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<mods:title id="BFDE8ADA1732FCB6158B787E1FED6470">Systematic Revision Of The Sand Scorpions, Genus Buthacus Birula, 1908 (Buthidae C. L. Koch, 1837) Of The Levant, With Redescription Of Buthacus Arenicola (Simon, 1885) From Algeria And Tunisia</mods:title>
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<mods:namePart id="2188816EEFB1385B79C085207743E85E">Cain, Shlomo</mods:namePart>
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<mods:namePart id="7B2A1CD722042191A453B8E2F78F2421">Gefen, Eran</mods:namePart>
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<taxonomicName id="4C3FF8E8FF817E0EB34F6D2B33E26DB4" ID-CoL="8LYX5" authority="Cain &amp; Gefen &amp; Prendini, 2021" authorityName="Cain &amp; Gefen &amp; Prendini" authorityYear="2021" box="[812,988,875,899]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="31" phylum="Arthropoda" rank="species" species="arava" status="sp. nov.">
<emphasis id="B94B5F79FF817E0EB34F6D2B33E26DB4" bold="true" box="[812,988,875,899]" italics="true" pageId="31" pageNumber="31">Buthacus arava</emphasis>
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<figureCitation id="13049FEEFF817E0EB2CC6DE5331A6D8A" box="[687,804,933,957]" captionStart="FIGURE 1" captionStartId="5.[109,194,1348,1369]" captionTargetBox="[109,1213,224,1318]" captionTargetId="figure-98@5.[109,1213,224,1318]" captionTargetPageId="5" captionText="FIGURE 1. Buthacus Birula, 1908, representative habitats. A. Mamshit [Kurnub], Israel, habitat of Buthacus amitaii, sp. nov. B. Naftah to Ong Jmal, Tunisia, habitat of Buthacus arenicola (Simon, 1885). C. Faiyum Oasis, Egypt, habitat of Buthacus leptochelys (Ehrenberg, 1829). D. Ashdod Nizzanim Nature Reserve, Israel, habitat of Buthacus nitzani Levy et al., 1973, stat. nov. E. Yotvata, Israel, habitat of Buthacus arava, sp. nov., and Buthacus yotvatensis Levy et al., 1973, stat. rev. F. Beer Milka, Israel, Holot Haluza sand dunes, Israel, habitat of Buthacus levyi, sp. nov., and B. nitzani." figureDoi="http://doi.org/10.5281/zenodo.5413471" httpUri="https://zenodo.org/record/5413471/files/figure.png" pageId="31" pageNumber="31">Figures 1E</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3536DE5336D6D8A" box="[816,851,933,957]" captionStart="FIGURE 3" captionStartId="7.[109,195,1394,1415]" captionTargetBox="[108,636,224,1364]" captionTargetId="figure-433@7.[108,636,224,1364]" captionTargetPageId="7" captionText="FIGURE 3. Buthacus Birula, 1908, habitus in life. A. Buthacus arava, sp. nov., ♂. B, C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (B), ♀ (C)." figureDoi="http://doi.org/10.5281/zenodo.5413477" httpUri="https://zenodo.org/record/5413477/files/figure.png" pageId="31" pageNumber="31">3A</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB33D6DE533536D8A" box="[862,877,933,957]" captionStart="FIGURE 4" captionStartId="8.[109,195,1546,1567]" captionTargetBox="[114,1208,223,1512]" captionTargetId="figure-9@8.[108,1212,213,1518]" captionTargetPageId="8" captionText="FIGURE 4. Map of southwestern Levant, plotting known locality records for Buthacus arava, sp. nov. in Israel and Jordan, based on material examined for the present study. White symbol denotes type locality." figureDoi="http://doi.org/10.5281/zenodo.5413485" httpUri="https://zenodo.org/record/5413485/files/figure.png" pageId="31" pageNumber="31">4</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB31A6DE533AB6D8A" box="[889,917,933,957]" captionStart="FIGURE 15" captionStartId="21.[108,194,1212,1233]" captionTargetBox="[151,1156,225,1183]" captionTargetPageId="21" captionText="FIGURE 15. Nonmetric multidimensional scaling (NMDS) plots for A, male and B, female Buthacus Birula, 1908, based on 24 morphometric ratios and counts, for 79 adults (each symbol represents an individual): Buthacus amitaii, sp. nov. (white triangles); Buthacus arava, sp. nov. (black triangles); Buthacus arenicola (Simon, 1885) (white inverted triangles); Buthacus leptochelys (Ehrenberg, 1829) (white circles); Buthacus levyi, sp. nov. (white squares); Buthacus nitzani Levy et al., 1973, stat. nov. (white diamonds); Buthacus tadmorensis (Simon, 1892), stat. rev. (black circles); Buthacus yotvatensis Levy et al., 1973, stat. rev. (black squares)." figureDoi="http://doi.org/10.5281/zenodo.5413513" httpUri="https://zenodo.org/record/5413513/files/figure.png" pageId="31" pageNumber="31">15</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3C36DE533D36D8B" box="[928,1005,933,957]" captionStart="FIGURE 16" captionStartId="36.[109,192,1628,1649]" captionTargetBox="[107,1211,226,1601]" captionTargetId="figure-9@36.[107,1211,226,1601]" captionTargetPageId="36" captionText="FIGURE 16. Buthacus Birula, 1908, carapace, dorsal aspect. A, B. Buthacus arava, sp. nov., A. Paratype ♂ (HUJ INVSC 3287),B. Paratype ♀ (HUJ INVSC 2561).C, D. Buthacus tadmorensis (Simon, 1892), stat.rev.,C. Lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912, D. Paralectotype ♀ (NHMW 2452). E, F. Buthacus yotvatensis Levy et al., 1973, stat. rev., E. ♂ (HUJ INVSC 3344), F. ♀ (HUJ INVSC 3318). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413521" httpUri="https://zenodo.org/record/5413521/files/figure.png" pageId="31" pageNumber="31">16A, B</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB39B6DE5347A6D8B" box="[1016,1092,933,957]" captionStart="FIGURE 19" captionStartId="40.[108,194,1626,1647]" captionTargetBox="[107,1211,224,1599]" captionTargetId="figure-90@40.[107,1211,224,1599]" captionTargetPageId="40" captionText="FIGURE 19. Buthacus Birula, 1908, sternum, genital opercula and pectines, ventral aspect. A, B. Buthacus arava, sp. nov., A. Paratype ♂ (HUJ INVSC 3287), B. Paratype ♀ (HUJ INVSC 2561). C, D. Buthacus tadmorensis (Simon, 1892), stat. rev., C. Lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912, D. Paralectotype ♀ (NHMW 2452). E, F. Buthacus yotvatensis Levy et al., 1973, stat. rev., E. ♂ (HUJ INVSC 3344), F. ♀ (HUJ INVSC 3318). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413529" httpUri="https://zenodo.org/record/5413529/files/figure.png" pageId="31" pageNumber="31">19A, B</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB42C6DE534456D8A" box="[1103,1147,933,957]" captionStart="FIGURE 21" captionStartId="42.[107,193,1288,1309]" captionTargetBox="[107,635,224,1260]" captionTargetId="figure-361@42.[107,635,224,1260]" captionTargetPageId="42" captionText="FIGURE 21. Buthacus Birula, 1908, dextral pedipalp chela movable finger, dorsal aspect, illustrating dentition. A. Buthacus amitaii, sp. nov., holotype ♂ (HUJ INVSC 3480). B. Buthacus arava, sp. nov., paratype ♂ (HUJ INVSC 3290). C. Buthacus arenicola (Simon, 1885), ♂ (AMNH). D. Buthacus leptochelys (Ehrenberg, 1829), ♂ (HUJ INVSC 859). E, F. Buthacus levyi, sp. nov., E. Holotype ♂ (HUJ INVSC 2082), F. Paratype ♀ (HUJ INVSC 3252). G. Buthacus nitzani Levy et al., 1973, stat. nov., ♂ (HUJ INVSC 3227). H. Buthacus tadmorensis (Simon, 1892), stat. rev., lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. I. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (HUJ INVSC 3302). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5413535" httpUri="https://zenodo.org/record/5413535/files/figure.png" pageId="31" pageNumber="31">21B</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB4E46DE5348B6D8A" box="[1159,1205,933,957]" captionStart="FIGURE 22" captionStartId="43.[109,192,1568,1589]" captionTargetBox="[113,1197,230,1541]" captionTargetId="figure-13@43.[108,1212,224,1541]" captionTargetPageId="43" captionText="FIGURE 22. Buthacus Birula, 1908, dextral legs IIV, basitarsus and telotarsus, ventral aspect. A. Buthacus arenicola (Simon, 1885), ♂ (AMNH). B. Buthacus levyi, sp. nov., holotype ♂ (HUJ INVSC 2082). C. Buthacus arava, sp. nov., paratype♂ (HUJ INVSC 3295). D. Buthacus leptochelys (Ehrenberg, 1829), ♀ (HUJ INVSC 1807). E. Buthacus tadmorensis (Simon, 1892), stat. rev., lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. F. Buthacus nitzani Levy et al., 1973, stat. nov., ♀ (HUJ INVSC 3226). G. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (HUJ INVSC 3302). H. Buthacus amitaii, sp. nov., holotype ♂ (HUJ INVSC 3480). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5413537" httpUri="https://zenodo.org/record/5413537/files/figure.png" pageId="31" pageNumber="31">22C</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB2A26D8632CE6DE9" box="[705,752,966,990]" captionStart="FIGURE 23" captionStartId="50.[108,194,1559,1580]" captionTargetBox="[108,1212,224,1533]" captionTargetId="figure-9@50.[108,1212,224,1533]" captionTargetPageId="50" captionText="FIGURE 23. Buthacus Birula, 1908, metasomal segments IV and telson, dorsal aspect. A. Buthacus arava, sp. nov., paratype ♀ (HUJ INVSC 2561). B. Buthacus tadmorensis (Simon, 1892), stat. rev., paralectotype ♀ (NHMW 2452) of Buthus pietschmanni Penther, 1912. C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INVSC 3318). D. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5413541" httpUri="https://zenodo.org/record/5413541/files/figure.png" pageId="31" pageNumber="31">23A</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB2986D8633146DE9" box="[763,810,966,990]" captionStart="FIGURE 25" captionStartId="52.[107,193,1562,1583]" captionTargetBox="[108,1212,224,1534]" captionTargetId="figure-73@52.[108,1212,224,1535]" captionTargetPageId="52" captionText="FIGURE 25. Buthacus Birula, 1908, metasomal segments IV and telson, lateral aspect. A. Buthacus arava, sp. nov., paratype ♀ (HUJ INVSC 2561). B. Buthacus tadmorensis (Simon, 1892), stat. rev., paralectotype ♀ (NHMW 2452) of Buthus pietschmanni Penther, 1912. C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INVSC 3318). D. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5413547" httpUri="https://zenodo.org/record/5413547/files/figure.png" pageId="31" pageNumber="31">25A</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3556D86335B6DE9" box="[822,869,966,990]" captionStart="FIGURE 27" captionStartId="54.[108,194,1566,1587]" captionTargetBox="[108,1212,224,1537]" captionTargetId="figure-9@54.[108,1212,224,1538]" captionTargetPageId="54" captionText="FIGURE 27. Buthacus Birula, 1908, metasomal segments IV and telson, ventral aspect. A. Buthacus arava, sp. nov., paratype ♀ (HUJ INVSC 2561). B. Buthacus tadmorensis (Simon, 1892), stat. rev., paralectotype ♀ (NHMW 2452) of Buthus pietschmanni Penther, 1912. C. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♀ (HUJ INVSC 3318). D. Buthacus arenicola (Simon, 1885), ♂ (AMNH). Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5413553" httpUri="https://zenodo.org/record/5413553/files/figure.png" pageId="31" pageNumber="31">27A</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3136D8633A06DE9" box="[880,926,966,990]" captionStart="FIGURE 29" captionStartId="60.[108,194,1439,1460]" captionTargetBox="[108,1212,224,1410]" captionTargetId="figure-113@60.[108,1212,224,1410]" captionTargetPageId="60" captionText="FIGURE 29. Buthacus Birula, 1908, telson, lateral aspect. A. Buthacus arenicola (Simon, 1885), ♂ (AMNH). B. Buthacus levyi, sp. nov., holotype ♂ (HUJ INVSC 2082). C. Buthacus arava, sp. nov., paratype ♂ (HUJ INVSC 2559). D. Buthacus leptochelys (Ehrenberg, 1829), ♂ (HUJ INVSC 859). E. Buthacus tadmorensis (Simon, 1892), stat. rev., lectotype ♂ (NHMW 2453) of Buthus pietschmanni Penther, 1912. F. Buthacus nitzani Levy et al., 1973, stat. nov., ♂ (HUJ INVSC 3227). G. Buthacus yotvatensis Levy et al., 1973, stat. rev., ♂ (HUJ INVSC 3302). H. Buthacus amitaii, sp. nov., holotype ♂ (HUJ INVSC 3480). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5413557" httpUri="https://zenodo.org/record/5413557/files/figure.png" pageId="31" pageNumber="31">29C</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3C96D8633F86DE9" box="[938,966,966,990]" captionStart="FIGURE 33" captionStartId="65.[108,194,1659,1680]" captionTargetBox="[119,1172,233,1626]" captionTargetId="figure-52@65.[109,1213,224,1629]" captionTargetPageId="65" captionText="FIGURE 33. Buthacus arava, sp. nov., habitus, dorsal (A, C) and ventral (B, D) aspects. A, B. Paratype ♂ (HUJ INVSC 2559). C, D. Paratype ♀ (HUJ INVSC 2561). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5413569" httpUri="https://zenodo.org/record/5413569/files/figure.png" pageId="31" pageNumber="31">33</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3B16D8633D06DE9" box="[978,1006,966,990]" captionStart="FIGURE 34" captionStartId="66.[108,194,1658,1679]" captionTargetBox="[338,981,223,1628]" captionTargetId="figure-9@66.[335,983,223,1629]" captionTargetPageId="66" captionText="FIGURE 34. Buthacus arava, sp. nov., ♂♂ paratypes (HUJ INVSC 2559, 3295), dextral pedipalp femur (A, B) and patella (C, E), prolateral (A, C), dorsal (B, D), and retrolateral (E) aspects. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413573" httpUri="https://zenodo.org/record/5413573/files/figure.png" pageId="31" pageNumber="31">34</figureCitation>
,
<figureCitation id="13049FEEFF817E0EB3996D8634286DE9" box="[1018,1046,966,990]" captionStart="FIGURE 35" captionStartId="67.[108,194,1354,1375]" captionTargetBox="[203,1119,227,1320]" captionTargetId="figure-13@67.[203,1119,227,1323]" captionTargetPageId="67" captionText="FIGURE 35. Buthacus arava, sp. nov., dextral pedipalp chela, dorsal (A, C) and retrolateral (B, D) aspects. A, B. Paratype ♂ (HUJ INVSC 3295). C, D. Paratype ♀ (HUJ INVSC 2561). Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5413575" httpUri="https://zenodo.org/record/5413575/files/figure.png" pageId="31" pageNumber="31">35</figureCitation>
; tables 1, 8, 9
</paragraph>
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<treatmentCitationGroup id="AB2FA445FF817E0EB2CF6DBC33F16A01" pageId="31" pageNumber="31">
<treatmentCitation id="0A9EA57AFF817E0EB2CF6DBC34B16A23" author="Amr, Z. S. &amp; M. A. A. Baker &amp; M. Al-Saraireh &amp; D. A. Warrell" box="[684,1167,1020,1044]" page="86" pageId="31" pageNumber="31" year="2021">
<taxonomicName id="4C3FF8E8FF817E0EB2CF6DBC34B16A23" ID-CoL="NYWL" authority=": Amr et al., 2021: 86" authorityName="Amr" authorityPageNumber="86" authorityYear="2021" box="[684,1167,1020,1044]" class="Arachnida" family="Buthidae" genus="Buthacus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="31" phylum="Arthropoda" rank="species" species="nigroaculeatus">
<emphasis id="B94B5F79FF817E0EB2CF6DBC338C6A23" box="[684,946,1020,1044]" italics="true" pageId="31" pageNumber="31">Buthacus nigroaculeatus:</emphasis>
<bibRefCitation id="EFAEFE9AFF817E0EB3DA6DBC34B16A23" author="Amr, Z. S. &amp; M. A. A. Baker &amp; M. Al-Saraireh &amp; D. A. Warrell" box="[953,1167,1020,1044]" pageId="31" pageNumber="31" pagination="83 - 103" refId="ref71217" refString="Amr, Z. S., M. A. A. Baker, M. Al-Saraireh, and D. A. Warrell. 2021. Scorpions and scorpion sting envenoming (scorpionism) in the Arab countries of the Middle East. Toxicon 191: 83 - 103." type="journal article" year="2021">Amr et al., 2021: 86</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 3B (misidentification).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C325D0E0FF817E0EB2AF6A1D348268F0" pageId="31" pageNumber="31" type="materials_examined">
<paragraph id="8B80836BFF817E0EB2AF6A1D348268F0" blockId="31.[684,1213,1115,1736]" pageId="31" pageNumber="31">
<materialsCitation id="3B578936FF817E0EB2AF6A1D32DB6ACF" collectingDate="2018-09-13" collectionCode="TYPE, MATERIAL, HUJ" collectorName="S. Cain" country="Israel" elevation="128" latitude="30.00289" location="Lotan" longLatPrecision="1" longitude="35.09558" municipality="Mehoz HaDarom" pageId="31" pageNumber="31" specimenCode="INVSC 3672" specimenCount="1" specimenCount-male="1" stateProvince="Southern" typeStatus="holotype">
<collectionCode id="ED2E1BAEFF817E0EB2AF6A1D33BC6A43" box="[716,898,1116,1140]" pageId="31" pageNumber="31">TYPE MATERIAL</collectionCode>
:
<typeStatus id="54843DC9FF817E0EB3F06A1C33C46A43" box="[915,1018,1116,1140]" pageId="31" pageNumber="31" type="holotype">Holotype</typeStatus>
<specimenCount id="9D3948E2FF817E0EB4676A1B34256A43" box="[1028,1051,1115,1140]" pageId="31" pageNumber="31" type="male"></specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB44E6A1C345F6A43" box="[1069,1121,1116,1140]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB40F6A1C32DA6AA2" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 3672</specimenCode>
),
<emphasis id="B94B5F79FF817E0EB2956A3D33666AA2" bold="true" box="[758,856,1149,1173]" pageId="31" pageNumber="31">
<collectingCountry id="F328C3FBFF817E0EB2956A3D336A6AA2" box="[758,852,1149,1173]" name="Israel" pageId="31" pageNumber="31">ISRAEL</collectingCountry>
:
</emphasis>
<emphasis id="B94B5F79FF817E0EB33C6A3D34266AA2" box="[863,1048,1149,1173]" italics="true" pageId="31" pageNumber="31">
<collectingMunicipality id="6BE41911FF817E0EB33C6A3D34266AA2" box="[863,1048,1149,1173]" pageId="31" pageNumber="31">Mehoz HaDarom</collectingMunicipality>
</emphasis>
(
<emphasis id="B94B5F79FF817E0EB44B6A3D32E36A81" italics="true" pageId="31" pageNumber="31">
<collectingRegion id="49FB4D89FF817E0EB44B6A3D32E36A81" country="Israel" name="Southern" pageId="31" pageNumber="31">Southern District</collectingRegion>
</emphasis>
):
<location id="8EE0D5B0FF817E0EB3646ADF33706A81" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB3646ADF33706A81" box="[775,846,1183,1206]" country="Israel" latitude="30.00289" longLatPrecision="1" longitude="35.09558" municipality="Mehoz HaDarom" name="Lotan" pageId="31" pageNumber="31" stateProvince="Southern">Lotan</location>
,
<locationDeviation id="9A329A49FF817E0EB30F6ADF34896A81" box="[876,1207,1182,1207]" location="Lotan Plantations" pageId="31" pageNumber="31">SE of Lotan Plantations</locationDeviation>
,
<geoCoordinate id="EE0BE5ACFF817E0EB2CF6A8033036AE0" box="[684,829,1214,1240]" degrees="30" direction="north" minutes="00" orientation="latitude" pageId="31" pageNumber="31" precision="1" seconds="10.4" value="30.00289">30°0010.4″N</geoCoordinate>
<geoCoordinate id="EE0BE5ACFF817E0EB3276AFF33EC6AE0" box="[836,978,1214,1240]" degrees="35" direction="east" minutes="05" orientation="longitude" pageId="31" pageNumber="31" precision="1" seconds="44.1" value="35.09558">35°0544.1″E</geoCoordinate>
,
<quantity id="4CC72E8EFF817E0EB3BE6A80341C6AE0" box="[989,1058,1216,1240]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="31" unit="m" value="128.0">
<elevation id="00126458FF817E0EB3BE6A80341C6AE0" box="[989,1058,1216,1240]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="31" unit="m" value="128.0">128 m</elevation>
</quantity>
,
<date id="FF81A5ABFF817E0EB44E6AFF34A16AE0" box="[1069,1183,1215,1239]" pageId="31" pageNumber="31" value="2018-09-13">
<collectingDate id="EFC55C43FF817E0EB44E6AFF34A16AE0" box="[1069,1183,1215,1239]" pageId="31" pageNumber="31" value="2018-09-13">13.ix.2018</collectingDate>
</date>
,
<collectorName id="26CAE6BDFF817E0EB4CA6A8032DB6ACF" pageId="31" pageNumber="31">S. Cain</collectorName>
</materialsCitation>
.
<materialsCitation id="3B578936FF817E0EB29A6AA1335C6B6B" collectingDate="2003-09-29" collectionCode="HUJ" collectorName="U. Shanas" country="Israel" elevation="123" latitude="29.983334" location="Lotan" longLatPrecision="1224" longitude="35.083332" municipality="Mehoz HaDarom" pageId="31" pageNumber="31" specimenCode="INVSC 2560" specimenCount="1" specimenCount-male="1" stateProvince="Southern" typeStatus="paratype">
<typeStatus id="54843DC9FF817E0EB29A6AA133546ACE" box="[761,874,1249,1273]" pageId="31" pageNumber="31" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B94B5F79FF817E0EB31E6AA033DD6ACF" bold="true" box="[893,995,1248,1272]" pageId="31" pageNumber="31">
<collectingCountry id="F328C3FBFF817E0EB31E6AA033E06ACF" box="[893,990,1248,1272]" name="Israel" pageId="31" pageNumber="31">ISRAEL</collectingCountry>
:
</emphasis>
<emphasis id="B94B5F79FF817E0EB3906AA034856ACF" box="[1011,1211,1248,1272]" italics="true" pageId="31" pageNumber="31">
<collectingMunicipality id="6BE41911FF817E0EB3906AA034856ACF" box="[1011,1211,1248,1272]" pageId="31" pageNumber="31">Mehoz HaDarom</collectingMunicipality>
</emphasis>
(
<emphasis id="B94B5F79FF817E0EB2D56B4233B76B2E" box="[694,905,1282,1306]" italics="true" pageId="31" pageNumber="31">
<collectingRegion id="49FB4D89FF817E0EB2D56B4233B76B2E" box="[694,905,1282,1306]" country="Israel" name="Southern" pageId="31" pageNumber="31">Southern District</collectingRegion>
</emphasis>
): near
<location id="8EE0D5B0FF817E0EB39A6B4234026B2E" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB39A6B4234026B2E" box="[1017,1084,1282,1305]" country="Israel" latitude="29.983334" longLatPrecision="1224" longitude="35.083332" municipality="Mehoz HaDarom" name="Lotan" pageId="31" pageNumber="31" stateProvince="Southern">Lotan</location>
[
<geoCoordinate id="EE0BE5ACFF817E0EB43B6B4134826B2E" box="[1112,1212,1281,1306]" degrees="29" direction="north" minutes="59" orientation="latitude" pageId="31" pageNumber="31" precision="925" value="29.983334">29°59N</geoCoordinate>
<geoCoordinate id="EE0BE5ACFF817E0EB2CF6B6333366B0D" box="[684,776,1314,1339]" degrees="35" direction="east" minutes="05" orientation="longitude" pageId="31" pageNumber="31" precision="925" value="35.083332">35°05E</geoCoordinate>
],
<quantity id="4CC72E8EFF817E0EB3436B6333526B0D" box="[800,876,1315,1339]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="31" pageNumber="31" unit="m" value="123.0">
<elevation id="00126458FF817E0EB3436B6333526B0D" box="[800,876,1315,1339]" metricMagnitude="2" metricUnit="m" metricValue="1.23" pageId="31" pageNumber="31" unit="m" value="123.0">123 m</elevation>
</quantity>
,
<date id="FF81A5ABFF817E0EB31E6B6333CE6B0C" box="[893,1008,1315,1339]" pageId="31" pageNumber="31" value="2003-09-29">
<collectingDate id="EFC55C43FF817E0EB31E6B6333CE6B0C" box="[893,1008,1315,1339]" pageId="31" pageNumber="31" value="2003-09-29">29.ix.2003</collectingDate>
</date>
,
<specimenCount id="9D3948E2FF817E0EB4626B6234266B0C" box="[1025,1048,1314,1339]" pageId="31" pageNumber="31" type="male"></specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB4486B63345E6B0C" box="[1067,1120,1315,1339]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB40F6B6332DA6B6B" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 2560</specimenCode>
),
<collectorName id="26CAE6BDFF817E0EB2956B04335C6B6B" box="[758,866,1348,1372]" pageId="31" pageNumber="31">U. Shanas</collectorName>
</materialsCitation>
;
<materialsCitation id="3B578936FF817E0EB30F6B0433C06815" collectingDate="2017-09-15" collectingDateMax="2018-03-27" collectingDateMin="2017-09-15" collectionCode="HUJ, AMCC" collectorName="Y. Olek &amp; S. Cain &amp; Olek &amp; Cain" country="Israel" elevation="128" latitude="30.00289" location="Lotan" longLatPrecision="1" longitude="35.09558" pageId="31" pageNumber="31" specimenCode="INVSC 3282, 3284- 3287, 3289-3291, LP 15042, INVSC 3283, LP 15043, INVSC 3288, INVSC3381" specimenCount="11" specimenCount-female="1" specimenCount-male="10" stateProvince="Southern" typeStatus="paratype">
<location id="8EE0D5B0FF817E0EB30F6B0433906B6C" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB30F6B0433906B6C" box="[876,942,1348,1371]" country="Israel" latitude="30.00289" longLatPrecision="1" longitude="35.09558" name="Lotan" pageId="31" pageNumber="31" stateProvince="Southern">Lotan</location>
,
<locationDeviation id="9A329A49FF817E0EB3DB6B0434866B6B" box="[952,1208,1348,1372]" location="Lotan Plantations" pageId="31" pageNumber="31">SE of Lotan Plantations</locationDeviation>
,
<geoCoordinate id="EE0BE5ACFF817E0EB2CF6B2533036B4B" box="[684,829,1380,1405]" degrees="30" direction="north" minutes="00" orientation="latitude" pageId="31" pageNumber="31" precision="1" seconds="10.4" value="30.00289">30°0010.4″N</geoCoordinate>
<geoCoordinate id="EE0BE5ACFF817E0EB3206B2533EF6B4B" box="[835,977,1380,1405]" degrees="35" direction="east" minutes="05" orientation="longitude" pageId="31" pageNumber="31" precision="1" seconds="44.1" value="35.09558">35°0544.1″E</geoCoordinate>
,
<quantity id="4CC72E8EFF817E0EB3BF6B25341C6B4B" box="[988,1058,1381,1405]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="31" unit="m" value="128.0">
<elevation id="00126458FF817E0EB3BF6B25341C6B4B" box="[988,1058,1381,1405]" metricMagnitude="2" metricUnit="m" metricValue="1.28" pageId="31" pageNumber="31" unit="m" value="128.0">128 m</elevation>
</quantity>
,
<date id="FF81A5ABFF817E0EB44F6B2534A36B4A" box="[1068,1181,1381,1405]" pageId="31" pageNumber="31" value="2017-09-15">
<collectingDate id="EFC55C43FF817E0EB44F6B2534A36B4A" box="[1068,1181,1381,1405]" pageId="31" pageNumber="31" value="2017-09-15">15.ix.2017</collectingDate>
</date>
,
<collectorName id="26CAE6BDFF817E0EB4CB6B2532DE6BA9" pageId="31" pageNumber="31">Y. Olek</collectorName>
and
<collectorName id="26CAE6BDFF817E0EB3756BC633586BA9" box="[790,870,1414,1438]" pageId="31" pageNumber="31">S. Cain</collectorName>
,
<specimenCount id="9D3948E2FF817E0EB3126BC633A26BA9" box="[881,924,1413,1438]" pageId="31" pageNumber="31" type="male">8 ♂</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB3CA6BC633E36BA9" box="[937,989,1414,1438]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB3806BC634516BA9" box="[995,1135,1414,1438]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 3282</specimenCode>
,
<specimenCode id="DB992B10FF817E0EB4196BC632DD6B88" collectionCode="HUJ" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">3284 3287</specimenCode>
,
<specimenCode id="DB992B10FF817E0EB2926BE733536B88" box="[753,877,1447,1471]" collectionCode="HUJ" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">32893291</specimenCode>
),
<specimenCount id="9D3948E2FF817E0EB3E16BE7338E6B88" box="[898,944,1446,1471]" pageId="31" pageNumber="31" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB3A26BE7342D6B88" box="[961,1043,1447,1471]" country="USA" httpUri="http://grbio.org/cool/m0b5-nwmg" name="Ambrose Monell Cryo Collection, American Museum of Natural History" pageId="31" pageNumber="31">AMCC</collectionCode>
[
<specimenCode id="DB992B10FF817E0EB4406BE734AD6B88" box="[1059,1171,1447,1471]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="31" pageNumber="31">LP 15042</specimenCode>
] ex HUJ
<specimenCode id="DB992B10FF817E0EB2846B88334B6BD7" box="[743,885,1480,1504]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 3283</specimenCode>
)
<specimenCount id="9D3948E2FF817E0EB3E56B88338A6BD7" box="[902,948,1479,1504]" pageId="31" pageNumber="31" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB3A76B88342B6BD7" box="[964,1045,1480,1504]" country="USA" httpUri="http://grbio.org/cool/m0b5-nwmg" name="Ambrose Monell Cryo Collection, American Museum of Natural History" pageId="31" pageNumber="31">AMCC</collectionCode>
[
<specimenCode id="DB992B10FF817E0EB4466B8934AA6BD7" box="[1061,1172,1480,1504]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="31" pageNumber="31">LP 15043</specimenCode>
] ex HUJ
<specimenCode id="DB992B10FF817E0EB28F6BA933BA6836" box="[748,900,1513,1537]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 3288</specimenCode>
),
<specimenCount id="9D3948E2FF817E0EB3FE6BA933F16836" box="[925,975,1512,1537]" pageId="31" pageNumber="31" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB3876BA934276836" box="[996,1049,1513,1537]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB4456BA9348E6836" box="[1062,1200,1513,1537]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC3381</specimenCode>
),
<date id="FF81A5ABFF817E0EB2CF684A33216815" box="[684,799,1546,1570]" pageId="31" pageNumber="31" value="2018-03-27">
<collectingDate id="EFC55C43FF817E0EB2CF684A33216815" box="[684,799,1546,1570]" pageId="31" pageNumber="31" value="2018-03-27">27.iii.2018</collectingDate>
</date>
, Y.
<collectorName id="26CAE6BDFF817E0EB327684A33466815" box="[836,888,1546,1570]" pageId="31" pageNumber="31">Olek</collectorName>
and S.
<collectorName id="26CAE6BDFF817E0EB3AB684A33C06815" box="[968,1022,1546,1570]" pageId="31" pageNumber="31">Cain</collectorName>
</materialsCitation>
;
<materialsCitation id="3B578936FF817E0EB46A684A33C768B2" collectingDate="2017-09-15" collectionCode="HUJ" collectorName="Y. Olek &amp; S. Cain" country="Israel" elevation="153" latitude="30.044611" location="Nahal Sha'alav" longLatPrecision="1" longitude="35.10936" pageId="31" pageNumber="31" specimenCode="INVSC 3299" specimenCount="1" specimenCount-male="1" stateProvince="Southern" typeStatus="paratype">
<location id="8EE0D5B0FF817E0EB46A684A349B6815" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB46A684A349B6815" box="[1033,1189,1546,1570]" country="Israel" latitude="30.044611" longLatPrecision="1" longitude="35.10936" name="Nahal Sha'alav" pageId="31" pageNumber="31" stateProvince="Southern">Nahal Shaalav</location>
,
<locationDeviation id="9A329A49FF817E0EB4CC684A343E6874" location="Kibbutz Yahel" pageId="31" pageNumber="31">
<quantity id="4CC72E8EFF817E0EB4CC684A32EF6874" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="31" unit="km" value="4.0">4 km</quantity>
S of Kibbutz Yahel
</locationDeviation>
,
<geoCoordinate id="EE0BE5ACFF817E0EB47D686B34826874" box="[1054,1212,1578,1603]" degrees="30" direction="north" minutes="02" orientation="latitude" pageId="31" pageNumber="31" precision="1" seconds="40.6" value="30.044611">30°0240.6″N</geoCoordinate>
<geoCoordinate id="EE0BE5ACFF817E0EB2CF680C33046853" box="[684,826,1611,1636]" degrees="35" direction="east" minutes="06" orientation="longitude" pageId="31" pageNumber="31" precision="1" seconds="33.7" value="35.10936">35°0633.7″E</geoCoordinate>
,
<quantity id="4CC72E8EFF817E0EB32B680C33AE6853" box="[840,912,1612,1636]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="31" pageNumber="31" unit="m" value="153.0">
<elevation id="00126458FF817E0EB32B680C33AE6853" box="[840,912,1612,1636]" metricMagnitude="2" metricUnit="m" metricValue="1.53" pageId="31" pageNumber="31" unit="m" value="153.0">153 m</elevation>
</quantity>
,
<date id="FF81A5ABFF817E0EB3FD680C34306853" box="[926,1038,1612,1636]" pageId="31" pageNumber="31" value="2017-09-15">
<collectingDate id="EFC55C43FF817E0EB3FD680C34306853" box="[926,1038,1612,1636]" pageId="31" pageNumber="31" value="2017-09-15">15.ix.2017</collectingDate>
</date>
,
<collectorName id="26CAE6BDFF817E0EB47F680D34506853" box="[1052,1134,1612,1636]" pageId="31" pageNumber="31">Y. Olek</collectorName>
and
<collectorName id="26CAE6BDFF817E0EB4CA680C32DC68B2" pageId="31" pageNumber="31">S. Cain</collectorName>
,
<specimenCount id="9D3948E2FF817E0EB28D682D332768B2" box="[750,793,1644,1669]" pageId="31" pageNumber="31" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB344682D336568B2" box="[807,859,1645,1669]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB300682D33CF68B2" box="[867,1009,1645,1669]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 3299</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B578936FF817E0EB467682D337B68FF" collectingDate="1974-10-08" collectionCode="HUJ" country="Israel" elevation="100" latitude="29.966667" location="Qetura" longLatPrecision="1224" longitude="35.05" pageId="31" pageNumber="31" specimenCode="INVSC 2329" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<location id="8EE0D5B0FF817E0EB467682D346E68B2" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB467682D346E68B2" box="[1028,1104,1645,1669]" country="Israel" latitude="29.966667" longLatPrecision="1224" longitude="35.05" name="Qetura" pageId="31" pageNumber="31">Qetura</location>
[
<geoCoordinate id="EE0BE5ACFF817E0EB43D682D348268B2" box="[1118,1212,1644,1669]" degrees="29" direction="north" minutes="58" orientation="latitude" pageId="31" pageNumber="31" precision="925" value="29.966667">29°58N</geoCoordinate>
<geoCoordinate id="EE0BE5ACFF817E0EB2CF68CE33386891" box="[684,774,1677,1702]" degrees="35" direction="east" minutes="03" orientation="longitude" pageId="31" pageNumber="31" precision="925" value="35.05">35°03E</geoCoordinate>
],
<quantity id="4CC72E8EFF817E0EB37868CF335D6891" box="[795,867,1679,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="31" unit="m" value="100.0">
<elevation id="00126458FF817E0EB37868CF335D6891" box="[795,867,1679,1703]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="31" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FF81A5ABFF817E0EB31368CF33F36890" box="[880,973,1679,1703]" pageId="31" pageNumber="31" value="1974-10-08">
<collectingDate id="EFC55C43FF817E0EB31368CF33F36890" box="[880,973,1679,1703]" pageId="31" pageNumber="31" value="1974-10-08">8.x.1974</collectingDate>
</date>
, M.
<collectingCountry id="F328C3FBFF817E0EB46268CE34006891" box="[1025,1086,1678,1702]" name="Israel" pageId="31" pageNumber="31">Israel</collectingCountry>
,
<specimenCount id="9D3948E2FF817E0EB42868CF34466891" box="[1099,1144,1677,1703]" pageId="31" pageNumber="31" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED2E1BAEFF817E0EB4EB68CF34826890" box="[1160,1212,1679,1703]" country="Israel" lsid="urn:lsid:biocol.org:col:15399" name="Hebrew University" pageId="31" pageNumber="31" type="Herbarium">HUJ</collectionCode>
<specimenCode id="DB992B10FF817E0EB2CF68F0330068FF" box="[684,830,1712,1736]" collectionCode="INVSC" pageId="31" pageNumber="31">INVSC 2329</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B578936FF817E0EB33168F0348268F0" box="[850,1212,1710,1736]" country="Israel" latitude="30.033333" location="Yahel" pageId="31" pageNumber="31" specimenCount="1" typeStatus="paratype">
S of
<location id="8EE0D5B0FF817E0EB3EB68EF33F868F0" LSID="urn:lsid:plazi:treatment:0396327DFF817E0EB34F6D2B348268F0:8EE0D5B0FF817E0EB3EB68EF33F868F0" box="[904,966,1711,1735]" country="Israel" latitude="30.033333" name="Yahel" pageId="31" pageNumber="31">Yahel</location>
, sand dunes,
<geoCoordinate id="EE0BE5ACFF817E0EB40268F0348268F0" box="[1121,1212,1710,1736]" degrees="30" direction="north" minutes="02" orientation="latitude" pageId="31" pageNumber="31" precision="925" value="30.033333">30°02N</geoCoordinate>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>