly, posterior margin truncate or weakly concave in the middle.
</paragraph>
<caption id="DF716624FFECE51F50B280AC6602F79B" ID-DOI="http://doi.org/10.5281/zenodo.5742367" ID-Zenodo-Dep="5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="45" pageNumber="162" startId="45.[151,191,1956,1978]" targetBox="[152,1389,232,1855]" targetPageId="45">
<paragraph id="8BB136ACFFECE51F50B280AC6602F79B" blockId="45.[151,1399,1956,2096]" pageId="45" pageNumber="162">
<emphasis id="B97AEABEFFECE51F50B280AC63FEF80E" bold="true" box="[151,245,1956,1978]" pageId="45" pageNumber="162">Figs 141</emphasis>
: Antennae of
<taxonomicName id="4C0E4D2FFFECE51F515080AD62DEF80E" box="[373,469,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97AEABEFFECE51F515080AD62A2F80E" box="[373,425,1957,1978]" italics="true" pageId="45" pageNumber="162">Zyras</emphasis>
spp.
</taxonomicName>
:
<taxonomicName id="4C0E4D2FFFECE51F51FE80AD6116F80E" authorityName="SCHUBERT" authorityYear="1908" box="[475,541,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="kraatzi">
<emphasis id="B97AEABEFFECE51F51FE80AD6116F80E" box="[475,541,1957,1978]" italics="true" pageId="45" pageNumber="162">kraatzi</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F520C80AD613EF80E" bold="true" box="[553,565,1957,1978]" pageId="45" pageNumber="162">1</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F526080AD61B8F80E" authorityName="SCHUBERT" authorityYear="1908" box="[581,691,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFECE51F526080AD61B8F80E" box="[581,691,1957,1978]" italics="true" pageId="45" pageNumber="162">exasperatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F529B80AD61C1F80E" bold="true" box="[702,714,1957,1978]" pageId="45" pageNumber="162">2</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F52FE80AD6025F80E" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[731,814,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFECE51F52FE80AD6025F80E" box="[731,814,1957,1978]" italics="true" pageId="45" pageNumber="162">pindarae</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F531F80AD6054F80E" bold="true" box="[826,863,1957,1978]" pageId="45" pageNumber="162">35</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F534A80AD60C4F80E" baseAuthorityName="Champion" baseAuthorityYear="1921" box="[879,975,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="perforatus">
<emphasis id="B97AEABEFFECE51F534A80AD60C4F80E" box="[879,975,1957,1978]" italics="true" pageId="45" pageNumber="162">perforatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F53FF80AD60EDF80E" bold="true" box="[986,998,1957,1978]" pageId="45" pageNumber="162">6</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F53D280AD6751F80E" authorityName="CAMERON" authorityYear="1939" box="[1015,1114,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B97AEABEFFECE51F53D280AD6751F80E" box="[1015,1114,1957,1978]" italics="true" pageId="45" pageNumber="162">championi</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F544080AD677AF80E" bold="true" box="[1125,1137,1957,1978]" pageId="45" pageNumber="162">7</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F54A780AD67C3F80E" authorityName="Assing" authorityYear="2017" box="[1154,1224,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B97AEABEFFECE51F54A780AD67C3F80E" box="[1154,1224,1957,1978]" italics="true" pageId="45" pageNumber="162">pallipes</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F54F180AD67EBF80E" bold="true" box="[1236,1248,1957,1978]" pageId="45" pageNumber="162">8</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F54D480AD6644F80E" authorityName="Last" authorityYear="1969" box="[1265,1359,1957,1978]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="condignus">
<emphasis id="B97AEABEFFECE51F54D480AD6644F80E" box="[1265,1359,1957,1978]" italics="true" pageId="45" pageNumber="162">condignus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F557880AD6662F80E" bold="true" box="[1373,1385,1957,1978]" pageId="45" pageNumber="162">9</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F50B280CB63C4F86C" authorityName="Cameron" authorityYear="1944" box="[151,207,1987,2008]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B97AEABEFFECE51F50B280CB63C4F86C" box="[151,207,1987,2008]" italics="true" pageId="45" pageNumber="162">nitens</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F50FA80CA63FCF863" bold="true" box="[223,247,1986,2007]" pageId="45" pageNumber="162">10</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F512880CB626CF86C" authorityName="Assing" authorityYear="2017" box="[269,359,1987,2008]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B97AEABEFFECE51F512880CB626CF86C" box="[269,359,1987,2008]" italics="true" pageId="45" pageNumber="162">truncatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F515380CA6285F863" bold="true" box="[374,398,1986,2007]" pageId="45" pageNumber="162">11</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F518180CA6102F863" authorityName="CAMERON" authorityYear="1939" box="[420,521,1986,2007]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="nilgiriensis">
<emphasis id="B97AEABEFFECE51F518180CA6102F863" box="[420,521,1986,2007]" italics="true" pageId="45" pageNumber="162">nilgiriensis</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F523C80CA613AF863" bold="true" box="[537,561,1986,2007]" pageId="45" pageNumber="162">12</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F526380CA61C9F863" box="[582,706,1986,2007]" class="Insecta" family="Staphylinidae" genus="Myrmedonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="hirsutiventris">
<emphasis id="B97AEABEFFECE51F526380CA61C9F863" box="[582,706,1986,2007]" italics="true" pageId="45" pageNumber="162">hirsutiventris</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F52F780CA61E1F863" bold="true" box="[722,746,1986,2007]" pageId="45" pageNumber="162">13</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F532580CB6059F86C" authorityName="Assing" authorityYear="2017" box="[768,850,1987,2008]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="russiceps">
<emphasis id="B97AEABEFFECE51F532580CB6059F86C" box="[768,850,1987,2008]" italics="true" pageId="45" pageNumber="162">russiceps</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F534780CA6071F863" bold="true" box="[866,890,1986,2007]" pageId="45" pageNumber="162">14</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F53AA80CA60D5F863" authorityName=", Pace" authorityYear="1986" box="[911,990,1986,2007]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="brignolii">
<emphasis id="B97AEABEFFECE51F53AA80CA60D5F863" box="[911,990,1986,2007]" italics="true" pageId="45" pageNumber="162">brignolii</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F53CB80CA670DF863" bold="true" box="[1006,1030,1986,2007]" pageId="45" pageNumber="162">15</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F543E80CA6762F863" authorityName="Fauvel" authorityYear="1904" box="[1051,1129,1986,2007]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFECE51F543E80CA6762F863" box="[1051,1129,1986,2007]" italics="true" pageId="45" pageNumber="162">hastatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F545F80CA6799F863" bold="true" box="[1146,1170,1986,2007]" pageId="45" pageNumber="162">16</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F548280CB6614F86C" authorityName="Pace" authorityYear="2010" box="[1191,1311,1987,2008]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="parageminus">
<emphasis id="B97AEABEFFECE51F548280CB6614F86C" box="[1191,1311,1987,2008]" italics="true" pageId="45" pageNumber="162">parageminus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F550880CA6663F863" bold="true" box="[1325,1384,1986,2007]" pageId="45" pageNumber="162">1718</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F50B280D763F2F840" authorityName="Assing" authorityYear="2017" box="[151,249,2015,2036]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="latilobatus">
<emphasis id="B97AEABEFFECE51F50B280D763F2F840" box="[151,249,2015,2036]" italics="true" pageId="45" pageNumber="162">latilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F512280D76214F840" bold="true" box="[263,287,2015,2036]" pageId="45" pageNumber="162">19</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F511780D76281F840" baseAuthorityName="Cameron" baseAuthorityYear="1925" box="[306,394,2015,2036]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="alternans">
<emphasis id="B97AEABEFFECE51F511780D76281F840" box="[306,394,2015,2036]" italics="true" pageId="45" pageNumber="162">alternans</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F51B280D762A4F840" bold="true" box="[407,431,2015,2036]" pageId="45" pageNumber="162">20</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F51E780D762F2F840" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[450,505,2015,2036]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B97AEABEFFECE51F51E780D762F2F840" box="[450,505,2015,2036]" italics="true" pageId="45" pageNumber="162">hirtus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F522280D76114F840" bold="true" box="[519,543,2015,2036]" pageId="45" pageNumber="162">21</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F521480E861A9F841" authorityName="Cameron" authorityYear="1939" box="[561,674,2016,2037]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B97AEABEFFECE51F521480E861A9F841" box="[561,674,2016,2037]" italics="true" pageId="45" pageNumber="162">nigroaeneus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F528A80D761CCF840" bold="true" box="[687,711,2015,2036]" pageId="45" pageNumber="162">22</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F52FF80D7602BF840" authorityName="Assing" authorityYear="2017" box="[730,800,2015,2036]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="luteipes">
<emphasis id="B97AEABEFFECE51F52FF80D7602BF840" box="[730,800,2015,2036]" italics="true" pageId="45" pageNumber="162">luteipes</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F530B80D7604DF840" bold="true" box="[814,838,2015,2036]" pageId="45" pageNumber="162">23</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F537D80D760C1F840" authorityName="Assing" authorityYear="2017" box="[856,970,2015,2036]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="longilobatus">
<emphasis id="B97AEABEFFECE51F537D80D760C1F840" box="[856,970,2015,2036]" italics="true" pageId="45" pageNumber="162">longilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F53FD80D760FBF840" bold="true" box="[984,1008,2015,2036]" pageId="45" pageNumber="162">24</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F542780E86759F841" authorityName="Pace" authorityYear="1986" box="[1026,1106,2016,2037]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B97AEABEFFECE51F542780E86759F841" box="[1026,1106,2016,2037]" italics="true" pageId="45" pageNumber="162">morvani</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F544580D76773F840" bold="true" box="[1120,1144,2015,2036]" pageId="45" pageNumber="162">25</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F54AF80D76648F841" authority="(Malaysia)" authorityName="Cameron" authorityYear="1939" baseAuthorityName="Malaysia" box="[1162,1347,2015,2037]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="gratellus">
<emphasis id="B97AEABEFFECE51F54AF80D767D2F840" box="[1162,1241,2015,2036]" italics="true" pageId="45" pageNumber="162">gratellus</emphasis>
(Malaysia)
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F557480D76662F840" bold="true" box="[1361,1385,2015,2036]" pageId="45" pageNumber="162">26</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F50B280F563FAF7A6" baseAuthorityName="Motschulsky" baseAuthorityYear="1861" box="[151,241,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="castaneus">
<emphasis id="B97AEABEFFECE51F50B280F563FAF7A6" box="[151,241,2045,2066]" italics="true" pageId="45" pageNumber="162">castaneus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B97AEABEFFECE51F517B80F5627DF7A6" bold="true" box="[350,374,2045,2066]" pageId="45" pageNumber="162">27</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F51A280F562D4F7A6" authorityName="CAMERON" authorityYear="1939" box="[391,479,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="proximus">
<emphasis id="B97AEABEFFECE51F51A280F562D4F7A6" box="[391,479,2045,2066]" italics="true" pageId="45" pageNumber="162">proximus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F51CE80F5612CF7A6" bold="true" box="[491,551,2045,2066]" pageId="45" pageNumber="162">2829</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F521D80F561ACF7A6" authorityName="Assing" authorityYear="2017" box="[568,679,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="novinversus">
<emphasis id="B97AEABEFFECE51F521D80F561ACF7A6" box="[568,679,2045,2066]" italics="true" pageId="45" pageNumber="162">novinversus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F529680F561E4F7A6" bold="true" box="[691,751,2045,2066]" pageId="45" pageNumber="162">3031</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F532480F5607FF7A6" authorityName="Assing" authorityYear="2017" box="[769,884,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="brevilobatus">
<emphasis id="B97AEABEFFECE51F532480F5607FF7A6" box="[769,884,2045,2066]" italics="true" pageId="45" pageNumber="162">brevilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F535A80F5609CF7A6" bold="true" box="[895,919,2045,2066]" pageId="45" pageNumber="162">32</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F538C80F56702F7A6" authorityName="Assing" authorityYear="2017" box="[937,1033,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="parvicollis">
<emphasis id="B97AEABEFFECE51F538C80F56702F7A6" box="[937,1033,2045,2066]" italics="true" pageId="45" pageNumber="162">parvicollis</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F543080F56726F7A6" bold="true" box="[1045,1069,2045,2066]" pageId="45" pageNumber="162">33</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F541B80F56791F7A6" authorityName="Assing" authorityYear="2017" box="[1086,1178,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="ambulans">
<emphasis id="B97AEABEFFECE51F541B80F56791F7A6" box="[1086,1178,2045,2066]" italics="true" pageId="45" pageNumber="162">ambulans</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F548380F567B5F7A6" bold="true" box="[1190,1214,2045,2066]" pageId="45" pageNumber="162">34</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F54EA80F56648F7A6" authorityName="Cameron" authorityYear="1943" box="[1231,1347,2045,2066]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B97AEABEFFECE51F54EA80F56648F7A6" box="[1231,1347,2045,2066]" italics="true" pageId="45" pageNumber="162">matangensis</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F557480F56662F7A6" bold="true" box="[1361,1385,2045,2066]" pageId="45" pageNumber="162">35</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F50B28F12620EF79B" authorityName="Assing" authorityYear="2017" box="[151,261,2074,2095]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="densissimus">
<emphasis id="B97AEABEFFECE51F50B28F12620EF79B" box="[151,261,2074,2095]" italics="true" pageId="45" pageNumber="162">densissimus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F51368F126220F79B" bold="true" box="[275,299,2074,2095]" pageId="45" pageNumber="162">36</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F511B8F1262ADF79B" authorityName="Assing" authorityYear="2017" box="[318,422,2074,2095]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="densihirtus">
<emphasis id="B97AEABEFFECE51F511B8F1262ADF79B" box="[318,422,2074,2095]" italics="true" pageId="45" pageNumber="162">densihirtus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F51918F1262C7F79B" bold="true" box="[436,460,2074,2095]" pageId="45" pageNumber="162">37</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F51FA8F126148F79B" authorityName="Assing" authorityYear="2017" box="[479,579,2074,2095]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="nigrihirtus">
<emphasis id="B97AEABEFFECE51F51FA8F126148F79B" box="[479,579,2074,2095]" italics="true" pageId="45" pageNumber="162">nigrihirtus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F52748F126162F79B" bold="true" box="[593,617,2074,2095]" pageId="45" pageNumber="162">38</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F52598F126039F784" authority="(Sulawesi)" authorityName="Cameron" authorityYear="1939" baseAuthorityName="Sulawesi" box="[636,818,2074,2096]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="gratellus">
<emphasis id="B97AEABEFFECE51F52598F1261C0F79B" box="[636,715,2074,2095]" italics="true" pageId="45" pageNumber="162">gratellus</emphasis>
(Sulawesi)
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F53658F126053F79B" bold="true" box="[832,856,2074,2095]" pageId="45" pageNumber="162">39</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F534E8F1260B1F79B" authorityName="Cameron" authorityYear="1939" box="[875,954,2074,2095]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="gardneri">
<emphasis id="B97AEABEFFECE51F534E8F1260B1F79B" box="[875,954,2074,2095]" italics="true" pageId="45" pageNumber="162">gardneri</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F53ED8F1260EBF79B" bold="true" box="[968,992,2074,2095]" pageId="45" pageNumber="162">40</emphasis>
),
<taxonomicName id="4C0E4D2FFFECE51F53D68F13672BF784" authorityName="Assing" authorityYear="2017" box="[1011,1056,2075,2096]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="162" phylum="Arthropoda" rank="species" species="titan">
<emphasis id="B97AEABEFFECE51F53D68F13672BF784" box="[1011,1056,2075,2096]" italics="true" pageId="45" pageNumber="162">titan</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFECE51F54088F12674EF79B" bold="true" box="[1069,1093,2074,2095]" pageId="45" pageNumber="162">41</emphasis>
). Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="DF716624FFEFE51C509880D46265F798" ID-DOI="http://doi.org/10.5281/zenodo.5742371" ID-Zenodo-Dep="5742371" httpUri="https://zenodo.org/record/5742371/files/figure.png" pageId="46" pageNumber="163" startId="46.[189,229,2012,2034]" targetBox="[191,1436,232,1943]" targetPageId="46">
<paragraph id="8BB136ACFFEFE51C509880D46265F798" blockId="46.[189,1436,2012,2093]" pageId="46" pageNumber="163">
<emphasis id="B97AEABEFFEFE51C509880D4622CF845" bold="true" box="[189,295,2012,2034]" pageId="46" pageNumber="163">Figs 4254</emphasis>
: Forebody of
<taxonomicName id="4C0E4D2FFFEFE51C519180D56111F846" box="[436,538,2013,2034]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B97AEABEFFEFE51C519180D562E3F846" box="[436,488,2013,2034]" italics="true" pageId="46" pageNumber="163">Zyras</emphasis>
spp.
</taxonomicName>
:
<taxonomicName id="4C0E4D2FFFEFE51C520180D4616DF845" authorityName="SCHUBERT" authorityYear="1908" box="[548,614,2012,2033]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="kraatzi">
<emphasis id="B97AEABEFFEFE51C520180D4616DF845" box="[548,614,2012,2033]" italics="true" pageId="46" pageNumber="163">kraatzi</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C525280D46184F845" bold="true" box="[631,655,2012,2033]" pageId="46" pageNumber="163">42</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C528080D56018F846" authorityName="SCHUBERT" authorityYear="1908" box="[677,787,2013,2034]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="exasperatus">
<emphasis id="B97AEABEFFEFE51C528080D56018F846" box="[677,787,2013,2034]" italics="true" pageId="46" pageNumber="163">exasperatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C530680D46030F845" bold="true" box="[803,827,2012,2033]" pageId="46" pageNumber="163">43</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C537480D460AFF845" baseAuthorityName="CHAMPION" baseAuthorityYear="1921" box="[849,932,2012,2033]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="pindarae">
<emphasis id="B97AEABEFFEFE51C537480D460AFF845" box="[849,932,2012,2033]" italics="true" pageId="46" pageNumber="163">pindarae</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C539080D460C6F845" bold="true" box="[949,973,2012,2033]" pageId="46" pageNumber="163">44</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C53C680D46748F845" baseAuthorityName="Champion" baseAuthorityYear="1921" box="[995,1091,2012,2033]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="perforatus">
<emphasis id="B97AEABEFFEFE51C53C680D46748F845" box="[995,1091,2012,2033]" italics="true" pageId="46" pageNumber="163">perforatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C547180D46767F845" bold="true" box="[1108,1132,2012,2033]" pageId="46" pageNumber="163">45</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C54A780D467EEF845" authorityName="CAMERON" authorityYear="1939" box="[1154,1253,2012,2033]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B97AEABEFFEFE51C54A780D467EEF845" box="[1154,1253,2012,2033]" italics="true" pageId="46" pageNumber="163">championi</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C54D080D46606F845" bold="true" box="[1269,1293,2012,2033]" pageId="46" pageNumber="163">46</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C550680D46662F845" authorityName="Assing" authorityYear="2017" box="[1315,1385,2012,2033]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="pallipes">
<emphasis id="B97AEABEFFEFE51C550680D46662F845" box="[1315,1385,2012,2033]" italics="true" pageId="46" pageNumber="163">pallipes</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C555280D46684F845" bold="true" box="[1399,1423,2012,2033]" pageId="46" pageNumber="163">47</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C509880F26210F7BB" authorityName="Last" authorityYear="1969" box="[189,283,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="condignus">
<emphasis id="B97AEABEFFEFE51C509880F26210F7BB" box="[189,283,2042,2063]" italics="true" pageId="46" pageNumber="163">condignus</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B97AEABEFFEFE51C51AE80F162A8F7BA" bold="true" box="[395,419,2041,2062]" pageId="46" pageNumber="163">48</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C519D80F2611DF7BB" authorityName="Last" authorityYear="1969" box="[440,534,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="condignus">
<emphasis id="B97AEABEFFEFE51C519D80F2611DF7BB" box="[440,534,2042,2063]" italics="true" pageId="46" pageNumber="163">condignus</emphasis>
</taxonomicName>
, Nepal (
<emphasis id="B97AEABEFFEFE51C524980F1618FF7BA" bold="true" box="[620,644,2041,2062]" pageId="46" pageNumber="163">49</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C52BC80F26001F7BB" authorityName="Cameron" authorityYear="1939" box="[665,778,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="nigroaeneus">
<emphasis id="B97AEABEFFEFE51C52BC80F26001F7BB" box="[665,778,2042,2063]" italics="true" pageId="46" pageNumber="163">nigroaeneus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C533F80F16039F7BA" bold="true" box="[794,818,2041,2062]" pageId="46" pageNumber="163">50</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C536D80F26093F7BB" authorityName="Pace" authorityYear="1986" box="[840,920,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B97AEABEFFEFE51C536D80F26093F7BB" box="[840,920,2042,2063]" italics="true" pageId="46" pageNumber="163">morvani</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C538D80F160CBF7BA" bold="true" box="[936,960,2041,2062]" pageId="46" pageNumber="163">51</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C53F080F26724F7BB" authorityName="Assing" authorityYear="2017" box="[981,1071,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="truncatus">
<emphasis id="B97AEABEFFEFE51C53F080F26724F7BB" box="[981,1071,2042,2063]" italics="true" pageId="46" pageNumber="163">truncatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C541B80F1675DF7BA" bold="true" box="[1086,1110,2041,2062]" pageId="46" pageNumber="163">52</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C544980F267D5F7BB" authorityName="Assing" authorityYear="2017" box="[1132,1246,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="longilobatus">
<emphasis id="B97AEABEFFEFE51C544980F267D5F7BB" box="[1132,1246,2042,2063]" italics="true" pageId="46" pageNumber="163">longilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C54CB80F1660DF7BA" bold="true" box="[1262,1286,2041,2062]" pageId="46" pageNumber="163">53</emphasis>
),
<taxonomicName id="4C0E4D2FFFEFE51C553E80F26662F7BB" authorityName="Fauvel" authorityYear="1904" box="[1307,1385,2042,2063]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="163" phylum="Arthropoda" rank="species" species="hastatus">
<emphasis id="B97AEABEFFEFE51C553E80F26662F7BB" box="[1307,1385,2042,2063]" italics="true" pageId="46" pageNumber="163">hastatus</emphasis>
</taxonomicName>
(
<emphasis id="B97AEABEFFEFE51C555280F16684F7BA" bold="true" box="[1399,1423,2041,2062]" pageId="46" pageNumber="163">54</emphasis>
). Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BB136ACFFEEE51D50B286C263D8FD15" blockId="47.[151,756,233,673]" pageId="47" pageNumber="164">
♂: sternite VIII (
<figureCitation id="13352A29FFEEE51D517B86C162BDFE55" box="[350,438,457,481]" captionStart="Figs 245259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245259: Zyras titan (245249), Z. densihirtus (250254), and Z. nigrihirtus (255259): median lobe of aedeagus in lateral and in ventral view (245246, 250251, 255256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 249</figureCitation>
) with numerous long black setae in posterior half, posterior margin convex; median lobe of aedeagus (
<figureCitation id="13352A29FFEEE51D517A850162FBFD95" box="[351,496,521,545]" captionStart="Figs 245259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245259: Zyras titan (245249), Z. densihirtus (250254), and Z. nigrihirtus (255259): median lobe of aedeagus in lateral and in ventral view (245246, 250251, 255256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Figs 245246</figureCitation>
) approximately
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long and of robust shape; ventral process basally with a pair of pronounced carinae; paramere (
<figureCitation id="13352A29FFEEE51D526485416193FDD5" box="[577,664,585,609]" captionStart="Figs 245259" captionStartId="62.[189,229,2016,2038]" captionTargetBox="[193,1430,242,1956]" captionTargetId="figure-101@62.[189,1436,232,1971]" captionTargetPageId="62" captionText="Figs 245259: Zyras titan (245249), Z. densihirtus (250254), and Z. nigrihirtus (255259): median lobe of aedeagus in lateral and in ventral view (245246, 250251, 255256); paramere (247, 252, 257); male tergite VIII (248, 253, 258); male sternite VIII (249, 254, 259). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742389" httpUri="https://zenodo.org/record/5742389/files/figure.png" pageId="47" pageNumber="164">Fig. 247</figureCitation>
) as long as median lobe and with very small apical lobe with long setae.
</paragraph>
<paragraph id="8BB136ACFFEEE51D50B285C16221FBD5" blockId="47.[151,756,713,1121]" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D50B285C16261FD55" bold="true" box="[151,362,713,737]" pageId="47" pageNumber="164">Comparative notes</emphasis>
: This species is characterized by its conspicuously large size, coarse punctation, its coloration, the punctation pattern of the pronotum, and by the male sexual characters. It is distinguished from the similarly large
<taxonomicName id="4C0E4D2FFFEEE51D50FD8442628BFCD5" authorityName="Cameron" authorityYear="1939" box="[216,384,842,865]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="preangeranus">
<emphasis id="B97AEABEFFEEE51D50FD8442628BFCD5" box="[216,384,842,865]" italics="true" pageId="47" pageNumber="164">Z. preangeranus</emphasis>
</taxonomicName>
by the coloration (
<taxonomicName id="4C0E4D2FFFEEE51D524D844263B7FC35" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="preangeranus">
<emphasis id="B97AEABEFFEEE51D524D844263B7FC35" italics="true" pageId="47" pageNumber="164">Z. preangeranus</emphasis>
</taxonomicName>
: antennomeres IVII blackish; head much darker than the pronotum; abdominal segments IIV reddish), much coarser punctation of the forebody, less slender antennomeres IVX, a completely different punctation pattern of the abdomen, and by a more robust aedeagus with a ventral process of completely different shape. For illustrations of
<taxonomicName id="4C0E4D2FFFEEE51D511B832262EDFBF5" authorityName="Cameron" authorityYear="1939" box="[318,486,1066,1089]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="preangeranus">
<emphasis id="B97AEABEFFEEE51D511B832262EDFBF5" box="[318,486,1066,1089]" italics="true" pageId="47" pageNumber="164">Z. preangeranus</emphasis>
</taxonomicName>
see
<bibRefCitation id="EF9F4B5DFFEEE51D523C832161C9FBF5" author="ASSING, V." box="[537,706,1065,1089]" pageId="47" pageNumber="164" pagination="87 - 175" refId="ref57174" refString="ASSING, V. 2016 a: A revision of Zyras STEPHENS sensu strictu of China, Taiwan, and Hong Kong, with records and (re-) descriptions of some species from other regions (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Stuttgarter Beitrage zur Naturkunde A, Neue Serie 9: 87 - 175." type="journal article" year="2016">ASSING (2016a)</bibRefCitation>
and
<figureCitation id="13352A29FFEEE51D50B28341622CFBD5" box="[151,295,1097,1121]" captionStart="Figs 260277" captionStartId="63.[151,191,1987,2009]" captionTargetBox="[157,1395,232,1954]" captionTargetId="figure-126@63.[151,1398,232,1971]" captionText="Figs 260277: Zyras morulus (260265), Z. variolatus (266270), Z. montanus (271275), and Z. preangeranus from Java (276277): forebody (260); antenna (261); abdomen (262); median lobe of aedeagus in lateral and in ventral view (263264, 266267, 271272, 276277); paramere (265, 268, 273); male tergite VIII (269, 274); male sternite VIII (270, 275). Scale bars: 260262: 1.0 mm; 263277: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888380" httpUri="https://zenodo.org/record/5888380/files/figure.png" pageId="47" pageNumber="164">Figs 276277</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3146527FFEEE51D50B2838162C6FAB5" pageId="47" pageNumber="164" type="distribution">
<paragraph id="8BB136ACFFEEE51D50B2838162C6FAB5" blockId="47.[151,756,1161,1281]" pageId="47" pageNumber="164">
<emphasis id="B97AEABEFFEEE51D50B283816102FB15" bold="true" box="[151,521,1161,1185]" pageId="47" pageNumber="164">Distribution and natural history</emphasis>
: The
<typeStatus id="54B5880EFFEEE51D527583826175FB16" box="[592,638,1162,1186]" pageId="47" pageNumber="164">type</typeStatus>
locality is identical to that of
<taxonomicName id="4C0E4D2FFFEEE51D515483A26103FB75" authorityName="Assing" authorityYear="2017" box="[369,520,1193,1217]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="164" phylum="Arthropoda" rank="species" species="densissimus">
<emphasis id="B97AEABEFFEEE51D515483A26103FB75" box="[369,520,1193,1217]" italics="true" pageId="47" pageNumber="164">Z. densissimus</emphasis>
</taxonomicName>
. The
<typeStatus id="54B5880EFFEEE51D526883A26170FB76" box="[589,635,1194,1218]" pageId="47" pageNumber="164">type</typeStatus>
specimens were collected with flight interception traps in lowland forest at an altitude of
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.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>