1026 lines
119 KiB
XML
1026 lines
119 KiB
XML
<document id="768A4CAADA027906FAE3A36EBAA20F04" ID-CLB-Dataset="46711" ID-DOI="10.11646/zootaxa.3753.5.3" ID-GBIF-Dataset="f39438f9-240f-4442-be2d-759640351dc2" ID-ISSN="1175-5326" ID-Zenodo-Dep="228774" ID-ZooBank="A039A90E-7A45-4E59-88B3-160E0C08DB76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460366207527" checkinUser="plazi" docAuthor="Matsui, Masafumi, Nishikawa, Kanto & Yambun, Paul" docDate="2014" docId="039AF439FF84A029FF24F963FF459CC3" docLanguage="en" docName="zt03753p452.pdf" docOrigin="Zootaxa 3753 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptolalax sabahmontanus Matsui, Nishikawa & Yambun, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="449" masterDocId="FFA38C41FF86A020FFB3FFCEFFB09926" masterDocTitle="A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)" masterLastPageNumber="452" masterPageNumber="440" pageNumber="442" updateTime="1698295234262" updateUser="ExternalLinkService">
|
||
<mods:mods id="2FB3703DCA3EF1CCD5AE009E9DC562C8" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1BEF6E7764CF852E3C2455C08C6829C5">
|
||
<mods:title id="14F932C6923D2E2E6CA51F7836E92FD2">A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="0D1ADEB4852BE15ED2A5A1FA00F0F12F" type="personal">
|
||
<mods:role id="B663493DAF36C9E7D4A95941E69A9A6A">
|
||
<mods:roleTerm id="FDE0E3FA0AB9B260EB95BB8481D9F1D9">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="34BAA3F597F688A660C1B6D04B2BA9FE">Matsui, Masafumi</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6254B43C8F44EE048593BF04C7D78246" type="personal">
|
||
<mods:role id="C3BAA87D9524CF5F1491F595974E09F0">
|
||
<mods:roleTerm id="EE099E42307B1337B12FF0D63870CF1C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="629E3B9EDE4B85B028C5ECAEEEC5423C">Nishikawa, Kanto</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A528C26161954CF82C79BAFA66EB9C79" type="personal">
|
||
<mods:role id="0D017F1A59605AADB9079DFB98CEA5BF">
|
||
<mods:roleTerm id="9C58B383B9F0813346A95395ED704D74">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2111BC2B81BB0E082897FC3EDD40A72A">Yambun, Paul</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C4779963E5A20C8338D834E7136102B8">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F81133CD3B7E9EA50A65AF35A1558591" type="host">
|
||
<mods:titleInfo id="5F33AD1B1CFF2BD060CCCFD4F5E2E14B">
|
||
<mods:title id="EE39CDC998787F9199D35EC4193681A2">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="F96828F1C81154DB0ADC21E42799D275">
|
||
<mods:date id="56E5DF81D376345710F84E1823300C7E">2014</mods:date>
|
||
<mods:detail id="085904643308FEA5B7E717CFFCA04DCC" type="volume">
|
||
<mods:number id="1A118381070EF5BE69854FD2219DA139">3753</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="86C1158378969AC20C59E58AF7A9FE66" type="issue">
|
||
<mods:number id="F83A3ADD64D53FEF55395C3B688C39F9">5</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="2EB37084074CC089D54FBCDA0D5D5428" unit="page">
|
||
<mods:start id="B17FD142F2116AF17AE65275940F91C0">440</mods:start>
|
||
<mods:end id="2D491EA366DDCB2CD0C48B66B2E55245">452</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="8D28BCDE0EDEAE9B450CADB15C43A3D3">journal article</mods:classification>
|
||
<mods:identifier id="6D358540225D6BB325D8A954ECB02BD6" type="CLB-Dataset">46711</mods:identifier>
|
||
<mods:identifier id="FE7172F94CC132DCCF0F376CBC1219CA" type="DOI">10.11646/zootaxa.3753.5.3</mods:identifier>
|
||
<mods:identifier id="D5181645FC74EFDCBA2CA166D2697F98" type="GBIF-Dataset">f39438f9-240f-4442-be2d-759640351dc2</mods:identifier>
|
||
<mods:identifier id="63B96A6359B90D6CBDF4FFB17F9D993E" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="16B622CED2813EDCD14D7B9B3D1F43D5" type="Zenodo-Dep">228774</mods:identifier>
|
||
<mods:identifier id="1CF6C49D04B84DE411BB24578F672227" type="ZooBank">A039A90E-7A45-4E59-88B3-160E0C08DB76</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="039AF439FF84A029FF24F963FF459CC3" ID-DOI="http://doi.org/10.5281/zenodo.5613489" ID-GBIF-Taxon="119410857" ID-Zenodo-Dep="5613489" LSID="urn:lsid:plazi:treatment:039AF439FF84A029FF24F963FF459CC3" httpUri="http://treatment.plazi.org/id/039AF439FF84A029FF24F963FF459CC3" lastPageId="9" lastPageNumber="449" pageId="2" pageNumber="442">
|
||
<subSubSection id="C32916A4FF84A022FF24F963FEFA9FCD" pageId="2" pageNumber="442" type="nomenclature">
|
||
<paragraph id="8B8C452FFF84A022FF24F963FD809FE1" blockId="2.[151,560,1709,1771]" box="[151,560,1709,1735]" pageId="2" pageNumber="442">
|
||
<heading id="D0C4F243FF84A022FF24F963FD809FE1" bold="true" box="[151,560,1709,1735]" fontSize="10" level="2" pageId="2" pageNumber="442" reason="2">
|
||
<emphasis id="B947993DFF84A022FF24F963FD809FE1" bold="true" box="[151,560,1709,1735]" pageId="2" pageNumber="442">
|
||
<taxonomicName id="4C333EACFF84A022FF24F963FE609FE0" ID-CoL="6PGBK" box="[151,464,1709,1734]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="2" pageNumber="442" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF84A022FF24F963FE609FE0" bold="true" box="[151,464,1709,1734]" italics="true" pageId="2" pageNumber="442">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF84A022FE65F960FD809FE1" box="[470,560,1710,1735]" pageId="2" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF84A022FF24F91CFEFA9FCD" blockId="2.[151,560,1709,1771]" box="[151,330,1746,1771]" pageId="2" pageNumber="442">
|
||
(
|
||
<figureCitation id="130859AAFF84A022FF2DF91CFEBB9FCD" box="[158,267,1746,1771]" captionStart="FIGURE 2" captionStartId="3.[151,250,1559,1581]" captionTargetBox="[340,1246,925,1536]" captionTargetId="figure@3.[340,1247,896,1537]" captionTargetPageId="3" captionText="FIGURE 2. Dorsolateral view of a male paratype of Leptolalax sabahmontanus sp. nov. (KUHE 48014) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park." httpUri="https://zenodo.org/record/228776/files/figure.png" pageId="2" pageNumber="442">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="130859AAFF84A022FEA4F91DFE979FCD" box="[279,295,1747,1771]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1218]" captionTargetBox="[249,1338,647,1176]" captionTargetId="figure@4.[241,1345,642,1180]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and ventral (B) views of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 10 mm." httpUri="https://zenodo.org/record/228777/files/figure.png" pageId="2" pageNumber="442">3</figureCitation>
|
||
,
|
||
<figureCitation id="130859AAFF84A022FE80F91DFEF29FCD" box="[307,322,1747,1771]" captionStart="FIGURE 4" captionStartId="6.[151,250,1135,1157]" captionTargetBox="[361,1225,195,1113]" captionTargetId="figure@6.[361,1226,195,1114]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of right hand (A) and foot (B) of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 5 mm." httpUri="https://zenodo.org/record/228778/files/figure.png" pageId="2" pageNumber="442">4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32916A4FF84A023FF24F8D3FE049A5E" lastPageId="3" lastPageNumber="443" pageId="2" pageNumber="442" type="materials_examined">
|
||
<paragraph id="8B8C452FFF84A022FF24F8D3FEA49E58" blockId="2.[151,1436,1817,2026]" pageId="2" pageNumber="442">
|
||
<emphasis id="B947993DFF84A022FF24F8D3FEBB9E10" bold="true" box="[151,267,1821,1846]" pageId="2" pageNumber="442">
|
||
<typeStatus id="5488FB8DFF84A022FF24F8D3FEB69E10" box="[151,262,1821,1846]" pageId="2" pageNumber="442" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
BORNEENSIS 22684, an adult male from Silau Silau trail along Sungai Silau-Silau (
|
||
<geoCoordinate id="EE0723E8FF84A022FB79F8D3FA969E13" box="[1226,1318,1817,1846]" direction="north" orientation="latitude" pageId="2" pageNumber="442" precision="925" value="6.0">06o00'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF84A022FA81F8D2FA279E13" box="[1330,1431,1817,1845]" direction="east" orientation="longitude" pageId="2" pageNumber="442" precision="925" value="116.53333">116o32'E</geoCoordinate>
|
||
, alt.
|
||
<quantity id="4CCBE8CAFF84A022FF71F88FFEA89E7F" box="[194,280,1857,1882]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="2" pageNumber="442" unit="m" value="1450.0">1450 m</quantity>
|
||
a.s.l.), Kinabalu National Park (KNP), Ranau District, Sabah,
|
||
<collectingCountry id="F32405BFFF84A022FC6DF88FFBF79E7C" box="[990,1095,1857,1882]" name="Malaysia" pageId="2" pageNumber="442">Malaysia</collectingCountry>
|
||
, collected on
|
||
<date id="FF8D63EFFF84A022FB56F88FFAC89E7C" box="[1253,1400,1857,1882]" pageId="2" pageNumber="442" value="2005-07-31">31 July 2005</date>
|
||
by M. Matsui.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF84A022FF74F847FA2C9ECF" blockId="2.[151,1436,1817,2026]" pageId="2" pageNumber="442">
|
||
<emphasis id="B947993DFF84A022FF74F847FEF69E84" bold="true" box="[199,326,1929,1954]" pageId="2" pageNumber="442">
|
||
<typeStatus id="5488FB8DFF84A022FF74F847FEF19E84" box="[199,321,1929,1954]" pageId="2" pageNumber="442" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
BORNEENSIS 22683, 22685, 22687–22691, 22701,
|
||
<collectionCode id="ED22DDEAFF84A022FB95F844FBC49E87" box="[1062,1140,1930,1953]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="2" pageNumber="442">KUHE</collectionCode>
|
||
48014–48016 (former BORNEENSIS 22681, 22682, 22686),
|
||
<specimenCount id="9D358EA6FF84A022FDE2F863FD049EE0" box="[593,692,1965,1990]" pageId="2" pageNumber="442" type="male">11 males</specimenCount>
|
||
data, same as the
|
||
<typeStatus id="5488FB8DFF84A022FC32F863FC579EE0" box="[897,999,1965,1990]" pageId="2" pageNumber="442" type="holotype">holotype</typeStatus>
|
||
. BORNEENSIS 22757, 22760,
|
||
<collectionCode id="ED22DDEAFF84A022FAFDF860FA2C9EE3" box="[1358,1436,1966,1989]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="2" pageNumber="442">KUHE</collectionCode>
|
||
48017 (former BORNEENSIS 22756), three males from the
|
||
<typeStatus id="5488FB8DFF84A022FCD0F81CFC239ECC" box="[867,915,2002,2026]" pageId="2" pageNumber="442">type</typeStatus>
|
||
locality, collected on
|
||
<date id="FF8D63EFFF84A022FB28F81FFAF99ECC" box="[1179,1353,2001,2026]" pageId="2" pageNumber="442" value="2005-08-02">2 August 2005</date>
|
||
by M.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF85A023FF24FF59FE6398F8" blockId="3.[151,1436,151,888]" pageId="3" pageNumber="443">
|
||
Matsui, P. Yambun, K. Nishikawa, and T. Shimada. SP 26736, 26737, two males from the
|
||
<typeStatus id="5488FB8DFF85A023FB2EFF56FB7D9996" box="[1181,1229,152,176]" pageId="3" pageNumber="443">type</typeStatus>
|
||
locality, collected on
|
||
<date id="FF8D63EFFF85A023FF0CFF71FECD99F1" box="[191,381,191,215]" pageId="3" pageNumber="443" value="2008-08-14">14 August 2008</date>
|
||
by Majin Silim. BORNEENSIS 22748, 22749, two males from Pandanus trail (
|
||
<geoCoordinate id="EE0723E8FF85A023FA88FF70FA2799F1" box="[1339,1431,187,215]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="6.0">06o00'N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FF24FF2BFF4C99D8" box="[151,252,226,254]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.53333">116o32'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FEBFFF2BFED699DB" box="[268,358,229,254]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="443" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l.), KNP, collected on
|
||
<date id="FF8D63EFFF85A023FD11FF28FCE199D8" box="[674,849,229,254]" pageId="3" pageNumber="443" value="2005-08-01">1 August 2005</date>
|
||
by M. Matsui, P. Yambun, K. Nishikawa, and T. Shimada.
|
||
<collectionCode id="ED22DDEAFF85A023FEBBFEC4FEE69807" box="[264,342,266,289]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="3" pageNumber="443">KUHE</collectionCode>
|
||
48018 (former BORNEENSIS 22784), BORNEENSIS 22785, 22786, three males from junction of Sg. Liwagu and Sg. Silau-Silau (
|
||
<geoCoordinate id="EE0723E8FF85A023FD9EFEFEFD27986E" box="[557,663,301,329]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="6.0">06o00’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FD17FEFEFCA6986E" box="[676,790,301,329]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.55">116o33’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FC97FEFFFCCA986E" box="[804,890,305,329]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="443" unit="m" value="1400.0">1400 m</quantity>
|
||
a.s.l.), KNP, collected on
|
||
<date id="FF8D63EFFF85A023FB14FEFEFAFF986F" box="[1191,1359,304,329]" pageId="3" pageNumber="443" value="2005-08-05">5 August 2005</date>
|
||
by M. Matsui, P. Yambun, K. Nishikawa, and T. Shimada. SP 26740, an adult female from Bundu Tuhan view trail, KNP, collected on
|
||
<date id="FF8D63EFFF85A023FE96FEB5FE6698B5" box="[293,470,379,403]" pageId="3" pageNumber="443" value="2008-08-14">14 August 2008</date>
|
||
by S. David. BORNEENSIS 22980, 23029, 23034, three males from Sayap (
|
||
<geoCoordinate id="EE0723E8FF85A023FA82FEB5FA2898B5" box="[1329,1432,375,404]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="6.1833334">06o11’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FF24FE6CFEBA989C" box="[151,266,414,443]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.566666">116o34’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FEA9FE6CFEEA989D" box="[282,346,418,443]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="3" pageNumber="443" unit="m" value="950.0">950m</quantity>
|
||
a.s.l.), KNP, Kota Belud District, collected on
|
||
<date id="FF8D63EFFF85A023FC30FE6CFB8D989D" box="[899,1085,418,443]" pageId="3" pageNumber="443" value="2005-08-12">12 August 2005</date>
|
||
by M. Matsui, Y. B. Paul, K. Nishikawa, and T. Shimada.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF85A023FF74FE22FE049A5E" blockId="3.[151,1436,151,888]" pageId="3" pageNumber="443">
|
||
<emphasis id="B947993DFF85A023FF74FE22FE099B23" bold="true" box="[199,441,492,517]" pageId="3" pageNumber="443">Referred specimens.</emphasis>
|
||
BORNEENSIS 8711, 8726, 8729, three males from Trail 4 (
|
||
<geoCoordinate id="EE0723E8FF85A023FBD1FE22FB7A9B22" box="[1122,1226,489,517]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="5.483333">05o29’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FB66FE22FAF99B22" box="[1237,1353,489,517]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.01667">116 o01’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FAE6FE22FA2C9B22" box="[1365,1436,492,517]" metricMagnitude="2" metricUnit="m" metricValue="7.54" pageId="3" pageNumber="443" unit="m" value="754.0">754 m</quantity>
|
||
a.s.l.) of
|
||
<collectionCode id="ED22DDEAFF85A023FEB6FDDFFEF19B0F" box="[261,321,529,553]" httpUri="http://grbio.org/cool/8va9-jhre" name="Universiti Malaysia Sabah" pageId="3" pageNumber="443">UMS</collectionCode>
|
||
2002 expedition to the Crocker Range, Ulu Kimanis, Crocker Range National Park (
|
||
<collectionCode id="ED22DDEAFF85A023FAF2FDDFFA209B0F" box="[1345,1424,529,553]" pageId="3" pageNumber="443">CRNP</collectionCode>
|
||
), Tambunan District, collected on
|
||
<date id="FF8D63EFFF85A023FDA2FDFBFD539B6B" box="[529,739,564,589]" pageId="3" pageNumber="443" value="2002-09-09">9 September 2002</date>
|
||
by M. Matsui. BORNEENSIS 8179, one male from Trail 5 (
|
||
<geoCoordinate id="EE0723E8FF85A023FF2CFD95FEBD9B55" box="[159,269,600,628]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="5.483333">05o29’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FEADFD95FE259B55" box="[286,405,600,628]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.01667">116o01’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FE15FD92FE439B55" box="[422,499,604,628]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="443" unit="m" value="820.0">820 m</quantity>
|
||
a.s.l.) of Ulu Kimanis,
|
||
<collectionCode id="ED22DDEAFF85A023FCABFD92FCD19B52" box="[792,865,604,628]" pageId="3" pageNumber="443">CRNP</collectionCode>
|
||
, collected on
|
||
<date id="FF8D63EFFF85A023FBBEFD92FB549B52" box="[1037,1252,603,628]" pageId="3" pageNumber="443" value="2002-09-01">1 September 2002</date>
|
||
by M. Matsui. BORNEENSIS 8673, one male collected on
|
||
<date id="FF8D63EFFF85A023FD0AFD4EFC249BBE" box="[697,916,639,664]" pageId="3" pageNumber="443" value="2002-09-07">7 September 2002</date>
|
||
by M. Matsui, and
|
||
<collectionCode id="ED22DDEAFF85A023FB25FD4EFB549BB1" box="[1174,1252,640,663]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="3" pageNumber="443">KUHE</collectionCode>
|
||
48020 (former BORNEENSIS 23470), one male collected on
|
||
<date id="FF8D63EFFF85A023FD1BFD69FCEC9B99" box="[680,860,678,703]" pageId="3" pageNumber="443" value="2005-08-19">19 August 2005</date>
|
||
by M. Matsui, from Trail 11 (
|
||
<geoCoordinate id="EE0723E8FF85A023FB02FD68FAAA9B98" box="[1201,1306,675,703]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="5.45">05o27’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FA95FD68FA279B98" box="[1318,1431,675,703]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.05">116o03’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FF24FD05FF409BC4" box="[151,240,715,739]" metricMagnitude="3" metricUnit="m" metricValue="1.427" pageId="3" pageNumber="443" unit="m" value="1427.0">1427 m</quantity>
|
||
a.s.l.) of Ulu Kimanis,
|
||
<collectionCode id="ED22DDEAFF85A023FDB9FD05FDE49BC5" box="[522,596,715,739]" pageId="3" pageNumber="443">CRNP</collectionCode>
|
||
.
|
||
<collectionCode id="ED22DDEAFF85A023FDD2FD05FD1F9BC4" box="[609,687,715,738]" httpUri="http://grbio.org/cool/jxgt-7i97" name="Kyoto University, Graduate School of Human and Environmental Studies" pageId="3" pageNumber="443">KUHE</collectionCode>
|
||
48019 (former BORNEENSIS 8855), one male collected on
|
||
<date id="FF8D63EFFF85A023FA33FD05FEE09A21" pageId="3" pageNumber="443" value="2003-12-12">12 December 2003</date>
|
||
, BORNEENSIS 12454, one male collected on
|
||
<date id="FF8D63EFFF85A023FC32FD20FB9F9A21" box="[897,1071,750,775]" pageId="3" pageNumber="443" value="2003-08-05">5 August 2003</date>
|
||
by M. Matsui, BORNEENSIS 12632–12634, three males collected on
|
||
<date id="FF8D63EFFF85A023FDC5FCDDFC989A0D" box="[630,808,787,811]" pageId="3" pageNumber="443" value="2003-08-09">9 August 2003</date>
|
||
by M. Matsui, K. Nishikawa, and T. Shimada, and BORNEENSIS 23482, one male collected on
|
||
<date id="FF8D63EFFF85A023FD15FCF7FCED9A74" box="[678,861,825,850]" pageId="3" pageNumber="443" value="2005-08-20">20 August 2005</date>
|
||
by M. Matsui, all from Mahua,
|
||
<collectionCode id="ED22DDEAFF85A023FB60FCF7FAAC9A77" box="[1235,1308,825,849]" pageId="3" pageNumber="443">CRNP</collectionCode>
|
||
(
|
||
<geoCoordinate id="EE0723E8FF85A023FA9DFCF7FA289A77" box="[1326,1432,822,850]" direction="north" orientation="latitude" pageId="3" pageNumber="443" precision="925" value="5.8">05o48’ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE0723E8FF85A023FF24FC91FEB79A51" box="[151,263,860,888]" direction="east" orientation="longitude" pageId="3" pageNumber="443" precision="925" value="116.4">116o24’ E</geoCoordinate>
|
||
,
|
||
<quantity id="4CCBE8CAFF85A023FEA0FC91FED89A51" box="[275,360,863,888]" metricMagnitude="3" metricUnit="m" metricValue="1.063" pageId="3" pageNumber="443" unit="m" value="1063.0">1063 m</quantity>
|
||
a.s.l.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4C15A7FF85A023FF24F9D9FCA49F6D" httpUri="https://zenodo.org/record/228776/files/figure.png" pageId="3" pageNumber="443" targetBox="[340,1246,925,1536]" targetPageId="3">
|
||
<paragraph id="8B8C452FFF85A023FF24F9D9FCA49F6D" blockId="3.[151,1436,1559,1611]" pageId="3" pageNumber="443">
|
||
<emphasis id="B947993DFF85A023FF24F9D9FEA29F0A" bold="true" box="[151,274,1559,1581]" pageId="3" pageNumber="443">FIGURE 2.</emphasis>
|
||
Dorsolateral view of a male paratype of
|
||
<taxonomicName id="4C333EACFF85A023FD1DF9D9FC0F9F0B" box="[686,959,1559,1581]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="443" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF85A023FD1DF9D9FC0F9F0B" box="[686,959,1559,1581]" italics="true" pageId="3" pageNumber="443">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF85A023FC76F9D9FBA59F0A" box="[965,1045,1559,1580]" pageId="3" pageNumber="443" rank="species">
|
||
<emphasis id="B947993DFF85A023FC76F9D9FBA59F0A" bold="true" box="[965,1045,1559,1580]" pageId="3" pageNumber="443">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(KUHE 48014) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32916A4FF85A023FF74F9BDFC629F89" pageId="3" pageNumber="443" type="etymology">
|
||
<paragraph id="8B8C452FFF85A023FF74F9BDFC629F89" blockId="3.[151,1436,1651,2036]" pageId="3" pageNumber="443">
|
||
<emphasis id="B947993DFF85A023FF74F9BDFEFE9FAA" bold="true" box="[199,334,1651,1676]" pageId="3" pageNumber="443">Etymology.</emphasis>
|
||
The specific epithet is from Sabah and a Latin adjective
|
||
<emphasis id="B947993DFF85A023FC7DF9BAFB8B9FAD" box="[974,1083,1652,1675]" italics="true" pageId="3" pageNumber="443">montanus</emphasis>
|
||
denoting mountain, alluding to the fact that the species inhabits on mountains of Sabah, northern Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32916A4FF85A024FF74F975FDAC99D1" lastPageId="4" lastPageNumber="444" pageId="3" pageNumber="443" type="diagnosis">
|
||
<paragraph id="8B8C452FFF85A024FF74F975FDAC99D1" blockId="3.[151,1436,1651,2036]" lastBlockId="4.[151,1436,151,608]" lastPageId="4" lastPageNumber="444" pageId="3" pageNumber="443">
|
||
<emphasis id="B947993DFF85A023FF74F975FEF19FF2" bold="true" box="[199,321,1723,1748]" pageId="3" pageNumber="443">Diagnosis.</emphasis>
|
||
The new species is assigned to
|
||
<taxonomicName id="4C333EACFF85A023FD06F975FC9D9FF5" box="[693,813,1723,1747]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="3" pageNumber="443" phylum="Chordata" rank="genus">
|
||
<emphasis id="B947993DFF85A023FD06F975FC9D9FF5" box="[693,813,1723,1747]" italics="true" pageId="3" pageNumber="443">Leptolalax</emphasis>
|
||
</taxonomicName>
|
||
on the basis of the following: lack of vomerine teeth; fingers rounded in tips and lacking webbing; inner palmar tubercle elevated, not extending to the first finger; subarticular tubercles indistinct, replaced by low callous tissue; lack of nuptial pad; toe webbing basal; lack of outer metatarsal tubercle; snout tip with white vertical bar (
|
||
<bibRefCitation id="EFA238DEFF85A023FC87F8E8FBF89E19" author="Dehling" box="[820,1096,1830,1855]" pageId="3" pageNumber="443" refString="Dehling, J. M. & Matsui, M. (2013) A new species of Leptolalax (Anura: Megophryidae) from Gunung Mulu National Park, Sarawak, East Malaysia (Borneo). Zootaxa, 3670 (1), 33 - 44. http: // dx. doi. org / 10.11646 / zootaxa. 3670.1.2" type="journal article" year="2013">Dehling & Matsui 2013</bibRefCitation>
|
||
). A medium-sized form of the genus, adult males about
|
||
<quantity id="4CCBE8CAFF85A023FE02F885FD9A9E45" box="[433,554,1867,1892]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="3" pageNumber="443" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
|
||
in SVL; snout rounded in both ventral view and lateral view; interorbital space narrower than upper eyelid; male vocal sac not bipartite; toe webbing basal; dorsal skin of trunk and head smooth with tiny tubercles; pectoral gland small and indistinct; supraaxillary gland and ventrolateral glandular ridges absent; venter spotted; advertisement call consisting of long series of 1–149 notes, each composed of one to two indistinct pulse groups, dominant frequency at 6.75–7.35 kHz, without prominent frequency modulation. Most similar to
|
||
<taxonomicName id="4C333EACFF82A024FEA6FF57FECC9996" box="[277,380,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="444" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF82A024FEA6FF57FECC9996" box="[277,380,152,176]" italics="true" pageId="4" pageNumber="444">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
, but differing in smaller body size, relatively smaller eye-tympanum distance and inner metatarsal tubercle, and larger interorbital distance, upper eyelid width, snout length, eye length, and forearm and hand length compared to
|
||
<taxonomicName id="4C333EACFF82A024FE00FF2FFDA599DE" box="[435,533,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="444" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF82A024FE00FF2FFDA599DE" box="[435,533,224,248]" italics="true" pageId="4" pageNumber="444">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32916A4FF82A029FF74FECAFF459CC3" lastPageId="9" lastPageNumber="449" pageId="4" pageNumber="444" type="description">
|
||
<paragraph id="8B8C452FFF82A024FF74FECAFC7F9B46" blockId="4.[151,1436,151,608]" pageId="4" pageNumber="444">
|
||
<emphasis id="B947993DFF82A024FF74FECAFCB0983B" bold="true" box="[199,768,260,285]" pageId="4" pageNumber="444">
|
||
Description of
|
||
<typeStatus id="5488FB8DFF82A024FECFFECAFE53983B" box="[380,483,260,285]" pageId="4" pageNumber="444" type="holotype">holotype</typeStatus>
|
||
(measurement in mm).
|
||
</emphasis>
|
||
SVL 26.3; head and body moderately stocky (
|
||
<figureCitation id="130859AAFF82A024FAADFECAFA3C983A" box="[1310,1420,260,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1218]" captionTargetBox="[249,1338,647,1176]" captionTargetId="figure@4.[241,1345,642,1180]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and ventral (B) views of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 10 mm." httpUri="https://zenodo.org/record/228777/files/figure.png" pageId="4" pageNumber="444">Figures 3</figureCitation>
|
||
); head longer (10.7) than broad (9.7) and wider than trunk; snout rounded in both ventral view and lateral view, slightly protruding beyond lower jaw; eye large (4.1) subequal to snout length (4.0); pupil vertical; canthus rostralis distinct, slightly concave in dorsal view between eye and nostril; convex between nostril and tip of snout; loreal region oblique, concave; nostril lateral, below canthus, distinctly closer to eye (2.1) than to tip of snout; internarial space (2.5) narrower than interorbital distance (3.0), latter slightly narrower than upper eyelid (3.2); pineal spot invisible; tympanum distinct, rounded, diameter (2.4) three-fifths that of eye, and separated from eye by about onethird tympanic diameter (0.9); symphysial knob on anteriormost part of mandible; vomerine ridge and teeth absent; tongue large, broad, notched distally, without papillae; a median, subgular vocal sac in the mid-ventral line; vocal openings small, just posterior to rictus, close to eustachian tube openings.
|
||
</paragraph>
|
||
<caption id="DF4C15A7FF82A024FF24FB62FEED9DC6" httpUri="https://zenodo.org/record/228777/files/figure.png" pageId="4" pageNumber="444" targetBox="[249,1338,647,1176]" targetPageId="4">
|
||
<paragraph id="8B8C452FFF82A024FF24FB62FEED9DC6" blockId="4.[151,1437,1196,1248]" pageId="4" pageNumber="444">
|
||
<emphasis id="B947993DFF82A024FF24FB62FEA29DE7" bold="true" box="[151,274,1196,1218]" pageId="4" pageNumber="444">FIGURE 3.</emphasis>
|
||
Dorsal (A) and ventral (B) views of male holotype of
|
||
<taxonomicName id="4C333EACFF82A024FC85FB62FBF79DE4" box="[822,1095,1196,1218]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="4" pageNumber="444" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF82A024FC85FB62FBF79DE4" box="[822,1095,1196,1218]" italics="true" pageId="4" pageNumber="444">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF82A024FBFEFB62FB2D9DE7" box="[1101,1181,1196,1217]" pageId="4" pageNumber="444" rank="species">
|
||
<emphasis id="B947993DFF82A024FBFEFB62FB2D9DE7" bold="true" box="[1101,1181,1196,1217]" pageId="4" pageNumber="444">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(BORNEENSIS 22684). Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C452FFF82A024FF74FAC3FCB09CFC" blockId="4.[151,1436,1293,2007]" pageId="4" pageNumber="444">
|
||
Forelimb slender, moderately long (19.7); hand and forearm long (15.0); fingers long and slender, without webbing or lateral fringe of skin (
|
||
<figureCitation id="130859AAFF82A024FDA7FAFFFDC69C6F" box="[532,630,1329,1354]" captionStart="FIGURE 4" captionStartId="6.[151,250,1135,1157]" captionTargetBox="[361,1225,195,1113]" captionTargetId="figure@6.[361,1226,195,1114]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of right hand (A) and foot (B) of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 5 mm." httpUri="https://zenodo.org/record/228778/files/figure.png" pageId="4" pageNumber="444">Figure 4</figureCitation>
|
||
A); relative length of fingers II<I<IV <III; tips rounded and thickened; nuptial pad absent; inner metacarpal tubercle large, subquadratic, and prominent in thenar and metacarpal region of fingers I and II, divided subequally in proximal and distal part by shallow superficial depression, separated by a distinct groove from much smaller, outer metacarpal tubercle in metacarpal region of fingers III and IV; subarticular tubercles indistinct, replaced by long, low callous tissue.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF82A024FF74FA2BFC859FFD" blockId="4.[151,1436,1293,2007]" pageId="4" pageNumber="444">
|
||
Hindlimb relatively long (43.7), more than two times length of forelimb; tibia (14.0) longer than foot (12.2); heels overlapping when limbs are held at right angles to body; tibiotarsal articulation of adpressed limb reaching level between nostril and snout; relative length of toes I<II<V<III<IV; toe tips rounded and thickened, smaller than finger tips; toe webbing basal, webbing formula
|
||
<date id="FF8D63EFFF82A024FDD4F99AFD369F4A" box="[615,646,1620,1645]" pageId="4" pageNumber="444">
|
||
<emphasis id="B947993DFF82A024FDD4F99AFDC29F4B" bold="true" box="[615,626,1620,1645]" pageId="4" pageNumber="444">I</emphasis>
|
||
1
|
||
</date>
|
||
3/4–2+
|
||
<date id="FF8D63EFFF82A024FD76F99AFCBB9F4A" box="[709,779,1620,1645]" pageId="4" pageNumber="444">
|
||
<emphasis id="B947993DFF82A024FD76F99AFD6A9F4B" bold="true" box="[709,730,1620,1645]" pageId="4" pageNumber="444">II</emphasis>
|
||
2–3
|
||
</date>
|
||
+
|
||
<emphasis id="B947993DFF82A024FCABF99AFC889F4B" bold="true" box="[792,824,1620,1645]" pageId="4" pageNumber="444">III</emphasis>
|
||
3–
|
||
<date id="FF8D63EFFF82A024FCE9F99AFCD89F4A" box="[858,872,1620,1644]" pageId="4" pageNumber="444">4</date>
|
||
<emphasis id="B947993DFF82A024FCC3F99AFC3E9F4B" bold="true" box="[880,910,1620,1645]" pageId="4" pageNumber="444">IV</emphasis>
|
||
4+–2 3/
|
||
<date id="FF8D63EFFF82A024FC64F9ADFC469F4B" box="[983,1014,1620,1649]" pageId="4" pageNumber="444">
|
||
4
|
||
<emphasis id="B947993DFF82A024FC51F99AFC469F4B" bold="true" box="[994,1014,1620,1645]" pageId="4" pageNumber="444">V</emphasis>
|
||
</date>
|
||
(
|
||
<figureCitation id="130859AAFF82A024FBB6F99AFBD69F4A" box="[1029,1126,1620,1645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1135,1157]" captionTargetBox="[361,1225,195,1113]" captionTargetId="figure@6.[361,1226,195,1114]" captionTargetPageId="6" captionText="FIGURE 4. Ventral view of right hand (A) and foot (B) of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 5 mm." httpUri="https://zenodo.org/record/228778/files/figure.png" pageId="4" pageNumber="444">Figure 4</figureCitation>
|
||
B); fringes of skin on lateral sides of toes very narrow, hardly discernible; inner metatarsal tubercle low, oval, large, length (1.4) slightly more than half length of first toe (2.0); no outer metatarsal tubercle; subarticular tubercles obscure, but elongate, replaced by low callous tissue from base to distal half of penultimate phalanx.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF82A024FF74F928FDD69EF1" blockId="4.[151,1436,1293,2007]" pageId="4" pageNumber="444">
|
||
Skin on dorsum and dorsal side of head scattered with tiny tubercles, weakly wrinkled on dorsal surfaces of limbs (
|
||
<figureCitation id="130859AAFF82A024FF2DF8C4FEB79E05" box="[158,263,1802,1827]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1218]" captionTargetBox="[249,1338,647,1176]" captionTargetId="figure@4.[241,1345,642,1180]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and ventral (B) views of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 10 mm." httpUri="https://zenodo.org/record/228777/files/figure.png" pageId="4" pageNumber="444">Figures 3</figureCitation>
|
||
A); lateral surface of trunk with an irregular row of several larger, low tubercles; weak wrinkles on dorsal surfaces of limbs forming indistinct, reticulated, predominantly longitudinal, low ridges; supratympanic fold low but distinct, slightly angular, running from posterior margin of eye to level of insertion of forelimb; several enlarged tubercles at posterior end of rictus; chest and abdomen smooth, skin of gular region not modified; pectoral gland very small and low, hardly discernible, at insertion of forelimb; supraaxillary gland hardly discernible, and femoral gland and ventrolateral glandular ridges absent.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF83A025FF74FF59FB8B9BD6" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="445">
|
||
<emphasis id="B947993DFF83A025FF74FF59FE909996" bold="true" box="[199,288,151,176]" pageId="5" pageNumber="445">Colour.</emphasis>
|
||
In life, olive brown dorsally on head and body, with large, dark brown, irregularly shaped markings, some with trace of light outline; medial dark brown markings comprising small reverse-triangle pattern on snout, much larger, similar-shaped one between eyes and continuous W-shaped pattern on shoulder region, and irregular pattern on sacral region; dorsolateral stripe-like pattern formed by dark spots from shoulder to groin, with series of smaller dark spots ventrolaterally on side of trunk; discontinuous black band below canthus rostralis surrounding nostril; light brown stripe from between nostrils to anteriormost edge of upper jaw; upper lip barred with dark and light brown; white spots on region posteroventral to eye and on rictus conspicuous; ventral edge of supratympanic fold and tympanum except for lower part dark brown; groin with dark brown spot extending onto thigh; limbs light brown dorsally marked with alternating light and dark brown crossbars; basic colouration on upper arm lighter than on lower arm; dark crossbars restricted to lower arm and fingers; dark brown crossbars on tibia incomplete, absent on dorsal surface; area of tibio-tarsal articulation not cream-coloured dorsally; throat and chest cream-coloured with fine, light brown mottling; posterolateral portion of throat without paired unpigmented areas; chin with dark brown mottling, denser along edge of lower jaw; abdomen cream-coloured with dark brown speckling, forming diffuse spots of various size; limbs ventrally cream-coloured, heavily dusted with brown on lower arm and hindlimb; posterior surface of thigh brown speckled with cream; iris red in upper half and along outer margin of pupil, silver with black reticulation in lower half. In alcohol, colour generally darker, although brown marking on upper part of tympanum lighter; pattern generally faded but still distinct (
|
||
<figureCitation id="130859AAFF83A025FC78FD19FB9D9BD6" box="[971,1069,727,752]" captionStart="FIGURE 3" captionStartId="4.[151,250,1196,1218]" captionTargetBox="[249,1338,647,1176]" captionTargetId="figure@4.[241,1345,642,1180]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and ventral (B) views of male holotype of Leptolalax sabahmontanus sp. nov. (BORNEENSIS 22684). Scale bar = 10 mm." httpUri="https://zenodo.org/record/228777/files/figure.png" pageId="5" pageNumber="445">Figure 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF83A025FF74FD32FEFB9ACA" blockId="5.[151,1436,151,1005]" pageId="5" pageNumber="445">
|
||
<emphasis id="B947993DFF83A025FF74FD32FE8F9A33" bold="true" box="[199,319,764,789]" pageId="5" pageNumber="445">Variation.</emphasis>
|
||
Individuals of the
|
||
<typeStatus id="5488FB8DFF83A025FDA9FD33FDFA9A33" box="[538,586,765,789]" pageId="5" pageNumber="445">type</typeStatus>
|
||
series are generally similar to each other in morphology. Variation in size and body proportions is given in
|
||
<tableCitation id="C6B17094FF83A025FDB4FCD1FDED9A1E" box="[519,605,799,824]" captionStart="TABLE 2" captionStartId="5.[151,239,1052,1075]" captionTargetBox="[159,1155,1140,1988]" captionTargetPageId="5" captionText="TABLE 2. Measurements in adults of Leptolalax sabahmontanus sp. nov. SVL (Mean ± SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations." httpUri="http://table.plazi.org/id/DF4C15A7FF83A025FF24FBD2FA909D71" pageId="5" pageNumber="445" tableUuid="DF4C15A7FF83A025FF24FBD2FA909D71">Table 2</tableCitation>
|
||
. Angle of supratympanic fold varies, and in six of eight
|
||
<typeStatus id="5488FB8DFF83A025FB63FCEEFA8E9A1E" box="[1232,1342,800,824]" pageId="5" pageNumber="445" type="paratype">paratypes</typeStatus>
|
||
, is more strongly angular than in the
|
||
<typeStatus id="5488FB8DFF83A025FE69FC8AFDF09A7B" box="[474,576,836,861]" pageId="5" pageNumber="445" type="holotype">holotype</typeStatus>
|
||
. Pectoral glands are more distinct in one
|
||
<typeStatus id="5488FB8DFF83A025FBAAFC8BFBCD9A7B" box="[1049,1149,837,861]" pageId="5" pageNumber="445" type="paratype">paratype</typeStatus>
|
||
. Dark brown marking on abdomen varies from scattered dots to large round spots. Male specimens from Kinabalu and Crocker do not differ in size and proportions of body. The single female available is larger (SVL = 35.0 mm) than the largest male (
|
||
<quantity id="4CCBE8CAFF83A025FADAFC42FF779AEE" metricMagnitude="-2" metricUnit="m" metricValue="2.79" pageId="5" pageNumber="445" unit="mm" value="27.9">27.9 mm</quantity>
|
||
), and has smaller values than males in HL, HW,
|
||
<collectingCountry id="F32405BFFF83A025FD40FC7EFC989AE1" box="[755,808,944,967]" name="India" pageId="5" pageNumber="445">IND</collectingCountry>
|
||
, IOD, UEW, SL, EL, LAL, FLL, TL, FL, and HLL, all relative to SVL.
|
||
</paragraph>
|
||
<caption id="DF4C15A7FF83A025FF24FBD2FA909D71" ID-Table-UUID="DF4C15A7FF83A025FF24FBD2FA909D71" httpUri="http://table.plazi.org/id/DF4C15A7FF83A025FF24FBD2FA909D71" pageId="5" pageNumber="445" targetBox="[159,1155,1140,1988]" targetIsTable="true" targetPageId="5">
|
||
<paragraph id="8B8C452FFF83A025FF24FBD2FA909D71" blockId="5.[151,1436,1052,1111]" pageId="5" pageNumber="445">
|
||
<emphasis id="B947993DFF83A025FF24FBD2FEBD9D15" bold="true" box="[151,269,1052,1075]" pageId="5" pageNumber="445">TABLE 2.</emphasis>
|
||
Measurements in adults of
|
||
<taxonomicName id="4C333EACFF83A025FDF6FBD2FCD89D12" box="[581,872,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="5" pageNumber="445" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF83A025FDF6FBD2FCD89D12" box="[581,872,1052,1076]" italics="true" pageId="5" pageNumber="445">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF83A025FCC0FBD2FC7C9D15" box="[883,972,1052,1075]" pageId="5" pageNumber="445" rank="species">
|
||
<emphasis id="B947993DFF83A025FCC0FBD2FC7C9D15" bold="true" box="[883,972,1052,1075]" pageId="5" pageNumber="445">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
SVL (Mean±SD, in mm) and medians of ratios (R) of other characters to SVL, followed by ranges in parenthesis. See text for character abbreviations.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C452FFF83A025FEF2FBBAFB939EE2" pageId="5" pageNumber="445">
|
||
<table id="F933B78FFF835FDFFF2CFBBAFB339EE2" box="[159,1155,1140,1988]" gridcols="3" gridrows="21" pageId="5" pageNumber="445">
|
||
<tr id="3503476DFF835FDFFF2CFBBAFB339DAC" box="[159,1155,1140,1162]" gridrow="0" pageId="5" pageNumber="445" rowspan-1="1">
|
||
<th id="76D22E11FF835FDFFF2CFBBAFD949DAC" box="[159,548,1140,1162]" gridcol="0" gridrow="0" pageId="5" pageNumber="445">Kinabalu</th>
|
||
<th id="76D22E11FF835FDFFC13FBBAFB339DAC" box="[928,1155,1140,1162]" gridcol="2" gridrow="0" pageId="5" pageNumber="445">Crocker</th>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFB53FB339D95" box="[159,1155,1181,1203]" gridrow="1" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFB53FD949D95" box="[159,548,1181,1203]" gridcol="0" gridrow="1" pageId="5" pageNumber="445">14 males</th>
|
||
<td id="76D22E11FF835FDFFD7FFB53FC989D95" box="[716,808,1181,1203]" gridcol="1" gridrow="1" pageId="5" pageNumber="445">1 female</td>
|
||
<td id="76D22E11FF835FDFFC13FB53FB339D95" box="[928,1155,1181,1203]" gridcol="2" gridrow="1" pageId="5" pageNumber="445">3 males</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFB08FB339DFA" box="[159,1155,1222,1244]" gridrow="2" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFB08FD949DFA" box="[159,548,1222,1244]" gridcol="0" gridrow="2" pageId="5" pageNumber="445">SVL 26.2±0.70 (25.4–27.7)</th>
|
||
<td id="76D22E11FF835FDFFD7FFB08FC989DFA" box="[716,808,1222,1244]" gridcol="1" gridrow="2" pageId="5" pageNumber="445">35.0 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FB08FB339DFA" box="[928,1155,1222,1244]" gridcol="2" gridrow="2" pageId="5" pageNumber="445">27.1±0.74 (26.5–27.9)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFB3EFB339C20" box="[159,1155,1264,1286]" gridrow="3" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFB3EFD949C20" box="[159,548,1264,1286]" gridcol="0" gridrow="3" pageId="5" pageNumber="445">RHL 39.6 (38.2–42.9)</th>
|
||
<td id="76D22E11FF835FDFFD7FFB3EFC989C20" box="[716,808,1264,1286]" gridcol="1" gridrow="3" pageId="5" pageNumber="445">36.3 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FB3EFB339C20" box="[928,1155,1264,1286]" gridcol="2" gridrow="3" pageId="5" pageNumber="445">38.7 (38.4–41.1)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFAD7FB339C09" box="[159,1155,1305,1327]" gridrow="4" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFAD7FD949C09" box="[159,548,1305,1327]" gridcol="0" gridrow="4" pageId="5" pageNumber="445">RHW 34.2 (32.6–36.9)</th>
|
||
<td id="76D22E11FF835FDFFD7FFAD7FC989C09" box="[716,808,1305,1327]" gridcol="1" gridrow="4" pageId="5" pageNumber="445">31.7 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FAD7FB339C09" box="[928,1155,1305,1327]" gridcol="2" gridrow="4" pageId="5" pageNumber="445">34.3 (33.6–34.4)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFA8CFB339C7E" box="[159,1155,1346,1368]" gridrow="5" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFA8CFD949C7E" box="[159,548,1346,1368]" gridcol="0" gridrow="5" pageId="5" pageNumber="445">RIND 9.4 (8.8–10.5)</th>
|
||
<td id="76D22E11FF835FDFFD7FFA8CFC989C7E" box="[716,808,1346,1368]" gridcol="1" gridrow="5" pageId="5" pageNumber="445">8.0 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FA8CFB339C7E" box="[928,1155,1346,1368]" gridcol="2" gridrow="5" pageId="5" pageNumber="445">9.4 (9.3–9.7)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFAA2FB339CA4" box="[159,1155,1388,1410]" gridrow="6" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFAA2FD949CA4" box="[159,548,1388,1410]" gridcol="0" gridrow="6" pageId="5" pageNumber="445">RIOD 11.4 (9.9–12.2)</th>
|
||
<td id="76D22E11FF835FDFFD7FFAA2FC989CA4" box="[716,808,1388,1410]" gridcol="1" gridrow="6" pageId="5" pageNumber="445">9.1 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FAA2FB339CA4" box="[928,1155,1388,1410]" gridcol="2" gridrow="6" pageId="5" pageNumber="445">10.1 (9.7–10.6)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFA5BFB339C8D" box="[159,1155,1429,1451]" gridrow="7" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFA5BFD949C8D" box="[159,548,1429,1451]" gridcol="0" gridrow="7" pageId="5" pageNumber="445">RUEW 12.1(11.0–13.0)</th>
|
||
<td id="76D22E11FF835FDFFD7FFA5BFC989C8D" box="[716,808,1429,1451]" gridcol="1" gridrow="7" pageId="5" pageNumber="445">10.3 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FA5BFB339C8D" box="[928,1155,1429,1451]" gridcol="2" gridrow="7" pageId="5" pageNumber="445">12.0 (11.6–12.5)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFA70FB339CF2" box="[159,1155,1470,1492]" gridrow="8" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFA70FD949CF2" box="[159,548,1470,1492]" gridcol="0" gridrow="8" pageId="5" pageNumber="445">RSL 15.2 (13.2–16.3)</th>
|
||
<td id="76D22E11FF835FDFFD7FFA70FC989CF2" box="[716,808,1470,1492]" gridcol="1" gridrow="8" pageId="5" pageNumber="445">13.1 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FA70FB339CF2" box="[928,1155,1470,1492]" gridcol="2" gridrow="8" pageId="5" pageNumber="445">15.1 (13.8–15.4)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CFA26FB339CD8" box="[159,1155,1512,1534]" gridrow="9" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CFA26FD949CD8" box="[159,548,1512,1534]" gridcol="0" gridrow="9" pageId="5" pageNumber="445">REL 15.7 (14.4–16.3)</th>
|
||
<td id="76D22E11FF835FDFFD7FFA26FC989CD8" box="[716,808,1512,1534]" gridcol="1" gridrow="9" pageId="5" pageNumber="445">13.7 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13FA26FB339CD8" box="[928,1155,1512,1534]" gridcol="2" gridrow="9" pageId="5" pageNumber="445">15.1 (14.2–16.2)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF9DFFB339F01" box="[159,1155,1553,1575]" gridrow="10" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF9DFFD949F01" box="[159,548,1553,1575]" gridcol="0" gridrow="10" pageId="5" pageNumber="445">RN-EL 8.1 (7.3–9.1)</th>
|
||
<td id="76D22E11FF835FDFFD7FF9DFFC989F01" box="[716,808,1553,1575]" gridcol="1" gridrow="10" pageId="5" pageNumber="445">–</td>
|
||
<td id="76D22E11FF835FDFFC13F9DFFB339F01" box="[928,1155,1553,1575]" gridcol="2" gridrow="10" pageId="5" pageNumber="445">8.2 (7.9–9.0)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF9F4FB339F76" box="[159,1155,1594,1616]" gridrow="11" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF9F4FD949F76" box="[159,548,1594,1616]" gridcol="0" gridrow="11" pageId="5" pageNumber="445">RTD 7.9 (6.9–10.1)</th>
|
||
<td id="76D22E11FF835FDFFD7FF9F4FC989F76" box="[716,808,1594,1616]" gridcol="1" gridrow="11" pageId="5" pageNumber="445">7.1 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F9F4FB339F76" box="[928,1155,1594,1616]" gridcol="2" gridrow="11" pageId="5" pageNumber="445">7.9 (7.9–8.6)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF9AAFB339F5C" box="[159,1155,1636,1658]" gridrow="12" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF9AAFD949F5C" box="[159,548,1636,1658]" gridcol="0" gridrow="12" pageId="5" pageNumber="445">RE-TL 3.0 (2.0–3.9)</th>
|
||
<td id="76D22E11FF835FDFFD7FF9AAFC989F5C" box="[716,808,1636,1658]" gridcol="1" gridrow="12" pageId="5" pageNumber="445">2.9 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F9AAFB339F5C" box="[928,1155,1636,1658]" gridcol="2" gridrow="12" pageId="5" pageNumber="445">3.0 (3.0–4.3)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF943FB339F85" box="[159,1155,1677,1699]" gridrow="13" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF943FD949F85" box="[159,548,1677,1699]" gridcol="0" gridrow="13" pageId="5" pageNumber="445">RLAL 55.2 (51.6–58.5)</th>
|
||
<td id="76D22E11FF835FDFFD7FF943FC989F85" box="[716,808,1677,1699]" gridcol="1" gridrow="13" pageId="5" pageNumber="445">50.6 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F943FB339F85" box="[928,1155,1677,1699]" gridcol="2" gridrow="13" pageId="5" pageNumber="445">53.7 (52.0–56.2)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF978FB339FEA" box="[159,1155,1718,1740]" gridrow="14" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF978FD949FEA" box="[159,548,1718,1740]" gridcol="0" gridrow="14" pageId="5" pageNumber="445">RFLL 73.6 (69.3–79.5)</th>
|
||
<td id="76D22E11FF835FDFFD7FF978FC989FEA" box="[716,808,1718,1740]" gridcol="1" gridrow="14" pageId="5" pageNumber="445">67.1 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F978FB339FEA" box="[928,1155,1718,1740]" gridcol="2" gridrow="14" pageId="5" pageNumber="445">73.4 (72.0–74.7)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF92EFB339FD0" box="[159,1155,1760,1782]" gridrow="15" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF92EFD949FD0" box="[159,548,1760,1782]" gridcol="0" gridrow="15" pageId="5" pageNumber="445">RIPL 7.3 (5.4–8.7)</th>
|
||
<td id="76D22E11FF835FDFFD7FF92EFC989FD0" box="[716,808,1760,1782]" gridcol="1" gridrow="15" pageId="5" pageNumber="445">6.0 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F92EFB339FD0" box="[928,1155,1760,1782]" gridcol="2" gridrow="15" pageId="5" pageNumber="445">7.5 (7.1–7.9)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF8C7FB339E39" box="[159,1155,1801,1823]" gridrow="16" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF8C7FD949E39" box="[159,548,1801,1823]" gridcol="0" gridrow="16" pageId="5" pageNumber="445">RTL 54.0 (50.4–56.5)</th>
|
||
<td id="76D22E11FF835FDFFD7FF8C7FC989E39" box="[716,808,1801,1823]" gridcol="1" gridrow="16" pageId="5" pageNumber="445">48.6 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F8C7FB339E39" box="[928,1155,1801,1823]" gridcol="2" gridrow="16" pageId="5" pageNumber="445">54.1 (52.2–55.8)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF8FCFB339E6E" box="[159,1155,1842,1864]" gridrow="17" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF8FCFD949E6E" box="[159,548,1842,1864]" gridcol="0" gridrow="17" pageId="5" pageNumber="445">RFL 46.5 (42.7–51.2)</th>
|
||
<td id="76D22E11FF835FDFFD7FF8FCFC989E6E" box="[716,808,1842,1864]" gridcol="1" gridrow="17" pageId="5" pageNumber="445">42.3 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F8FCFB339E6E" box="[928,1155,1842,1864]" gridcol="2" gridrow="17" pageId="5" pageNumber="445">47.4 (47.3–52.1)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF892FB339E54" box="[159,1155,1884,1906]" gridrow="18" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF892FD949E54" box="[159,548,1884,1906]" gridcol="0" gridrow="18" pageId="5" pageNumber="445">RHLL 170.3 (162.2–186.0)</th>
|
||
<td id="76D22E11FF835FDFFD7FF892FC989E54" box="[716,808,1884,1906]" gridcol="1" gridrow="18" pageId="5" pageNumber="445">160.6 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F892FB339E54" box="[928,1155,1884,1906]" gridcol="2" gridrow="18" pageId="5" pageNumber="445">173.9 (169.0–178.9)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF84BFB339EBD" box="[159,1155,1925,1947]" gridrow="19" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF84BFD949EBD" box="[159,548,1925,1947]" gridcol="0" gridrow="19" pageId="5" pageNumber="445">RIMTL 5.1 (4.7–6.2)</th>
|
||
<td id="76D22E11FF835FDFFD7FF84BFC989EBD" box="[716,808,1925,1947]" gridcol="1" gridrow="19" pageId="5" pageNumber="445">6.0 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F84BFB339EBD" box="[928,1155,1925,1947]" gridcol="2" gridrow="19" pageId="5" pageNumber="445">6.0 (4.9–7.5)</td>
|
||
</tr>
|
||
<tr id="3503476DFF835FDFFF2CF860FB339EE2" box="[159,1155,1966,1988]" gridrow="20" pageId="5" pageNumber="445">
|
||
<th id="76D22E11FF835FDFFF2CF860FD949EE2" box="[159,548,1966,1988]" gridcol="0" gridrow="20" pageId="5" pageNumber="445">R1TOEL 8.2 (7.1–9.0)</th>
|
||
<td id="76D22E11FF835FDFFD7FF860FC989EE2" box="[716,808,1966,1988]" gridcol="1" gridrow="20" pageId="5" pageNumber="445">7.1 (–)</td>
|
||
<td id="76D22E11FF835FDFFC13F860FB339EE2" box="[928,1155,1966,1988]" gridcol="2" gridrow="20" pageId="5" pageNumber="445">8.8 (8.7–9.0)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF4C15A7FF80A026FF24FBA1FDE09D82" httpUri="https://zenodo.org/record/228778/files/figure.png" pageId="6" pageNumber="446" targetBox="[361,1225,195,1113]" targetPageId="6">
|
||
<paragraph id="8B8C452FFF80A026FF24FBA1FDE09D82" blockId="6.[151,1435,1135,1188]" pageId="6" pageNumber="446">
|
||
<emphasis id="B947993DFF80A026FF24FBA1FEAA9DA2" bold="true" box="[151,282,1135,1157]" pageId="6" pageNumber="446">FIGURE 4.</emphasis>
|
||
Ventral view of right hand (A) and foot (B) of male holotype of
|
||
<taxonomicName id="4C333EACFF80A026FBA8FBA1FA849DA3" box="[1051,1332,1135,1157]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF80A026FBA8FBA1FA849DA3" box="[1051,1332,1135,1157]" italics="true" pageId="6" pageNumber="446">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF80A026FAF0FBBEFA2B9DA3" box="[1347,1435,1136,1157]" pageId="6" pageNumber="446" rank="species">
|
||
<emphasis id="B947993DFF80A026FAF0FBBEFA2B9DA3" bold="true" box="[1347,1435,1136,1157]" pageId="6" pageNumber="446">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(BORNEENSIS 22684). Scale bar = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C452FFF80A026FF74FB05FABA9CC5" blockId="6.[151,1437,1227,2014]" pageId="6" pageNumber="446">
|
||
<emphasis id="B947993DFF80A026FF74FB05FE2C9DC2" bold="true" box="[199,412,1227,1252]" pageId="6" pageNumber="446">Call characterics.</emphasis>
|
||
The advertisement call of four
|
||
<typeStatus id="5488FB8DFF80A026FCA2FB02FCCC9DC2" box="[785,892,1228,1252]" pageId="6" pageNumber="446" type="paratype">paratypes</typeStatus>
|
||
(BORNEENSIS 22690, 22691; KUHE 48015, 48016) from KNP, recorded at the air temperatures of 17.2–17.7o C were 0.02 to 11.6 (mean±SD = 1.8±2.1, n = 54) s in duration, with a call interval of 0.1–4.0 (0.5±0.6, n = 48) s (
|
||
<figureCitation id="130859AAFF80A026FCD0FAD8FC759C09" box="[867,965,1302,1327]" captionStart="FIGURE 5" captionStartId="8.[151,250,1810,1832]" captionTargetBox="[221,1361,269,1782]" captionTargetId="figure@8.[219,1368,265,1789]" captionTargetPageId="8" captionText="FIGURE 5. Sonagram (A, C, E) and wave form (B, D, F) showing advertisement calls of a paratype of Leptolalax sabahmontanus sp. nov. (KUHE 48016) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park, recorded at an air temperature of 17.2 ° C." httpUri="https://zenodo.org/record/228779/files/figure.png" pageId="6" pageNumber="446">Figure 5</figureCitation>
|
||
,
|
||
<tableCitation id="C6B17094FF80A026FC62FAD8FB969C09" box="[977,1062,1302,1327]" captionStart="TABLE 3" captionStartId="7.[151,239,512,535]" captionTargetBox="[159,1417,762,869]" captionTargetPageId="7" captionText="TABLE 3. Parameters of the advertisement call of six males of Leptolalax sabahmontanus sp. nov., recorded at Kinabalu National Park (KNP) and Crocker Range National Park (CRNP). Parameter values given as mean ± SD, followed by sample size (n) and range in parentheses. AT: air temperature; Call L: call length; Call ITV: call interval; Note RR: note repetition rate; Note L: note length; Note ITV: note interval; Df at beginning: dominant frequency at the beginning of call; Df at end: dominant frequency at the end of call; Df decrease: decrease in dominant frequency in a call." httpUri="http://table.plazi.org/id/DF4C15A7FF81A027FF24FDCEFA2C9BB1" pageId="6" pageNumber="446" tableUuid="DF4C15A7FF81A027FF24FDCEFA2C9BB1">Table 3</tableCitation>
|
||
). A call consisted of a series of 1 to 149 (mean±SD = 24.2±26.9, n = 54) distinct notes. Each note lasted 3–23 (11.7±4.6, n = 46) ms with a note interval of 54–78 (65.4±6.0, n = 32) ms, and a note repetition rate of 12.5–17.3 (13.4±0.9, n = 51) notes per second. Each note was composed of 1–2 (1.5±0.5, n = 46) indistinct pulse groups. There was no marked frequency modulation within a note, and the dominant frequency at the beginning of the call, 6.90–7.35 (7.10±0.11, n = 46) kHz, ended at 6.75–7.10 (6.94±0.09) kHz, with the decrease of only 0.0–0.4 (0.16±0.08, n = 46) kHz.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF80A026FF74FA20FD169FC4" blockId="6.[151,1437,1227,2014]" pageId="6" pageNumber="446">The advertisement calls of two males from CRNP (BORNEENSIS 8673 from Trail 11 and 12454 from Mahua) recorded at the air temperature of 19.8 o C and 21.3o C, respectively, were basically similar to Kinabalu calls. However, calls of BORNEENSIS 12454 consisted of longer notes (22–25 [24.0±1.1, n = 11] ms) with shorter intervals (39–44 [41.9±1.4, n = 10] ms) and higher frequency (initial dominant 7.40–7.60 [7.53±0.06, n = 11] kHz and final dominant 7.20–7.35 [7.29±0.05] kHz) than in others. Additionally, a harmonic frequency was detected at 20.25–20.45 (20.33±0.08, n = 11) kHz in the calls of BORNEENSIS 12454, probably because of the quieter environment in which the recording was made.</paragraph>
|
||
<paragraph id="8B8C452FFF80A027FF74F923FD1098F6" blockId="6.[151,1437,1227,2014]" lastBlockId="7.[151,1436,151,464]" lastPageId="7" lastPageNumber="447" pageId="6" pageNumber="446">
|
||
<emphasis id="B947993DFF80A026FF74F923FEDD9E20" bold="true" box="[199,365,1773,1798]" pageId="6" pageNumber="446">Comparisons.</emphasis>
|
||
<taxonomicName id="4C333EACFF80A026FEC4F923FD1A9E23" box="[375,682,1773,1797]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF80A026FEC4F923FD1A9E23" box="[375,682,1773,1797]" italics="true" pageId="6" pageNumber="446">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
differs from congeneric species occurring outside Borneo in the same manner as in
|
||
<taxonomicName id="4C333EACFF80A026FEC6F8DCFC869E0C" authority="Dehling & Matsui 2013" authorityName="Dehling & Matsui" authorityYear="2013" box="[373,822,1809,1834]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF80A026FEC6F8DCFDB79E0F" box="[373,519,1809,1833]" italics="true" pageId="6" pageNumber="446">L. fritinniens</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA238DEFF80A026FDABF8DFFC9E9E0C" author="Dehling" box="[536,814,1809,1834]" pageId="6" pageNumber="446" refString="Dehling, J. M. & Matsui, M. (2013) A new species of Leptolalax (Anura: Megophryidae) from Gunung Mulu National Park, Sarawak, East Malaysia (Borneo). Zootaxa, 3670 (1), 33 - 44. http: // dx. doi. org / 10.11646 / zootaxa. 3670.1.2" type="journal article" year="2013">Dehling & Matsui 2013</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, and comparisons with Bornean species are pertinent.
|
||
<taxonomicName id="4C333EACFF80A026FF24F8FBFE779E6B" box="[151,455,1845,1869]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF80A026FF24F8FBFE779E6B" box="[151,455,1845,1869]" italics="true" pageId="6" pageNumber="446">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
morphologically differs from all the described Bornean species in the following way. By the presence of ventral markings, it resembles
|
||
<taxonomicName id="4C333EACFF80A026FD7BF894FC999E57" box="[712,809,1881,1905]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF80A026FD7BF894FC999E57" box="[712,809,1881,1905]" italics="true" pageId="6" pageNumber="446">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF80A026FC8BF894FC1B9E57" box="[824,939,1881,1905]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF80A026FC8BF894FC1B9E57" box="[824,939,1881,1905]" italics="true" pageId="6" pageNumber="446">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C333EACFF80A026FC52F894FBC29E57" box="[993,1138,1881,1905]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF80A026FC52F894FBC29E57" box="[993,1138,1881,1905]" italics="true" pageId="6" pageNumber="446">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
. Of these, the new species is most similar to
|
||
<taxonomicName id="4C333EACFF80A026FED9F8B0FE7E9EB3" box="[362,462,1917,1941]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF80A026FED9F8B0FE7E9EB3" box="[362,462,1917,1941]" italics="true" pageId="6" pageNumber="446">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
, but results of statistical tests in males indicated that the new species (Kinabalu and Crocker combined, n = 17) is smaller than
|
||
<taxonomicName id="4C333EACFF80A026FDC6F86CFD689E9F" box="[629,728,1953,1977]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF80A026FDC6F86CFD689E9F" box="[629,728,1953,1977]" italics="true" pageId="6" pageNumber="446">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
from Mulu (n = 7) in SVL (mean =
|
||
<quantity id="4CCBE8CAFF80A026FBDEF86FFB619E9F" box="[1133,1233,1953,1978]" metricMagnitude="-2" metricUnit="m" metricValue="2.6399999999999997" pageId="6" pageNumber="446" unit="mm" value="26.4">26.4 mm</quantity>
|
||
vs.
|
||
<quantity id="4CCBE8CAFF80A026FB4DF86CFAD29E9F" box="[1278,1378,1954,1978]" metricMagnitude="-2" metricUnit="m" metricValue="2.91" pageId="6" pageNumber="446" unit="mm" value="29.1">29.1 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C333EACFF80A026FA36F86CFF6B9EFB" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="6" pageNumber="446" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF80A026FA36F86CFF6B9EFB" italics="true" pageId="6" pageNumber="446">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
), and that it has smaller eye-tympanum distance (median = 3.0% vs. 4.1%) and inner metatarsal tubercle (5.1% vs. 5.8%), and larger interorbital distance (11.3% vs. 9.9%), upper eyelid width (12.1% vs. 11.2%), snout length (15.2% vs. 13.1%), eye length (15.6% vs. 14.0%), and forearm and hand length (55.1% vs. 52.4%), all relative to SVL, than
|
||
<taxonomicName id="4C333EACFF81A027FE35FF2FFE5899DE" box="[390,488,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF81A027FE35FF2FFE5899DE" box="[390,488,224,248]" italics="true" pageId="7" pageNumber="447">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
(Mann-Whitney U-test, p <0.05).
|
||
<taxonomicName id="4C333EACFF81A027FCDFFF2EFB2A99DE" box="[876,1178,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF81A027FCDFFF2EFB2A99DE" box="[876,1178,224,248]" italics="true" pageId="7" pageNumber="447">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C333EACFF81A027FA9AFF2FFA2B99DE" box="[1321,1435,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF81A027FA9AFF2FFA2B99DE" box="[1321,1435,224,248]" italics="true" pageId="7" pageNumber="447">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
in the absence of dark cross bars on the upper arm, but differs from it by a smaller (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FB63FECAFAC4983A" box="[1232,1396,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
vs.
|
||
<quantity id="4CCBE8CAFF81A027FF24FEE6FE8B9819" box="[151,315,296,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.5150000000000006" metricValueMax="3.9200000000000004" metricValueMin="3.1100000000000003" pageId="7" pageNumber="447" unit="mm" value="35.15" valueMax="39.2" valueMin="31.1">31.1–39.2 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C333EACFF81A027FED3FEE7FE649866" box="[352,468,296,320]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF81A027FED3FEE7FE649866" box="[352,468,296,320]" italics="true" pageId="7" pageNumber="447">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
) and stockier body, and less developed ventral dark marking (body more slender and dark marking usually conspicuous in
|
||
<taxonomicName id="4C333EACFF81A027FD8AFE83FD1C9842" box="[569,684,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF81A027FD8AFE83FD1C9842" box="[569,684,332,356]" italics="true" pageId="7" pageNumber="447">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C333EACFF81A027FD77FE82FC429842" box="[708,1010,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF81A027FD77FE82FC429842" box="[708,1010,332,356]" italics="true" pageId="7" pageNumber="447">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C333EACFF81A027FB9BFE83FB2B9842" box="[1064,1179,332,356]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF81A027FB9BFE83FB2B9842" box="[1064,1179,332,356]" italics="true" pageId="7" pageNumber="447">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
are markedly different in advertisement calls (see below). From
|
||
<taxonomicName id="4C333EACFF81A027FDD6FEBFFD4598AE" box="[613,757,368,392]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF81A027FDD6FEBFFD4598AE" box="[613,757,368,392]" italics="true" pageId="7" pageNumber="447">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs by the smaller body size (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FF24FE5AFE8B988A" box="[151,315,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
SVL vs.
|
||
<quantity id="4CCBE8CAFF81A027FE1AFE5AFDFD988A" box="[425,589,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="3.11" metricValueMax="3.4799999999999995" metricValueMin="2.7399999999999998" pageId="7" pageNumber="447" unit="mm" value="31.099999999999998" valueMax="34.8" valueMin="27.4">27.4–34.8 mm</quantity>
|
||
SVL in
|
||
<taxonomicName id="4C333EACFF81A027FD03FE5BFCF2988A" box="[688,834,404,428]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF81A027FD03FE5BFCF2988A" box="[688,834,404,428]" italics="true" pageId="7" pageNumber="447">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
) and having a single, medially arranged vocal sac in males (vs. bipartite vocal sac in
|
||
<taxonomicName id="4C333EACFF81A027FE4CFE77FD3F98F6" box="[511,655,440,464]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF81A027FE4CFE77FD3F98F6" box="[511,655,440,464]" italics="true" pageId="7" pageNumber="447">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4C15A7FF81A027FF24FDCEFA2C9BB1" ID-Table-UUID="DF4C15A7FF81A027FF24FDCEFA2C9BB1" httpUri="http://table.plazi.org/id/DF4C15A7FF81A027FF24FDCEFA2C9BB1" pageId="7" pageNumber="447" targetBox="[159,1417,762,869]" targetIsTable="true" targetPageId="7">
|
||
<paragraph id="8B8C452FFF81A027FF24FDCEFA2C9BB1" blockId="7.[151,1436,511,663]" pageId="7" pageNumber="447">
|
||
<emphasis id="B947993DFF81A027FF24FDCEFEBE9B31" bold="true" box="[151,270,512,535]" pageId="7" pageNumber="447">TABLE 3.</emphasis>
|
||
Parameters of the advertisement call of six males of
|
||
<taxonomicName id="4C333EACFF81A027FCCCFE31FB149B31" box="[895,1188,511,535]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF81A027FCCCFE31FB149B31" box="[895,1188,511,535]" italics="true" pageId="7" pageNumber="447">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF81A027FB02FDCEFABC9B31" box="[1201,1292,512,535]" pageId="7" pageNumber="447" rank="species">
|
||
<emphasis id="B947993DFF81A027FB02FDCEFABC9B31" bold="true" box="[1201,1292,512,535]" pageId="7" pageNumber="447">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, recorded at Kinabalu National Park (KNP) and Crocker Range National Park (CRNP). Parameter values given as mean ± SD, followed by sample size (n) and range in parentheses. AT: air temperature; Call L: call length; Call ITV: call interval; Note RR: note repetition rate; Note L: note length; Note ITV: note interval; Df at beginning: dominant frequency at the beginning of call; Df at end: dominant frequency at the end of call; Df decrease: decrease in dominant frequency in a call.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C452FFF81A027FF2CFD7CFA309BEE" blockId="7.[159,1408,690,743]" box="[159,1408,690,712]" pageId="7" pageNumber="447">Voucher BORNENSIS BORNENSIS BORNENSIS BORNENSIS BORNENSIS BORNENSIS</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FED5FD1FFAFF9AA9" blockId="7.[159,1408,690,743]" lastBlockId="7.[159,1417,889,1283]" pageId="7" pageNumber="447">22686 22691 22690 22682 8673 12454 (0.2–2.6) (0.8–11.6) (0.02–3.8) (0.02–4.0) (0.3–5.0) (0.1–2.4)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFD35FAC09A43" pageId="7" pageNumber="447">
|
||
<table id="F933B78FFF815FDFFF2CFD34FA399A43" box="[159,1417,762,869]" gridcols="6" gridrows="3" pageId="7" pageNumber="447">
|
||
<tr id="3503476DFF815FDFFF2CFD34FA399A36" box="[159,1417,762,784]" gridrow="0" pageId="7" pageNumber="447">
|
||
<th id="76D22E11FF815FDFFF2CFD34FF4F9A36" box="[159,255,762,784]" gridcol="0" gridrow="0" pageId="7" pageNumber="447">Loc</th>
|
||
<th id="76D22E11FF815FDFFED5FD34FE539A36" box="[358,483,762,784]" gridcol="1" gridrow="0" pageId="7" pageNumber="447">KNP</th>
|
||
<th id="76D22E11FF815FDFFDA8FD34FD3C9A36" box="[539,652,762,784]" gridcol="2" gridrow="0" pageId="7" pageNumber="447">KNP</th>
|
||
<th id="76D22E11FF815FDFFD62FD34FCFF9A36" box="[721,847,762,784]" gridcol="3" gridrow="0" pageId="7" pageNumber="447">KNP</th>
|
||
<th id="76D22E11FF815FDFFC34FD34FBB49A36" box="[903,1028,762,784]" gridcol="4" gridrow="0" pageId="7" pageNumber="447">KNP</th>
|
||
<th id="76D22E11FF815FDFFB8EFD34FA399A36" box="[1085,1417,762,784]" gridcol="5" gridrow="0" pageId="7" pageNumber="447">CRNP Trail 11 CRNP Mahua</th>
|
||
</tr>
|
||
<tr id="3503476DFF815FDFFF2CFCEDFA399A1A" box="[159,1417,803,828]" gridrow="1" pageId="7" pageNumber="447">
|
||
<th id="76D22E11FF815FDFFF2CFCEDFF4F9A1A" box="[159,255,803,828]" gridcol="0" gridrow="1" pageId="7" pageNumber="447">AT (o C)</th>
|
||
<td id="76D22E11FF815FDFFED5FCEDFE539A1A" box="[358,483,803,828]" gridcol="1" gridrow="1" pageId="7" pageNumber="447">17.2</td>
|
||
<td id="76D22E11FF815FDFFDA8FCEDFD3C9A1A" box="[539,652,803,828]" gridcol="2" gridrow="1" pageId="7" pageNumber="447">17.7</td>
|
||
<td id="76D22E11FF815FDFFD62FCEDFCFF9A1A" box="[721,847,803,828]" gridcol="3" gridrow="1" pageId="7" pageNumber="447">17.7</td>
|
||
<td id="76D22E11FF815FDFFC34FCEDFBB49A1A" box="[903,1028,803,828]" gridcol="4" gridrow="1" pageId="7" pageNumber="447">17.7</td>
|
||
<td id="76D22E11FF815FDFFB8EFCEDFA399A1A" box="[1085,1417,803,828]" gridcol="5" gridrow="1" pageId="7" pageNumber="447">19.8 21.3</td>
|
||
</tr>
|
||
<tr id="3503476DFF815FDFFF2CFC81FA399A43" box="[159,1417,847,869]" gridrow="2" pageId="7" pageNumber="447">
|
||
<th id="76D22E11FF815FDFFF2CFC81FF4F9A43" box="[159,255,847,869]" gridcol="0" gridrow="2" pageId="7" pageNumber="447">Call L (s)</th>
|
||
<td id="76D22E11FF815FDFFED5FC81FE539A43" box="[358,483,847,869]" gridcol="1" gridrow="2" pageId="7" pageNumber="447">1.3±0.7 (17)</td>
|
||
<td id="76D22E11FF815FDFFDA8FC81FD3C9A43" box="[539,652,847,869]" gridcol="2" gridrow="2" pageId="7" pageNumber="447">4.1±4.1 (9)</td>
|
||
<td id="76D22E11FF815FDFFD62FC81FCFF9A43" box="[721,847,847,869]" gridcol="3" gridrow="2" pageId="7" pageNumber="447">1.4±1.0 (14)</td>
|
||
<td id="76D22E11FF815FDFFC34FC81FBB49A43" box="[903,1028,847,869]" gridcol="4" gridrow="2" pageId="7" pageNumber="447">1.4±1.2 (15)</td>
|
||
<td id="76D22E11FF815FDFFB8EFC81FA399A43" box="[1085,1417,847,869]" gridcol="5" gridrow="2" pageId="7" pageNumber="447">2.2±1.4 (11) 1.1±0.7 (15)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFC6CFAFF9AC4" blockId="7.[159,1417,889,1283]" pageId="7" pageNumber="447">Call ITV (s) 0.5±0.4 (16) 0.8±1.4 (8) 0.3±0.2 (13) 0.4±0.1 (13) 0.3±0.1 (10) 0.3±0.1 (14) (0.2–2.0) (0.2–4.0) (0.1–0.6) (0.1–0.6) (0.2–0.4) (0.1–0.5)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFC3BFA399D2D" blockId="7.[159,1417,889,1283]" box="[159,1417,1013,1035]" pageId="7" pageNumber="447">Notes/Call 17.3±8.7 (17) 54.0±53.1 (9) 18.6±12.8 (14) 19.0±15.0 (15) 32.0±20.7 (11) 18.2±10.4 (15)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FED5FBD0FA869D12" blockId="7.[159,1417,889,1283]" box="[358,1334,1054,1076]" pageId="7" pageNumber="447">(3–33) (10–149) (1–48) (1–53) (5–72) (2–37)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFB86FAD89DA1" blockId="7.[159,1417,889,1283]" pageId="7" pageNumber="447">Note RR (/s) 13.3±0.8 (17) 13.2±0.2 (9) 13.3±1.0 (14) 13.8±1.2 (15) 14.8±0.9 (11) 16.4±1.0 (15) (12.7–16.2) (12.8–13.4) (12.5–16.4) (13.0–17.3) (14.2–17.2) (15.6–18.9)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFB55FACB9D96" blockId="7.[159,1417,889,1283]" box="[159,1403,1178,1200]" pageId="7" pageNumber="447">Note L (ms) 6.0±1.3 (11) 18.2±2.4 (11) 10.3±1.0 (13) 12.6±1.2 (11) 15.0±3.0 (10) 24.0±1.1 (11)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FED5FB0AFAF29DFC" blockId="7.[159,1417,889,1283]" box="[358,1346,1220,1242]" pageId="7" pageNumber="447">(3–8) (14–23) (9–12) (11–15) (10–18) (22–25)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFB20FAD99CF4" blockId="7.[159,1417,889,1283]" lastBlockId="7.[159,1416,1468,1572]" pageId="7" pageNumber="447">Note ITV (ms) 73.1±1.4 (10) 58.7±2.2 (10) 71.9±2.5 (12) 64.3±1.5 (10) 54.3±2.6 (9) 41.9±1.4 (10) (6.80–7.00) (7.00–7.10) (6.95–7.05) (6.75–6.90) (6.70–6.85) (7.20–7.35)</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FED5FAD8FA389C8E" pageId="7" pageNumber="447">
|
||
<table id="F933B78FFF815FDFFF2CFAD8FA399C8E" box="[159,1417,1302,1448]" gridcols="7" gridrows="3" pageId="7" pageNumber="447">
|
||
<tr id="3503476DFF815FDFFF2CFAD8FA399C0A" box="[159,1417,1302,1324]" gridrow="0" pageId="7" pageNumber="447" rowspan-0="1">
|
||
<th id="76D22E11FF815FDFFED5FAD8FE4B9C0A" box="[358,507,1302,1324]" gridcol="1" gridrow="0" pageId="7" pageNumber="447">(71–75)</th>
|
||
<th id="76D22E11FF815FDFFDA8FAD8FD019C0A" box="[539,689,1302,1324]" gridcol="2" gridrow="0" pageId="7" pageNumber="447">(54–61)</th>
|
||
<th id="76D22E11FF815FDFFD62FAD8FCD89C0A" box="[721,872,1302,1324]" gridcol="3" gridrow="0" pageId="7" pageNumber="447">(68–78)</th>
|
||
<th id="76D22E11FF815FDFFC34FAD8FBAD9C0A" box="[903,1053,1302,1324]" gridcol="4" gridrow="0" pageId="7" pageNumber="447">(62–66)</th>
|
||
<th id="76D22E11FF815FDFFB8EFAD8FB639C0A" box="[1085,1235,1302,1324]" gridcol="5" gridrow="0" pageId="7" pageNumber="447">(51–59)</th>
|
||
<th id="76D22E11FF815FDFFB41FAD8FA399C0A" box="[1266,1417,1302,1324]" gridcol="6" gridrow="0" pageId="7" pageNumber="447">(39–44)</th>
|
||
</tr>
|
||
<tr id="3503476DFF815FDFFF2CFAF1FA399C59" box="[159,1417,1343,1407]" gridrow="1" pageId="7" pageNumber="447">
|
||
<th id="76D22E11FF815FDFFF2CFAF1FE8E9C59" box="[159,318,1343,1407]" gridcol="0" gridrow="1" pageId="7" pageNumber="447">Df at beginning (kHz)</th>
|
||
<td id="76D22E11FF815FDFFED5FAF1FE4B9C59" box="[358,507,1343,1407]" gridcol="1" gridrow="1" pageId="7" pageNumber="447">7.02±0.06 (11) (6.95–7.10)</td>
|
||
<td id="76D22E11FF815FDFFDA8FAF1FD019C59" box="[539,689,1343,1407]" gridcol="2" gridrow="1" pageId="7" pageNumber="447">7.15±0.05 (11) (7.10–7.20)</td>
|
||
<td id="76D22E11FF815FDFFD62FAF1FCD89C59" box="[721,872,1343,1407]" gridcol="3" gridrow="1" pageId="7" pageNumber="447">7.23±0.07 (13) (7.10–7.35)</td>
|
||
<td id="76D22E11FF815FDFFC34FAF1FBAD9C59" box="[903,1053,1343,1407]" gridcol="4" gridrow="1" pageId="7" pageNumber="447">6.99±0.05 (11) (6.95–7.10)</td>
|
||
<td id="76D22E11FF815FDFFB8EFAF1FB639C59" box="[1085,1235,1343,1407]" gridcol="5" gridrow="1" pageId="7" pageNumber="447">6.80±0.04 (10) (6.75–6.85)</td>
|
||
<td id="76D22E11FF815FDFFB41FAF1FA399C59" box="[1266,1417,1343,1407]" gridcol="6" gridrow="1" pageId="7" pageNumber="447">7.53±0.06 (11) (7.40–7.60)</td>
|
||
</tr>
|
||
<tr id="3503476DFF815FDFFF2CFA5CFA399C8E" box="[159,1417,1426,1448]" gridrow="2" pageId="7" pageNumber="447">
|
||
<th id="76D22E11FF815FDFFF2CFA5CFE8E9C8E" box="[159,318,1426,1448]" gridcol="0" gridrow="2" pageId="7" pageNumber="447">Df at end (kHz)</th>
|
||
<td id="76D22E11FF815FDFFED5FA5CFE4B9C8E" box="[358,507,1426,1448]" gridcol="1" gridrow="2" pageId="7" pageNumber="447">6.90±0.07 (11)</td>
|
||
<td id="76D22E11FF815FDFFDA8FA5CFD019C8E" box="[539,689,1426,1448]" gridcol="2" gridrow="2" pageId="7" pageNumber="447">7.03±0.03 (11)</td>
|
||
<td id="76D22E11FF815FDFFD62FA5CFCD89C8E" box="[721,872,1426,1448]" gridcol="3" gridrow="2" pageId="7" pageNumber="447">7.01±0.04 (13)</td>
|
||
<td id="76D22E11FF815FDFFC34FA5CFBAD9C8E" box="[903,1053,1426,1448]" gridcol="4" gridrow="2" pageId="7" pageNumber="447">6.83±0.04 (11)</td>
|
||
<td id="76D22E11FF815FDFFB8EFA5CFB639C8E" box="[1085,1235,1426,1448]" gridcol="5" gridrow="2" pageId="7" pageNumber="447">6.75±0.05 (10)</td>
|
||
<td id="76D22E11FF815FDFFB41FA5CFA399C8E" box="[1266,1417,1426,1448]" gridcol="6" gridrow="2" pageId="7" pageNumber="447">7.29±0.05 (11)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF81A027FF2CFA2BFA389CDD" blockId="7.[159,1416,1468,1572]" box="[159,1416,1509,1531]" pageId="7" pageNumber="447">Df decrease 0.13±0.07 (11) 0.13±0.06 (11) 0.22±0.08 (13) 0.16±0.07 (11) 0.05±0.04 (10) 0.24±0.07 (11)</paragraph>
|
||
<paragraph id="8B8C452FFF81A028FF2CF9CAFCC399F2" blockId="7.[159,1416,1468,1572]" lastBlockId="8.[151,1436,151,213]" lastPageId="8" lastPageNumber="448" pageId="7" pageNumber="447">
|
||
(kHz) (0.05–0.20) (0.05–0.25) (0.10–0.40) (0.10–0.35) (0–0.10) (0.10–0.35) The presence of ventral markings clearly differentiates
|
||
<taxonomicName id="4C333EACFF81A027FCF5F9A4FBA69FA7" box="[838,1046,1641,1665]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF81A027FCF5F9A4FBA69FA7" box="[838,1046,1641,1665]" italics="true" pageId="7" pageNumber="447">L. sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C333EACFF81A027FBD3F9A4FB739FA7" box="[1120,1219,1642,1665]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF81A027FBD3F9A4FB739FA7" box="[1120,1219,1642,1665]" italics="true" pageId="7" pageNumber="447">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF81A027FB67F9A4FAF29FA7" box="[1236,1346,1641,1665]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="hamidi">
|
||
<emphasis id="B947993DFF81A027FB67F9A4FAF29FA7" box="[1236,1346,1641,1665]" italics="true" pageId="7" pageNumber="447">L. hamidi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C333EACFF81A027FA36F9A4FF509F83" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF81A027FA36F9A4FF509F83" italics="true" pageId="7" pageNumber="447">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
. Furthermore, from
|
||
<taxonomicName id="4C333EACFF81A027FE79F940FD9C9F83" box="[458,556,1678,1701]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF81A027FE79F940FD9C9F83" box="[458,556,1678,1701]" italics="true" pageId="7" pageNumber="447">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
, the new species differs by having a smaller body size (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FA9FF943FF739FEF" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
vs. 31.1–34.0 mm SVL in
|
||
<taxonomicName id="4C333EACFF81A027FE41F97CFDE39FEF" box="[498,595,1714,1737]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF81A027FE41F97CFDE39FEF" box="[498,595,1714,1737]" italics="true" pageId="7" pageNumber="447">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
), olive brown dorsum with black pattern without distinct light outlines (vs. dorsum pinkish brown with dark brown markings with conspicuous thin light outlines in
|
||
<taxonomicName id="4C333EACFF81A027FB08F918FA909FCB" box="[1211,1312,1750,1773]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF81A027FB08F918FA909FCB" box="[1211,1312,1750,1773]" italics="true" pageId="7" pageNumber="447">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
). From
|
||
<emphasis id="B947993DFF81A027FA36F918FE719E37" italics="true" pageId="7" pageNumber="447">
|
||
<taxonomicName id="4C333EACFF81A027FA36F918FF589E37" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF81A027FF40F934FE719E37" box="[243,449,1785,1809]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus">L. sabahmontanus</taxonomicName>
|
||
</emphasis>
|
||
also differs by having a smaller body size (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FB94F937FB7B9E37" box="[1063,1227,1785,1810]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
vs.
|
||
<quantity id="4CCBE8CAFF81A027FB4AF937FA2C9E37" box="[1273,1436,1785,1809]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" metricValueMax="2.88" metricValueMin="2.84" pageId="7" pageNumber="447" unit="mm" value="28.6" valueMax="28.8" valueMin="28.4">28.4–28.8 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C333EACFF81A027FF05F8D0FE939E13" box="[182,291,1821,1845]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="hamidi">
|
||
<emphasis id="B947993DFF81A027FF05F8D0FE939E13" box="[182,291,1821,1845]" italics="true" pageId="7" pageNumber="447">L. hamidi</emphasis>
|
||
</taxonomicName>
|
||
) and small dorsal markings without light outlines (vs. large dark brown dorsal markings with light outlines in
|
||
<taxonomicName id="4C333EACFF81A027FEA7F88CFE309E7F" box="[276,384,1857,1881]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="hamidi">
|
||
<emphasis id="B947993DFF81A027FEA7F88CFE309E7F" box="[276,384,1857,1881]" italics="true" pageId="7" pageNumber="447">L. hamidi</emphasis>
|
||
</taxonomicName>
|
||
). From
|
||
<taxonomicName id="4C333EACFF81A027FE68F88CFDF29E7F" box="[475,578,1858,1881]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF81A027FE68F88CFDF29E7F" box="[475,578,1858,1881]" italics="true" pageId="7" pageNumber="447">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF81A027FDE2F88CFCAF9E7F" box="[593,799,1857,1881]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="sabahmontanus">
|
||
<emphasis id="B947993DFF81A027FDE2F88CFCAF9E7F" box="[593,799,1857,1881]" italics="true" pageId="7" pageNumber="447">L. sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
clearly differs by having a smaller body size (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FF24F8ABFE8A9E5B" box="[151,314,1893,1918]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
vs. 29.2–32.0 mm in
|
||
<taxonomicName id="4C333EACFF81A027FD9DF8A8FD259E5B" box="[558,661,1894,1917]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF81A027FD9DF8A8FD259E5B" box="[558,661,1894,1917]" italics="true" pageId="7" pageNumber="447">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
), not strongly shagreened back, presence of dorsal markings, usually white venter, and absence of flush colour on ventral side of legs and groin (vs. dorsum tuberculated, without conspicuous markings, venter yellow, and groin and ventral sides of limbs orange in
|
||
<taxonomicName id="4C333EACFF81A027FBD0F860FB7C9EE3" box="[1123,1228,1966,1989]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF81A027FBD0F860FB7C9EE3" box="[1123,1228,1966,1989]" italics="true" pageId="7" pageNumber="447">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
). Finally, the new species is similar to
|
||
<taxonomicName id="4C333EACFF81A027FECCF81CFE419ECF" box="[383,497,2002,2025]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="B947993DFF81A027FECCF81CFE419ECF" box="[383,497,2002,2025]" italics="true" pageId="7" pageNumber="447">L. maurus</emphasis>
|
||
</taxonomicName>
|
||
in body size (male SVL
|
||
<quantity id="4CCBE8CAFF81A027FCA2F81FFC069ECF" box="[785,950,2001,2026]" metricMagnitude="-2" metricUnit="m" metricValue="2.665" metricValueMax="2.79" metricValueMin="2.54" pageId="7" pageNumber="447" unit="mm" value="26.65" valueMax="27.9" valueMin="25.4">25.4–27.9 mm</quantity>
|
||
vs. 25.0–
|
||
<quantity id="4CCBE8CAFF81A027FB96F81FFB3A9ECF" box="[1061,1162,2001,2026]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="7" pageNumber="447" unit="mm" value="26.9">26.9 mm</quantity>
|
||
SVL in
|
||
<taxonomicName id="4C333EACFF81A027FB5EF81CFAD09ECF" box="[1261,1376,2002,2025]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="7" pageNumber="447" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="B947993DFF81A027FB5EF81CFAD09ECF" box="[1261,1376,2002,2025]" italics="true" pageId="7" pageNumber="447">L. maurus</emphasis>
|
||
</taxonomicName>
|
||
), but differs from it by having an olive brown dorsum with black pattern, and maculated light-colored ventrum (vs. having a dark brown to black dorsum and ventrum in
|
||
<taxonomicName id="4C333EACFF8EA028FD42FF73FCD299F2" box="[753,866,189,212]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="448" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="B947993DFF8EA028FD42FF73FCD299F2" box="[753,866,189,212]" italics="true" pageId="8" pageNumber="448">L. maurus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF4C15A7FF8EA028FF24F8DCFD869E43" httpUri="https://zenodo.org/record/228779/files/figure.png" pageId="8" pageNumber="448" targetBox="[221,1361,269,1782]" targetPageId="8">
|
||
<paragraph id="8B8C452FFF8EA028FF24F8DCFD869E43" blockId="8.[151,1436,1810,1893]" pageId="8" pageNumber="448">
|
||
<emphasis id="B947993DFF8EA028FF24F8DCFEA99E0E" bold="true" box="[151,281,1810,1832]" pageId="8" pageNumber="448">FIGURE 5.</emphasis>
|
||
Sonagram (A, C, E) and wave form (B, D, F) showing advertisement calls of a paratype of
|
||
<taxonomicName id="4C333EACFF8EA028FA83F8DCFE869E61" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="448" phylum="Chordata" rank="species" species="sabahmontanus" status="sp. nov.">
|
||
<emphasis id="B947993DFF8EA028FA83F8DCFE869E61" italics="true" pageId="8" pageNumber="448">Leptolalax sabahmontanus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2742446FF8EA028FE8FF8FFFE3D9E60" box="[316,397,1841,1862]" pageId="8" pageNumber="448" rank="species">
|
||
<emphasis id="B947993DFF8EA028FE8FF8FFFE3D9E60" bold="true" box="[316,397,1841,1862]" pageId="8" pageNumber="448">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(KUHE 48016) from the type locality, Silau Silau trail along Sg. Silau-Silau, Kinabalu National Park, recorded at an air temperature of 17.2°C.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8C452FFF8EA028FF74F85CFA2C9ED3" blockId="8.[151,1436,1938,2038]" pageId="8" pageNumber="448">
|
||
The new species also differs from the known Bornean members acoustically. By lacking clear frequency modulation within a note, it differs from
|
||
<taxonomicName id="4C333EACFF8EA028FDEFF879FD5C9EE9" box="[604,748,1975,1999]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="448" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF8EA028FDEFF879FD5C9EE9" box="[604,748,1975,1999]" italics="true" pageId="8" pageNumber="448">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8EA028FD49F879FCD49EE8" box="[762,868,1974,1998]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="448" phylum="Chordata" rank="species" species="hamidi">
|
||
<emphasis id="B947993DFF8EA028FD49F879FCD49EE8" box="[762,868,1974,1998]" italics="true" pageId="8" pageNumber="448">L. hamidi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C333EACFF8EA028FC13F879FBB09EE8" box="[928,1024,1975,1998]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="8" pageNumber="448" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF8EA028FC13F879FBB09EE8" box="[928,1024,1975,1998]" italics="true" pageId="8" pageNumber="448">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
, although the dominant frequency of the call of the new species (6.90–7.35 kHz at 17.2–17.7o C) overlaps (7.25–9.20 kHz at 24.3–24.9°C in
|
||
<emphasis id="B947993DFF8EA028FA36F810FA2C9ED3" box="[1413,1436,2014,2037]" italics="true" pageId="8" pageNumber="448">L.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF8FA029FF24FF56FF4D9866" blockId="9.[151,1436,151,1509]" pageId="9" pageNumber="449">
|
||
<taxonomicName id="4C333EACFF8FA029FF24FF56FEBA9996" box="[151,266,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF8FA029FF24FF56FEBA9996" box="[151,266,152,176]" italics="true" pageId="9" pageNumber="449">fritinniens</emphasis>
|
||
</taxonomicName>
|
||
, 6.70–7.30 kHz at 22.9–24.1°C in
|
||
<taxonomicName id="4C333EACFF8FA029FD16FF57FCA49996" box="[677,788,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="hamidi">
|
||
<emphasis id="B947993DFF8FA029FD16FF57FCA49996" box="[677,788,152,176]" italics="true" pageId="9" pageNumber="449">L. hamidi</emphasis>
|
||
</taxonomicName>
|
||
, and 6.80–7.15 kHz at 19–22°C in
|
||
<taxonomicName id="4C333EACFF8FA029FB0AFF57FAAC9989" box="[1209,1308,152,176]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF8FA029FB0AFF57FAAC9989" box="[1209,1308,152,176]" italics="true" pageId="9" pageNumber="449">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
). The new species is similar to other species without conspicuous frequency modulation, but the dominant frequency (6.90– 7.35 kHz) of the new species is much higher (6.05–6.40 kHz at unknown temperature in
|
||
<taxonomicName id="4C333EACFF8FA029FB37FF2FFB5699DE" box="[1156,1254,224,248]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF8FA029FB37FF2FFB5699DE" box="[1156,1254,224,248]" italics="true" pageId="9" pageNumber="449">L. dringi</emphasis>
|
||
</taxonomicName>
|
||
; 5.40–5.90 kHz at 17.4°C in
|
||
<taxonomicName id="4C333EACFF8FA029FE81FECBFE2D983A" box="[306,413,261,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF8FA029FE81FECBFE2D983A" box="[306,413,261,284]" italics="true" pageId="9" pageNumber="449">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
; 5.15 kHz at unknown temperature in
|
||
<taxonomicName id="4C333EACFF8FA029FCDFFECBFC52983A" box="[876,994,261,284]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="maurus">
|
||
<emphasis id="B947993DFF8FA029FCDFFECBFC52983A" box="[876,994,261,284]" italics="true" pageId="9" pageNumber="449">L. maurus</emphasis>
|
||
</taxonomicName>
|
||
; 2.60–2.80 kHz at 20.0–26.2°C in
|
||
<taxonomicName id="4C333EACFF8FA029FA36FECBFF5C9866" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="gracilis">
|
||
<emphasis id="B947993DFF8FA029FA36FECBFF5C9866" italics="true" pageId="9" pageNumber="449">L. gracilis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF8FA029FF74FE82FCC598D3" blockId="9.[151,1436,151,1509]" pageId="9" pageNumber="449">
|
||
<emphasis id="B947993DFF8FA029FF74FE82FEAA9843" bold="true" box="[199,282,332,357]" pageId="9" pageNumber="449">Range.</emphasis>
|
||
The species appears to be restricted to Sabah, and has been found around the Headquarters (HQ) of KNP (Silau Silau trail, Pandanus trail, Bundu Tuhan view trail, and along Sg. Liwagu), Ranau District, and Sg. Wario, Sayap, Kota Belud District. It also occurs in CRNP, Tambunan District (Trails 4, 5, and 11 of UMS 2002 expedition to the Crocker Range, Ulu Kimanis, and along Sg. Mahua, Mahua). The known localities are all in mountain regions, ranging in altitude from
|
||
<quantity id="4CCBE8CAFF8FA029FDC9FE12FD7198D2" box="[634,705,476,501]" metricMagnitude="2" metricUnit="m" metricValue="7.54" pageId="9" pageNumber="449" unit="m" value="754.0">754 m</quantity>
|
||
to
|
||
<quantity id="4CCBE8CAFF8FA029FD57FE12FC8998D2" box="[740,825,476,501]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="449" unit="m" value="1500.0">1500 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF8FA029FF74FE31FF599BC9" blockId="9.[151,1436,151,1509]" pageId="9" pageNumber="449">
|
||
<emphasis id="B947993DFF8FA029FF74FE31FE399B3E" bold="true" box="[199,393,511,536]" pageId="9" pageNumber="449">Natural history.</emphasis>
|
||
Specimens for which data are available were caught in primary rain forest at night 2–50 (median = 5) m away from banks of streams with width of 1–12 (median = 5.5) m. Of the 29 frogs recorded, 20 were found on leaves of short grasses, two on stones at stream banks, two on the trail, three among litters on the forest floor, and two hidden under rotten log or among root of grasses. The height of the grass at which frogs perched is 10–100 (median = 30) cm from above the ground. Reproductive details are unknown, but males were actively calling from late July to mid-August and in mid-December, and these periods must be within the breeding season.
|
||
</paragraph>
|
||
<paragraph id="8B8C452FFF8FA029FF74FD32FF459CC3" blockId="9.[151,1436,151,1509]" pageId="9" pageNumber="449">
|
||
The new species is found sympatric with
|
||
<taxonomicName id="4C333EACFF8FA029FD25FD33FD4D9A32" box="[662,765,765,788]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="arayai">
|
||
<emphasis id="B947993DFF8FA029FD25FD33FD4D9A32" box="[662,765,765,788]" italics="true" pageId="9" pageNumber="449">L. arayai</emphasis>
|
||
</taxonomicName>
|
||
around Headquarters of Kinabalu National Park (KNPHQ),
|
||
<taxonomicName id="4C333EACFF8FA029FF24FCEFFF4A9A11" box="[151,250,800,824]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF8FA029FF24FCEFFF4A9A11" box="[151,250,800,824]" italics="true" pageId="9" pageNumber="449">L. pictus</emphasis>
|
||
</taxonomicName>
|
||
at Sayap, KNP, and Mahua, CRNP, and
|
||
<taxonomicName id="4C333EACFF8FA029FD67FCEFFCD89A1E" box="[724,872,800,824]" class="Amphibia" family="Megophryidae" genus="Leptolalax" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="fritinniens">
|
||
<emphasis id="B947993DFF8FA029FD67FCEFFCD89A1E" box="[724,872,800,824]" italics="true" pageId="9" pageNumber="449">L. fritinniens</emphasis>
|
||
</taxonomicName>
|
||
at Trails of Ulu Kimanis, CRNP. The associated anuran species observed were:
|
||
<taxonomicName id="4C333EACFF8FA029FDB4FC8AFC0B9A7B" authority="Fischer" authorityName="Fischer" box="[519,955,836,861]" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="montanum">
|
||
<emphasis id="B947993DFF8FA029FDB4FC8AFD029A7A" box="[519,690,836,860]" italics="true" pageId="9" pageNumber="449">Leptobrachium</emphasis>
|
||
cf.
|
||
<emphasis id="B947993DFF8FA029FD55FC8BFCEC9A7A" box="[742,860,837,860]" italics="true" pageId="9" pageNumber="449">montanum</emphasis>
|
||
Fischer
|
||
</taxonomicName>
|
||
(at KNPHQ),
|
||
<taxonomicName id="4C333EACFF8FA029FBDEFC8AFEB99AA6" authority="Cochran" authorityName="Cochran" class="Amphibia" family="Megophryidae" genus="Leptobrachium" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="abbotti">
|
||
<emphasis id="B947993DFF8FA029FBDEFC8AFAA89A7A" box="[1133,1304,836,860]" italics="true" pageId="9" pageNumber="449">Leptobrachium</emphasis>
|
||
cf.
|
||
<emphasis id="B947993DFF8FA029FAFFFC8AFA2B9A7A" box="[1356,1435,836,860]" italics="true" pageId="9" pageNumber="449">abbotti</emphasis>
|
||
(Cochran)
|
||
</taxonomicName>
|
||
(at Ulu Kimanis),
|
||
<taxonomicName id="4C333EACFF8FA029FE6EFCA6FC429AA6" authority="Smith (Ulu Kimanis)" authorityName="Smith (Ulu Kimanis" box="[477,1010,871,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="B947993DFF8FA029FE6EFCA6FD4A9AA6" box="[477,762,872,896]" italics="true" pageId="9" pageNumber="449">Leptobrachella baluensis</emphasis>
|
||
Smith (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FC4DFCA6FE319A83" authority="(Inger, Stuebing & Tan) (Ulu Kimanis)" authorityName="(Inger, Stuebing & Tan) (Ulu Kimanis" class="Amphibia" family="Megophryidae" genus="Xenophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="dringi">
|
||
<emphasis id="B947993DFF8FA029FC4DFCA6FB719AA6" box="[1022,1217,872,896]" italics="true" pageId="9" pageNumber="449">Xenophrys dringi</emphasis>
|
||
(Inger, Stuebing & Tan) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FE21FC42FCE79A83" authority="(Schlegel) (Mahua)" authorityName="(Schlegel) (Mahua" box="[402,855,908,933]" class="Amphibia" family="Megophryidae" genus="Megophrys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="nasuta">
|
||
<emphasis id="B947993DFF8FA029FE21FC42FDD89A82" box="[402,616,908,932]" italics="true" pageId="9" pageNumber="449">Megophrys nasuta</emphasis>
|
||
(Schlegel) (Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FCDBFC42FA289A83" authority="(Tshudi) (Ulu Kimanis)" authorityName="(Tshudi) (Ulu Kimanis" box="[872,1432,908,933]" class="Amphibia" family="Bufonidae" genus="Leptophryne" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="borbonica">
|
||
<emphasis id="B947993DFF8FA029FCDBFC42FBC59A82" box="[872,1141,908,932]" italics="true" pageId="9" pageNumber="449">Leptophryne borbonica</emphasis>
|
||
(Tshudi) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FF24FC7EFE129AEE" authority="Inger" authorityName="Inger" box="[151,418,944,968]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="hanitschi">
|
||
<emphasis id="B947993DFF8FA029FF24FC7EFED09AEE" box="[151,352,944,968]" italics="true" pageId="9" pageNumber="449">Ansonia hanitschi</emphasis>
|
||
Inger
|
||
</taxonomicName>
|
||
(KNPHQ),
|
||
<taxonomicName id="4C333EACFF8FA029FD99FC7EFBE99AEE" authority="Inger (Sayap, Ulu Kimanis, Mahua)" authorityName="Inger (Sayap, Ulu Kimanis, Mahua" box="[554,1113,943,968]" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="platysoma">
|
||
<emphasis id="B947993DFF8FA029FD99FC7EFD0C9AEE" box="[554,700,944,968]" italics="true" pageId="9" pageNumber="449">A. platysoma</emphasis>
|
||
Inger (Sayap, Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FBD7FC7EFE3E9ACB" authority="Inger (Sayap, Ulu Kimanis, Mahua)" authorityName="Inger (Sayap, Ulu Kimanis, Mahua" class="Amphibia" family="Bufonidae" genus="Ansonia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="longidigita">
|
||
<emphasis id="B947993DFF8FA029FBD7FC7EFB4E9AEE" box="[1124,1278,944,968]" italics="true" pageId="9" pageNumber="449">A. longidigita</emphasis>
|
||
Inger (Sayap, Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FE2AFC1AFCC59ACB" authority="(Peters) (Ulu Kimanis)" authorityName="(Peters) (Ulu Kimanis" box="[409,885,980,1005]" class="Amphibia" family="Ranidae" genus="Hylarana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="raniceps">
|
||
<emphasis id="B947993DFF8FA029FE2AFC1AFDDB9ACA" box="[409,619,980,1004]" italics="true" pageId="9" pageNumber="449">Hylarana raniceps</emphasis>
|
||
(Peters) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FC33FC1AFACC9ACB" authority="Inger & Haile (Ulu Kimanis)" authorityName="Inger & Haile (Ulu Kimanis" box="[896,1404,980,1005]" class="Amphibia" family="Ranidae" genus="Staurois" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="parvus">
|
||
<emphasis id="B947993DFF8FA029FC33FC1AFB9E9ACA" box="[896,1070,980,1004]" italics="true" pageId="9" pageNumber="449">Staurois parvus</emphasis>
|
||
Inger & Haile (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FA34FC1AFDC49D36" authority="(Gunther) (Ulu Kimanis, Mahua)" authorityName="(Gunther) (Ulu Kimanis, Mahua" class="Amphibia" family="Ranidae" genus="Staurois" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="guttatus">
|
||
<emphasis id="B947993DFF8FA029FA34FC1AFF419D36" italics="true" pageId="9" pageNumber="449">S. guttatus</emphasis>
|
||
(Günther) (Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FD33FC36FBDE9D36" authority="(Boulenger) (Ulu Kimanis)" authorityName="(Boulenger) (Ulu Kimanis" box="[640,1134,1015,1040]" class="Amphibia" family="Ranidae" genus="Staurois" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="latopalmatus">
|
||
<emphasis id="B947993DFF8FA029FD33FC36FC9F9D36" box="[640,815,1016,1040]" italics="true" pageId="9" pageNumber="449">S. latopalmatus</emphasis>
|
||
(Boulenger) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FBC9FC36FA2B9D36" authority="Boulenger" authorityName="Boulenger" box="[1146,1435,1015,1040]" class="Amphibia" family="Ranidae" genus="Staurois" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="tuberilingus">
|
||
<emphasis id="B947993DFF8FA029FBC9FC36FAAF9D36" box="[1146,1311,1016,1040]" italics="true" pageId="9" pageNumber="449">S. tuberilingus</emphasis>
|
||
Boulenger
|
||
</taxonomicName>
|
||
(KNPHQ, Sayap, Ulu Kimanis),
|
||
<taxonomicName id="4C333EACFF8FA029FD9DFBD3FAD89D12" authority="Shimada, Matsui, Yambun & Sudin (Sayap)" authorityName="Shimada, Matsui, Yambun & Sudin (Sayap" box="[558,1384,1052,1077]" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="dyscritus">
|
||
<emphasis id="B947993DFF8FA029FD9DFBD3FC8A9D12" box="[558,826,1052,1076]" italics="true" pageId="9" pageNumber="449">Meristogenys dyscritus</emphasis>
|
||
Shimada, Matsui, Yambun & Sudin (Sayap)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FACEFBD3FD069D7E" authority="(Matsui) (Ulu Kimanis, Mahua)" authorityName="(Matsui) (Ulu Kimanis, Mahua" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="orphnocnemis">
|
||
<emphasis id="B947993DFF8FA029FACEFBD3FE869D7E" italics="true" pageId="9" pageNumber="449">M. orphnocnemis</emphasis>
|
||
(Matsui) (Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FD76FB8FFBD19D7E" authority="(Boulenger) (Sayap)" authorityName="(Boulenger) (Sayap" box="[709,1121,1087,1112]" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="whiteheadi">
|
||
<emphasis id="B947993DFF8FA029FD76FB8FFCDA9D7E" box="[709,874,1088,1112]" italics="true" pageId="9" pageNumber="449">M. whiteheadi</emphasis>
|
||
(Boulenger) (Sayap)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FBC2FB8FFDAD9D5B" authority="Shimada, Matsui, Yambun & Sudin (Mahua)" authorityName="Shimada, Matsui, Yambun & Sudin (Mahua" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="stigmachilus">
|
||
<emphasis id="B947993DFF8FA029FBC2FB8FFB389D7E" box="[1137,1160,1089,1112]" italics="true" pageId="9" pageNumber="449">M</emphasis>
|
||
.
|
||
<emphasis id="B947993DFF8FA029FB29FB8EFA979D7E" box="[1178,1319,1088,1112]" italics="true" pageId="9" pageNumber="449">stigmachilus</emphasis>
|
||
Shimada, Matsui, Yambun & Sudin (Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FD9BFBABFC8B9D5A" authority="Inger" authorityName="Inger" box="[552,827,1124,1148]" class="Amphibia" family="Ranidae" genus="Meristogenys" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="kinabaluensis">
|
||
<emphasis id="B947993DFF8FA029FD9BFBABFD579D5A" box="[552,743,1124,1148]" italics="true" pageId="9" pageNumber="449">M. kinabaluensis</emphasis>
|
||
(Inger)
|
||
</taxonomicName>
|
||
(KNPHQ, Ulu Kimanis, Mahua),
|
||
<taxonomicName id="4C333EACFF8FA029FB09FBABFE349D86" authority="(Boulenger) (Sayap)" authorityName="(Boulenger) (Sayap" class="Amphibia" family="Ranidae" genus="Huia" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="cavitympanum">
|
||
<emphasis id="B947993DFF8FA029FB09FBABFA2B9D5A" box="[1210,1435,1125,1148]" italics="true" pageId="9" pageNumber="449">Huia cavitympanum</emphasis>
|
||
(Boulenger) (Sayap)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FE26FB46FC6B9D86" authority="(Boulenger) (Ulu Kimanis)" authorityName="(Boulenger) (Ulu Kimanis" box="[405,987,1159,1184]" class="Amphibia" family="Dicroglossidae" genus="Occidozyga" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="B947993DFF8FA029FE26FB46FD209D86" box="[405,656,1160,1184]" italics="true" pageId="9" pageNumber="449">Occidozyga baluensis</emphasis>
|
||
(Boulenger) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FC58FB47FEB29DE3" authority="(Boulenger) (Ulu Kimanis)" authorityName="(Boulenger) (Ulu Kimanis" class="Amphibia" family="Dicroglossidae" genus="Ingerana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="B947993DFF8FA029FC58FB47FB789D86" box="[1003,1224,1160,1184]" italics="true" pageId="9" pageNumber="449">Ingerana baluensis</emphasis>
|
||
(Boulenger) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FEA2FB63FC349DE3" authority="(Boulenger) (Ulu Kimanis)" authorityName="(Boulenger) (Ulu Kimanis" box="[273,900,1196,1221]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="palavanensis">
|
||
<emphasis id="B947993DFF8FA029FEA2FB63FD8C9DE2" box="[273,572,1196,1220]" italics="true" pageId="9" pageNumber="449">Limnonectes palavanensis</emphasis>
|
||
(Boulenger) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FC27FB63FB239DE3" authority="Tschudi" authorityName="Tschudi" box="[916,1171,1196,1221]" class="Amphibia" family="Dicroglossidae" genus="Limnonectes" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="kuhlii">
|
||
<emphasis id="B947993DFF8FA029FC27FB63FC1B9DE2" box="[916,939,1197,1220]" italics="true" pageId="9" pageNumber="449">L.</emphasis>
|
||
cf.
|
||
<emphasis id="B947993DFF8FA029FC6EFB62FBAD9DE2" box="[989,1053,1196,1220]" italics="true" pageId="9" pageNumber="449">kuhlii</emphasis>
|
||
(Tschudi)
|
||
</taxonomicName>
|
||
(KNPHQ, Sayap, Ulu Kimanis),
|
||
<taxonomicName id="4C333EACFF8FA029FEA4FB1EFCFD9DCE" authority="(Gravenhorst) (Mahua)" authorityName="(Gravenhorst) (Mahua" box="[279,845,1231,1256]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="limnocharis">
|
||
<emphasis id="B947993DFF8FA029FEA4FB1EFD9A9DCE" box="[279,554,1232,1256]" italics="true" pageId="9" pageNumber="449">Fejervarya limnocharis</emphasis>
|
||
(Gravenhorst) (Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FCD1FB1EFB249DCE" authority="Peters" authorityName="Peters" box="[866,1172,1232,1256]" class="Amphibia" family="Rhacophoridae" genus="Nyctixalus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="pictus">
|
||
<emphasis id="B947993DFF8FA029FCD1FB1EFB9C9DC1" box="[866,1068,1232,1256]" italics="true" pageId="9" pageNumber="449">Nyctixalus pictus</emphasis>
|
||
(Peters)
|
||
</taxonomicName>
|
||
(KNPHQ),
|
||
<taxonomicName id="4C333EACFF8FA029FA82FB1EFEFB9C2B" authority="Smith" authorityName="Smith" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="mjobergi">
|
||
<emphasis id="B947993DFF8FA029FA82FB1EFF4B9C2A" italics="true" pageId="9" pageNumber="449">Philautus mjobergi</emphasis>
|
||
Smith
|
||
</taxonomicName>
|
||
(KNPHQ, Sayap, Ulu Kimanis),
|
||
<taxonomicName id="4C333EACFF8FA029FD53FB3AFAD59C2B" authority="(Boulenger) (Sayap, Ulu Kimanis, Mahua)" authorityName="(Boulenger) (Sayap, Ulu Kimanis, Mahua" box="[736,1381,1268,1293]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="petersi">
|
||
<emphasis id="B947993DFF8FA029FD53FB3AFCEF9C2A" box="[736,863,1268,1292]" italics="true" pageId="9" pageNumber="449">Ph. petersi</emphasis>
|
||
(Boulenger) (Sayap, Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FAC5FB3AFD0B9C16" authority="Inger (Sayap, Ulu Kimanis, Mahua)" authorityName="Inger (Sayap, Ulu Kimanis, Mahua" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="aurantium">
|
||
<emphasis id="B947993DFF8FA029FAC5FB3AFEBC9C09" italics="true" pageId="9" pageNumber="449">Ph. aurantium</emphasis>
|
||
Inger (Sayap, Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FD79FAD6FAD79C16" authority="Inger, Stuebing & Tan (Sayap, Ulu Kimanis)" authorityName="Inger, Stuebing & Tan (Sayap, Ulu Kimanis" box="[714,1383,1303,1328]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="bunitus">
|
||
<emphasis id="B947993DFF8FA029FD79FAD6FCFE9C16" box="[714,846,1304,1328]" italics="true" pageId="9" pageNumber="449">Ph. bunitus</emphasis>
|
||
Inger, Stuebing & Tan (Sayap, Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FAC5FAD6FCB29C73" authority="Stuebing & Wong (Ulu Kimanis)" authorityName="Stuebing & Wong (Ulu Kimanis" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="erythrophthalmus">
|
||
<emphasis id="B947993DFF8FA029FAC5FAD6FEED9C72" italics="true" pageId="9" pageNumber="449">Ph. erythrophthalmus</emphasis>
|
||
Stuebing & Wong (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FCA7FAF2FBD59C73" authority="Boulenger" authorityName="Boulenger" box="[788,1125,1340,1365]" class="Amphibia" family="Rhacophoridae" genus="Philautus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="macroscelis">
|
||
<emphasis id="B947993DFF8FA029FCA7FAF2FC7E9C72" box="[788,974,1340,1364]" italics="true" pageId="9" pageNumber="449">Ph. macroscelis</emphasis>
|
||
(Boulenger)
|
||
</taxonomicName>
|
||
(KNPHQ, Ulu Kimanis),
|
||
<taxonomicName id="4C333EACFF8FA029FF24FAAEFE429C5E" authority="Ahl" authorityName="Ahl" box="[151,498,1375,1400]" class="Diplopoda" family="Chelodesmidae" genus="Rhacophorus" kingdom="Animalia" order="Polydesmida" pageId="9" pageNumber="449" phylum="Arthropoda" rank="species" species="angulirostris">
|
||
<emphasis id="B947993DFF8FA029FF24FAAEFE739C5E" box="[151,451,1376,1400]" italics="true" pageId="9" pageNumber="449">Rhacophorus angulirostris</emphasis>
|
||
Ahl
|
||
</taxonomicName>
|
||
(KNPHQ, Ulu Kimanis, Mahua),
|
||
<taxonomicName id="4C333EACFF8FA029FCC2FAAEFA289C5E" authority="(Gravenhorst) (Mahua)" authorityName="(Gravenhorst) (Mahua" box="[881,1432,1375,1400]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="leucomystax">
|
||
<emphasis id="B947993DFF8FA029FCC2FAAEFB3B9C5E" box="[881,1163,1376,1400]" italics="true" pageId="9" pageNumber="449">Polypedates leucomystax</emphasis>
|
||
(Gravenhorst) (Mahua)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FF24FA4BFDEC9CBB" authority="(Boulenger) (Ulu Kimanis)" authorityName="(Boulenger) (Ulu Kimanis" box="[151,604,1412,1437]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="otilophus">
|
||
<emphasis id="B947993DFF8FA029FF24FA4BFEAC9CBA" box="[151,284,1412,1436]" italics="true" pageId="9" pageNumber="449">P. otilophus</emphasis>
|
||
(Boulenger) (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FDD9FA4AFB0D9CBB" authority="Boulenger (Ulu Kimanis)" authorityName="Boulenger (Ulu Kimanis" box="[618,1213,1412,1437]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="heterochirus">
|
||
<emphasis id="B947993DFF8FA029FDD9FA4AFC209CBA" box="[618,912,1412,1436]" italics="true" pageId="9" pageNumber="449">Kalophrynus heterochirus</emphasis>
|
||
Boulenger (Ulu Kimanis)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C333EACFF8FA029FB79FA4BFA2C9CBB" authority="Kiew" authorityName="Kiew" box="[1226,1436,1412,1437]" class="Amphibia" family="Dicroglossidae" genus="Ingerana" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="baluensis">
|
||
<emphasis id="B947993DFF8FA029FB79FA4BFAE69CBA" box="[1226,1366,1412,1436]" italics="true" pageId="9" pageNumber="449">K. baluensis</emphasis>
|
||
Kiew
|
||
</taxonomicName>
|
||
(KNPHQ),
|
||
<taxonomicName id="4C333EACFF8FA029FEADFA67FCB89CE6" authority="Tschudi (Crocker Trails)" authorityName="Tschudi (Crocker Trails" box="[286,776,1447,1472]" class="Amphibia" family="Microhylidae" genus="Kalophrynus" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="pleurostigma">
|
||
<emphasis id="B947993DFF8FA029FEADFA67FE689CE6" box="[286,472,1448,1472]" italics="true" pageId="9" pageNumber="449">K. pleurostigma</emphasis>
|
||
Tschudi (Crocker Trails)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C333EACFF8FA029FCFCFA66FF429CC3" authority="Mocquard (Sayap, Ulu Kimanis, Mahua)" authorityName="Mocquard (Sayap, Ulu Kimanis, Mahua" class="Amphibia" family="Microhylidae" genus="Chaperina" kingdom="Animalia" order="Anura" pageId="9" pageNumber="449" phylum="Chordata" rank="species" species="fusca">
|
||
<emphasis id="B947993DFF8FA029FCFCFA66FBBF9CE6" box="[847,1039,1448,1472]" italics="true" pageId="9" pageNumber="449">Chaperina fusca</emphasis>
|
||
Mocquard (Sayap, Ulu Kimanis, Mahua)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |