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<document id="37C051C47316C5302A054BEA15C6F5FF" ID-CLB-Dataset="46304" ID-DOI="10.5281/zenodo.205904" ID-GBIF-Dataset="62376e09-f30b-4fb0-a635-8f8f43e008fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="205904" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391967158" checkinUser="plazi" docAuthor="Sasakawa, Mitsuhiro" docDate="2011" docId="03A787A1FFD3FFBAFF5EFF22EB231F0A" docLanguage="en" docName="zt03038p028.pdf" docOrigin="Zootaxa 3038" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Heteromeringia spinulifera Sasakawa, 2011, sp. nov" docType="treatment" docVersion="14" lastPageNumber="24" masterDocId="FF9EFFD9FFC4FFADFFC9FFB5EF5D1878" masterDocTitle="Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="24" updateTime="1698298315006" updateUser="ExternalLinkService">
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<mods:title id="D4E060F26018DC731D08424348322987">Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)</mods:title>
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<mods:namePart id="EBACF9D99315A0935A1FC97B06B28C4B">Sasakawa, Mitsuhiro</mods:namePart>
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<mods:date id="D203FF3BB644724BA4CC4C9443CE898C">2011</mods:date>
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<treatment id="03A787A1FFD3FFBAFF5EFF22EB231F0A" ID-DOI="http://doi.org/10.5281/zenodo.3506871" ID-GBIF-Taxon="119416976" ID-Zenodo-Dep="3506871" LSID="urn:lsid:plazi:treatment:03A787A1FFD3FFBAFF5EFF22EB231F0A" httpUri="http://treatment.plazi.org/id/03A787A1FFD3FFBAFF5EFF22EB231F0A" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="C314653CFFD3FFBAFF5EFF22EE7A18AC" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="8BB136B7FFD3FFBAFF5EFF22ED7318C8" blockId="23.[151,558,151,212]" box="[151,558,151,176]" pageId="23" pageNumber="24">
<heading id="D0F981DBFFD3FFBAFF5EFF22ED7318C8" bold="true" box="[151,558,151,176]" fontSize="10" level="2" pageId="23" pageNumber="24" reason="2">
<emphasis id="B97AEAA5FFD3FFBAFF5EFF22ED7318C8" bold="true" box="[151,558,151,176]" pageId="23" pageNumber="24">
<taxonomicName id="4C0E4D34FFD3FFBAFF5EFF22EE8E18C8" ID-CoL="8CBWV" box="[151,467,151,176]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="spinulifera" status="sp. nov.">
<emphasis id="B97AEAA5FFD3FFBAFF5EFF22EE8E18C8" bold="true" box="[151,467,151,176]" italics="true" pageId="23" pageNumber="24">Heteromeringia spinulifera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DEFFD3FFBAFE13FF2DED7318C8" box="[474,558,152,176]" pageId="23" pageNumber="24" rank="species">sp. nov</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF5EFF09EE7A18AC" blockId="23.[151,558,151,212]" box="[151,295,188,212]" pageId="23" pageNumber="24">
(
<figureCitation id="13352A32FFD3FFBAFF56FF09EE4318AC" box="[159,286,188,212]" captionStart="FIGURES 36 41" captionStartId="22.[151,264,1924,1947]" captionTargetBox="[179,1422,598,1853]" captionTargetId="figure@22.[151,1436,589,1903]" captionTargetPageId="22" captionText="FIGURES 36 41. Male genitalia of Heteromeringia pectinata sp. nov. (36 38, paratype) and H. spinulifera sp. nov. (39 41, paratype). 37, pre- and postgonite. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205910/files/figure.png" pageId="23" pageNumber="24">Figs 3941</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF5EFEB1EDEE19FF" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="8BB136B7FFD3FFBAFF5EFEB1EDEE19FF" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF5EFEB1EE181964" bold="true" box="[151,325,259,284]" pageId="23" pageNumber="24">
<typeStatus id="54B58815FFD3FFBAFF5EFEB1EF8D1964" box="[151,208,260,284]" pageId="23" pageNumber="24">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B58815FFD3FFBAFE85FEB1EEE91964" box="[332,436,260,284]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED1FAE72FFD3FFBAFE36FEB1ED121964" LSID="urn:lsid:biocol.org:col:34697" box="[511,591,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="23" pageNumber="24">BPBM</collectionCode>
17252),
<collectingCountry id="F3197627FFD3FFBAFD7AFEB1EC691964" box="[691,820,260,284]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
, Luzon, Nueva Vizcaya, Dalton Pass,
<quantity id="4CF69B52FFD3FFBAFB29FEB1EA741964" box="[1248,1321,260,284]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="23" pageNumber="24" unit="m" value="915.0">915 m</quantity>
,
<date id="FFB01077FFD3FFBAFAFCFEB1EF8F1947" pageId="23" pageNumber="24" value="1968-04-09" valueMax="1968-04-10" valueMin="1968-04-09">910. IV. 1968</date>
, D. E. Hardy.
<typeStatus id="54B58815FFD3FFBAFEB8FE9DEEBF1938" box="[369,482,296,320]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
: 23,2Ƥ, same data as
<typeStatus id="54B58815FFD3FFBAFD14FE92EC1F1947" box="[733,834,295,319]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
, but coll. M. Delfinado, and D. E. Hardy and M. Delfinado; 3, Zamboanga del Norte, Manucan,
<quantity id="4CF69B52FFD3FFBAFD43FEF9ED8E191C" box="[650,723,332,356]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="23" pageNumber="24" unit="m" value="420.0">420 m</quantity>
,
<date id="FFB01077FFD3FFBAFD29FEF9EC35191C" box="[736,872,332,356]" pageId="23" pageNumber="24" value="1959-10-12">12. X. 1959</date>
, L. W. Quate; 3, Camarines Sur, Mt. Isarog,
<quantity id="4CF69B52FFD3FFBAFAADFEF9EFBD19FF" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="23" pageNumber="24" unit="m" value="550.0" valueMax="600.0" valueMin="500.0">500 600 m</quantity>
,
<date id="FFB01077FFD3FFBAFF22FEDAEE3419FF" box="[235,361,367,391]" pageId="23" pageNumber="24" value="1963-04-09">9. IV. 1963</date>
, H. M. Torrevillas, light trap.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF0EFE26EC60198C" pageId="23" pageNumber="24" type="diagnosis">
<paragraph id="8BB136B7FFD3FFBAFF0EFE26EC60198C" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EFE26EE1C19D4" bold="true" box="[199,321,403,428]" pageId="23" pageNumber="24">Diagnosis.</emphasis>
Largely black; face and gena yellow in male and blackish in female; palpus brown to black; postpronotal lobe and katepisternum yellow; arista distinctly pubescent; dc two; surstylus narrow; cerci united; mesophallus with dense microscopic spinules on membrane.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF0EFE4AECBF1D9F" pageId="23" pageNumber="24" type="description">
<paragraph id="8BB136B7FFD3FFBAFF0EFE4AEA511AB4" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EFE4AEE041A60" bold="true" box="[199,345,511,536]" pageId="23" pageNumber="24">Description.</emphasis>
MALE. Head black but vertical angle and dorsal orbit brown, face, gena, and ventral half of postgena yellow; frons shining; gena whitish tomentose. Antenna yellow, arista brown; palpus black but yellow on apical 1/3. Thorax black but postpronotal lobe and katepisternum yellow; mesoscutum weakly shining, pleura strongly shining. Wing hyaline, faintly tinged with yellowish brown anteroapically; calypter with margin brown, fringe pale brown; halter yellow. Legs yellow; mid coxa brownish at base, fore femur with brown ring on apical 1/5, fore tibia and tarsomeres 14 brown. Abdomen shining black; cercus yellowish brown. Bristles brown to black.
</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFD6CEBD11B07" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">Frons almost as wide as eye, slightly diverging ventrally; first and second or equal in length, third or about 2/3 as long as second or, distance between first and second or 33.5 times as long as that between second and third or; oh minute; frontalia with sparse hairs ventrolaterally; oc minute; poc slightly shorter than third or; gena 1/61/7 as high as eye: pm three or four. First antennal flagellomere round, slightly broader than long, distinctly pilose; arista sparsely plumose (longest hair about 1/2 as long as width of flagellomere) except for base.</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFC38EADB1C77" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">Mesoscutum with two dc, anterior dc far before level of sa, distance between two dc 2.5 times as long as that between transverse suture and anterior dc, eight rows of acr before transverse suture but six rows behind that; scutellum with bsc 1/4 as long as asc. Wing 2.53.1 (2.9) mm long; three costal sections in proportion of 3840: 10: 4.55.0, r-m at distal 1/3 of cell dm, M1 ratio 5.05.8, ultimate section of CuA1 about 4/5 length of penultimate.</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFBAAEC371C92" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
Epandrium (
<figureCitation id="13352A32FFD3FFBAFE9AFBAAEE8B1C4F" box="[339,470,1055,1079]" captionStart="FIGURES 36 41" captionStartId="22.[151,264,1924,1947]" captionTargetBox="[179,1422,598,1853]" captionTargetId="figure@22.[151,1436,589,1903]" captionTargetPageId="22" captionText="FIGURES 36 41. Male genitalia of Heteromeringia pectinata sp. nov. (36 38, paratype) and H. spinulifera sp. nov. (39 41, paratype). 37, pre- and postgonite. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205910/files/figure.png" pageId="23" pageNumber="24">Figs 39, 40</figureCitation>
) with surstylus and cercus long, surstylus bearing three inner apical spinules and one or two pale, minute spinules, cerci united with each other; proctiger small, strongly chitinized. Hypandrium with two setae; pregonite knob-like, with several short setae; postgonite conical, well chitinized, with three strong apical setae. Phallapodeme (
<figureCitation id="13352A32FFD3FFBAFE58FB3FEEB81CDA" box="[401,485,1162,1186]" captionStart="FIGURES 36 41" captionStartId="22.[151,264,1924,1947]" captionTargetBox="[179,1422,598,1853]" captionTargetId="figure@22.[151,1436,589,1903]" captionTargetPageId="22" captionText="FIGURES 36 41. Male genitalia of Heteromeringia pectinata sp. nov. (36 38, paratype) and H. spinulifera sp. nov. (39 41, paratype). 37, pre- and postgonite. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205910/files/figure.png" pageId="23" pageNumber="24">Fig. 41</figureCitation>
) 160 µm long; mesophallus 840 µm long and microscopically spinulose on membrane, paraphallus as long as hypophallus and right sclerite distinctly serrated at base, distiphallus curved at middle; ejaculatory apodeme 330350 µm long, 120140 µm broad.
</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFB42ED7F1D77" blockId="23.[151,1437,259,1907]" box="[199,546,1271,1295]" pageId="23" pageNumber="24">Body length 3.13.7 (3.5) mm.</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFAAEED1B1DE7" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
FEMALE. Similar to male, but face and gena dark brown to black; antenna brownish yellow to pale brown, with scape and pedicel distinctly darker; palpus entirely pale brown; fore tarsomeres 1 and 2 brown, 13 rather broad, almost as wide as distal width of tibia; wing with apical infuscation darker than that of male; wing length
<quantity id="4CF69B52FFD3FFBAFF5EFA32EE7C1DE7" box="[151,289,1415,1439]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="23" pageNumber="24" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.23.6 mm</quantity>
, body length
<quantity id="4CF69B52FFD3FFBAFE7EFA32ED1C1DE7" box="[439,577,1415,1439]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="23" pageNumber="24" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.84.2 mm</quantity>
.
</paragraph>
<paragraph id="8BB136B7FFD3FFBAFF0EFA19ECBF1D9F" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EFA19EE721DBB" box="[199,303,1452,1475]" italics="true" pageId="23" pageNumber="24">Variation</emphasis>
. A male, collected on Mt. Isarog, has brownish basal and apical rings on the hind tibia (each about 1/ 4 as long as whole tibial length), but the genitalia are typical of the species.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF0EFA47EEB01E72" box="[199,493,1522,1547]" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8BB136B7FFD3FFBAFF0EFA47EEB01E72" blockId="23.[151,1437,259,1907]" box="[199,493,1522,1547]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EFA47EE3D1E73" bold="true" box="[199,352,1522,1547]" pageId="23" pageNumber="24">Distribution.</emphasis>
<collectingCountry id="F3197627FFD3FFBAFEAEFA47EEB41E72" box="[359,489,1522,1546]" name="Philippines" pageId="23" pageNumber="24">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF0EF9A2EF8B1F37" pageId="23" pageNumber="24" type="discussion">
<paragraph id="8BB136B7FFD3FFBAFF0EF9A2EF8B1F37" blockId="23.[151,1437,259,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EF9A2EE661E57" bold="true" box="[199,315,1559,1583]" pageId="23" pageNumber="24">Remarks.</emphasis>
This new species is similar to New Guinean
<taxonomicName id="4C0E4D34FFD3FFBAFC86F9ADEA141E57" authority="Sasakawa, 1966" authorityName="Sasakawa" authorityYear="1966" box="[847,1353,1559,1583]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="stenygralis">
<emphasis id="B97AEAA5FFD3FFBAFC86F9ADEBDB1E57" box="[847,1158,1559,1583]" italics="true" pageId="23" pageNumber="24">Heteromeringia stenygralis</emphasis>
<bibRefCitation id="EF9F4B46FFD3FFBAFB46F9A2EA141E57" author="Sasakawa" box="[1167,1353,1559,1583]" pageId="23" pageNumber="24" refString="Sasakawa, M. (1966) Studies on the Oriental and Pacific Clusiidae (Diptera). Part 1. Genus Heteromeringia Czerny, with one new related genus. Pacific Insects, 8, 61 - 100." type="journal article" year="1966">Sasakawa, 1966</bibRefCitation>
</taxonomicName>
, which also has a yellow postpronotal lobe and katepisternum, and
<taxonomicName id="4C0E4D34FFD3FFBAFC89F989EBC81E2A" authority="Sasakawa, 1966" authorityName="Sasakawa" authorityYear="1966" box="[832,1173,1594,1619]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B97AEAA5FFD3FFBAFC89F989EC891E2B" box="[832,980,1596,1619]" italics="true" pageId="23" pageNumber="24">H. papuensis</emphasis>
<bibRefCitation id="EF9F4B46FFD3FFBAFC15F98FEBC81E2A" author="Sasakawa" box="[988,1173,1594,1618]" pageId="23" pageNumber="24" refString="Sasakawa, M. (1966) Studies on the Oriental and Pacific Clusiidae (Diptera). Part 1. Genus Heteromeringia Czerny, with one new related genus. Pacific Insects, 8, 61 - 100." type="journal article" year="1966">Sasakawa, 1966</bibRefCitation>
</taxonomicName>
, which also has microscopic spinules on the membrane of the mesophallus. The new species differs from
<taxonomicName id="4C0E4D34FFD3FFBAFB89F9D5EB801E0F" box="[1088,1245,1631,1655]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="stenygralis">
<emphasis id="B97AEAA5FFD3FFBAFB89F9D5EB091E0F" box="[1088,1108,1632,1655]" italics="true" pageId="23" pageNumber="24">H</emphasis>
.
<emphasis id="B97AEAA5FFD3FFBAFBABF9EAEB801E0F" box="[1122,1245,1631,1655]" italics="true" pageId="23" pageNumber="24">stenygralis</emphasis>
</taxonomicName>
in the coloration of the ventral pleura and hind tibia, and the structures of the male genitalia; in
<taxonomicName id="4C0E4D34FFD3FFBAFBC4F931EBF31EE3" box="[1037,1198,1667,1691]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="stenygralis">
<emphasis id="B97AEAA5FFD3FFBAFBC4F931EB7C1EE3" box="[1037,1057,1668,1691]" italics="true" pageId="23" pageNumber="24">H</emphasis>
.
<emphasis id="B97AEAA5FFD3FFBAFBF9F936EBF31EE3" box="[1072,1198,1667,1691]" italics="true" pageId="23" pageNumber="24">stenygralis</emphasis>
</taxonomicName>
, the katepisternum is only yellow on the posterior half, the meron is yellow, the hind tibia is brown excluding the extremities, the surstylus is lobate, and the left hypophallus is crescent-shaped (
<bibRefCitation id="EF9F4B46FFD3FFBAFCDDF97FEB121E9A" author="Sasakawa" box="[788,1103,1738,1762]" pageId="23" pageNumber="24" refString="Sasakawa, M. (1966) Studies on the Oriental and Pacific Clusiidae (Diptera). Part 1. Genus Heteromeringia Czerny, with one new related genus. Pacific Insects, 8, 61 - 100." type="journal article" year="1966">Sasakawa 1966: fig. 11A, B</bibRefCitation>
). Also, it differs from
<taxonomicName id="4C0E4D34FFD3FFBAFA8EF979EF801F7F" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B97AEAA5FFD3FFBAFA8EF979EF801F7F" italics="true" pageId="23" pageNumber="24">H. papuensis</emphasis>
</taxonomicName>
in the coloration of the thorax and the structure of the phallus; in
<taxonomicName id="4C0E4D34FFD3FFBAFC1EF945EB361F7F" box="[983,1131,1776,1799]" class="Insecta" family="Clusiidae" genus="Heteromeringia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="papuensis">
<emphasis id="B97AEAA5FFD3FFBAFC1EF945EB361F7F" box="[983,1131,1776,1799]" italics="true" pageId="23" pageNumber="24">H. papuensis</emphasis>
</taxonomicName>
, the postpronotal lobe and katepisternum are brown, the paraphallus is longer, and the distiphallus is almost straight (
<bibRefCitation id="EF9F4B46FFD3FFBAFB7BF8A7EF941F37" author="Sasakawa" pageId="23" pageNumber="24" refString="Sasakawa, M. (1966) Studies on the Oriental and Pacific Clusiidae (Diptera). Part 1. Genus Heteromeringia Czerny, with one new related genus. Pacific Insects, 8, 61 - 100." type="journal article" year="1966">Sasakawa 1966: fig. 10B</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFD3FFBAFF0EF8EEEB231F0A" box="[199,1150,1882,1907]" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BB136B7FFD3FFBAFF0EF8EEEB231F0A" blockId="23.[151,1437,259,1907]" box="[199,1150,1882,1907]" pageId="23" pageNumber="24">
<emphasis id="B97AEAA5FFD3FFBAFF0EF8EEEE1B1F0B" bold="true" box="[199,326,1883,1907]" pageId="23" pageNumber="24">Etymology</emphasis>
. The specific name refers to the spinulose membrane of the mesophallus.
</paragraph>
</subSubSection>
</treatment>
</document>