127 lines
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127 lines
17 KiB
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<document id="A734812C8F196B0868CB0EDACF59FB0F" ID-CLB-Dataset="7383" ID-DOI="10.11646/zootaxa.4951.3.7" ID-GBIF-Dataset="a6ee038f-4cda-4f38-8fde-8ad0efec8f0f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4668453" ID-ZooBank="EB876346-4282-4B4E-8585-ED50F1A616E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617782094822" checkinUser="plazi" docAuthor="Martens, Jochen, Maghradze, Eter & Barjadze, Shalva" docDate="2021" docId="03A787A4FFF3FFDA3584F6FF188EF8D7" docLanguage="en" docName="zootaxa.4951.3.7.pdf" docOrigin="Zootaxa 4951 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nemastomatidae" docType="treatment" docVersion="8" lastPageNumber="553" masterDocId="FF9EFFDCFFFFFFD63513F30F1D1AFFA4" masterDocTitle="Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)" masterLastPageNumber="558" masterPageNumber="541" pageNumber="553" updateTime="1698936134096" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="093922193AE7A1941BF3358DED3100DC">
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<mods:title id="3A997990475B85F3369BA2BFAA40369C">Two new species of the genus Nemaspela Šilhavý from caves in Georgia (Opiliones Nemastomatidae)</mods:title>
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<mods:name id="12B4F59D5E2734E930FC74DECD5A0CFE" type="personal">
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<mods:roleTerm id="8BBF2E216966C7A876230D7E4A37AA80">Author</mods:roleTerm>
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<mods:namePart id="CD8E4A0353827679558C350BCE3AF357">Martens, Jochen</mods:namePart>
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<mods:affiliation id="F4DFF64438C73855582EDF99E3D28E54">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany; Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany</mods:affiliation>
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</mods:role>
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<mods:namePart id="5C635D72C2D828B83CADD71D863D2D6C">Maghradze, Eter</mods:namePart>
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<mods:affiliation id="AD4CB2973535FF4E73C8E47B90EDA634">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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<mods:name id="24323E50DF33FE69B89A51F86F7699BD" type="personal">
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<mods:roleTerm id="35D8FCF258536DC949742F97AEC759F7">Author</mods:roleTerm>
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<mods:namePart id="E0E75013DA6169C07480F9E188C261D6">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="9902320CADA90FB69531A26528267EB3">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Georgia</mods:affiliation>
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<mods:typeOfResource id="ED072788701305F044ED2694D26AE3C6">text</mods:typeOfResource>
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<mods:title id="ACE8EF07A3CB18FE4B15D440BEAC868A">Zootaxa</mods:title>
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<mods:part id="06A4867F3BD04B499DCEFFB025D72AF8">
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<mods:date id="EC497A79000B38288CA84E77F439717A">2021</mods:date>
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<mods:number id="EB6D436F71FEC0F3DF95367C86F0063B">2021-04-07</mods:number>
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<mods:number id="0E299680776A9AB2F62EFDED1C326EFF">4951</mods:number>
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<treatment id="03A787A4FFF3FFDA3584F6FF188EF8D7" ID-DOI="http://doi.org/10.5281/zenodo.4685059" ID-GBIF-Taxon="181003510" ID-Zenodo-Dep="4685059" LSID="urn:lsid:plazi:treatment:03A787A4FFF3FFDA3584F6FF188EF8D7" httpUri="http://treatment.plazi.org/id/03A787A4FFF3FFDA3584F6FF188EF8D7" lastPageNumber="553" pageId="12" pageNumber="553">
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<subSubSection id="C3146539FFF3FFDA3584F6FF1F3AF9AE" box="[151,544,1520,1547]" pageId="12" pageNumber="553" type="nomenclature">
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<paragraph id="8BB136B2FFF3FFDA3584F6FF1F3AF9AE" blockId="12.[151,1437,1520,1907]" box="[151,544,1520,1547]" pageId="12" pageNumber="553">
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<heading id="D0F981DEFFF3FFDA3584F6FF1F3AF9AE" bold="true" box="[151,544,1520,1547]" fontSize="11" level="1" pageId="12" pageNumber="553" reason="1">
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<emphasis id="B97AEAA0FFF3FFDA3584F6FF1F3AF9AE" bold="true" box="[151,544,1520,1547]" pageId="12" pageNumber="553">
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<taxonomicName id="4C0E4D31FFF3FFDA3584F6FF1C7AF9AE" ID-CoL="D7W" box="[151,352,1520,1546]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="553" phylum="Arthropoda" rank="family">Nemastomatidae</taxonomicName>
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in the Holarctic
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3146539FFF3FFDA35D4F51A188EF8D7" pageId="12" pageNumber="553" type="description">
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<paragraph id="8BB136B2FFF3FFDA35D4F51A188EF8D7" blockId="12.[151,1437,1520,1907]" pageId="12" pageNumber="553">
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This opilionid family of predominantly small species shows a holarctic distribution. The nominate subfamily,
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<taxonomicName id="4C0E4D31FFF3FFDA3584F5361CF6F9F7" authority="Simon, 1879" authorityName="Simon" authorityYear="1879" box="[151,492,1593,1619]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="553" phylum="Arthropoda" rank="subFamily" subFamily="Nemastomatinae">Nemastomatinae Simon, 1879</taxonomicName>
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has its strongholds in the West Palearctic with a distributional focus in central and southern Europe,
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<collectingCountry id="F3197622FFF3FFDA3473F5521CAAF9D3" box="[352,432,1629,1655]" name="Turkey" pageId="12" pageNumber="553">Turkey</collectingCountry>
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and the Caucasus (
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<bibRefCitation id="EF9F4B43FFF3FFDA379FF5521E4FF9D3" author="Schonhofer, A. L." box="[652,853,1629,1655]" pageId="12" pageNumber="553" pagination="1 - 68" refId="ref10322" refString="Schonhofer, A. L. (2013) A taxonomic catalogue of the Dyspnoi Hansen and Sorensen, 1904 (Arachnida: Opiliones). Zootaxa, 3679 (1), 1 - 68. https: // doi. org / 10.11646 / zootaxa. 3679.1.1" type="journal article" year="2013">Schönhofer 2013</bibRefCitation>
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); a few recently discovered outliers extend the area to Central Asia and western
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<collectingCountry id="F3197622FFF3FFDA34C0F58E1F0CF93F" box="[467,534,1665,1691]" name="China" pageId="12" pageNumber="553">China</collectingCountry>
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(
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<bibRefCitation id="EF9F4B43FFF3FFDA3737F58E1FFEF93F" author="Snegovaya, N. Yu." box="[548,740,1665,1691]" pageId="12" pageNumber="553" pagination="351 - 354" refId="ref10578" refString="Snegovaya, N. Yu. (2010) New harvestman genus and species from Kyrgyz Republic (Kyrgyzstan) (Aravhnida: Opiliones: Nemastomatidae). Acta Zoologica Bulgarica, 62, 351 - 354." type="journal article" year="2010">Snegovaya 2010</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA37FDF58E1E96F93F" author="Martens, J." box="[750,908,1665,1691]" pageId="12" pageNumber="553" pagination="444 - 450" refId="ref9890" refString="Martens, J. (2016) Sinostoma yunnanicum, the first nemastomatine harvestman in China (Arachnida: Opiliones: Nemastomatidae). Zootaxa, 4126 (3), 444 - 450. https: // doi. org / 10.11646 / zootaxa. 4126.3.9." type="journal article" year="2016">Martens 2016</bibRefCitation>
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,
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<bibRefCitation id="EF9F4B43FFF3FFDA3684F58E1EC8F93F" author="Martens, J." box="[919,978,1665,1691]" pageId="12" pageNumber="553" pagination="187 - 201" refId="ref9938" refString="Martens, J. (2017) On Starengovia Snegovaya, a genus of Asian nemastomatines (Arachnida: Opiliones: Nemastomatidae). Revue suisse de Zoologie, 124, 187 - 201. https: // doi. org / 10.5281 / zenodo. 893462" type="journal article" year="2017">2017</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA36CDF58E19F2F93F" author="Zhang, C. & Martens, J." box="[990,1256,1665,1691]" pageId="12" pageNumber="553" pagination="105 - 115" refId="ref10983" refString="Zhang, C. & Martens, J. (2018) Ancient home or in exile? The easternmost species of genus Starengovia Snegovaya, 2010 found in China (Opiliones, Nemastomatidae, Nemastomatinae). ZooKeys, 770, 105 - 115. https: // doi. org / 10.3897 / zookeys. 770.25491" type="journal article" year="2018">Zhang & Martens 2018</bibRefCitation>
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). The subfamily comprises about 120 species (
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<bibRefCitation id="EF9F4B43FFF3FFDA34F8F5AA1FA8F91B" author="Schonhofer, A. L." box="[491,690,1701,1727]" pageId="12" pageNumber="553" pagination="1 - 68" refId="ref10322" refString="Schonhofer, A. L. (2013) A taxonomic catalogue of the Dyspnoi Hansen and Sorensen, 1904 (Arachnida: Opiliones). Zootaxa, 3679 (1), 1 - 68. https: // doi. org / 10.11646 / zootaxa. 3679.1.1" type="journal article" year="2013">Schönhofer 2013</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA37ACF5AA1E7BF964" author="Martens, J." box="[703,865,1701,1728]" pageId="12" pageNumber="553" pagination="444 - 450" refId="ref9890" refString="Martens, J. (2016) Sinostoma yunnanicum, the first nemastomatine harvestman in China (Arachnida: Opiliones: Nemastomatidae). Zootaxa, 4126 (3), 444 - 450. https: // doi. org / 10.11646 / zootaxa. 4126.3.9." type="journal article" year="2016">Martens 2016</bibRefCitation>
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). The second subfamily,
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<taxonomicName id="4C0E4D31FFF3FFDA316FF5AA1C31F947" authority="Shear & Gruber, 1983" authorityName="Shear & Gruber" authorityYear="1983" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="553" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">
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Ortholasmatinae
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<bibRefCitation id="EF9F4B43FFF3FFDA302DF5AA1C31F947" author="Shear, W. A. & Gruber, J." pageId="12" pageNumber="553" pagination="1 - 65" refId="ref10511" refString="Shear, W. A. & Gruber, J. (1983) The opilionid subfamily Ortholasmatinae (Opiliones, Troguloidea, Nemastomatidae). American Museum Novitates, 2757, 1 - 65." type="journal article" year="1983">Shear & Gruber, 1983</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="EF9F4B43FFF3FFDA3429F5C61F2BF947" author="Shear, W. A. & Gruber, J." box="[314,561,1737,1763]" pageId="12" pageNumber="553" pagination="1 - 65" refId="ref10511" refString="Shear, W. A. & Gruber, J. (1983) The opilionid subfamily Ortholasmatinae (Opiliones, Troguloidea, Nemastomatidae). American Museum Novitates, 2757, 1 - 65." type="journal article" year="1983">Shear & Gruber 1983</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA372EF5C61FA5F947" author="Shear, W. A." box="[573,703,1737,1763]" pageId="12" pageNumber="553" pagination="191 - 198" refId="ref10409" refString="Shear, W. A. (2006) Martensolasma jocheni, a new genus and species of harvestman from Mexico (Opiliones, Nemastomatidae, Ortholasmatinae). Zootaxa, 1325, 191 - 198. https: // doi. org / 10.11646 / zootaxa. 1325.1.11" type="journal article" year="2006">Shear 2006</bibRefCitation>
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,
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<bibRefCitation id="EF9F4B43FFF3FFDA37D9F5C61E1EF947" author="Shear, W. A." box="[714,772,1737,1763]" pageId="12" pageNumber="553" pagination="9 - 45" refId="ref10458" refString="Shear, W. A. (2010) New species and records of Ortholasmatine harvestmen from Mexico, Honduras and the western United States (Opiliones, Nemastomatidae, Ortholasmatinae). ZooKeys, 52, 9 - 45. https: // doi. org / 10.3897 / zookeys. 52.471" type="journal article" year="2010">2010</bibRefCitation>
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) is distributed in
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<collectingCountry id="F3197622FFF3FFDA36DBF5C6192BF947" box="[968,1073,1737,1763]" name="China" pageId="12" pageNumber="553">East Asia</collectingCountry>
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including
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<collectingCountry id="F3197622FFF3FFDA31BAF5C619F3F947" box="[1193,1257,1737,1763]" name="Japan" pageId="12" pageNumber="553">Japan</collectingCountry>
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(
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<bibRefCitation id="EF9F4B43FFF3FFDA31EBF5C61882F947" author="Martens, J." box="[1272,1432,1737,1763]" pageId="12" pageNumber="553" pagination="79 - 110" refId="ref9985" refString="Martens, J. (2019) An ancient radiation: Ortholasmatine harvestmen in Asia - a new genus, three new species and a revision of the known species (Arachnida, Opiliones, Nemastomatidae). Revue suisse de Zoologie, 126 (1), 79 - 110. https: // doi. org / 10.5281 / zenodo. 2619524" type="journal article" year="2019">Martens 2019</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA3584F5E21CF7F8A3" author="Schwendinger, P. J. & Gruber, J." box="[151,493,1773,1799]" pageId="12" pageNumber="553" pagination="57 - 60" refId="ref10371" refString="Schwendinger, P. J. & Gruber, J. (1992) A new Dendrolasma (Opiliones, Nemastomatidae) from Thailand. Bulletin of the British Arachnological Society, 9, 57 - 60." type="journal article" year="1992">Schwendinger & Gruber 1992</bibRefCitation>
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), and the west coast of North America. Its area extends southward to Central America, the Mexican highlands and the Honduran tropics (
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<bibRefCitation id="EF9F4B43FFF3FFDA37FEF41E1E6BF88F" author="Shear, W. A." box="[749,881,1809,1835]" pageId="12" pageNumber="553" pagination="9 - 45" refId="ref10458" refString="Shear, W. A. (2010) New species and records of Ortholasmatine harvestmen from Mexico, Honduras and the western United States (Opiliones, Nemastomatidae, Ortholasmatinae). ZooKeys, 52, 9 - 45. https: // doi. org / 10.3897 / zookeys. 52.471" type="journal article" year="2010">Shear 2010</bibRefCitation>
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;
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<bibRefCitation id="EF9F4B43FFF3FFDA366FF41E199EF88F" author="Cruz-Lopez, J. A. & Cruz-Bonilla, A. & Francke, O. F." box="[892,1156,1809,1835]" pageId="12" pageNumber="553" pagination="714 - 729" refId="ref9188" refString="Cruz-Lopez, J. A., Cruz-Bonilla, A. & Francke, O. F. (2018) Molecules and morphology reveal a new aberrant harvestman genus of Ortholasmatinae (Opiliones, Dypsnoi, Nemastomatidae) from Mexico. Systematics and Biodiversity, 16, 714 - 729. https: // doi. org / 10.1080 / 14772000.2018.1476416" type="journal article" year="2018">
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Cruz-Lopéz
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<emphasis id="B97AEAA0FFF3FFDA311BF41E195AF88F" box="[1032,1088,1809,1835]" italics="true" pageId="12" pageNumber="553">et al.</emphasis>
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2018
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</bibRefCitation>
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). This subfamily is much poorer in genera and species than
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<taxonomicName id="4C0E4D31FFF3FFDA3708F43A1FC1F8EB" authorityName="Simon" authorityYear="1879" box="[539,731,1845,1871]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="553" phylum="Arthropoda" rank="subFamily" subFamily="Nemastomatinae">Nemastomatinae</taxonomicName>
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; seven genera of the former are described and 18 of the latter. The majority of
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<taxonomicName id="4C0E4D31FFF3FFDA345DF4561F1DF8D7" authorityName="Shear & Gruber" authorityYear="1983" box="[334,519,1881,1907]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="553" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
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species occur in western North America and in Central America (
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<bibRefCitation id="EF9F4B43FFF3FFDA31F4F4561893F8D7" author="Martens, J." box="[1255,1417,1881,1907]" pageId="12" pageNumber="553" pagination="79 - 110" refId="ref9985" refString="Martens, J. (2019) An ancient radiation: Ortholasmatine harvestmen in Asia - a new genus, three new species and a revision of the known species (Arachnida, Opiliones, Nemastomatidae). Revue suisse de Zoologie, 126 (1), 79 - 110. https: // doi. org / 10.5281 / zenodo. 2619524" type="journal article" year="2019">Martens 2019</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |