257 lines
36 KiB
XML
257 lines
36 KiB
XML
<document id="D4AF41F06A67F88939AEC640DF9AA181" ID-DOI="10.5281/zenodo.2619524" ID-GBIF-Dataset="a9fc01a1-cd41-459f-b9d5-addaf795dce4" ID-ISSN="0035-418" ID-Zenodo-Dep="2619524" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583846249953" checkinUser="carolina" docAuthor="Martens, Jochen" docDate="2019" docId="039AE028BA19FFBCB7CAF9C7FAB0FD0A" docLanguage="en" docName="RevSuisseZool.126.1.79-110.pdf.imf" docOrigin="Revue suisse de Zoologie 126 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Asiolasma Martens 2019, gen. nov." docType="treatment" docVersion="6" lastPageNumber="86" masterDocId="FFA39850BA1DFFBBB412FFE6F935FF93" masterDocTitle="An ancient radiation: Ortholasmatine harvestmen in Asia - a new genus, three new species and a revision of the known species (Arachnida, Opiliones, Nemastomatidae)" masterLastPageNumber="110" masterPageNumber="79" pageNumber="83" updateTime="1698726360747" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9D786AD3D585F3FD9BBE0E92CA5A8EBB">An ancient radiation: Ortholasmatine harvestmen in Asia - a new genus, three new species and a revision of the known species (Arachnida, Opiliones, Nemastomatidae)</mods:title>
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<mods:name id="1DE39382DF029A34D211C0C74666D766" type="personal">
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<mods:namePart id="914E7EC7850795E3C9C344F75107A6E7">Martens, Jochen</mods:namePart>
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<mods:typeOfResource id="DE548272A16261F93AF1E637FB726411">text</mods:typeOfResource>
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<mods:title id="DBF0C4DF1EB56056F4A524341C76FCAE">Revue suisse de Zoologie</mods:title>
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<mods:part id="0CECEE375253A88096136386AFD58532">
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<mods:date id="1F51CBE9E5A98B76E8A45BD8FF9430AA">2019</mods:date>
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<mods:number id="93026CE2C67ACEEFEEB4C3EE53BCFED1">2019-03-31</mods:number>
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<mods:number id="3B3B888ACBC747FDE5970ACACA733911">126</mods:number>
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<mods:start id="1C43B04BB2060D48C7274D37E04AC56B">79</mods:start>
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<mods:identifier id="1F0B84F8B4CB688307B38D7CB1F0D040" type="DOI">10.5281/zenodo.2619524</mods:identifier>
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<mods:identifier id="1E1FA36971FD35C9A16FC16DEA4B7240" type="ISSN">0035-418</mods:identifier>
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<mods:identifier id="E10801D01E0D43812001C3780765B3AC" type="Zenodo-Dep">2619524</mods:identifier>
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<treatment id="039AE028BA19FFBCB7CAF9C7FAB0FD0A" ID-DOI="http://doi.org/10.5281/zenodo.3706053" ID-GBIF-Taxon="162481146" ID-Zenodo-Dep="3706053" LSID="urn:lsid:plazi:treatment:039AE028BA19FFBCB7CAF9C7FAB0FD0A" httpUri="http://treatment.plazi.org/id/039AE028BA19FFBCB7CAF9C7FAB0FD0A" lastPageId="7" lastPageNumber="86" pageId="4" pageNumber="83">
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<subSubSection id="C32902B5BA19FFBFB7CAF9C7FC31F9AB" box="[984,1284,1568,1592]" pageId="4" pageNumber="83" type="nomenclature">
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<paragraph id="8B8C513EBA19FFBFB7CAF9C7FC31F9AB" blockId="4.[984,1284,1568,1592]" box="[984,1284,1568,1592]" pageId="4" pageNumber="83">
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<heading id="D0C4E652BA19FFBFB7CAF9C7FC31F9AB" bold="true" box="[984,1284,1568,1592]" centered="true" level="2" pageId="4" pageNumber="83" reason="2">
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<emphasis id="B9478D2CBA19FFBFB7CAF9C7FC31F9AB" bold="true" box="[984,1284,1568,1592]" pageId="4" pageNumber="83">
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Genus
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<taxonomicName id="4C332ABDBA19FFBFB035F9C6FDACF9AB" ID-CoL="7NRDM" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[1063,1177,1568,1592]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="83" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9478D2CBA19FFBFB035F9C6FDACF9AB" bold="true" box="[1063,1177,1568,1592]" italics="true" pageId="4" pageNumber="83">Asiolasma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2743057BA19FFBFB0B2F9C7FC31F9AB" box="[1184,1284,1569,1592]" pageId="4" pageNumber="83" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32902B5BA19FFBFB752F9B7FAE1F91B" pageId="4" pageNumber="83" type="type_taxon">
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<paragraph id="8B8C513EBA19FFBFB752F9B7FAE1F91B" blockId="4.[832,1436,1617,1672]" pageId="4" pageNumber="83">
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<emphasis id="B9478D2CBA19FFBFB752F9B7FAEAF9FB" bold="true" box="[832,991,1617,1640]" pageId="4" pageNumber="83">
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<typeStatus id="5488EF9CBA19FFBFB752F9B7FA4DF9FB" box="[832,888,1617,1640]" pageId="4" pageNumber="83">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C332ABDBA19FFBFB7FCF9B7FAE5F91B" ID-CoL="7VPKN" authority="Schwendinger & Gruber, 1992" authorityName="Schwendinger & Gruber" authorityYear="1992" class="Arachnida" family="Nemastomatidae" genus="Dendrolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="83" phylum="Arthropoda" rank="species" species="angka">
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<emphasis id="B9478D2CBA19FFBFB7FCF9B7FDE5F9FB" box="[1006,1232,1617,1640]" italics="true" pageId="4" pageNumber="83">Dendrolasma angka</emphasis>
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<bibRefCitation id="EFA22CCFBA19FFBFB0CDF9B7FAE5F91B" author="Schwendinger & Gruber" firstAuthor="Schwendinger" pageId="4" pageNumber="83" pagination="57 - 60" refId="ref18545" refString="Schwendinger P. J., Gruber J. 1992. A new Dendrolasma (Opiliones, Nemastomatidae) from Thailand. Bulletin of the British Arachnological Society 9: 57 - 60." type="journal article" year="1992">Schwendinger & Gruber, 1992</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32902B5BA19FFBEB752F947F9D9F981" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83" type="diagnosis">
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<paragraph id="8B8C513EBA19FFBEB752F947F9D9F981" blockId="4.[832,1437,1697,2077]" lastBlockId="5.[151,756,1498,1554]" lastPageId="5" lastPageNumber="84" pageId="4" pageNumber="83">
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<emphasis id="B9478D2CBA19FFBFB752F947FA8CF92B" bold="true" box="[832,953,1697,1720]" pageId="4" pageNumber="83">Diagnosis:</emphasis>
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Asian
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<taxonomicName id="4C332ABDBA19FFBFB008F947FDFBF92B" authorityName="Shear & Gruber" authorityYear="1983" box="[1050,1230,1697,1720]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="83" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
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with free metapeltidium not joined to prosoma or to opisthosoma. Characterized by an unspecialized penis with a rather short, slightly twisted stylus, opening of seminal duct sub-terminal; spicules on apex of penis rather long on do, ve and la sides of glans (rarely on distal part of truncus), in most cases symmetrically arranged in three (rarely four) groups on glans and on distal part of truncus, rather uniform in all known species; network of anvil-shaped denticles on do side of body present, cells not strongly symmetrical, appearing somewhat disorderly, individual cell-forming denticles low in most species, thus without marked sculpture on opisthosomal scutum.
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</paragraph>
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</subSubSection>
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<caption id="DF4C01B6BA18FFBEB48BFADFFAF4FAF8" ID-DOI="http://doi.org/10.5281/zenodo.3702601" ID-Zenodo-Dep="3702601" httpUri="https://zenodo.org/record/3702601/files/figure.png" pageId="5" pageNumber="84" startId="5.[153,194,1337,1358]" targetBox="[153,1438,255,1290]" targetPageId="5">
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<paragraph id="8B8C513EBA18FFBEB48BFADFFAF4FAF8" blockId="5.[153,1437,1337,1387]" pageId="5" pageNumber="84">
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Figs 2-5.
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<taxonomicName id="4C332ABDBA18FFBEB4EAFADFF8FAFADD" authorityName="Suzuki" authorityYear="1963" box="[248,463,1337,1358]" class="Arachnida" family="Nemastomatidae" genus="Cladolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="84" phylum="Arthropoda" rank="species" species="parvulum">
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<emphasis id="B9478D2CBA18FFBEB4EAFADFF8FAFADD" box="[248,463,1337,1358]" italics="true" pageId="5" pageNumber="84">Cladolasma parvulum</emphasis>
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</taxonomicName>
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. (2) Body of male in dorsal view. (3) Body of female in dorsal view. (4) Prosoma of male in dorsal view. (5) Opisthosoma of male in dorsal view. Photographs by J. Schmidt.
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</paragraph>
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</caption>
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<subSubSection id="C32902B5BA18FFBEB485F9CDF8F9F975" pageId="5" pageNumber="84" type="etymology">
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<paragraph id="8B8C513EBA18FFBEB485F9CDF8F9F975" blockId="5.[151,756,1579,1766]" pageId="5" pageNumber="84">
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<emphasis id="B9478D2CBA18FFBEB485F9CDF9D6F9D1" bold="true" box="[151,227,1579,1602]" pageId="5" pageNumber="84">Name:</emphasis>
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It is derived from Asia, the continent where the species of this genus live, and the Greek noun “elasma”, a plate, mainly of metal, referring to the hood on the anterior margin of the prosoma. The name is in analogy of the other generic names of
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<taxonomicName id="4C332ABDBA18FFBEB616F948FB8FF956" authorityName="Shear & Gruber" authorityYear="1983" box="[516,698,1710,1733]" class="Arachnida" family="Nemastomatidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="84" phylum="Arthropoda" rank="subFamily" subFamily="Ortholasmatinae">Ortholasmatinae</taxonomicName>
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. The gender of the name is neuter.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32902B5BA18FFBCB485F919FAB0FD0A" lastPageId="7" lastPageNumber="86" pageId="5" pageNumber="84" type="distribution">
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<paragraph id="8B8C513EBA18FFBEB485F919FDDCF90B" blockId="5.[151,757,1791,2077]" lastBlockId="5.[831,1436,1496,1688]" pageId="5" pageNumber="84">
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<emphasis id="B9478D2CBA18FFBEB485F919F816F885" bold="true" box="[151,291,1791,1814]" pageId="5" pageNumber="84">Distribution</emphasis>
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(
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<figureCitation id="13084DBBBA18FFBEB524F919F84FF885" box="[310,378,1791,1814]" captionStart="Fig" captionStartId="3.[151,186,1964,1985]" captionTargetBox="[151,1436,1216,1921]" captionTargetId="figure@3.[151,1436,1216,1921]" captionTargetPageId="3" captionText="Fig. 1. Distributional records of ortholasmatine species in East and Southeast Asia. (1) Asiolasma billsheari sp. nov., China, Gansu, Qinling range. (2) A. juergengruberi sp. nov., China, Yunnan, above Lugu Hu. (3) Same species as before, China, north of Lijiang. (4) A. ailaoshan, China, Yunnan, Ailao Shan. (5) A. damingshan, China, Guangxi, Daming Shan. (6) A. schwendingeri sp. nov., Vietnam, Mt Ba Vi. (7) A. angka, Thailand, Doi Inthanon. (8) Dendrolasma parvulum, Japan, Shikoku Island." figureDoi="http://doi.org/10.5281/zenodo.3702599" httpUri="https://zenodo.org/record/3702599/files/figure.png" pageId="5" pageNumber="84">Fig. 1</figureCitation>
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): The six hitherto known species occur in northern
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<collectingCountry id="F32411AEBA18FFBEB57FF8C6F8F8F8A4" box="[365,461,1824,1847]" name="Thailand" pageId="5" pageNumber="84">Thailand</collectingCountry>
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(Doi Inthanon), northern
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<collectingCountry id="F32411AEBA18FFBEB485F8A7F9C0F8CB" box="[151,245,1857,1880]" name="Vietnam" pageId="5" pageNumber="84">Vietnam</collectingCountry>
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, as well as southwestern and southern
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<collectingCountry id="F32411AEBA18FFBEB6A6F8A7FBC1F8CB" box="[692,756,1857,1880]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
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(
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<collectingRegion id="49F79FDCBA18FFBEB48CF887F9C1F8EB" box="[158,244,1889,1912]" country="China" name="Yunnan" pageId="5" pageNumber="84">Yunnan</collectingRegion>
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,
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<collectingRegion id="49F79FDCBA18FFBEB511F887F86AF8EB" box="[259,351,1889,1912]" country="China" name="Guangxi" pageId="5" pageNumber="84">Guangxi</collectingRegion>
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and
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<collectingRegion id="49F79FDCBA18FFBEB589F887F8EAF8EB" box="[411,479,1889,1912]" country="China" name="Gansu" pageId="5" pageNumber="84">Gansu</collectingRegion>
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provinces, respectively). The few available records are highly scattered and span a geographically and climatically large and diverse area from the tropics/subtropics in
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<collectingCountry id="F32411AEBA18FFBEB5F6F822FB74F848" box="[484,577,1988,2011]" name="Vietnam" pageId="5" pageNumber="84">Vietnam</collectingCountry>
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, southern
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<collectingCountry id="F32411AEBA18FFBEB6A6F822FBC1F848" box="[692,756,1988,2011]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
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and northern
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<collectingCountry id="F32411AEBA18FFBEB521F803F8A6F86F" box="[307,403,2021,2044]" name="Thailand" pageId="5" pageNumber="84">Thailand</collectingCountry>
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to high altitudes of temperate mountain ranges in central southwestern and partly southern
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<collectingCountry id="F32411AEBA18FFBEB7A0FA3EFAC2FA7C" box="[946,1015,1496,1519]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
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. According to current knowledge,
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<taxonomicName id="4C332ABDBA18FFBEB72DFA1CFA9AF982" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[831,943,1530,1553]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="108" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9478D2CBA18FFBEB72DFA1CFA9AF982" box="[831,943,1530,1553]" italics="true" pageId="5" pageNumber="84">Asiolasma</emphasis>
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</taxonomicName>
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species are confined to mountainous areas between
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<quantity id="4CCBFCDBBA18FFBEB7BBF9FAFD36F9A0" box="[937,1027,1564,1587]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="5" pageNumber="84" unit="m" value="1200.0">1200 m</quantity>
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and
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<quantity id="4CCBFCDBBA18FFBEB05AF9FAFD97F9A0" box="[1096,1186,1564,1587]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="5" pageNumber="84" unit="m" value="3300.0">3300 m</quantity>
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in
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<collectingCountry id="F32411AEBA18FFBEB0C7F9FAFC2DF9A0" box="[1237,1304,1564,1587]" name="China" pageId="5" pageNumber="84">China</collectingCountry>
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,
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<quantity id="4CCBFCDBBA18FFBEB13EF9FAFC4DF9A0" box="[1324,1400,1564,1587]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="84" unit="m" value="900.0">900 m</quantity>
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in northern
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<collectingCountry id="F32411AEBA18FFBEB7B7F9DBFD36F9C7" box="[933,1027,1597,1620]" name="Vietnam" pageId="5" pageNumber="84">Vietnam</collectingCountry>
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,
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<quantity id="4CCBFCDBBA18FFBEB000F9DBFD93F9C7" box="[1042,1190,1597,1620]" metricMagnitude="3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="5" pageNumber="84" unit="m" value="2450.0" valueMax="2500.0" valueMin="2400.0">2400-2500 m</quantity>
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in northern
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<collectingCountry id="F32411AEBA18FFBEB124F9DBFCADF9C7" box="[1334,1432,1597,1620]" name="Thailand" pageId="5" pageNumber="84">Thailand</collectingCountry>
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, and the sole
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<taxonomicName id="4C332ABDBA18FFBEB7C3F9B9FD66F9E5" authorityName="Suzuki" authorityYear="1963" box="[977,1107,1631,1654]" class="Arachnida" family="Nemastomatidae" genus="Cladolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus">
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<emphasis id="B9478D2CBA18FFBEB7C3F9B9FD66F9E5" box="[977,1107,1631,1654]" italics="true" pageId="5" pageNumber="84">Cladolasma</emphasis>
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</taxonomicName>
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species has been recorded at
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<quantity id="4CCBFCDBBA18FFBEB72DF967FAFAF90B" box="[831,975,1665,1688]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="5" pageNumber="84" unit="m" value="1350.0" valueMax="1500.0" valueMin="1200.0">1200-1500 m</quantity>
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on Shikoku Island,
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<collectingCountry id="F32411AEBA18FFBEB0B7F967FDD0F90B" box="[1189,1253,1665,1688]" name="Japan" pageId="5" pageNumber="84">Japan</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8C513EBA18FFBCB72DF954FB84FD0A" blockId="5.[831,1437,1714,2077]" lastBlockId="7.[151,756,258,921]" lastPageId="7" lastPageNumber="86" pageId="5" pageNumber="84">
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<emphasis id="B9478D2CBA18FFBEB72DF954FAD6F95A" bold="true" box="[831,995,1714,1737]" pageId="5" pageNumber="84">Relationships:</emphasis>
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Judging from male genital morphology and formation of the hood, all
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<taxonomicName id="4C332ABDBA18FFBEB0A7F932FC10F978" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[1205,1317,1748,1771]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="84" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B9478D2CBA18FFBEB0A7F932FC10F978" box="[1205,1317,1748,1771]" italics="true" pageId="5" pageNumber="84">Asiolasma</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2743057BA18FFBEB121F932FCA8F978" box="[1331,1437,1748,1771]" pageId="5" pageNumber="84" rank="genus">gen. nov.</taxonomicNameLabel>
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species known at present seem to be closely related with each other. These characters seem to be rather uniform, although differences between species are present. Specific distinctions are also found in body shape, body size, equipment of body with a lattice work of anvil-shaped tubercles, size and shape of pedipalps and pattern of setae on pedipalpal articles, presence or absence of pedipalpal glands and shape of apophyses on chelicerae. These need to be carefully identified. The
|
|
<taxonomicName id="4C332ABDBA1AFFBCB4DFFEE4F808FE8A" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[205,317,258,281]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="B9478D2CBA1AFFBCB4DFFEE4F808FE8A" box="[205,317,258,281]" italics="true" pageId="7" pageNumber="86">Asiolasma</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2743057BA1AFFBCB55BFEE4F884FE8A" box="[329,433,258,281]" pageId="7" pageNumber="86" rank="genus">gen. nov.</taxonomicNameLabel>
|
|
species can be divided into two groups. Three of the southern species display only a pointed hook on the upper side of the 2nd cheliceral article (
|
|
<taxonomicName id="4C332ABDBA1AFFBCB4E2FE84F860FEEA" authorityName="Schwendinger & Gruber" authorityYear="1992" box="[240,341,354,377]" class="Arachnida" family="Nemastomatidae" genus="Dendrolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="angka">
|
|
<emphasis id="B9478D2CBA1AFFBCB4E2FE84F860FEEA" box="[240,341,354,377]" italics="true" pageId="7" pageNumber="86">A. angka</emphasis>
|
|
</taxonomicName>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB575FE84F88AFEEA" box="[359,447,354,377]" captionStart="Figs 27-30" captionStartId="9.[149,190,1394,1415]" captionTargetBox="[149,1415,259,1338]" captionTargetId="figure@9.[149,1434,256,1355]" captionTargetPageId="9" captionText="Figs 27-30. Asiolasma angka, male (27, 29), female (28, 30). (27-28) Right chelicera in prolateral view. (29-30) Right pedipalp in prolateral view. Scales: 1.2 mm (27-28); 0.3 mm (29-30)." figureDoi="http://doi.org/10.5281/zenodo.3702609" httpUri="https://zenodo.org/record/3702609/files/figure.png" pageId="7" pageNumber="86">Fig. 27</figureCitation>
|
|
],
|
|
<taxonomicName id="4C332ABDBA1AFFBCB5C5FE84FB48FEEA" authorityName="Zhang & Zhang" authorityYear="2013" box="[471,637,354,377]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="damingshan">
|
|
<emphasis id="B9478D2CBA1AFFBCB5C5FE84FB48FEEA" box="[471,637,354,377]" italics="true" pageId="7" pageNumber="86">A. damingshan</emphasis>
|
|
</taxonomicName>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB682FE84FBDDFEEA" box="[656,744,354,377]" captionStart="Figs 33-37" captionStartId="12.[151,192,1996,2017]" captionTargetBox="[176,1426,731,1933]" captionTargetId="figure@12.[176,1433,724,1933]" captionTargetPageId="12" captionText="Figs 33-37. Asiolasma damingshan, male holotype. (33) Body in dorsal view. (34) Same in lateral view. (35) Chelicera in prolateral view. (36) Apophsis of 2nd cheliceral article in prolateral view. (37) Left pedipalp in retrolateral view. Figs 33-34 reproduced from Zhang & Zhang (2013). Scales: 0.03 mm (36); 0.2 mm (35); 0.4 (37); 0.5 mm (33-34)." figureDoi="http://doi.org/10.5281/zenodo.3702613" httpUri="https://zenodo.org/record/3702613/files/figure.png" pageId="7" pageNumber="86">Fig. 35</figureCitation>
|
|
],
|
|
<taxonomicName id="4C332ABDBA1AFFBCB485FE64F86EFE0A" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[151,347,386,409]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="schwendingeri" status="sp. nov.">
|
|
<emphasis id="B9478D2CBA1AFFBCB485FE64F86EFE0A" box="[151,347,386,409]" italics="true" pageId="7" pageNumber="86">A. schwendingeri</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2743057BA1AFFBCB578FE64F8F3FE0A" box="[362,454,386,409]" pageId="7" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB5C9FE64FB71FE0A" box="[475,580,386,409]" captionStart="Figs 94-103" captionStartId="24.[151,192,1949,1970]" captionTargetBox="[196,1359,259,1912]" captionTargetId="figure@24.[172,1449,254,1916]" captionTargetPageId="24" captionText="Figs 94-103. Asiolasma schwendingeri sp. nov., male holotype (94-100, 102), female paratype (101, 103). (94) Glans penis in lateral view. (95) Same in ventral view. (96) Same in dorsal view. (97) Whole penis in lateral view. (98) Same in ventral view. (99) Same in dorsal view. (100-101) Right chelicera in prolateral view. (102-103) Right pedipalp in prolateral view. Scales: 0.1 mm (94-96); 0.3 mm (100-101); 0.25 mm (97-99); 0.5 mm (102-103)." figureDoi="http://doi.org/10.5281/zenodo.3702633" httpUri="https://zenodo.org/record/3702633/files/figure.png" pageId="7" pageNumber="86">Fig. 100</figureCitation>
|
|
]; in the single northern species (
|
|
<taxonomicName id="4C332ABDBA1AFFBCB54CFE44F8D2FE2A" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[350,487,418,441]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="billsheari" status="sp. nov.">
|
|
<emphasis id="B9478D2CBA1AFFBCB54CFE44F8D2FE2A" box="[350,487,418,441]" italics="true" pageId="7" pageNumber="86">A. billsheari</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2743057BA1AFFBCB5E0FE44FB7FFE2A" box="[498,586,418,441]" pageId="7" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB649FE44FB89FE2A" box="[603,700,418,441]" captionStart="Figs 112-117" captionStartId="27.[151,192,1288,1309]" captionTargetBox="[176,1395,267,1244]" captionTargetId="figure@27.[176,1439,255,1255]" captionTargetPageId="27" captionText="Figs 112-117. Asiolasma billsheari sp. nov., male holotype. (112) Right chelicera in prolateral view. (113) Same in retrolateral view. (114) Apophysis of 2nd cheliceral article in prolateral view. (115) Pedipalp in prolateral view. (116) Distal part of pedipalpal femur, whole patella and proximal part of tibia in retrolateral view. (117) Apophysis on ventro-distal part of pedipalpal patella in prolateral view. Scales: 0.1 mm (114); 0.2 mm (112-113); 0.3 mm (115-116); no scale (117)." figureDoi="http://doi.org/10.5281/zenodo.3702637" httpUri="https://zenodo.org/record/3702637/files/figure.png" pageId="7" pageNumber="86">Fig. 112</figureCitation>
|
|
] and in two of the southern species this hook is present as well but situated on a massive rounded apophysis (
|
|
<taxonomicName id="4C332ABDBA1AFFBCB4B2FDE4F851FD8A" authority="Martens, 2019" authorityName="Martens" authorityYear="2019" box="[160,356,514,537]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="juergengruberi" status="sp. nov.">
|
|
<emphasis id="B9478D2CBA1AFFBCB4B2FDE4F851FD8A" box="[160,356,514,537]" italics="true" pageId="7" pageNumber="86">A. juergengruberi</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A2743057BA1AFFBCB57CFDE4F8F0FD8A" box="[366,453,514,537]" pageId="7" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB5C7FDE4FB66FD8A" box="[469,595,514,537]" captionStart="Figs 72-77" captionStartId="20.[151,192,1554,1575]" captionTargetBox="[176,1411,263,1509]" captionTargetId="figure@20.[166,1433,257,1532]" captionTargetPageId="20" captionText="Figs 72-77. Asiolasma juergengruberi sp. nov.; male (72-75, 77), female (76). (72, 74, 76) Right chelicera in prolateral view. (73, 75) Same in retrolateral view. (77) Apophysis of 1st cheliceral article. (72-73, 76-77) specimen from Lugu Hu. (74-75) specimen from Lijiang. Scale: 0.3 mm (72-76); no scale (77)." figureDoi="http://doi.org/10.5281/zenodo.3702625" httpUri="https://zenodo.org/record/3702625/files/figure.png" pageId="7" pageNumber="86">Figs 72-74</figureCitation>
|
|
],
|
|
<taxonomicName id="4C332ABDBA1AFFBCB67BFDE4FBC1FD8A" authorityName="Zhang, Zhao & Zhang" authorityYear="2018" box="[617,756,514,537]" class="Arachnida" family="Nemastomatidae" genus="Asiolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="ailaoshan">
|
|
<emphasis id="B9478D2CBA1AFFBCB67BFDE4FBC1FD8A" box="[617,756,514,537]" italics="true" pageId="7" pageNumber="86">A. ailaoshan</emphasis>
|
|
</taxonomicName>
|
|
[
|
|
<figureCitation id="13084DBBBA1AFFBCB48CFDC4F813FDAA" box="[158,294,546,569]" captionStart="Figs 55-59" captionStartId="16.[151,192,1807,1828]" captionTargetBox="[186,1437,262,1766]" captionTargetId="figure@16.[152,1440,254,1795]" captionTargetPageId="16" captionText="Figs 55-59. Asiolasma ailaoshan, male(55-56, 58), female (57, 59). (55, 57) Right chelicera in prolateral view. (56) Same in retrolateral view. (58-59) Left pedipalp in prolateral view. Scale: 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.3702619" httpUri="https://zenodo.org/record/3702619/files/figure.png" pageId="7" pageNumber="86">Figs 55-56</figureCitation>
|
|
]). The Japanese
|
|
<taxonomicName id="4C332ABDBA1AFFBCB5EAFDC4FBC1FDAA" authorityName="Suzuki" authorityYear="1963" box="[504,756,546,569]" class="Arachnida" family="Nemastomatidae" genus="Cladolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="species" species="parvulum">
|
|
<emphasis id="B9478D2CBA1AFFBCB5EAFDC4FBC1FDAA" box="[504,756,546,569]" italics="true" pageId="7" pageNumber="86">Cladolasma parvulum</emphasis>
|
|
</taxonomicName>
|
|
stands apart and differs markedly in genital morphology, in a little developed do network of anvil-shaped tubercles and in a small anterior hood (see above).
|
|
</paragraph>
|
|
<caption id="DF4C01B6BA1BFFBDB485F84AFA2FF78B" ID-DOI="http://doi.org/10.5281/zenodo.3702603" ID-Zenodo-Dep="3702603" httpUri="https://zenodo.org/record/3702603/files/figure.png" pageId="6" pageNumber="85" startId="6.[151,192,1964,1985]" targetBox="[156,1424,259,1920]" targetPageId="6">
|
|
<paragraph id="8B8C513EBA1BFFBDB485F84AFC53F868" blockId="6.[151,1437,1964,2072]" pageId="6" pageNumber="85">
|
|
Figs 6-15.
|
|
<emphasis id="B9478D2CBA1BFFBDB511F84AF8D5F852" box="[259,480,1964,1985]" italics="true" pageId="6" pageNumber="85">
|
|
<taxonomicName id="4C332ABDBA1BFFBDB511F84AF8E8F852" authorityName="Suzuki" authorityYear="1963" box="[259,477,1964,1985]" class="Arachnida" family="Nemastomatidae" genus="Cladolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="85" phylum="Arthropoda" rank="species" species="parvulum">Cladolasma parvulum</taxonomicName>
|
|
,
|
|
</emphasis>
|
|
male (6-12, 14), female (13, 15). (6) Glans penis in lateral view. (7) Same in dorsal view. (8) Same in ventral view. (9) Truncus penis in lateral view. (10) Same in dorsal view. (11) Same in ventral view. (12) Right chelicera of male in prolateral view. (13) Same of female. (14) Right pedipalp of male in prolateral view. (15) Same of female.
|
|
</paragraph>
|
|
<paragraph id="8B8C513EBA1BFFBDB511F7E5FA2FF78B" blockId="6.[151,1437,1964,2072]" box="[259,794,2051,2072]" pageId="6" pageNumber="85">Scales: 0.25 mm (9-11); 0.1 mm (6-8); 0.3 mm (12-15).</paragraph>
|
|
</caption>
|
|
<paragraph id="8B8C513EBA1AFFBCB485FD44FAB0FD0A" blockId="7.[151,756,258,921]" lastBlockId="7.[831,1437,258,665]" pageId="7" pageNumber="86">
|
|
All species of the American ortholasmatine genera
|
|
<taxonomicName id="4C332ABDBA1AFFBCB485FD24F881FD4A" authority="Banks, 1894 a" authorityName="Banks" authorityYear="1894" box="[151,436,706,729]" class="Arachnida" family="Nemastomatidae" genus="Ortholasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB485FD24F823FD4A" box="[151,278,706,729]" italics="true" pageId="7" pageNumber="86">Ortholasma</emphasis>
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB50DFD24F881FD4A" author="Banks" box="[287,436,706,729]" firstAuthor="Banks" pageId="7" pageNumber="86" pagination="11 - 12" refId="ref18169" refString="Banks N. 1894 a. The Nemastomatidae and Trogulidae of the United States. Psyche 7: 11 - 12." type="journal article" year="1894" yearSuffix="a">Banks, 1894a</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C332ABDBA1AFFBCB5D3FD24FBC5FD4A" authority="Banks, 1894 b" authorityName="Banks" authorityYear="1894" box="[449,752,706,729]" class="Arachnida" family="Nemastomatidae" genus="Dendrolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB5D3FD24FB64FD4A" box="[449,593,706,729]" italics="true" pageId="7" pageNumber="86">Dendrolasma</emphasis>
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB64BFD24FBC5FD4A" author="Banks" box="[601,752,706,729]" firstAuthor="Banks" pageId="7" pageNumber="86" pagination="160 - 164" refId="ref18191" refString="Banks N. 1894 b. Washington Phalangida, with a description of a new southern Leiobunum. Canadian Entomologist 26: 160 - 164." type="journal article" year="1894" yearSuffix="b">Banks, 1894b</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C332ABDBA1AFFBCB485FD04FB71FD6A" authority="Goodnight & Goodnight, 1942" authorityName="Goodnight & Goodnight" authorityYear="1942" box="[151,580,738,761]" class="Arachnida" family="Nemastomatidae" genus="Trilasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB485FD04F9C3FD6A" box="[151,246,738,761]" italics="true" pageId="7" pageNumber="86">Trilasma</emphasis>
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB4EEFD04FB71FD6A" author="Goodnight & Goodnight" box="[252,580,738,761]" firstAuthor="Goodnight" pageId="7" pageNumber="86" pagination="1 - 18" refId="ref18298" refString="Goodnight C. J., Goodnight M. 1942. Phalangida from Mexico. American Museum Novitates 1164: 1 - 18." type="journal article" year="1942">Goodnight & Goodnight, 1942</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C332ABDBA1AFFBCB65CFD04F823FC8A" authority="Shear, 2006" authorityName="Shear" authorityYear="2006" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB65CFD04FBC1FD6A" box="[590,756,738,761]" italics="true" pageId="7" pageNumber="86">Martensolasma</emphasis>
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB485FCE4F823FC8A" author="Shear" box="[151,278,770,793]" firstAuthor="Shear" pageId="7" pageNumber="86" pagination="191 - 198" refId="ref18579" refString="Shear W. A. 2006. Martensolasma jocheni, a new genus and species of harvestman from Mexico (Opiliones, Nemastomatidae, Ortholasmatinae). Zootaxa 1325: 191 - 198." type="journal article" year="2006">Shear, 2006</bibRefCitation>
|
|
</taxonomicName>
|
|
and
|
|
<emphasis id="B9478D2CBA1AFFBCB559FCE4F8E0FC8A" box="[331,469,770,793]" italics="true" pageId="7" pageNumber="86">Cryptolasma</emphasis>
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB5CEFCE4F87FFCAA" author="Cruz-Lopez, Cruz-Bonilla & Francke" firstAuthor="Cruz-Lopez" pageId="7" pageNumber="86" pagination="714 - 729" refId="ref18251" refString="Cruz-Lopez J. A., Cruz-Bonilla A., Francke O. F. 2018. Molecules and morphology reveal a new aberrant harvestman genus of Ortholasmatinae (Opiliones, Dypsnoi, Nemastomatidae) from Mexico. Systematics and Biodiversity 16: 714 - 729." type="journal article" year="2018">Cruz-López, Cruz-Bonilla & Francke, 2018</bibRefCitation>
|
|
clearly differ from Asian species by the following characters: i) the penis is much more slender, the inflated base shorter, the stylus formed like a barbed hook (not so in
|
|
<taxonomicName id="4C332ABDBA1AFFBCB553FC64F8E4FC0A" authorityName="Banks" authorityYear="1894" box="[321,465,898,921]" class="Arachnida" family="Nemastomatidae" genus="Dendrolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB553FC64F8E4FC0A" box="[321,465,898,921]" italics="true" pageId="7" pageNumber="86">Dendrolasma</emphasis>
|
|
</taxonomicName>
|
|
), the armature of the glans is more diversified and the spicules often of different size and pattern (uniform in
|
|
<taxonomicName id="4C332ABDBA1AFFBCB040FEC4FDE4FEAA" authorityName="Banks" authorityYear="1894" box="[1106,1233,290,313]" class="Arachnida" family="Nemastomatidae" genus="Ortholasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB040FEC4FDE4FEAA" box="[1106,1233,290,313]" italics="true" pageId="7" pageNumber="86">Ortholasma</emphasis>
|
|
</taxonomicName>
|
|
); ii) many (up to twelve on each side) apophyses are present on the hood, the single distal and all following lateral apophyses of the hood are interconnected by fine bridges only at the distal end of the apophyses (hood lacking in
|
|
<taxonomicName id="4C332ABDBA1AFFBCB0F4FE44FCB9FE2A" authorityName="Shear" authorityYear="2006" box="[1254,1420,418,441]" class="Arachnida" family="Nemastomatidae" genus="Martensolasma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="86" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B9478D2CBA1AFFBCB0F4FE44FCB9FE2A" box="[1254,1420,418,441]" italics="true" pageId="7" pageNumber="86">Martensolasma</emphasis>
|
|
</taxonomicName>
|
|
); iii) the network of closed keel cells formed by anvilshaped tubercles on the do scutum is regular, distinctly symmetrical and ornamental, the individual tubercles markedly elevated, resulting in a distinct and elaborate structure of the network (for all characters compare multiple details in
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB008FD84FC2CFDEA" author="Shear & Gruber" box="[1050,1305,610,633]" firstAuthor="Shear" pageId="7" pageNumber="86" pagination="1 - 65" refId="ref18651" refString="Shear W. A., Gruber J. 1983. The opilionid subfamily Ortholasmatinae (Opiliones, Troguloidea, Nemastomatidae). American Museum Novitates 2757: 1 - 65." type="journal article" year="1983">Shear & Gruber, 1983</bibRefCitation>
|
|
and
|
|
<bibRefCitation id="EFA22CCFBA1AFFBCB148FD84FA4CFD0A" author="Shear" firstAuthor="Shear" pageId="7" pageNumber="86" pagination="9 - 45" refId="ref18613" refString="Shear W. A. 2010. New species and records of Ortholasmatine harvestmen from Mexico, Honduras and the western United States (Opiliones, Nemastomatidae, Ortholasmatinae). ZooKeys 52: 9 - 45." type="journal article" year="2010">Shear, 2010</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |