376 lines
53 KiB
XML
376 lines
53 KiB
XML
<document id="6A2E821C01CFE3A77E05A72FC3F4F5A3" ID-CLB-Dataset="2472" ID-DOI="10.21248/contrib.entomol.67.1.117-192" ID-GBIF-Dataset="39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742363" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1638267576315" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03A787BAFFE3E51150C686FB623AFA75" docLanguage="en" docName="ContribEntomol.67.1.117-192.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) novinversus Assing, 2017, nom. nov." docType="treatment" docUuid="3BD2568F-509A-4132-9F55-131D59FE9405" docUuidSource="ZooBank" docVersion="12" lastPageNumber="152" masterDocId="FF9EFFC2FFC1E53250258708630BFFB4" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="192" masterPageNumber="117" pageNumber="151" updateTime="1699471407760" updateUser="ExternalLinkService">
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<mods:title id="F357C1210B6BB6B389CE8514F05B1E4C">On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="CEC315775EC89F90F48E7E981466F8C8">Assing, Volker</mods:namePart>
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<mods:title id="37BEA6825843FB5EB439DDF63D187008">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="4818828FCF99894104FF43F5BFC6206C">
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<mods:date id="32668363A39F6F9CBE79E3897FF7515C">2017</mods:date>
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<mods:number id="D616DC57D24DC8547B78C1607DE3C7B7">2017-06-30</mods:number>
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<mods:number id="62EBB115DB606B6EF141684FE4CB1A8A">67</mods:number>
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<treatment id="03A787BAFFE3E51150C686FB623AFA75" ID-DOI="http://doi.org/10.5281/zenodo.5888479" ID-GBIF-Taxon="192498190" ID-Zenodo-Dep="5888479" LSID="urn:lsid:zoobank.org:act:3BD2568F-509A-4132-9F55-131D59FE9405" httpUri="http://treatment.plazi.org/id/03A787BAFFE3E51150C686FB623AFA75" lastPageId="35" lastPageNumber="152" pageId="34" pageNumber="151">
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<subSubSection id="C3146527FFE3E51050C686FB61E8FD9B" pageId="34" pageNumber="151" type="nomenclature">
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<paragraph id="8BB136ACFFE3E51050C686FB615BFDBA" blockId="34.[227,739,499,591]" box="[227,592,499,526]" pageId="34" pageNumber="151">
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<heading id="D0F981C0FFE3E51050C686FB615BFDBA" box="[227,592,499,526]" fontSize="12" level="2" pageId="34" pageNumber="151" reason="2">
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<taxonomicName id="4C0E4D2FFFE3E51050C686FB62E8FDBA" ID-CoL="7GHZ5" authorityName="Assing" authorityYear="2017" box="[227,483,499,526]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="novinversus" status="nom. nov." subGenus="Zyras">
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<emphasis id="B97AEABEFFE3E51050C686FB6212FDBA" box="[227,281,499,526]" italics="true" pageId="34" pageNumber="151">Zyras</emphasis>
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(
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<emphasis id="B97AEABEFFE3E510510286FB6256FDBA" box="[295,349,499,526]" italics="true" pageId="34" pageNumber="151">Zyras</emphasis>
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)
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<emphasis id="B97AEABEFFE3E510514F86FB62E8FDBA" box="[362,483,499,526]" italics="true" pageId="34" pageNumber="151">novinversus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24957C5FFE3E51051CC86FB615BFDBA" box="[489,592,499,526]" pageId="34" pageNumber="151">nom. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="8BB136ACFFE3E51050C6851361E8FD9B" blockId="34.[227,739,499,591]" box="[227,739,539,559]" pageId="34" pageNumber="151">
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<uri id="FF9F3AAEFFE3E51050C6851361E8FD9B" box="[227,739,539,559]" pageId="34" pageNumber="151">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA80C79FFE3E51051B6851361E8FD9B" box="[403,739,539,559]" pageId="34" pageNumber="151">3BD2568F-509A-4132-9F55-131D59FE9405</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146527FFE3E51050C685336103FDFB" box="[227,520,571,591]" pageId="34" pageNumber="151" type="description">
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<paragraph id="8BB136ACFFE3E51050C685336103FDFB" blockId="34.[227,739,499,591]" box="[227,520,571,591]" pageId="34" pageNumber="151">
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(
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<figureCitation id="13352A29FFE3E51050C285336236FDFB" box="[231,317,571,591]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="34" pageNumber="151">Figs 30–31</figureCitation>
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,
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<figureCitation id="13352A29FFE3E510516185336252FDFB" box="[324,345,571,591]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="34" pageNumber="151">67</figureCitation>
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,
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<figureCitation id="13352A29FFE3E510514585336275FDFB" box="[352,382,571,591]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="34" pageNumber="151">106</figureCitation>
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,
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<figureCitation id="13352A29FFE3E51051A0853362C1FDFB" box="[389,458,571,591]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="34" pageNumber="151">198–202</figureCitation>
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,
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||
<figureCitation id="13352A29FFE3E51051F485336108FDFB" box="[465,515,571,591]" captionStart="Map 5" captionStartId="65.[151,196,1829,1850]" captionTargetBox="[151,1398,827,1821]" captionTargetId="figure-87@65.[151,1398,827,1821]" captionTargetPageId="65" captionText="Map 5: Revised distributions of Zyras castaneus (black circles) and Z. novinversus (white circles) in the southern East Palaearctic" figureDoi="http://doi.org/10.5281/zenodo.5742405" httpUri="https://zenodo.org/record/5742405/files/figure.png" pageId="34" pageNumber="151">Map 5</figureCitation>
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||
)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE3E5105098856461B6FD35" box="[189,701,619,641]" pageId="34" pageNumber="151" type="reference_group">
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<paragraph id="8BB136ACFFE3E5105098856461B6FD35" blockId="34.[189,701,619,641]" box="[189,701,619,641]" pageId="34" pageNumber="151">
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<treatmentCitationGroup id="AB1E1182FFE3E5105098856461B6FD35" box="[189,701,619,641]" pageId="34" pageNumber="151">
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<treatmentCitation id="0AAF10BDFFE3E51050988564613CFD34" author="PACE, R." box="[189,567,619,641]" page="339" pageId="34" pageNumber="151" year="2012">
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||
<taxonomicName id="4C0E4D2FFFE3E51050988564613CFD34" ID-CoL="5DCTM" authority="PACE, 2012 b: 339" authorityName="PACE" authorityPageNumber="339" authorityYear="2012" box="[189,567,619,641]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="inversus" subGenus="Zyras">
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||
<emphasis id="B97AEABEFFE3E5105098856463FAFD35" box="[189,241,620,641]" italics="true" pageId="34" pageNumber="151">Zyras</emphasis>
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||
(
|
||
<emphasis id="B97AEABEFFE3E51050DA85646238FD35" box="[255,307,620,641]" italics="true" pageId="34" pageNumber="151">Zyras</emphasis>
|
||
)
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||
<emphasis id="B97AEABEFFE3E510516485646286FD35" box="[321,397,620,641]" italics="true" pageId="34" pageNumber="151">inversus</emphasis>
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||
<bibRefCitation id="EF9F4B5DFFE3E51051B78563613CFD34" author="PACE, R." box="[402,567,619,641]" pageId="34" pageNumber="151" pagination="319 - 360" refId="ref59001" refString="PACE, R. 2012 b: New data, new species, and two new genera of Aleocharinae from the Oriental Region (Insecta: Coleoptera: Staphylinidae). - Vernate 31: 319 - 360." type="journal article" year="2012">PACE, 2012b: 339</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
; preoccupied.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3146527FFE3E510509885A162B1FCF5" pageId="34" pageNumber="151" type="materials_examined">
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||
<paragraph id="8BB136ACFFE3E510509885A162B1FCF5" blockId="34.[189,794,681,833]" pageId="34" pageNumber="151">
|
||
<emphasis id="B97AEABEFFE3E510509885A162DFFD75" bold="true" box="[189,468,681,705]" pageId="34" pageNumber="151">Type material examined</emphasis>
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||
:
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||
<materialsCitation id="3B663CF1FFE3E51051CD85A162B1FCF5" ID-GBIF-Occurrence="3450162362" collectingDate="2010-10-24" collectorName="G. de Rougemont & V. Assing" country="Thailand" location="Doi Angkhang" municipality="Doi Angkhang" pageId="34" pageNumber="151" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B5880EFFE3E51051CD85A16140FD75" box="[488,587,681,705]" pageId="34" pageNumber="151" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D08FD25FFE3E510527C85A26165FD75" box="[601,622,682,705]" pageId="34" pageNumber="151">♀</specimenCount>
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: “
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<collectingCountry id="F319763CFFE3E51052AE85A1601DFD75" box="[651,790,681,705]" name="Thailand" pageId="34" pageNumber="151">THAILAND</collectingCountry>
|
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,
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||
<collectingMunicipality id="6BD5ACD6FFE3E510509885C16256FD55" box="[189,349,713,737]" pageId="34" pageNumber="151">Doi Angkhang</collectingMunicipality>
|
||
,
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||
<date id="FFB0106CFFE3E510514385C162D9FD55" box="[358,466,713,737]" pageId="34" pageNumber="151" value="2010-10-24">
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<collectingDate id="EFF4E984FFE3E510514385C162D9FD55" box="[358,466,713,737]" pageId="34" pageNumber="151" value="2010-10-24">24.X.2010</collectingDate>
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||
</date>
|
||
,
|
||
<collectorName id="26FB537AFFE3E51051F985C1619CFD56" box="[476,663,713,738]" pageId="34" pageNumber="151">G. de Rougemont</collectorName>
|
||
/ Holotypus
|
||
<taxonomicName id="4C0E4D2FFFE3E510509885E2625DFCB5" authorityName="PACE" authorityYear="2012" box="[189,342,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="inversus">
|
||
<emphasis id="B97AEABEFFE3E510509885E2625DFCB5" box="[189,342,746,769]" italics="true" pageId="34" pageNumber="151">Zyras inversus</emphasis>
|
||
</taxonomicName>
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||
mihi, det. R.
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||
<bibRefCitation id="EF9F4B5DFFE3E51051D485E2616FFCB5" author="PACE, R." box="[497,612,745,769]" pageId="34" pageNumber="151" pagination="23 - 40" refId="ref58918" refString="PACE, R. 2011: Aleocharinae from India gathered by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Lavori della Societa Veneziana di Scienze Naturali 36: 23 - 40." type="journal article" year="2011">Pace, 2011</bibRefCitation>
|
||
/
|
||
<taxonomicName id="4C0E4D2FFFE3E51052A485E26011FCB5" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[641,794,746,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="inversus" status="sp. nov.">
|
||
<emphasis id="B97AEABEFFE3E51052A485E26011FCB5" box="[641,794,746,769]" italics="true" pageId="34" pageNumber="151">Zyras inversus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24957C5FFE3E5105098840263FDFC95" box="[189,246,778,801]" pageId="34" pageNumber="151" rank="species">n. sp.</taxonomicNameLabel>
|
||
, det.
|
||
<originalDetermination id="A8108E50FFE3E510510A84026274FC95" box="[303,383,778,801]" pageId="34" pageNumber="151">
|
||
R.
|
||
<bibRefCitation id="EF9F4B5DFFE3E510516984026274FC95" author="PACE, R." box="[332,383,778,801]" pageId="34" pageNumber="151" pagination="23 - 40" refId="ref58918" refString="PACE, R. 2011: Aleocharinae from India gathered by Guillaume de Rougemont (Coleoptera, Staphylinidae). - Lavori della Societa Veneziana di Scienze Naturali 36: 23 - 40." type="journal article" year="2011">Pace</bibRefCitation>
|
||
</originalDetermination>
|
||
, 2011 /
|
||
<taxonomicName id="4C0E4D2FFFE3E51051F784026186FC95" authorityName="Assing" authorityYear="2017" box="[466,653,778,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="novinversus" status="nom. nov.">
|
||
<emphasis id="B97AEABEFFE3E51051F784026186FC95" box="[466,653,778,801]" italics="true" pageId="34" pageNumber="151">Zyras novinversus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24957C5FFE3E51052B6840261EDFC95" box="[659,742,778,801]" pageId="34" pageNumber="151">nom. n.</taxonomicNameLabel>
|
||
, det.
|
||
<collectorName id="26FB537AFFE3E510509884226214FCF5" box="[189,287,809,833]" pageId="34" pageNumber="151">V. Assing</collectorName>
|
||
2016” (cRou).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE3E510509884616293F955" pageId="34" pageNumber="151" type="discussion">
|
||
<paragraph id="8BB136ACFFE3E51050988461614FFB95" blockId="34.[189,794,873,1409]" pageId="34" pageNumber="151">
|
||
<emphasis id="B97AEABEFFE3E510509884616227FC35" bold="true" box="[189,300,873,897]" pageId="34" pageNumber="151">Comment</emphasis>
|
||
: The original description is based on
|
||
<specimenCount id="9D08FD25FFE3E51052D784626206FC15" pageId="34" pageNumber="151" type="female">two females</specimenCount>
|
||
from “
|
||
<collectingCountry id="F319763CFFE3E510517A848162CAFC15" box="[351,449,905,929]" name="Thailand" pageId="34" pageNumber="151">Thailand</collectingCountry>
|
||
Doi Angkhang” (
|
||
<bibRefCitation id="EF9F4B5DFFE3E51052AF84816004FC15" author="PACE, R." box="[650,783,905,929]" pageId="34" pageNumber="151" pagination="319 - 360" refId="ref59001" refString="PACE, R. 2012 b: New data, new species, and two new genera of Aleocharinae from the Oriental Region (Insecta: Coleoptera: Staphylinidae). - Vernate 31: 319 - 360." type="journal article" year="2012">PACE 2012b</bibRefCitation>
|
||
).
|
||
<location id="8ED16077FFE3E510509884A163ECFC75" LSID="urn:lsid:plazi:treatment:03A787BAFFE3E51150C686FB623AFA75:8ED16077FFE3E510509884A163ECFC75" box="[189,231,937,961]" country="Thailand" name="The" pageId="34" pageNumber="151">The</location>
|
||
examined
|
||
<typeStatus id="54B5880EFFE3E510514E84A162C2FC75" box="[363,457,937,961]" pageId="34" pageNumber="151" type="holotype">holotype</typeStatus>
|
||
is evidently subject to postmortem darkening and has the eyes deformed, most likely as a result of the effect of chemicals or other improper treatment during original dissection.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E5105098832262B5FA35" blockId="34.[189,794,873,1409]" pageId="34" pageNumber="151">
|
||
<taxonomicName id="4C0E4D2FFFE3E5105098832262C1FBF5" authority="PACE, 2012" authorityName="PACE" authorityYear="2012" box="[189,458,1065,1089]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="inversus">
|
||
<emphasis id="B97AEABEFFE3E51050988322625AFBF5" box="[189,337,1066,1089]" italics="true" pageId="34" pageNumber="151">Zyras inversus</emphasis>
|
||
PACE, 2012
|
||
</taxonomicName>
|
||
is a primary homonym of
|
||
<taxonomicName id="4C0E4D2FFFE3E51052C583226137FBD5" authority="PACE, 2001" authorityName="PACE" authorityYear="2001" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="inversus" subGenus="Glossacantha">
|
||
<emphasis id="B97AEABEFFE3E51052C583226011FBF5" box="[736,794,1066,1089]" italics="true" pageId="34" pageNumber="151">Zyras</emphasis>
|
||
(
|
||
<emphasis id="B97AEABEFFE3E51050E38341625BFBD5" box="[198,336,1097,1121]" italics="true" pageId="34" pageNumber="151">Glossacantha</emphasis>
|
||
)
|
||
<emphasis id="B97AEABEFFE3E5105146834262B3FBD5" box="[355,440,1098,1121]" italics="true" pageId="34" pageNumber="151">inversus</emphasis>
|
||
PACE, 2001
|
||
</taxonomicName>
|
||
, a species described based on
|
||
<specimenCount id="9D08FD25FFE3E510510D836262A2FB35" box="[296,425,1129,1153]" pageId="34" pageNumber="151" type="female">two females</specimenCount>
|
||
from
|
||
<collectingRegion id="49CAF84EFFE3E51051D483616179FB35" box="[497,626,1129,1153]" country="India" name="Tamil Nadu" pageId="34" pageNumber="151">Tamil Nadu</collectingRegion>
|
||
in South
|
||
<collectingCountry id="F319763CFFE3E51052C783616011FB35" box="[738,794,1129,1153]" name="India" pageId="34" pageNumber="151">India</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EF9F4B5DFFE3E51050E183816241FB15" author="PACE, R." box="[196,330,1161,1185]" pageId="34" pageNumber="151" pagination="31 - 47" refId="ref58592" refString="PACE, R. 2001 b: Aleocharinae nuove o poco note dell'India (Coleoptera, Staphylinidae). - Nouvelle Revue d'Entomologie (N. S.) 18 (1): 31 - 47." type="journal article" year="2001">Pace 2001b</bibRefCitation>
|
||
). The junior primary homonym is here replaced with the nomen novum
|
||
<taxonomicName id="4C0E4D2FFFE3E510527183A26011FB75" authorityName="Assing" authorityYear="2017" box="[596,794,1194,1217]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="novinversus">
|
||
<emphasis id="B97AEABEFFE3E510527183A26011FB75" box="[596,794,1194,1217]" italics="true" pageId="34" pageNumber="151">Zyras novinversus</emphasis>
|
||
</taxonomicName>
|
||
(a combination of novus and inversus). It is, however, with some hesitation that I propose a replacement name. This species is most likely widespread and the possibility that one of the unrevised names described based on material from Java and Sumatra refers to this name cannot be ruled out with certainty.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E510509882A16293F955" blockId="34.[189,794,1449,1761]" pageId="34" pageNumber="151">
|
||
<emphasis id="B97AEABEFFE3E510509882A1611DFA75" bold="true" box="[189,534,1449,1473]" pageId="34" pageNumber="151">Additional material examined</emphasis>
|
||
:
|
||
<materialsCitation id="3B663CF1FFE3E510520E82A1601EF995" ID-GBIF-Occurrence="3450162389" collectingDate="1988-04-18" collectingDateMax="1988-04-27" collectingDateMin="1988-04-18" collectionCode="BMNH" collectorName="Brendell" country="Thailand" elevation="300" latitude="15.466666" location="Umphang district" longLatPrecision="1285" longitude="98.8" pageId="34" pageNumber="151" specimenCount="7" stateProvince="Tak province">
|
||
<emphasis id="B97AEABEFFE3E510520E82A16184FA75" bold="true" box="[555,655,1449,1473]" pageId="34" pageNumber="151">
|
||
<collectingCountry id="F319763CFFE3E510520E82A16184FA75" box="[555,655,1449,1473]" name="Thailand" pageId="34" pageNumber="151">Thailand</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D08FD25FFE3E510528682A261EBFA75" box="[675,736,1450,1473]" pageId="34" pageNumber="151" type="generic">7 exs.</specimenCount>
|
||
,
|
||
<collectingRegion id="49CAF84EFFE3E51052D682A16217FA55" country="Thailand" name="Tak" pageId="34" pageNumber="151">Tak province</collectingRegion>
|
||
,
|
||
<location id="8ED16077FFE3E510510382C162EFFA55" LSID="urn:lsid:plazi:treatment:03A787BAFFE3E51150C686FB623AFA75:8ED16077FFE3E510510382C162EFFA55" box="[294,484,1481,1505]" country="Thailand" latitude="15.466666" longLatPrecision="1285" longitude="98.8" name="Umphang district" pageId="34" pageNumber="151" stateProvince="Tak province">Umphang district</location>
|
||
,
|
||
<taxonomicName id="4C0E4D2FFFE3E51051CB82C16282F9B5" authority="Bae Stream, Thung Yai Wildlife Sanctuary" authorityName="Bae Stream, Thung Yai Wildlife Sanctuary" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="151" phylum="Arthropoda" rank="species" species="song">Song Bae Stream, Thung Yai Wildlife Sanctuary</taxonomicName>
|
||
,
|
||
<geoCoordinate id="EE3A506BFFE3E51051BD82E162E5F9B5" box="[408,494,1513,1537]" degrees="15" direction="north" minutes="28" orientation="latitude" pageId="34" pageNumber="151" precision="925" value="15.466666">15°28'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3A506BFFE3E51051D882E16144F9B5" box="[509,591,1513,1537]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="34" pageNumber="151" precision="925" value="98.8">98°48'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CF69B49FFE3E510527B82E161A8F9B5" box="[606,675,1513,1537]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="151" unit="m" value="300.0">
|
||
<elevation id="0023D19FFFE3E510527B82E161A8F9B5" box="[606,675,1513,1537]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="151" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
, evergreen rain forest,
|
||
<date id="FFB0106CFFE3E510511E810162DEF995" box="[315,469,1545,1569]" pageId="34" pageNumber="151" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">
|
||
<collectingDate id="EFF4E984FFE3E510511E810162DEF995" box="[315,469,1545,1569]" pageId="34" pageNumber="151" value="1988-04-18" valueMax="1988-04-27" valueMin="1988-04-18">18–27.IV.1988</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FB537AFFE3E510523581016161F995" box="[528,618,1545,1569]" pageId="34" pageNumber="151">Brendell</collectorName>
|
||
(
|
||
<collectionCode id="ED1FAE69FFE3E510525C810161C6F995" box="[633,717,1545,1569]" pageId="34" pageNumber="151">BMNH</collectionCode>
|
||
, cAss)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B663CF1FFE3E510509881216179F915" ID-GBIF-Occurrence="3450162322" collectingDate="1988-04-27" collectingDateMax="1988-05-06" collectingDateMin="1988-04-27" collectionCode="BMNH" collectorName="Karen & Brendell" country="Thailand" county="Umphang district" elevation="300" latitude="15.5" location="Thung Yai Wildlife Sanctuary" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" pageId="34" pageNumber="151" specimenCount="12" stateProvince="Tak province">
|
||
<specimenCount id="9D08FD25FFE3E51050988121620CF9F5" box="[189,263,1577,1601]" pageId="34" pageNumber="151" type="generic">12 exs.</specimenCount>
|
||
,
|
||
<collectingRegion id="49CAF84EFFE3E5105133812162A3F9F5" box="[278,424,1577,1601]" country="Thailand" name="Tak" pageId="34" pageNumber="151">Tak province</collectingRegion>
|
||
,
|
||
<collectingCounty id="62D04E20FFE3E510519D81216177F9F5" box="[440,636,1577,1601]" pageId="34" pageNumber="151">Umphang district</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BD5ACD6FFE3E51052A98122600BF9F5" box="[652,768,1577,1601]" pageId="34" pageNumber="151">Mae Chan</collectingMunicipality>
|
||
– Mae Klong confluence,
|
||
<location id="8ED16077FFE3E51051EF8141601DF9D5" LSID="urn:lsid:plazi:treatment:03A787BAFFE3E51150C686FB623AFA75:8ED16077FFE3E51051EF8141601DF9D5" box="[458,790,1609,1633]" country="Thailand" county="Umphang district" latitude="15.5" longLatPrecision="1284" longitude="98.8" municipality="Mae Chan" name="Thung Yai Wildlife Sanctuary" pageId="34" pageNumber="151" stateProvince="Tak province">Thung Yai Wildlife Sanctuary</location>
|
||
,
|
||
<geoCoordinate id="EE3A506BFFE3E510509881616218F935" box="[189,275,1641,1665]" degrees="15" direction="north" minutes="30" orientation="latitude" pageId="34" pageNumber="151" precision="925" value="15.5">15°30'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3A506BFFE3E510513A8161627AF935" box="[287,369,1641,1665]" degrees="98" direction="east" minutes="48" orientation="longitude" pageId="34" pageNumber="151" precision="925" value="98.8">98°48'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CF69B49FFE3E5105158816162C9F935" box="[381,450,1641,1665]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="151" unit="m" value="300.0">
|
||
<elevation id="0023D19FFFE3E5105158816162C9F935" box="[381,450,1641,1665]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="151" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
, edge of
|
||
<collectorName id="26FB537AFFE3E51052068162616FF935" box="[547,612,1642,1665]" pageId="34" pageNumber="151">Karen</collectorName>
|
||
clearing,
|
||
<date id="FFB0106CFFE3E51052EB81616211F915" pageId="34" pageNumber="151" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">
|
||
<collectingDate id="EFF4E984FFE3E51052EB81616211F915" pageId="34" pageNumber="151" value="1988-04-27" valueMax="1988-05-06" valueMin="1988-04-27">27.IV.– 6.V.1988</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26FB537AFFE3E5105145818162B1F915" box="[352,442,1673,1697]" pageId="34" pageNumber="151">Brendell</collectorName>
|
||
(
|
||
<collectionCode id="ED1FAE69FFE3E51051F58181612FF915" box="[464,548,1673,1697]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="34" pageNumber="151">BMNH</collectionCode>
|
||
, cAss)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B663CF1FFE3E51052A181816298F955" ID-GBIF-Occurrence="3450162334" collectingDate="2004-05" country="Laos" elevation="1400" location="Phongsaly province" pageId="34" pageNumber="151" specimenCount="15" stateProvince="Phongsaly province">
|
||
<emphasis id="B97AEABEFFE3E51052A1818161BCF915" bold="true" box="[644,695,1673,1697]" pageId="34" pageNumber="151">
|
||
<collectingCountry id="F319763CFFE3E51052A1818161BCF915" box="[644,695,1673,1697]" name="Laos" pageId="34" pageNumber="151">Laos</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D08FD25FFE3E51052E98181601DF915" box="[716,790,1673,1697]" pageId="34" pageNumber="151" type="generic">15 exs.</specimenCount>
|
||
,
|
||
<collectingRegion id="49CAF84EFFE3E510509881A162ABF975" box="[189,416,1705,1729]" country="Laos" name="Phongsali" pageId="34" pageNumber="151">Phongsaly province</collectingRegion>
|
||
,
|
||
<collectingRegion id="49CAF84EFFE3E510519F81A16123F975" box="[442,552,1705,1729]" country="Laos" name="Phongsali" pageId="34" pageNumber="151">Phongsaly</collectingRegion>
|
||
env.,
|
||
<quantity id="4CF69B49FFE3E51052A681A1601DF975" box="[643,790,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="34" pageNumber="151" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">
|
||
<elevation id="0023D19FFFE3E51052A681A1601DF975" box="[643,790,1705,1729]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="34" pageNumber="151" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFB0106CFFE3E510509881C1620EF955" box="[189,261,1737,1761]" pageId="34" pageNumber="151" value="2004-05">
|
||
<collectingDate id="EFF4E984FFE3E510509881C1620EF955" box="[189,261,1737,1761]" pageId="34" pageNumber="151" value="2004-05">V.2004</collectingDate>
|
||
</date>
|
||
(cMar, cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE3E5115098800163F4FC55" lastPageId="35" lastPageNumber="152" pageId="34" pageNumber="151" type="description">
|
||
<paragraph id="8BB136ACFFE3E510509880016729FE75" blockId="34.[189,794,1801,2081]" lastBlockId="34.[832,1437,233,2081]" pageId="34" pageNumber="151">
|
||
<emphasis id="B97AEABEFFE3E51050988001625DF895" bold="true" box="[189,342,1801,1825]" pageId="34" pageNumber="151">Redescription</emphasis>
|
||
: Body length
|
||
<quantity id="4CF69B49FFE3E51051D580016179F895" box="[496,626,1801,1825]" metricMagnitude="-3" metricUnit="m" metricValue="5.55" metricValueMax="6.6" metricValueMin="4.5" pageId="34" pageNumber="151" unit="mm" value="5.55" valueMax="6.6" valueMin="4.5">4.5–6.6 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CF69B49FFE3E51051218021628EF8F5" box="[260,389,1833,1857]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="34" pageNumber="151" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1–2.9 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="13352A29FFE3E510520280216195F8F5" box="[551,670,1833,1857]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="34" pageNumber="151">Figs 30–31</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE3E5105296802161C5F8F5" box="[691,718,1833,1857]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="34" pageNumber="151">67</figureCitation>
|
||
,
|
||
<figureCitation id="13352A29FFE3E51052C180216005F8F5" box="[740,782,1833,1857]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="34" pageNumber="151">106</figureCitation>
|
||
): head and pronotum black; elytra reddish-yellow with the postero-lateral portions blackish, rarely nearly uniformly dark with only the humeral angles indistinctly paler; abdomen with segments II–V reddish (rarely reddishbrown with reddish margins), segment VI reddish or brown with reddish margins, segment VII usually darkbrown to blackish-brown with the anterior portion reddish, rarely completely reddish, segment VIII darkbrown to blackish-brown, segment IX dark-brown, and tergite X reddish; legs pale-yellowish; antennae blackishbrown to black with antennomeres I–II reddish-yellow to brown and the apex of antennomere XI dark-reddish; maxillary palpi yellowish-brown, with the terminal palpomere yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E510536586C160F2FC95" blockId="34.[832,1437,233,2081]" pageId="34" pageNumber="151">
|
||
Head (
|
||
<figureCitation id="13352A29FFE3E51053AF86C160DEFE55" box="[906,981,457,481]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="34" pageNumber="151">Fig. 67</figureCitation>
|
||
) moderately transverse, broadly impunctate along middle; punctation in lateral dorsal portions moderately coarse and sparse. Eyes large, twice as long as postocular region in dorsal view, or nearly so. Antenna (
|
||
<figureCitation id="13352A29FFE3E5105362854160B6FDD5" box="[839,957,585,609]" captionStart="Figs 1–41" captionStartId="45.[151,191,1956,1978]" captionTargetBox="[152,1389,232,1855]" captionTargetId="figure-203@45.[151,1398,232,1874]" captionTargetPageId="45" captionText="Figs 1–41:Antennae of Zyrasspp.:kraatzi(1),exasperatus(2),pindarae (3–5),perforatus(6),championi(7),pallipes (8),condignus (9), nitens (10), truncatus (11), nilgiriensis (12), hirsutiventris (13), russiceps (14), brignolii (15), hastatus (16), parageminus (17–18), latilobatus (19), alternans (20), hirtus (21), nigroaeneus (22), luteipes (23), longilobatus (24), morvani (25), gratellus (Malaysia) (26), castaneus, lectotype (27),proximus (28–29),novinversus (30–31), brevilobatus(32), parvicollis(33), ambulans (34),matangensis (35), densissimus (36), densihirtus (37), nigrihirtus (38), gratellus (Sulawesi) (39), gardneri (40), titan (41). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742367" httpUri="https://zenodo.org/record/5742367/files/figure.png" pageId="34" pageNumber="151">Figs 30–31</figureCitation>
|
||
)
|
||
<quantity id="4CF69B49FFE3E51053EB85416746FDD5" box="[974,1101,585,609]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="34" pageNumber="151" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
|
||
long; antennomeres IV and V weakly oblong, VI approximately as long as broad, VII approximately as long as broad or weakly transverse, VIII–X of gradually increasing width and increasingly transverse, X approximately 1.5 times as broad as long, and XI elongate, approximately as long as the combined length of VIII–X.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E5105365842260B7FBD5" blockId="34.[832,1437,233,2081]" pageId="34" pageNumber="151">
|
||
Pronotum (
|
||
<figureCitation id="13352A29FFE3E51053E68421671BFCF5" box="[963,1040,809,833]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="34" pageNumber="151">Fig. 67</figureCitation>
|
||
) approximately 1.2 times as broad as long and 1.07–1.15 times as broad as head, broadest near anterior angles, distinctly tapering posteriad; lateral margins straight or weakly sinuate in posterior two-thirds (dorsal view); punctation very coarse, very irregularly distributed, arranged in more or less distinct clusters, these clusters often situated in very shallow impressions, middle and lateral portion with extensive impunctate patches; lateral margins each with four long, black setae.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E51053658361666CFAB5" blockId="34.[832,1437,233,2081]" pageId="34" pageNumber="151">
|
||
Elytra (
|
||
<figureCitation id="13352A29FFE3E51053B1836160D4FB35" box="[916,991,1129,1153]" captionStart="Figs 55–67" captionStartId="49.[151,191,1991,2013]" captionTargetBox="[151,1397,232,1962]" captionTargetId="figure-119@49.[151,1398,232,1971]" captionTargetPageId="49" captionText="Figs 55–67: Forebody of Zyrasspp.:parageminus (55–56; 55:holotype;56:holotype ofZ. nameriensis),latilobatus (57),alternans (58), hirtus (59), nilgiriensis (60), russiceps (61), gardneri (62), hirsutiventris (63),luteipes (64), proximus, holotype (65), brignolii, holotype of Z. thainiger (66), novinversus (67). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742375" httpUri="https://zenodo.org/record/5742375/files/figure.png" pageId="34" pageNumber="151">Fig. 67</figureCitation>
|
||
) 0.85–0.93 times as long as pronotum; punctation coarse, defined, moderately dense to dense, and almost regularly distributed, somewhat less dense in posterior portion. Hind wings fully developed. Metatarsomere I shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E51053658201671FF835" blockId="34.[832,1437,233,2081]" pageId="34" pageNumber="151">
|
||
Abdomen (
|
||
<figureCitation id="13352A29FFE3E510539E82016719FA95" box="[955,1042,1289,1313]" captionStart="Figs 100–116" captionStartId="54.[189,229,2007,2029]" captionTargetBox="[189,1436,231,1959]" captionTargetId="figure-125@54.[189,1436,231,1970]" captionTargetPageId="54" captionText="Figs 100–116: Abdomen of Zyras spp.: luteipes (100), nilgiriensis (101), brevilobatus(102), brignolii (103), parvicollis (104), proximus, holotype (105), novinversus (106), ambulans (107), matangensis (108), nitens (109), gratellus (Malaysia) (110), densissimus (111), gratellus (Sulawesi) (112), titan (113), russiceps (114), densihirtus (115), nigrihirtus (116). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742387" httpUri="https://zenodo.org/record/5742387/files/figure.png" pageId="34" pageNumber="151">Fig. 106</figureCitation>
|
||
) narrower than elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V with dense and rather coarse non-setiferous punctation; tergite III with a lateral setiferous puncture on either side and with four setiferous punctures at posterior margin; tergites IV–V with a lateral setiferous puncture on either side (often accompanied by few non-setiferous punctures) and with six setiferous punctures at posterior margin; tergite VI anteriorly with more or less extensive non-setiferous punctation, with a lateral setiferous puncture on either side (often accompanied by few non-setiferous punctures) and with six setiferous punctures at posterior margin; tergite VII with a transverse band of sparse and rather fine non-setiferous punctures anteriorly, with a transverse series of approximately eight setiferous punctures at posterior fourth, and with scattered setiferous punctures at posterior margin, palisade fringe present; tergite VIII with sparse setiferous punctures in posterior portion, posterior margin with sexual dimorphism.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE3E51153658082624FFE95" blockId="34.[832,1437,233,2081]" lastBlockId="35.[151,756,233,385]" lastPageId="35" lastPageNumber="152" pageId="34" pageNumber="151">
|
||
♂: posterior margin of tergite VIII (
|
||
<figureCitation id="13352A29FFE3E51054EB8081662DF815" box="[1230,1318,1929,1953]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="34" pageNumber="151">Fig. 201</figureCitation>
|
||
) with four weakly pronounced blunt projections; posterior margin of sternite VIII (
|
||
<figureCitation id="13352A29FFE3E51053CB80C1674EF855" box="[1006,1093,1993,2017]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="34" pageNumber="151">Fig. 202</figureCitation>
|
||
) weakly convex, nearly truncate in the middle; median lobe of aedeagus approximately
|
||
<quantity id="4CF69B49FFE3E51053658F016095F795" box="[832,926,2057,2081]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="34" pageNumber="151" unit="mm" value="0.75">0.75 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="13352A29FFE3E51054B88F016620F795" box="[1181,1323,2057,2081]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="34" pageNumber="151">Figs 198–199</figureCitation>
|
||
; paramere (
|
||
<figureCitation id="13352A29FFE2E51150BA87E163FDFEB5" box="[159,246,233,257]" captionStart="Figs 191–207" captionStartId="59.[151,191,2011,2033]" captionTargetBox="[153,1385,236,1953]" captionTargetId="figure-115@59.[151,1398,231,1970]" captionText="Figs 191–207: Zyras proximus (191–197; 191–192: Thailand; 193–197: India), Z. novinversus (198–202), Z. brevilobatus (203–205), and Z. parvicollis (206–207): median lobe of aedeagus in lateral and in ventral view (191–194, 198–199, 203–204); paramere (195, 200, 205); male tergite VIII (196, 201, 206); male sternite VIII (197, 202, 207). Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5888374" httpUri="https://zenodo.org/record/5888374/files/figure.png" pageId="35" pageNumber="152">Fig. 200</figureCitation>
|
||
) approximately
|
||
<quantity id="4CF69B49FFE2E511518887E162F4FEB5" box="[429,511,233,257]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="35" pageNumber="152" unit="mm" value="0.8">0.8 mm</quantity>
|
||
long and with rather long apical lobe.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE2E51150B2862262ADFE35" blockId="35.[151,756,233,385]" pageId="35" pageNumber="152">♀: posterior margin of tergite VIII smoothly convex or with shallow median concavity; posterior margin of sternite VIII weakly concave.</paragraph>
|
||
<paragraph id="8BB136ACFFE2E51150B286A163F5FD95" blockId="35.[151,756,425,545]" pageId="35" pageNumber="152">
|
||
<emphasis id="B97AEABEFFE2E51150B286A16276FE75" bold="true" box="[151,381,425,449]" pageId="35" pageNumber="152">Intraspefic variation</emphasis>
|
||
: Body size, the coloration of the elytra and abdomen, as well as the shapes of tergite and sternite VIII are subject to rather pronounced intraspefic variation.
|
||
</paragraph>
|
||
<paragraph id="8BB136ACFFE2E51150B2854163F4FC55" blockId="35.[151,756,585,993]" pageId="35" pageNumber="152">
|
||
<emphasis id="B97AEABEFFE2E51150B28541626DFDD5" bold="true" box="[151,358,585,609]" pageId="35" pageNumber="152">Comparative notes</emphasis>
|
||
: As can be inferred from the generally similar external characters, particularly the conspicuously irregular punctation and the often slightly uneven surface of the pronotum, and the punctation pattern of the abdomen, as well as from the sexual dimorphism of tergite VIII, the similar modifications of the male tergite VIII, and from the relatively long apical lobe of the paramere,
|
||
<taxonomicName id="4C0E4D2FFFE2E51150FF8422627AFCF5" authorityName="Assing" authorityYear="2017" box="[218,369,810,833]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="novinversus">
|
||
<emphasis id="B97AEABEFFE2E51150FF8422627AFCF5" box="[218,369,810,833]" italics="true" pageId="35" pageNumber="152">Z. novinversus</emphasis>
|
||
</taxonomicName>
|
||
is very closely allied to
|
||
<taxonomicName id="4C0E4D2FFFE2E5115255842261E6FCF5" authorityName="CAMERON" authorityYear="1939" box="[624,749,810,833]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis id="B97AEABEFFE2E5115255842261E6FCF5" box="[624,749,810,833]" italics="true" pageId="35" pageNumber="152">Z. proximus</emphasis>
|
||
</taxonomicName>
|
||
, from which it is distinguished by the different coloration of the abdomen, an on average less uneven disc of the pronotum, less pronounced modifications of the male tergite VIII, and by the shape of the ventral process of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3146527FFE2E51150B28301623AFA75" pageId="35" pageNumber="152" type="distribution">
|
||
<paragraph id="8BB136ACFFE2E51150B28301623AFA75" blockId="35.[151,756,1033,1473]" pageId="35" pageNumber="152">
|
||
<emphasis id="B97AEABEFFE2E51150B283016106FB95" bold="true" box="[151,525,1033,1057]" pageId="35" pageNumber="152">Distribution and natural history</emphasis>
|
||
: Although currently known only from
|
||
<collectingCountry id="F319763CFFE2E5115150832162DCFBF5" box="[373,471,1065,1089]" name="Thailand" pageId="35" pageNumber="152">Thailand</collectingCountry>
|
||
and
|
||
<collectingCountry id="F319763CFFE2E51152388322615BFBF5" box="[541,592,1066,1089]" name="Laos" pageId="35" pageNumber="152">Laos</collectingCountry>
|
||
(
|
||
<figureCitation id="13352A29FFE2E5115243832161A4FBF5" box="[614,687,1065,1089]" captionStart="Map 5" captionStartId="65.[151,196,1829,1850]" captionTargetBox="[151,1398,827,1821]" captionTargetId="figure-87@65.[151,1398,827,1821]" captionTargetPageId="65" captionText="Map 5: Revised distributions of Zyras castaneus (black circles) and Z. novinversus (white circles) in the southern East Palaearctic" figureDoi="http://doi.org/10.5281/zenodo.5742405" httpUri="https://zenodo.org/record/5742405/files/figure.png" pageId="35" pageNumber="152">Map 5</figureCitation>
|
||
), this species is most likely more widespread in the Oriental region, as is suggested by the vast distribution of the similar and closely related
|
||
<taxonomicName id="4C0E4D2FFFE2E51151D38382617DFB15" authorityName="CAMERON" authorityYear="1939" box="[502,630,1162,1185]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis id="B97AEABEFFE2E51151D38382617DFB15" box="[502,630,1162,1185]" italics="true" pageId="35" pageNumber="152">Z. proximus</emphasis>
|
||
</taxonomicName>
|
||
, as well as by the fact that at least the examined non-type material was evidently collected on the wing (practically all the specimens have the hind wings fully unfolded). The additional material from
|
||
<collectingCountry id="F319763CFFE2E51151978201611FFA95" box="[434,532,1289,1313]" name="Thailand" pageId="35" pageNumber="152">Thailand</collectingCountry>
|
||
was collected in an evergreen rain forest and at the edge of a clearing at an altitude of
|
||
<quantity id="4CF69B49FFE2E511512882416258FAD5" box="[269,339,1353,1377]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="35" pageNumber="152" unit="m" value="300.0">300 m</quantity>
|
||
, together with several other species of
|
||
<taxonomicName id="4C0E4D2FFFE2E51150B28262621EFA35" authority="sensu" authorityName="SenSU" box="[151,277,1386,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="186" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B97AEABEFFE2E51150B2826263D9FA35" box="[151,210,1386,1409]" italics="true" pageId="35" pageNumber="152">Zyras</emphasis>
|
||
sensu
|
||
</taxonomicName>
|
||
strictu, among them numerous
|
||
<taxonomicName id="4C0E4D2FFFE2E511524B826261E6FA35" authorityName="CAMERON" authorityYear="1939" box="[622,749,1386,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="152" phylum="Arthropoda" rank="species" species="proximus">
|
||
<emphasis id="B97AEABEFFE2E511524B826261E6FA35" box="[622,749,1386,1409]" italics="true" pageId="35" pageNumber="152">Z. proximus</emphasis>
|
||
</taxonomicName>
|
||
. The specimens from
|
||
<collectingCountry id="F319763CFFE2E51151A1828262BCFA15" box="[388,439,1418,1441]" name="Laos" pageId="35" pageNumber="152">Laos</collectingCountry>
|
||
were taken at an altitude of
|
||
<quantity id="4CF69B49FFE2E51150B282A16226FA75" box="[151,301,1449,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="35" pageNumber="152" unit="m" value="1400.0" valueMax="1500.0" valueMin="1300.0">1300–1500 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |