treatments-xml/data/03/96/2D/03962D077778424CFF49FF16FC4EFA8B.xml

<document id="2215E25B75766F3D27C95D5F155C1F7B" ID-DOI="10.5281/zenodo.199333" ID-GBIF-Dataset="db8f4069-a3c7-4b2b-9d03-46f9826e17c0" ID-ISSN="1175-5326" ID-Zenodo-Dep="199333" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460334135379" checkinUser="plazi" docAuthor="Reboleira, Ana Sofia P. S., Zaragoza, Juan A., Gonçalves, Fernando &amp; Oromí, Pedro" docDate="2010" docId="03962D077778424CFF49FF16FC4EFA8B" docLanguage="en" docName="zt02681p019.pdf" docOrigin="Zootaxa 2681" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Titanobochica magna Zaragoza &amp; Reboleira, sp. nov." docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FFAF557F777C4241FFDEFF81FFB3FFBC" masterDocTitle="Titanobochica, surprising discovery of a new cave-dwelling genus from southern Portugal (Arachnida: Pseudoscorpiones: Bochicidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="5" updateTime="1698259348710" updateUser="plazi">
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<mods:title id="9DFF32EC1896BCCF82293737BC9CE94F">Titanobochica, surprising discovery of a new cave-dwelling genus from southern Portugal (Arachnida: Pseudoscorpiones: Bochicidae)</mods:title>
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<mods:namePart id="B7F0481EFCDA23B46C916B20EECCD256">Reboleira, Ana Sofia P. S.</mods:namePart>
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<mods:namePart id="B2C0BF9DE0B95D58458018742A699E2E">Zaragoza, Juan A.</mods:namePart>
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<mods:namePart id="1118AAE8B58E59E98831BE7E9AFDFBCE">Oromí, Pedro</mods:namePart>
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<taxonomicName id="4C3FE79277784245FF49FF16FD0EFF0D" ID-CoL="577Y9" authority="Zaragoza &amp; Reboleira" authorityName="Zaragoza &amp; Reboleira" box="[151,701,151,177]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="magna" status="sp. nov.">
<emphasis id="B94B400377784245FF49FF16FE2DFF0D" bold="true" box="[151,414,151,177]" italics="true" pageId="4" pageNumber="5">Titanobochica magna</emphasis>
Zaragoza &amp; Reboleira
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(
<figureCitation id="1304809477784245FF41FF3CFEA5FF6B" box="[159,278,189,215]" captionStart-0="FIGURE 2" captionStart-1="FIGURES 3 – 12" captionStart-2="FIGURES 13 – 17" captionStart-3="FIGURES 18 – 23" captionStart-4="FIGURES 24 – 27" captionStart-5="FIGURES 28 – 31" captionStartId-0="4.[151,260,1596,1620]" captionStartId-1="8.[151,273,1846,1870]" captionStartId-2="9.[151,269,1880,1904]" captionStartId-3="10.[151,269,1950,1974]" captionStartId-4="11.[151,270,1919,1943]" captionStartId-5="12.[151,270,1797,1821]" captionTargetBox-0="[188,1398,642,1574]" captionTargetBox-1="[232,1360,200,1810]" captionTargetBox-2="[205,1386,203,1839]" captionTargetBox-3="[210,1361,194,1922]" captionTargetBox-4="[176,1411,194,1896]" captionTargetBox-5="[151,1436,194,1773]" captionTargetId-0="figure@4.[188,1398,642,1574]" captionTargetId-1="figure@8.[221,1369,197,1822]" captionTargetId-2="figure@9.[191,1389,194,1859]" captionTargetId-3="figure@10.[200,1370,194,1929]" captionTargetId-4="figure@11.[176,1412,194,1896]" captionTargetId-5="figure@12.[151,1436,194,1775]" captionTargetPageId-0="4" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionTargetPageId-4="11" captionTargetPageId-5="12" captionText-0="FIGURE 2. Titanobochica magna gen. nov., sp. nov., general appearance of living specimen from Algarão do Remexido." captionText-1="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." captionText-2="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." captionText-3="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." captionText-4="FIGURES 24 – 27. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 24. carapace; 25. opisthosoma, tergite VII partially teratological; 26. chelicerae; 27. distal portion of fingers of left chelicera. Scale bars (mm): 0.12 (Fig. 27), 0.30 (Fig 26), 1.00 (Figs 24, 25)." captionText-5="FIGURES 28 – 31. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 28. fingers of the left chela, dorsal view, A: glandular pore magnified; 29. tip of the chelal movable finger, with venedens, A: magnification of a tooth; 30. right chelal hand, partial dorsal view; 31. leg IV, detail of claws and subterminal feathery setae. Scale bars (mm): 0.009 (Fig. 28 A), 0.01 (Fig. 29 A), 0.075 (Figs 29, 31), 0.75 (Fig. 30), 1.00 (Fig. 28)." httpUri-0="https://zenodo.org/record/199335/files/figure.png" httpUri-1="https://zenodo.org/record/199336/files/figure.png" httpUri-2="https://zenodo.org/record/199337/files/figure.png" httpUri-3="https://zenodo.org/record/199338/files/figure.png" httpUri-4="https://zenodo.org/record/199339/files/figure.png" httpUri-5="https://zenodo.org/record/199340/files/figure.png" pageId="4" pageNumber="5">Figs 2–31</figureCitation>
)
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material.
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Male 
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: 
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: 
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, Algarve, Algarão do Remexido, 
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(
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: 
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: 2D (
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, 
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, 
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(
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: 
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(
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); Gruta da Senhora, 
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: 
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<date id="FF81BAD177784245FD69FD8CFC13FD9B" box="[695,928,525,551]" pageId="4" pageNumber="5" value="2009-12-29">29 December 2009</date>
: 
<specimenCount id="9D39579877784245FC73FD8CFC4DFD9B" box="[941,1022,525,551]" pageId="4" pageNumber="5" type="male">1 male</specimenCount>
(
<collectionCode id="ED2E04D477784245FBD0FD8CFBD7FD9B" box="[1038,1124,525,551]" httpUri="http://grbio.org/cool/x2t0-vyyc" name="Departamento de Zoologia, Universidad de La Laguna" pageId="4" pageNumber="5">DZUL</collectionCode>
); all collected by A.S.P.S. Reboleira.
</paragraph>
</subSubSection>
<caption id="DF40CC9977784245FF49F9BDFEBFF9CF" httpUri="https://zenodo.org/record/199335/files/figure.png" pageId="4" pageNumber="5" targetBox="[188,1398,642,1574]" targetPageId="4">
<paragraph id="8B809C1177784245FF49F9BDFEBFF9CF" blockId="4.[151,1436,1596,1651]" pageId="4" pageNumber="5">
<emphasis id="B94B400377784245FF49F9BDFE91F9E8" bold="true" box="[151,290,1596,1620]" pageId="4" pageNumber="5">FIGURE 2.</emphasis>
<taxonomicName id="4C3FE79277784245FEF4F9BDFD97F9EF" box="[298,548,1596,1619]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B400377784245FEF4F9BDFD97F9EF" box="[298,548,1596,1619]" italics="true" pageId="4" pageNumber="5">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B400377784245FDF2F9BDFCB5F9E8" bold="true" box="[556,774,1596,1620]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A278FD7877784245FDF2F9BDFD29F9E8" box="[556,666,1596,1620]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, general appearance of living specimen from Algarão do Remexido.
</paragraph>
</caption>
<subSubSection id="C325CF9A77784245FF1BF920FDA7F900" box="[197,532,1697,1724]" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8B809C1177784245FF1BF920FDA7F900" blockId="4.[151,1437,1697,1910]" box="[197,532,1697,1724]" pageId="4" pageNumber="5">
<emphasis id="B94B400377784245FF1BF920FEF6F907" bold="true" box="[197,325,1697,1723]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C325CF9A77784245FF1BF946FA99F95D" box="[197,1322,1735,1761]" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8B809C1177784245FF1BF946FA99F95D" blockId="4.[151,1437,1697,1910]" box="[197,1322,1735,1761]" pageId="4" pageNumber="5">
<emphasis id="B94B400377784245FF1BF946FEE1F95D" bold="true" box="[197,338,1735,1761]" pageId="4" pageNumber="5">Etymology.</emphasis>
From the Latin adjective 
<emphasis id="B94B400377784245FD5FF946FC82F95C" box="[641,817,1735,1760]" italics="true" pageId="4" pageNumber="5">magnus,-a,-um</emphasis>
, meaning big, in reference to its large size.
</paragraph>
</subSubSection>
<subSubSection id="C325CF9A7778424CFF1BF96DFCB0FDCE" lastPageId="13" lastPageNumber="14" pageId="4" pageNumber="5" type="description">
<paragraph id="8B809C1177784245FF1BF96DFC6AF890" blockId="4.[151,1437,1697,1910]" pageId="4" pageNumber="5">
<emphasis id="B94B400377784245FF1BF96DFED6F8BA" bold="true" box="[197,357,1772,1798]" pageId="4" pageNumber="5">Description.</emphasis>
The data correspond to the male 
<typeStatus id="548422B377784245FCD6F96DFCCBF8BA" box="[776,888,1772,1798]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
, followed by 
<typeStatus id="548422B377784245FBFDF96DFB3EF8BA" box="[1059,1165,1772,1798]" pageId="4" pageNumber="5" type="paratype">paratype</typeStatus>
males and females in parentheses. Measurements and ratios of adults and nymphs in 
<tableCitation id="C6BDA9AA77784245FCA5F893FC66F890" box="[891,981,1810,1836]" captionStart="TABLE 2" captionStartId="5.[151,240,375,399]" captionText="TABLE 2. Titanobochica magna gen. nov., sp. nov.: measurements and ratios. M: male; F: female; T: tritonymph., D:" pageId="4" pageNumber="5">Table 2</tableCitation>
.
</paragraph>
<paragraph id="8B809C1177784245FF1BF8B6FB3BF8CA" blockId="4.[151,1437,1697,1910]" pageId="4" pageNumber="5">
A large pseudoscorpion (habitus: 
<figureCitation id="1304809477784245FDA2F8B6FD7FF8ED" box="[636,716,1847,1873]" captionStart="FIGURE 2" captionStartId="4.[151,260,1596,1620]" captionTargetBox="[188,1398,642,1574]" captionTargetId="figure@4.[188,1398,642,1574]" captionTargetPageId="4" captionText="FIGURE 2. Titanobochica magna gen. nov., sp. nov., general appearance of living specimen from Algarão do Remexido." httpUri="https://zenodo.org/record/199335/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
), body length about 
<quantity id="4CC731F477784245FC04F8B6FB90F8ED" box="[986,1059,1847,1873]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="5" unit="mm" value="6.0">6 mm</quantity>
(
<tableCitation id="C6BDA9AA77784245FBEBF8B6FB2FF8ED" box="[1077,1180,1847,1873]" captionStart="TABLE 2" captionStartId="5.[151,240,375,399]" captionText="TABLE 2. Titanobochica magna gen. nov., sp. nov.: measurements and ratios. M: male; F: female; T: tritonymph., D:" pageId="4" pageNumber="5">Table 2</tableCitation>
). Desclerotized and depigmented. Opisthosoma and legs pale. Carapace, chelicerae and pedipalps reddish.
</paragraph>
<paragraph id="8B809C1177794244FF1BFF19FEB6FEFB" blockId="5.[151,1436,152,327]" pageId="5" pageNumber="6">
Carapace (
<figureCitation id="1304809477794244FE9FFF19FE3FFF0E" box="[321,396,152,178]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="5" pageNumber="6">Figs 3</figureCitation>
, 
<figureCitation id="1304809477794244FE47FF19FE09FF0E" box="[409,442,152,178]" captionStart="FIGURES 24 – 27" captionStartId="11.[151,270,1919,1943]" captionTargetBox="[176,1411,194,1896]" captionTargetId="figure@11.[176,1412,194,1896]" captionTargetPageId="11" captionText="FIGURES 24 – 27. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 24. carapace; 25. opisthosoma, tergite VII partially teratological; 26. chelicerae; 27. distal portion of fingers of left chelicera. Scale bars (mm): 0.12 (Fig. 27), 0.30 (Fig 26), 1.00 (Figs 24, 25)." httpUri="https://zenodo.org/record/199339/files/figure.png" pageId="5" pageNumber="6">24</figureCitation>
). Elongate, maximum width at median portion. Without eyes or eyes spots. Epistome lacking (
<figureCitation id="1304809477794244FEDCFF3CFEE3FF6B" box="[258,336,189,215]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
). Chaetotaxy: 40 (M: 33–41; F: 34–40) setae, formula (total number of setae in each row followed by number of lateral setae in parentheses): 6:20(8):8:6 [M: 6:16–20(4–7):5–8:4–7; F: 6:18–21(4– 9):4–7:6–7]. One pair of lyrifissures on each side of pre-ocular zone. Numerous diminutive pores over entire carapace.
</paragraph>
<caption id="DF40CC9977794244FF49FEF6FA2FFE33" box="[151,1436,375,399]" pageId="5" pageNumber="6">
<paragraph id="8B809C1177794244FF49FEF6FA2FFE33" blockId="5.[151,1436,375,1112]" box="[151,1436,375,399]" pageId="5" pageNumber="6">
<emphasis id="B94B400377794244FF49FEF6FEB8FE33" bold="true" box="[151,267,375,399]" pageId="5" pageNumber="6">TABLE 2.</emphasis>
<taxonomicName id="4C3FE79277794244FECDFEF9FE4EFE33" box="[275,509,376,399]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B400377794244FECDFEF9FE4EFE33" box="[275,509,376,399]" italics="true" pageId="5" pageNumber="6">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B400377794244FDDAFEF6FD7CFE33" bold="true" box="[516,719,375,399]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A278FD7877794244FDDAFEF6FDDFFE33" box="[516,620,375,399]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
: measurements and ratios. M: male; F: female; T: tritonymph., D:
</paragraph>
</caption>
<paragraph id="8B809C1177794244FF49FE19FE97FE13" blockId="5.[151,1436,375,1112]" box="[151,292,408,431]" pageId="5" pageNumber="6">deutonymph.</paragraph>
<paragraph id="8B809C1177794244FEB4FE4BFAACFE61" blockId="5.[151,1436,375,1112]" box="[362,1311,458,477]" pageId="5" pageNumber="6">
M 
<typeStatus id="548422B377794244FE5CFE4BFE7FFE61" box="[386,460,458,477]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
M 
<typeStatus id="548422B377794244FDF1FE4BFD32FE61" box="[559,641,458,477]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
F 
<typeStatus id="548422B377794244FCD3FE4BFCECFE61" box="[781,863,458,477]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
T 
<typeStatus id="548422B377794244FC33FE4BFB8DFE61" box="[1005,1086,458,477]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
D 
<typeStatus id="548422B377794244FB13FE4BFAACFE61" box="[1229,1311,458,477]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
</paragraph>
<paragraph id="8B809C1177794244FF41FE6EFABEFDBE" blockId="5.[151,1436,375,1112]" box="[159,1293,495,514]" pageId="5" pageNumber="6">Body 6.16 4.83–5.76 5.26–6.06 4.56–4.82 3.06–3.10</paragraph>
<paragraph id="8B809C1177794244FF41FD94FAAAFD94" blockId="5.[151,1436,375,1112]" box="[159,1305,533,552]" pageId="5" pageNumber="6">Carapace 1.36× 1.27–1.37× 1.24–1.37× 1.16–1.20× 1.19–1.32×</paragraph>
<paragraph id="8B809C1177794244FEB4FDBBFAC4FDF1" blockId="5.[151,1436,375,1112]" box="[362,1399,570,589]" pageId="5" pageNumber="6">(1.66/1.22) (1.64–1.70/1.20–1.34) (1.66–1.78/1.24–1.40) (1.20–1.34/1.02–1.12) (0.80–1.00/0.67–0.76)</paragraph>
<paragraph id="8B809C1177794244FF41FDDEFF42FDCE" blockId="5.[151,1436,375,1112]" box="[159,241,607,626]" pageId="5" pageNumber="6">Chelicera</paragraph>
<paragraph id="8B809C1177794244FF41FD04FB59FD24" blockId="5.[151,1436,375,1112]" box="[159,1258,645,664]" pageId="5" pageNumber="6">Palm 2.33× 2.27–2.32× 2.04–2.17× 1.95–2.12× 2.00×</paragraph>
<paragraph id="8B809C1177794244FEB4FD2BFAC4FD01" blockId="5.[151,1436,375,1112]" box="[362,1399,682,701]" pageId="5" pageNumber="6">(0.98/0.42) (0.96–1.05/0.42–0.45) (1.00–1.08/0.46–0.50) (0.74–0.89/0.38–0.42) (0.57–0.60/0.29–0.30)</paragraph>
<paragraph id="8B809C1177794244FF41FD4EFABEFD5E" blockId="5.[151,1436,375,1112]" box="[159,1293,719,738]" pageId="5" pageNumber="6">Finger 0.59 0.58–0.61 0.58–0.68 0.45–0.52 0.30–0.36</paragraph>
<paragraph id="8B809C1177794244FF41FD74FF59FCB4" blockId="5.[151,1436,375,1112]" box="[159,234,757,776]" pageId="5" pageNumber="6">Pedipalp</paragraph>
<paragraph id="8B809C1177794244FF41FC9BFAAAFC91" blockId="5.[151,1436,375,1112]" box="[159,1305,794,813]" pageId="5" pageNumber="6">Trochanter 2.00× 1.87–2.10× 1.89–1.96× 1.88–1.95× 1.78–1.94×</paragraph>
<paragraph id="8B809C1177794244FEB4FCBEFAC4FCEE" blockId="5.[151,1436,375,1112]" box="[362,1399,831,850]" pageId="5" pageNumber="6">(1.00/0.50) (1.00–1.13/0.48–0.56) (1.02–1.12/0.54–0.57) (0.75–0.84/0.40–0.43) (0.52–0.55/0.28–0.29)</paragraph>
<paragraph id="8B809C1177794244FF41FCE4FAAAFCC4" blockId="5.[151,1436,375,1112]" box="[159,1305,869,888]" pageId="5" pageNumber="6">Femur 11.04× 10.18–11.24× 10.72–10.98× 7.52–8.19× 7.58–8.38×</paragraph>
<paragraph id="8B809C1177794244FEB4FC0BFAC4FC21" blockId="5.[151,1436,375,1112]" box="[362,1399,906,925]" pageId="5" pageNumber="6">(4.09/0.37) (4.07–4.28/0.37–0.42) (4.21–4.40/0.38–0.40) (2.58–3.12/0.32–0.41) (1.76–1.82/0.21–0.24)</paragraph>
<paragraph id="8B809C1177794244FF41FC2EFAAAFC7E" blockId="5.[151,1436,375,1112]" box="[159,1305,943,962]" pageId="5" pageNumber="6">Patella 9.00× 8.60–9.13× 8.81–9.18× 6.96–7.56× 5.93–6.42×</paragraph>
<paragraph id="8B809C1177794244FEB4FC54FAC4FC54" blockId="5.[151,1436,375,1112]" box="[362,1399,981,1000]" pageId="5" pageNumber="6">(3.94/0.44) (3.94–4.20/0.43–0.47) (4.04–4.32/0.44–0.49) (2.46–2.84/0.35–0.41) (1.54–1.60/0.24–0.27)</paragraph>
<paragraph id="8B809C1177794244FF41FC7BFABEFBB1" blockId="5.[151,1436,375,1112]" box="[159,1293,1018,1037]" pageId="5" pageNumber="6">Patella pedicel 0.62 0.60–0.67 0.65–0.70 0.48–0.57 0.30–0.32</paragraph>
<paragraph id="8B809C1177794244FF41FB9EFAAAFB8E" blockId="5.[151,1436,375,1112]" box="[159,1305,1055,1074]" pageId="5" pageNumber="6">Hand 2.92× 2.55–2.81× 2.68–2.82× 2.26–2.43× 2.45–2.55×</paragraph>
<paragraph id="8B809C1177794247FEB4FBC4FA8EFACE" blockId="5.[151,1436,375,1112]" lastBlockId="6.[151,1437,1256,1991]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
(2.10/0.72) (2.11–2.19/0.76–0.84) (2.08–2.31/0.75/0.82) (1.43–1.78/0.63–0.73) (1.08–1.16/0.44–0.46) Coxal area (
<figureCitation id="13048094777A4247FE8DFB69FE19FABE" box="[339,426,1256,1282]" captionStart="FIGURES 18 – 23" captionStartId="10.[151,269,1950,1974]" captionTargetBox="[210,1361,194,1922]" captionTargetId="figure@10.[200,1370,194,1929]" captionTargetPageId="10" captionText="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="6" pageNumber="7">Fig. 18</figureCitation>
). Apex of pedipalpal coxa long and acute, with 2 distal setae (one female with 3 setae on one apex lobe and 2 on the other). Pedipalpal coxa with 15–19 setae (M: 15–19; F: 14–19), pedal coxa I: 21–24 (M: 18–26; F: 19–24), II: 18–19 (M: 15–21; F: 16–20), III: 16 (M: 12–20; F: 13–17), IV: 25 (M: 19–26; F: 17–26). Anterolateral process of coxa I rounded and moderately prominent; medial process absent.
</paragraph>
<paragraph id="8B809C1177794244FF41FBEBFA2FF90D" pageId="5" pageNumber="6">
<table id="F93F6EB17779BDBEFF49FBEBFA2FF90D" box="[151,1436,1130,1713]" gridcols="6" gridrows="6" pageId="5" pageNumber="6">
<tr id="350F9E537779BDBEFF49FBEBFA2FFBC1" box="[151,1436,1130,1149]" gridrow="0" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49FBEBFEF3FBC1" box="[151,320,1130,1149]" gridcol="0" gridrow="0" pageId="5" pageNumber="6">Hand pedicel</th>
<th id="76DEF72F7779BDBEFEB4FBEBFE7AFBC1" box="[362,457,1130,1149]" gridcol="1" gridrow="0" pageId="5" pageNumber="6">0.30</th>
<th id="76DEF72F7779BDBEFDC9FBEBFD65FBC1" box="[535,726,1130,1149]" gridcol="2" gridrow="0" pageId="5" pageNumber="6">0.30–0.32</th>
<th id="76DEF72F7779BDBEFD22FBEBFC08FBC1" box="[764,955,1130,1149]" gridcol="3" gridrow="0" pageId="5" pageNumber="6">0.31–0.32</th>
<th id="76DEF72F7779BDBEFC05FBEBFB29FBC1" box="[987,1178,1130,1149]" gridcol="4" gridrow="0" pageId="5" pageNumber="6">0.18–0.22</th>
<th id="76DEF72F7779BDBEFB66FBEBFA2FFBC1" box="[1208,1436,1130,1149]" gridcol="5" gridrow="0" pageId="5" pageNumber="6">0.15</th>
</tr>
<tr id="350F9E537779BDBEFF49FB11FA2FFB51" box="[151,1436,1168,1261]" gridrow="1" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49FB11FEF3FB51" box="[151,320,1168,1261]" gridcol="0" gridrow="1" pageId="5" pageNumber="6">Fixed finger Movable finger Chela</th>
<td id="76DEF72F7779BDBEFEB4FB11FE7AFB51" box="[362,457,1168,1261]" gridcol="1" gridrow="1" pageId="5" pageNumber="6">3.72 3.25 8.00×</td>
<td id="76DEF72F7779BDBEFDC9FB11FD65FB51" box="[535,726,1168,1261]" gridcol="2" gridrow="1" pageId="5" pageNumber="6">3.66–3.82 3.30–3.42 6.98–7.53×</td>
<td id="76DEF72F7779BDBEFD22FB11FC08FB51" box="[764,955,1168,1261]" gridcol="3" gridrow="1" pageId="5" pageNumber="6">3.83–4.08 3.46–3.68 7.37–7.73×</td>
<td id="76DEF72F7779BDBEFC05FB11FB29FB51" box="[987,1178,1168,1261]" gridcol="4" gridrow="1" pageId="5" pageNumber="6">2.69–3.09 2.33–2.70 6.29–6.51×</td>
<td id="76DEF72F7779BDBEFB66FB11FA2FFB51" box="[1208,1436,1168,1261]" gridcol="5" gridrow="1" pageId="5" pageNumber="6">1.83–1.92 1.65–1.66 6.61–6.73×</td>
</tr>
<tr id="350F9E537779BDBEFF49FA81FA2FFAE1" box="[151,1436,1280,1373]" gridrow="2" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49FA81FEF3FAE1" box="[151,320,1280,1373]" gridcol="0" gridrow="2" pageId="5" pageNumber="6">Chela/carapace Chela/femur</th>
<td id="76DEF72F7779BDBEFEB4FA81FE7AFAE1" box="[362,457,1280,1373]" gridcol="1" gridrow="2" pageId="5" pageNumber="6">(5.76/0.72) 3.47× 1.42×</td>
<td id="76DEF72F7779BDBEFDC9FA81FD65FAE1" box="[535,726,1280,1373]" gridcol="2" gridrow="2" pageId="5" pageNumber="6">(5.72–5.96/0.76–0.84) 3.45–3.59× 1.37–1.44×</td>
<td id="76DEF72F7779BDBEFD22FA81FC08FAE1" box="[764,955,1280,1373]" gridcol="3" gridrow="2" pageId="5" pageNumber="6">(5.80–6.29/0.75–0.82) 3.41–3.64× 1.38–1.43×</td>
<td id="76DEF72F7779BDBEFC05FA81FB29FAE1" box="[987,1178,1280,1373]" gridcol="4" gridrow="2" pageId="5" pageNumber="6">(4.09–4.77/0.63–0.73) 3.20–3.67× 1.50–1.59×</td>
<td id="76DEF72F7779BDBEFB66FA81FA2FFAE1" box="[1208,1436,1280,1373]" gridcol="5" gridrow="2" pageId="5" pageNumber="6">(2.91–3.06/0.44–0.46) 3.06–3.64× 1.65–1.68×</td>
</tr>
<tr id="350F9E537779BDBEFF49FAF1FA2FFA71" box="[151,1436,1392,1485]" gridrow="3" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49FAF1FEF3FA71" box="[151,320,1392,1485]" gridcol="0" gridrow="3" pageId="5" pageNumber="6">Femur/carapace Femur/fixed finger Femur/patella</th>
<td id="76DEF72F7779BDBEFEB4FAF1FE7AFA71" box="[362,457,1392,1485]" gridcol="1" gridrow="3" pageId="5" pageNumber="6">2.45× 1.09× 1.03×</td>
<td id="76DEF72F7779BDBEFDC9FAF1FD65FA71" box="[535,726,1392,1485]" gridcol="2" gridrow="3" pageId="5" pageNumber="6">2.45–2.55× 1.08–1.13× 1.02–1.05×</td>
<td id="76DEF72F7779BDBEFD22FAF1FC08FA71" box="[764,955,1392,1485]" gridcol="3" gridrow="3" pageId="5" pageNumber="6">2.47–2.58× 1.08–1.10× 1.02–1.04×</td>
<td id="76DEF72F7779BDBEFC05FAF1FB29FA71" box="[987,1178,1392,1485]" gridcol="4" gridrow="3" pageId="5" pageNumber="6">2.02–2.45× 0.96–1.01× 1.05–1.10×</td>
<td id="76DEF72F7779BDBEFB66FAF1FA2FFA71" box="[1208,1436,1392,1485]" gridcol="5" gridrow="3" pageId="5" pageNumber="6">1.82–2.20× 0.95–0.96× 1.14×</td>
</tr>
<tr id="350F9E537779BDBEFF49FA61FA2FF981" box="[151,1436,1504,1597]" gridrow="4" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49FA61FEF3F981" box="[151,320,1504,1597]" gridcol="0" gridrow="4" pageId="5" pageNumber="6">Patella/hand Fixed Finger/hand Leg I</th>
<td id="76DEF72F7779BDBEFEB4FA61FE7AF981" box="[362,457,1504,1597]" gridcol="1" gridrow="4" pageId="5" pageNumber="6">1.81× 1.77×</td>
<td id="76DEF72F7779BDBEFDC9FA61FD65F981" box="[535,726,1504,1597]" gridcol="2" gridrow="4" pageId="5" pageNumber="6">1.82–1.93× 1.72–1.79×</td>
<td id="76DEF72F7779BDBEFD22FA61FC08F981" box="[764,955,1504,1597]" gridcol="3" gridrow="4" pageId="5" pageNumber="6">1.87–1.94× 1.77–1.84×</td>
<td id="76DEF72F7779BDBEFC05FA61FB29F981" box="[987,1178,1504,1597]" gridcol="4" gridrow="4" pageId="5" pageNumber="6">1.60–1.72× 1.73–1.88×</td>
<td id="76DEF72F7779BDBEFB66FA61FA2FF981" box="[1208,1436,1504,1597]" gridcol="5" gridrow="4" pageId="5" pageNumber="6">1.38–1.43× 1.66–1.70×</td>
</tr>
<tr id="350F9E537779BDBEFF49F9D1FA2FF90D" box="[151,1436,1616,1713]" gridrow="5" pageId="5" pageNumber="6">
<th id="76DEF72F7779BDBEFF49F9D1FEF3F90D" box="[151,320,1616,1713]" gridcol="0" gridrow="5" pageId="5" pageNumber="6">Trochanter</th>
<td id="76DEF72F7779BDBEFEB4F9D1FE7AF90D" box="[362,457,1616,1713]" gridcol="1" gridrow="5" pageId="5" pageNumber="6">1.68× (0.61/0.36)</td>
<td id="76DEF72F7779BDBEFDC9F9D1FD65F90D" box="[535,726,1616,1713]" gridcol="2" gridrow="5" pageId="5" pageNumber="6">1.63–1.84× (0.61–0.68/0.37–0.38)</td>
<td id="76DEF72F7779BDBEFD22F9D1FC08F90D" box="[764,955,1616,1713]" gridcol="3" gridrow="5" pageId="5" pageNumber="6">1.68–1.76× (0.62–0.70/0.37–0.40)</td>
<td id="76DEF72F7779BDBEFC05F9D1FB29F90D" box="[987,1178,1616,1713]" gridcol="4" gridrow="5" pageId="5" pageNumber="6">1.59–1.64× (0.46–0.50/0.28–0.34)</td>
<td id="76DEF72F7779BDBEFB66F9D1FA2FF90D" box="[1208,1436,1616,1713]" gridcol="5" gridrow="5" pageId="5" pageNumber="6">1.64–1.65× (0.34/0.21) continued next page</td>
</tr>
</table>
</paragraph>
<paragraph id="8B809C11777A4247FF41FF1EFAAAFB14" pageId="6" pageNumber="7">
<table id="F93F6EB1777ABDBEFF41FF1EFAC4FB14" box="[159,1399,159,1192]" gridcols="5" gridrows="28" pageId="6" pageNumber="7">
<tr id="350F9E53777ABDBEFF41FF1EFAC4FF0B" box="[159,1399,159,183]" gridrow="0" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DEF72F777ABDBEFF41FF1EFE7FFF0B" box="[159,460,159,183]" gridcol="0" gridrow="0" pageId="6" pageNumber="7">
<emphasis id="B94B4003777A4247FF41FF1EFEA3FF0B" bold="true" box="[159,272,159,183]" pageId="6" pageNumber="7">TABLE 2.</emphasis>
(continued)
</th>
</tr>
<tr id="350F9E53777ABDBEFF41FF4BFAC4FF61" box="[159,1399,202,221]" gridrow="1" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FF4BFE7FFF61" box="[159,460,202,221]" gridcol="0" gridrow="1" pageId="6" pageNumber="7">M holotype</th>
<td id="76DEF72F777ABDBEFDC9FF4BFD65FF61" box="[535,726,202,221]" gridcol="1" gridrow="1" pageId="6" pageNumber="7">M paratypes</td>
<td id="76DEF72F777ABDBEFD22FF4BFC08FF61" box="[764,955,202,221]" gridcol="2" gridrow="1" pageId="6" pageNumber="7">F paratypes</td>
<td id="76DEF72F777ABDBEFC05FF4BFB29FF61" box="[987,1178,202,221]" gridcol="3" gridrow="1" pageId="6" pageNumber="7">T paratypes</td>
<td id="76DEF72F777ABDBEFB66FF4BFAC4FF61" box="[1208,1399,202,221]" gridcol="4" gridrow="1" pageId="6" pageNumber="7">D paratypes</td>
</tr>
<tr id="350F9E53777ABDBEFF41FF6EFAC4FEBE" box="[159,1399,239,258]" gridrow="2" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FF6EFE7FFEBE" box="[159,460,239,258]" gridcol="0" gridrow="2" pageId="6" pageNumber="7">Femur 7.56×</th>
<td id="76DEF72F777ABDBEFDC9FF6EFD65FEBE" box="[535,726,239,258]" gridcol="1" gridrow="2" pageId="6" pageNumber="7">7.48–8.14×</td>
<td id="76DEF72F777ABDBEFD22FF6EFC08FEBE" box="[764,955,239,258]" gridcol="2" gridrow="2" pageId="6" pageNumber="7">7.57–7.77×</td>
<td id="76DEF72F777ABDBEFC05FF6EFB29FEBE" box="[987,1178,239,258]" gridcol="3" gridrow="2" pageId="6" pageNumber="7">6.20–6.93×</td>
<td id="76DEF72F777ABDBEFB66FF6EFAC4FEBE" box="[1208,1399,239,258]" gridcol="4" gridrow="2" pageId="6" pageNumber="7">5.90×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FE94FAC4FE94" box="[159,1399,277,296]" gridrow="3" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FE94FE7FFE94" box="[159,460,277,296]" gridcol="0" gridrow="3" pageId="6" pageNumber="7">(1.97/0.26)</th>
<td id="76DEF72F777ABDBEFDC9FE94FD65FE94" box="[535,726,277,296]" gridcol="1" gridrow="3" pageId="6" pageNumber="7">(1.98–2.10/0.25–0.27)</td>
<td id="76DEF72F777ABDBEFD22FE94FC08FE94" box="[764,955,277,296]" gridcol="2" gridrow="3" pageId="6" pageNumber="7">(2.02–2.14/0.26–0.28)</td>
<td id="76DEF72F777ABDBEFC05FE94FB29FE94" box="[987,1178,277,296]" gridcol="3" gridrow="3" pageId="6" pageNumber="7">(1.26–1.49/0.20–0.24)</td>
<td id="76DEF72F777ABDBEFB66FE94FAC4FE94" box="[1208,1399,277,296]" gridcol="4" gridrow="3" pageId="6" pageNumber="7">(0.83–0.92/0.14–0.16)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FEBBFAC4FEF1" box="[159,1399,314,333]" gridrow="4" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FEBBFE7FFEF1" box="[159,460,314,333]" gridcol="0" gridrow="4" pageId="6" pageNumber="7">Patella 3.23×</th>
<td id="76DEF72F777ABDBEFDC9FEBBFD65FEF1" box="[535,726,314,333]" gridcol="1" gridrow="4" pageId="6" pageNumber="7">3.19–3.36×</td>
<td id="76DEF72F777ABDBEFD22FEBBFC08FEF1" box="[764,955,314,333]" gridcol="2" gridrow="4" pageId="6" pageNumber="7">3.20–3.36×</td>
<td id="76DEF72F777ABDBEFC05FEBBFB29FEF1" box="[987,1178,314,333]" gridcol="3" gridrow="4" pageId="6" pageNumber="7">2.81–3.06×</td>
<td id="76DEF72F777ABDBEFB66FEBBFAC4FEF1" box="[1208,1399,314,333]" gridcol="4" gridrow="4" pageId="6" pageNumber="7">1.83–1.96×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FEDEFAC4FECE" box="[159,1399,351,370]" gridrow="5" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FEDEFE7FFECE" box="[159,460,351,370]" gridcol="0" gridrow="5" pageId="6" pageNumber="7">(0.71/0.22)</th>
<td id="76DEF72F777ABDBEFDC9FEDEFD65FECE" box="[535,726,351,370]" gridcol="1" gridrow="5" pageId="6" pageNumber="7">(0.74–0.79/0.22–0.25)</td>
<td id="76DEF72F777ABDBEFD22FEDEFC08FECE" box="[764,955,351,370]" gridcol="2" gridrow="5" pageId="6" pageNumber="7">(0.74–0.77/0.22–0.24)</td>
<td id="76DEF72F777ABDBEFC05FEDEFB29FECE" box="[987,1178,351,370]" gridcol="3" gridrow="5" pageId="6" pageNumber="7">(0.50–0.59/0.17–0.21)</td>
<td id="76DEF72F777ABDBEFB66FEDEFAC4FECE" box="[1208,1399,351,370]" gridcol="4" gridrow="5" pageId="6" pageNumber="7">(0.33–0.37/0.18–0.19)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FE04FAC4FE24" box="[159,1399,389,408]" gridrow="6" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FE04FE7FFE24" box="[159,460,389,408]" gridcol="0" gridrow="6" pageId="6" pageNumber="7">Tibia 13.51×</th>
<td id="76DEF72F777ABDBEFDC9FE04FD65FE24" box="[535,726,389,408]" gridcol="1" gridrow="6" pageId="6" pageNumber="7">12.46–14.10×</td>
<td id="76DEF72F777ABDBEFD22FE04FC08FE24" box="[764,955,389,408]" gridcol="2" gridrow="6" pageId="6" pageNumber="7">12.93–13.55×</td>
<td id="76DEF72F777ABDBEFC05FE04FB29FE24" box="[987,1178,389,408]" gridcol="3" gridrow="6" pageId="6" pageNumber="7">9.33–10.69×</td>
<td id="76DEF72F777ABDBEFB66FE04FAC4FE24" box="[1208,1399,389,408]" gridcol="4" gridrow="6" pageId="6" pageNumber="7">8.95–9.46×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FE2BFAC4FE01" box="[159,1399,426,445]" gridrow="7" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FE2BFE7FFE01" box="[159,460,426,445]" gridcol="0" gridrow="7" pageId="6" pageNumber="7">(2.00/0.15)</th>
<td id="76DEF72F777ABDBEFDC9FE2BFD65FE01" box="[535,726,426,445]" gridcol="1" gridrow="7" pageId="6" pageNumber="7">(1.97–2.02/0.14–0.17)</td>
<td id="76DEF72F777ABDBEFD22FE2BFC08FE01" box="[764,955,426,445]" gridcol="2" gridrow="7" pageId="6" pageNumber="7">(1.94–2.10/0.15–0.16)</td>
<td id="76DEF72F777ABDBEFC05FE2BFB29FE01" box="[987,1178,426,445]" gridcol="3" gridrow="7" pageId="6" pageNumber="7">(1.12–1.46/0.12–0.15)</td>
<td id="76DEF72F777ABDBEFB66FE2BFAC4FE01" box="[1208,1399,426,445]" gridcol="4" gridrow="7" pageId="6" pageNumber="7">(0.80–0.85/0.08–0.10)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FE4EFAC4FE5E" box="[159,1399,463,482]" gridrow="8" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FE4EFE7FFE5E" box="[159,460,463,482]" gridcol="0" gridrow="8" pageId="6" pageNumber="7">Metatarsus 1.68×</th>
<td id="76DEF72F777ABDBEFDC9FE4EFD65FE5E" box="[535,726,463,482]" gridcol="1" gridrow="8" pageId="6" pageNumber="7">1.63–1.79×</td>
<td id="76DEF72F777ABDBEFD22FE4EFC08FE5E" box="[764,955,463,482]" gridcol="2" gridrow="8" pageId="6" pageNumber="7">1.57–1.80×</td>
<td id="76DEF72F777ABDBEFC05FE4EFB29FE5E" box="[987,1178,463,482]" gridcol="3" gridrow="8" pageId="6" pageNumber="7">1.55–1.67×</td>
<td id="76DEF72F777ABDBEFB66FE4EFAC4FE5E" box="[1208,1399,463,482]" gridcol="4" gridrow="8" pageId="6" pageNumber="7">1.67–1.75×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FE74FAC4FDB4" box="[159,1399,501,520]" gridrow="9" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FE74FE7FFDB4" box="[159,460,501,520]" gridcol="0" gridrow="9" pageId="6" pageNumber="7">(0.22/0.13)</th>
<td id="76DEF72F777ABDBEFDC9FE74FD65FDB4" box="[535,726,501,520]" gridcol="1" gridrow="9" pageId="6" pageNumber="7">(0.21–0.24/0.13–0.14)</td>
<td id="76DEF72F777ABDBEFD22FE74FC08FDB4" box="[764,955,501,520]" gridcol="2" gridrow="9" pageId="6" pageNumber="7">(0.24/0.13–0.16)</td>
<td id="76DEF72F777ABDBEFC05FE74FB29FDB4" box="[987,1178,501,520]" gridcol="3" gridrow="9" pageId="6" pageNumber="7">(0.17–0.21/0.11–0.13)</td>
<td id="76DEF72F777ABDBEFB66FE74FAC4FDB4" box="[1208,1399,501,520]" gridcol="4" gridrow="9" pageId="6" pageNumber="7">(0.14–0.15/0.08–0.09)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FD9BFAC4FD91" box="[159,1399,538,557]" gridrow="10" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FD9BFE7FFD91" box="[159,460,538,557]" gridcol="0" gridrow="10" pageId="6" pageNumber="7">Tarsus 14.17×</th>
<td id="76DEF72F777ABDBEFDC9FD9BFD65FD91" box="[535,726,538,557]" gridcol="1" gridrow="10" pageId="6" pageNumber="7">13.23–14.96×</td>
<td id="76DEF72F777ABDBEFD22FD9BFC08FD91" box="[764,955,538,557]" gridcol="2" gridrow="10" pageId="6" pageNumber="7">13.60–14.07×</td>
<td id="76DEF72F777ABDBEFC05FD9BFB29FD91" box="[987,1178,538,557]" gridcol="3" gridrow="10" pageId="6" pageNumber="7">9.47–10.00×</td>
<td id="76DEF72F777ABDBEFB66FD9BFAC4FD91" box="[1208,1399,538,557]" gridcol="4" gridrow="10" pageId="6" pageNumber="7">10.33–10.50×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FDBEFAC4FDEE" box="[159,1399,575,594]" gridrow="11" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FDBEFE7FFDEE" box="[159,460,575,594]" gridcol="0" gridrow="11" pageId="6" pageNumber="7">(1.70/0.12)</th>
<td id="76DEF72F777ABDBEFDC9FDBEFD65FDEE" box="[535,726,575,594]" gridcol="1" gridrow="11" pageId="6" pageNumber="7">(1.68–1.75/0.12–0.13)</td>
<td id="76DEF72F777ABDBEFD22FDBEFC08FDEE" box="[764,955,575,594]" gridcol="2" gridrow="11" pageId="6" pageNumber="7">(1.74–1.90/0.13–0.14)</td>
<td id="76DEF72F777ABDBEFC05FDBEFB29FDEE" box="[987,1178,575,594]" gridcol="3" gridrow="11" pageId="6" pageNumber="7">(1.12–1.42/0.12–0.15)</td>
<td id="76DEF72F777ABDBEFB66FDBEFAC4FDEE" box="[1208,1399,575,594]" gridcol="4" gridrow="11" pageId="6" pageNumber="7">(0.84–0.93/0.08–0.09)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FDE4FAC4FDC4" box="[159,1399,613,632]" gridrow="12" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FDE4FE7FFDC4" box="[159,460,613,632]" gridcol="0" gridrow="12" pageId="6" pageNumber="7">Femur/patella 2.77×</th>
<td id="76DEF72F777ABDBEFDC9FDE4FD65FDC4" box="[535,726,613,632]" gridcol="1" gridrow="12" pageId="6" pageNumber="7">2.64–2.73×</td>
<td id="76DEF72F777ABDBEFD22FDE4FC08FDC4" box="[764,955,613,632]" gridcol="2" gridrow="12" pageId="6" pageNumber="7">2.73–2.83×</td>
<td id="76DEF72F777ABDBEFC05FDE4FB29FDC4" box="[987,1178,613,632]" gridcol="3" gridrow="12" pageId="6" pageNumber="7">2.42–2.55×</td>
<td id="76DEF72F777ABDBEFB66FDE4FAC4FDC4" box="[1208,1399,613,632]" gridcol="4" gridrow="12" pageId="6" pageNumber="7">2.50–2.51×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FD0BFAC4FD21" box="[159,1399,650,669]" gridrow="13" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FD0BFE7FFD21" box="[159,460,650,669]" gridcol="0" gridrow="13" pageId="6" pageNumber="7">Tarsus/Metatarsus 7.80×</th>
<td id="76DEF72F777ABDBEFDC9FD0BFD65FD21" box="[535,726,650,669]" gridcol="1" gridrow="13" pageId="6" pageNumber="7">7.28–8.25×</td>
<td id="76DEF72F777ABDBEFD22FD0BFC08FD21" box="[764,955,650,669]" gridcol="2" gridrow="13" pageId="6" pageNumber="7">7.25–8.09×</td>
<td id="76DEF72F777ABDBEFC05FD0BFB29FD21" box="[987,1178,650,669]" gridcol="3" gridrow="13" pageId="6" pageNumber="7">6.59–6.76×</td>
<td id="76DEF72F777ABDBEFB66FD0BFAC4FD21" box="[1208,1399,650,669]" gridcol="4" gridrow="13" pageId="6" pageNumber="7">6.00–6.20×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FD2EFAC4FD7E" box="[159,1399,687,706]" gridrow="14" pageId="6" pageNumber="7" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
<th id="76DEF72F777ABDBEFF41FD2EFE7FFD7E" box="[159,460,687,706]" gridcol="0" gridrow="14" pageId="6" pageNumber="7">Leg IV</th>
</tr>
<tr id="350F9E53777ABDBEFF41FD54FAC4FD54" box="[159,1399,725,744]" gridrow="15" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FD54FE7FFD54" box="[159,460,725,744]" gridcol="0" gridrow="15" pageId="6" pageNumber="7">Trochanter 2.65×</th>
<td id="76DEF72F777ABDBEFDC9FD54FD65FD54" box="[535,726,725,744]" gridcol="1" gridrow="15" pageId="6" pageNumber="7">2.43–2.63×</td>
<td id="76DEF72F777ABDBEFD22FD54FC08FD54" box="[764,955,725,744]" gridcol="2" gridrow="15" pageId="6" pageNumber="7">2.55–2.58×</td>
<td id="76DEF72F777ABDBEFC05FD54FB29FD54" box="[987,1178,725,744]" gridcol="3" gridrow="15" pageId="6" pageNumber="7">2.62–2.75×</td>
<td id="76DEF72F777ABDBEFB66FD54FAC4FD54" box="[1208,1399,725,744]" gridcol="4" gridrow="15" pageId="6" pageNumber="7">2.05–2.50×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FD7BFAC4FCB1" box="[159,1399,762,781]" gridrow="16" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FD7BFE7FFCB1" box="[159,460,762,781]" gridcol="0" gridrow="16" pageId="6" pageNumber="7">(0.90/0.34)</th>
<td id="76DEF72F777ABDBEFDC9FD7BFD65FCB1" box="[535,726,762,781]" gridcol="1" gridrow="16" pageId="6" pageNumber="7">(0.82–0.94/0.34–0.36)</td>
<td id="76DEF72F777ABDBEFD22FD7BFC08FCB1" box="[764,955,762,781]" gridcol="2" gridrow="16" pageId="6" pageNumber="7">(0.90–0.98/0.35–0.38)</td>
<td id="76DEF72F777ABDBEFC05FD7BFB29FCB1" box="[987,1178,762,781]" gridcol="3" gridrow="16" pageId="6" pageNumber="7">(0.64–0.80/0.26–0.30)</td>
<td id="76DEF72F777ABDBEFB66FD7BFAC4FCB1" box="[1208,1399,762,781]" gridcol="4" gridrow="16" pageId="6" pageNumber="7">(0.38–0.43/0.17–0.19)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FC9EFAC4FC8E" box="[159,1399,799,818]" gridrow="17" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FC9EFE7FFC8E" box="[159,460,799,818]" gridcol="0" gridrow="17" pageId="6" pageNumber="7">Femur 1.86×</th>
<td id="76DEF72F777ABDBEFDC9FC9EFD65FC8E" box="[535,726,799,818]" gridcol="1" gridrow="17" pageId="6" pageNumber="7">1.67–1.86×</td>
<td id="76DEF72F777ABDBEFD22FC9EFC08FC8E" box="[764,955,799,818]" gridcol="2" gridrow="17" pageId="6" pageNumber="7">1.86–1.93×</td>
<td id="76DEF72F777ABDBEFC05FC9EFB29FC8E" box="[987,1178,799,818]" gridcol="3" gridrow="17" pageId="6" pageNumber="7">1.52–1.73×</td>
<td id="76DEF72F777ABDBEFB66FC9EFAC4FC8E" box="[1208,1399,799,818]" gridcol="4" gridrow="17" pageId="6" pageNumber="7">1.75–1.77×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FCC4FAC4FCE4" box="[159,1399,837,856]" gridrow="18" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FCC4FE7FFCE4" box="[159,460,837,856]" gridcol="0" gridrow="18" pageId="6" pageNumber="7">(0.54/0.29)</th>
<td id="76DEF72F777ABDBEFDC9FCC4FD65FCE4" box="[535,726,837,856]" gridcol="1" gridrow="18" pageId="6" pageNumber="7">(0.50–0.57/0.29–0.31)</td>
<td id="76DEF72F777ABDBEFD22FCC4FC08FCE4" box="[764,955,837,856]" gridcol="2" gridrow="18" pageId="6" pageNumber="7">(0.56–0.60/0.30–0.32)</td>
<td id="76DEF72F777ABDBEFC05FCC4FB29FCE4" box="[987,1178,837,856]" gridcol="3" gridrow="18" pageId="6" pageNumber="7">(0.37–0.46/0.23–0.27)</td>
<td id="76DEF72F777ABDBEFB66FCC4FAC4FCE4" box="[1208,1399,837,856]" gridcol="4" gridrow="18" pageId="6" pageNumber="7">(0.27–0.32/0.15–0.18)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FCEBFAC4FCC1" box="[159,1399,874,893]" gridrow="19" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FCEBFE7FFCC1" box="[159,460,874,893]" gridcol="0" gridrow="19" pageId="6" pageNumber="7">Patella 10.17×</th>
<td id="76DEF72F777ABDBEFDC9FCEBFD65FCC1" box="[535,726,874,893]" gridcol="1" gridrow="19" pageId="6" pageNumber="7">8.94–9.87×</td>
<td id="76DEF72F777ABDBEFD22FCEBFC08FCC1" box="[764,955,874,893]" gridcol="2" gridrow="19" pageId="6" pageNumber="7">9.22–9.98×</td>
<td id="76DEF72F777ABDBEFC05FCEBFB29FCC1" box="[987,1178,874,893]" gridcol="3" gridrow="19" pageId="6" pageNumber="7">7.21–7.91×</td>
<td id="76DEF72F777ABDBEFB66FCEBFAC4FCC1" box="[1208,1399,874,893]" gridcol="4" gridrow="19" pageId="6" pageNumber="7">6.79–6.84×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FC0EFAC4FC1E" box="[159,1399,911,930]" gridrow="20" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FC0EFE7FFC1E" box="[159,460,911,930]" gridcol="0" gridrow="20" pageId="6" pageNumber="7">(3.05/0.30)</th>
<td id="76DEF72F777ABDBEFDC9FC0EFD65FC1E" box="[535,726,911,930]" gridcol="1" gridrow="20" pageId="6" pageNumber="7">(2.95–3.12/0.31–0.33)</td>
<td id="76DEF72F777ABDBEFD22FC0EFC08FC1E" box="[764,955,911,930]" gridcol="2" gridrow="20" pageId="6" pageNumber="7">(3.05–3.21/0.32–0.35)</td>
<td id="76DEF72F777ABDBEFC05FC0EFB29FC1E" box="[987,1178,911,930]" gridcol="3" gridrow="20" pageId="6" pageNumber="7">(1.86–2.24/0.24–0.29)</td>
<td id="76DEF72F777ABDBEFB66FC0EFAC4FC1E" box="[1208,1399,911,930]" gridcol="4" gridrow="20" pageId="6" pageNumber="7">(1.16–1.30/0.17–0.19)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FC34FAC4FC74" box="[159,1399,949,968]" gridrow="21" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FC34FE7FFC74" box="[159,460,949,968]" gridcol="0" gridrow="21" pageId="6" pageNumber="7">Tibia 15.78×</th>
<td id="76DEF72F777ABDBEFDC9FC34FD65FC74" box="[535,726,949,968]" gridcol="1" gridrow="21" pageId="6" pageNumber="7">15.23–16.71×</td>
<td id="76DEF72F777ABDBEFD22FC34FC08FC74" box="[764,955,949,968]" gridcol="2" gridrow="21" pageId="6" pageNumber="7">14.62–15.98×</td>
<td id="76DEF72F777ABDBEFC05FC34FB29FC74" box="[987,1178,949,968]" gridcol="3" gridrow="21" pageId="6" pageNumber="7">11.44–13.09×</td>
<td id="76DEF72F777ABDBEFB66FC34FAC4FC74" box="[1208,1399,949,968]" gridcol="4" gridrow="21" pageId="6" pageNumber="7">10.70–10.83×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FC5BFAC4FC51" box="[159,1399,986,1005]" gridrow="22" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FC5BFE7FFC51" box="[159,460,986,1005]" gridcol="0" gridrow="22" pageId="6" pageNumber="7">(2.84/0.18)</th>
<td id="76DEF72F777ABDBEFDC9FC5BFD65FC51" box="[535,726,986,1005]" gridcol="1" gridrow="22" pageId="6" pageNumber="7">(2.81–2.88/0.17–0.18)</td>
<td id="76DEF72F777ABDBEFD22FC5BFC08FC51" box="[764,955,986,1005]" gridcol="2" gridrow="22" pageId="6" pageNumber="7">(2.84–2.97/0.18–0.20)</td>
<td id="76DEF72F777ABDBEFC05FC5BFB29FC51" box="[987,1178,986,1005]" gridcol="3" gridrow="22" pageId="6" pageNumber="7">(1.67–2.12/0.15–0.16)</td>
<td id="76DEF72F777ABDBEFB66FC5BFAC4FC51" box="[1208,1399,986,1005]" gridcol="4" gridrow="22" pageId="6" pageNumber="7">(1.12–1.22/0.10–0.11)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FC7EFAC4FBAE" box="[159,1399,1023,1042]" gridrow="23" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FC7EFE7FFBAE" box="[159,460,1023,1042]" gridcol="0" gridrow="23" pageId="6" pageNumber="7">Metatarsus 1.70×</th>
<td id="76DEF72F777ABDBEFDC9FC7EFD65FBAE" box="[535,726,1023,1042]" gridcol="1" gridrow="23" pageId="6" pageNumber="7">1.77–1.94×</td>
<td id="76DEF72F777ABDBEFD22FC7EFC08FBAE" box="[764,955,1023,1042]" gridcol="2" gridrow="23" pageId="6" pageNumber="7">1.81–1.94×</td>
<td id="76DEF72F777ABDBEFC05FC7EFB29FBAE" box="[987,1178,1023,1042]" gridcol="3" gridrow="23" pageId="6" pageNumber="7">1.89–2.02×</td>
<td id="76DEF72F777ABDBEFB66FC7EFAC4FBAE" box="[1208,1399,1023,1042]" gridcol="4" gridrow="23" pageId="6" pageNumber="7">2.00×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FBA4FAC4FB84" box="[159,1399,1061,1080]" gridrow="24" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FBA4FE7FFB84" box="[159,460,1061,1080]" gridcol="0" gridrow="24" pageId="6" pageNumber="7">(0.27/0.16)</th>
<td id="76DEF72F777ABDBEFDC9FBA4FD65FB84" box="[535,726,1061,1080]" gridcol="1" gridrow="24" pageId="6" pageNumber="7">(0.28–0.33/0.15–0.17)</td>
<td id="76DEF72F777ABDBEFD22FBA4FC08FB84" box="[764,955,1061,1080]" gridcol="2" gridrow="24" pageId="6" pageNumber="7">(0.30–0.31/0.16–0.17)</td>
<td id="76DEF72F777ABDBEFC05FBA4FB29FB84" box="[987,1178,1061,1080]" gridcol="3" gridrow="24" pageId="6" pageNumber="7">(0.23–0.26/0.12–0.14)</td>
<td id="76DEF72F777ABDBEFB66FBA4FAC4FB84" box="[1208,1399,1061,1080]" gridcol="4" gridrow="24" pageId="6" pageNumber="7">(0.18–0.19/0.09–0.10)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FBCBFAC4FBE1" box="[159,1399,1098,1117]" gridrow="25" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FBCBFE7FFBE1" box="[159,460,1098,1117]" gridcol="0" gridrow="25" pageId="6" pageNumber="7">Tarsus 17.29×</th>
<td id="76DEF72F777ABDBEFDC9FBCBFD65FBE1" box="[535,726,1098,1117]" gridcol="1" gridrow="25" pageId="6" pageNumber="7">17.36–18.66×</td>
<td id="76DEF72F777ABDBEFD22FBCBFC08FBE1" box="[764,955,1098,1117]" gridcol="2" gridrow="25" pageId="6" pageNumber="7">17.97–18.37×</td>
<td id="76DEF72F777ABDBEFC05FBCBFB29FBE1" box="[987,1178,1098,1117]" gridcol="3" gridrow="25" pageId="6" pageNumber="7">12.60–13.53×</td>
<td id="76DEF72F777ABDBEFB66FBCBFAC4FBE1" box="[1208,1399,1098,1117]" gridcol="4" gridrow="25" pageId="6" pageNumber="7">12.56–13.37×</td>
</tr>
<tr id="350F9E53777ABDBEFF41FBF1FAC4FB3F" box="[159,1399,1136,1155]" gridrow="26" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FBF1FE7FFB3F" box="[159,460,1136,1155]" gridcol="0" gridrow="26" pageId="6" pageNumber="7">(2.42/0.14)</th>
<td id="76DEF72F777ABDBEFDC9FBF1FD65FB3F" box="[535,726,1136,1155]" gridcol="1" gridrow="26" pageId="6" pageNumber="7">(2.46–2.54/0.13–0.14)</td>
<td id="76DEF72F777ABDBEFD22FBF1FC08FB3F" box="[764,955,1136,1155]" gridcol="2" gridrow="26" pageId="6" pageNumber="7">(2.57–2.75/0.14–0.15)</td>
<td id="76DEF72F777ABDBEFC05FBF1FB29FB3F" box="[987,1178,1136,1155]" gridcol="3" gridrow="26" pageId="6" pageNumber="7">(1.55–2.01/0.12–0.15)</td>
<td id="76DEF72F777ABDBEFB66FBF1FAC4FB3F" box="[1208,1399,1136,1155]" gridcol="4" gridrow="26" pageId="6" pageNumber="7">(0.18–0.19/0.09–0.10)</td>
</tr>
<tr id="350F9E53777ABDBEFF41FB14FAC4FB14" box="[159,1399,1173,1192]" gridrow="27" pageId="6" pageNumber="7">
<th id="76DEF72F777ABDBEFF41FB14FE7FFB14" box="[159,460,1173,1192]" gridcol="0" gridrow="27" pageId="6" pageNumber="7">Tarsus/Metatarsus 8.64×</th>
<td id="76DEF72F777ABDBEFDC9FB14FD65FB14" box="[535,726,1173,1192]" gridcol="1" gridrow="27" pageId="6" pageNumber="7">7.64–9.20×</td>
<td id="76DEF72F777ABDBEFD22FB14FC08FB14" box="[764,955,1173,1192]" gridcol="2" gridrow="27" pageId="6" pageNumber="7">8.43–8.96×</td>
<td id="76DEF72F777ABDBEFC05FB14FB29FB14" box="[987,1178,1173,1192]" gridcol="3" gridrow="27" pageId="6" pageNumber="7">6.46–7.62×</td>
<td id="76DEF72F777ABDBEFB66FB14FAC4FB14" box="[1208,1399,1173,1192]" gridcol="4" gridrow="27" pageId="6" pageNumber="7">6.28–6.68×</td>
</tr>
</table>
</paragraph>
<paragraph id="8B809C11777A4247FF1BFAFCFB4EF87B" blockId="6.[151,1437,1256,1991]" pageId="6" pageNumber="7">
Opisthosoma (
<figureCitation id="13048094777A4247FEA6FAFCFDBBFA2B" box="[376,520,1405,1431]" captionStart="FIGURES 18 – 23" captionStartId="10.[151,269,1950,1974]" captionTargetBox="[210,1361,194,1922]" captionTargetId="figure@10.[200,1370,194,1929]" captionTargetPageId="10" captionText="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="6" pageNumber="7">Figs 22–23</figureCitation>
, 
<figureCitation id="13048094777A4247FDCBFAFCFD8BFA2B" box="[533,568,1405,1431]" captionStart="FIGURES 24 – 27" captionStartId="11.[151,270,1919,1943]" captionTargetBox="[176,1411,194,1896]" captionTargetId="figure@11.[176,1412,194,1896]" captionTargetPageId="11" captionText="FIGURES 24 – 27. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 24. carapace; 25. opisthosoma, tergite VII partially teratological; 26. chelicerae; 27. distal portion of fingers of left chelicera. Scale bars (mm): 0.12 (Fig. 27), 0.30 (Fig 26), 1.00 (Figs 24, 25)." httpUri="https://zenodo.org/record/199339/files/figure.png" pageId="6" pageNumber="7">25</figureCitation>
). Elongate. Pleural membranes smoothly, longitudinally striate. Some tergites with one or two median subdiscal microsetae; tergal chaetotaxy I–X (totals, followed by number of median microsetae in parentheses): 6: 6: 6: 10(1): 9(2): 9(1): 9(1): 9(1): 10(1): 7 [M: 4–6: 5–7: 5–6: 7–8(0–1): 7–10(1–2): 8–9(1–2): 7–10(1–2): 6–9(0–1): 7–9(0–1): 6–9; F: 4–7: 6–7: 6: 8–9(0–1): 8–12(1–2): 8–12(1–2): 7–12(1–2): 7–10(1–2): 7–9(0–1): 7–8]. Male genital area (
<figureCitation id="13048094777A4247FC82F993FC04F990" box="[860,951,1554,1580]" captionStart="FIGURES 18 – 23" captionStartId="10.[151,269,1950,1974]" captionTargetBox="[210,1361,194,1922]" captionTargetId="figure@10.[200,1370,194,1929]" captionTargetPageId="10" captionText="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="6" pageNumber="7">Fig. 19</figureCitation>
): 18 (22–26) setae on sternite II plus 15 (19–23) small setae on anterior genital opening, posterior genital opening with 15 (12–17) small setae, sternite III with 10 (7–12) discal and 11 (9–12) marginal setae. Female 
<typeStatus id="548422B3777A4247FCA1F9DCFC47F9CB" box="[895,1012,1629,1655]" pageId="6" pageNumber="7" type="paratype">paratypes</typeStatus>
: genital area 18–21 very small setae on sternite II, 13–17 marginal setae and 8–10 small discal setae on sternite III. Sternites IV-VII, in addition to the marginal setae, with one or two rows of discal setae, most reduced to microsetae, resulting sternal plates biseriate or triseriate; sternites VIII-IX with some dispersed discal setae; sternal chaetotaxy IV-X (expressed as discal setae/posterior setae): 9/10: 8/14: 10/12: 9/12: 5/9: 3/11: 10; [M: 9/10: 8/14: 10/12: 9/12: 5/9: 3/11: 10; [M: 9–11/8–14: 7–10/8–12: 7–8/9–14: 5–7/7–12: 4–7/8–10: 3–6/7–11: 0–1/7–10; F: 6–11/6–11: 6–8/9– 15: 6–9/10–14: 6–10/8–13: 5–9/8–13: 2–5/8–13: 0–1/7–10]; segment 
<date id="FF81BAD1777A4247FC34F8BCFB92F8EB" box="[1002,1057,1853,1879]" pageId="6" pageNumber="7">XI 7</date>
setae (7–8), segment XII (anal cone) bears 2 tergal and 3 sternal setae (teratology) (2–2). Tergites and sternites without tactile setae. Stigmata on sternite III (
<figureCitation id="13048094777A4247FE99F809FE2EF81E" box="[327,413,1928,1954]" captionStart="FIGURES 18 – 23" captionStartId="10.[151,269,1950,1974]" captionTargetBox="[210,1361,194,1922]" captionTargetId="figure@10.[200,1370,194,1929]" captionTargetPageId="10" captionText="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="6" pageNumber="7">Fig. 20</figureCitation>
) with 12-14 (M: 9–10; F: 11–14) setae, stigmata on sternite IV (
<figureCitation id="13048094777A4247FB4FF809FB55F81E" box="[1169,1254,1928,1954]" captionStart="FIGURES 18 – 23" captionStartId="10.[151,269,1950,1974]" captionTargetBox="[210,1361,194,1922]" captionTargetId="figure@10.[200,1370,194,1929]" captionTargetPageId="10" captionText="FIGURES 18 – 23. Titanobochica magna gen. nov., sp. nov., male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II – VII, ventral view; 23. tergites I – VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 – 21), 0.50 (Figs 18, 22, 23)." httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="6" pageNumber="7">Fig. 21</figureCitation>
) with 9-10 (M: 5–8; F: 5–9) setae. Two lyrifissures in medial zone, one more on each side of anterolateral zone.
</paragraph>
<paragraph id="8B809C11777B4246FF1BFF19FC93FE60" blockId="7.[151,1437,152,1935]" pageId="7" pageNumber="8">
Chelicera (
<figureCitation id="13048094777B4246FE94FF19FE0AFF0E" box="[330,441,152,178]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Figs 5–6</figureCitation>
, 
<figureCitation id="13048094777B4246FE18FF19FDA4FF0E" box="[454,535,152,178]" captionStart="FIGURES 24 – 27" captionStartId="11.[151,270,1919,1943]" captionTargetBox="[176,1411,194,1896]" captionTargetId="figure@11.[176,1412,194,1896]" captionTargetPageId="11" captionText="FIGURES 24 – 27. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 24. carapace; 25. opisthosoma, tergite VII partially teratological; 26. chelicerae; 27. distal portion of fingers of left chelicera. Scale bars (mm): 0.12 (Fig. 27), 0.30 (Fig 26), 1.00 (Figs 24, 25)." httpUri="https://zenodo.org/record/199339/files/figure.png" pageId="7" pageNumber="8">26–27</figureCitation>
). Four setae on hand (4, seldom 5–6; when variation occurs one chelicera always with 4) and one seta on movable finger, 0.77 (M: 0.69–075; F: 0.74–0.79) from base. Galea diminutive and simple, dagger-blade shaped, tip distinctly proximal of apex of finger, length: 0.03 (M: 0.03–0.05; F: 0.03) mm (
<figureCitation id="13048094777B4246FEFEFE89FED8FE9E" box="[288,363,264,290]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
, ga). Fixed finger with 8 teeth, distal tooth larger than the rest and two reduced to tiny protuberances (8–10). Movable finger with an extremely large distal tooth, turned slightly laterally to lodge the fixed finger, moreover with a ridge joining 4 (3–6) teeth and 3 (3–5) separate teeth up to beginning of basal third, total 8 (8–13) teeth. Rallum (
<figureCitation id="13048094777B4246FDA2FEF9FD70FE2E" box="[636,707,376,402]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Fig. 7</figureCitation>
) with five externally serrate blades (5, one teratological male with only 
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one rallum and 
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the other), basal blade slightly shorter than others. Serrula exterior with 35 (36–40) blades, serrula interior with 34 (33–37) blades.
</paragraph>
<paragraph id="8B809C11777B4246FF1BFE69FC90FB18" blockId="7.[151,1437,152,1935]" pageId="7" pageNumber="8">
Pedipalps (
<figureCitation id="13048094777B4246FE99FE69FE7DFDBE" box="[327,462,488,514]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Figs 13–17</figureCitation>
, 
<figureCitation id="13048094777B4246FE05FE69FD9AFDBE" box="[475,553,488,514]" captionStart="FIGURES 28 – 31" captionStartId="12.[151,270,1797,1821]" captionTargetBox="[151,1436,194,1773]" captionTargetId="figure@12.[151,1436,194,1775]" captionTargetPageId="12" captionText="FIGURES 28 – 31. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 28. fingers of the left chela, dorsal view, A: glandular pore magnified; 29. tip of the chelal movable finger, with venedens, A: magnification of a tooth; 30. right chelal hand, partial dorsal view; 31. leg IV, detail of claws and subterminal feathery setae. Scale bars (mm): 0.009 (Fig. 28 A), 0.01 (Fig. 29 A), 0.075 (Figs 29, 31), 0.75 (Fig. 30), 1.00 (Fig. 28)." httpUri="https://zenodo.org/record/199340/files/figure.png" pageId="7" pageNumber="8">28–30</figureCitation>
). Articles smooth, except fingers of chela; some micropores dispersed over whole pedipalp and in groups of 9–10 at bases of femur, patella and hand. Trochanter short. Femur and patella very slender, patella only slightly shorter than femur; femur distinctly longer than fixed chelal finger and without glandular tubercle at base of antiaxial face. Chela smooth, except distal portion of hand and the fingers, which are weakly granulated; fixed finger with irregular outline at the distal portion and distinctly longer than movable finger. Hand with oval outline at paraxial face and flattened at antiaxial face; trichobothrium 
<emphasis id="B94B4003777B4246FE8AFD49FED8FD5D" box="[340,363,712,737]" italics="true" pageId="7" pageNumber="8">ib</emphasis>
located in basal half of dorsum of hand, 
<emphasis id="B94B4003777B4246FCBEFD49FCCFFD5D" box="[864,892,712,737]" italics="true" pageId="7" pageNumber="8">eb</emphasis>
in distal portion of hand; 10 (9–14) thin and small chemosensory setae (
<figureCitation id="13048094777B4246FE00FD6CFD84FCBB" box="[478,567,749,775]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
, cs) on dorsum of hand, in a row from 
<emphasis id="B94B4003777B4246FBD0FD6CFB96FCBA" box="[1038,1061,749,774]" italics="true" pageId="7" pageNumber="8">ib</emphasis>
to base of finger (
<figureCitation id="13048094777B4246FADDFD6CFAECFCBB" box="[1283,1375,749,775]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Figs 13</figureCitation>
, 
<figureCitation id="13048094777B4246FAB3FD6CFA3CFCBB" box="[1389,1423,749,775]" captionStart="FIGURES 28 – 31" captionStartId="12.[151,270,1797,1821]" captionTargetBox="[151,1436,194,1773]" captionTargetId="figure@12.[151,1436,194,1775]" captionTargetPageId="12" captionText="FIGURES 28 – 31. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 28. fingers of the left chela, dorsal view, A: glandular pore magnified; 29. tip of the chelal movable finger, with venedens, A: magnification of a tooth; 30. right chelal hand, partial dorsal view; 31. leg IV, detail of claws and subterminal feathery setae. Scale bars (mm): 0.009 (Fig. 28 A), 0.01 (Fig. 29 A), 0.075 (Figs 29, 31), 0.75 (Fig. 30), 1.00 (Fig. 28)." httpUri="https://zenodo.org/record/199340/files/figure.png" pageId="7" pageNumber="8">30</figureCitation>
). Fixed finger with tip rounded (
<figureCitation id="13048094777B4246FDDBFC93FD3DFC90" box="[517,654,786,812]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Figs 15–16</figureCitation>
), venedens and venom apparatus completely absent; trichobothria 
<emphasis id="B94B4003777B4246FF49FCB9FF0DFCED" box="[151,190,824,849]" italics="true" pageId="7" pageNumber="8">esb</emphasis>
and 
<emphasis id="B94B4003777B4246FF29FCB9FEAAFCED" box="[247,281,824,849]" italics="true" pageId="7" pageNumber="8">isb</emphasis>
at base of finger, medial portion of finger without trichobothria, 
<emphasis id="B94B4003777B4246FBC8FCB9FB81FCED" box="[1046,1074,824,849]" italics="true" pageId="7" pageNumber="8">ist</emphasis>
slightly proximal of 
<emphasis id="B94B4003777B4246FAF4FCB9FAF9FCED" box="[1322,1354,824,849]" italics="true" pageId="7" pageNumber="8">est</emphasis>
, 
<emphasis id="B94B4003777B4246FA84FCB9FAD9FCED" box="[1370,1386,824,849]" italics="true" pageId="7" pageNumber="8">it</emphasis>
and 
<emphasis id="B94B4003777B4246FF49FCDCFF1EFCCA" box="[151,173,861,886]" italics="true" pageId="7" pageNumber="8">et</emphasis>
at tip of finger; with one apical sublateral tooth and 112 (M: 107–121; F: 111–116) teeth up to base of finger, apically uniseriate, basally decreasing in size (
<figureCitation id="13048094777B4246FC80FC04FC72FC23" box="[862,961,901,927]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
); 26th to 45th teeth from the apex are heterodentate, alternately weakly different in shape and with a slightly different orientation (
<figureCitation id="13048094777B4246FB16FC2BFAACFC78" box="[1224,1311,938,964]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
). Movable finger (
<figureCitation id="13048094777B4246FF32FC51FEC2FC56" box="[236,369,976,1002]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Figs 14, 17</figureCitation>
) with functional venom apparatus, 
<emphasis id="B94B4003777B4246FCCEFC51FC72FC55" box="[784,961,976,1001]" italics="true" pageId="7" pageNumber="8">nodus ramosus</emphasis>
(nr) proximal of trichobothria 
<emphasis id="B94B4003777B4246FAF4FC51FA8DFC55" box="[1322,1342,976,1001]" italics="true" pageId="7" pageNumber="8">st</emphasis>
and not expanded (
<figureCitation id="13048094777B4246FEC9FC74FEDEFBB3" box="[279,365,1013,1039]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Fig. 14</figureCitation>
); distance trichobothria 
<emphasis id="B94B4003777B4246FD56FC77FD76FBB3" box="[648,709,1013,1039]" italics="true" pageId="7" pageNumber="8">sb–st</emphasis>
1.82× (1.17–1.92) longer than distance 
<emphasis id="B94B4003777B4246FB49FC77FB7CFBB3" box="[1175,1231,1013,1039]" italics="true" pageId="7" pageNumber="8">b–sb</emphasis>
; with 77 (M: 68– 81; F: 74–77) teeth, reaching to distal of trichobothrium 
<emphasis id="B94B4003777B4246FC89FB9AFCDDFB88" box="[855,878,1051,1076]" italics="true" pageId="7" pageNumber="8">b,</emphasis>
rounded and progressively flattened in basal portion, last 6 (7–8) basal teeth without dental canal; one male 
<typeStatus id="548422B3777B4246FC5AFBC1FC5AFBE6" box="[900,1001,1088,1114]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
with teratological movable finger of one chela shorter than normal, length 
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, only trichobothria 
<emphasis id="B94B4003777B4246FC76FBE7FC04FBC3" box="[936,951,1126,1151]" italics="true" pageId="7" pageNumber="8">b</emphasis>
and 
<emphasis id="B94B4003777B4246FC2FFBE7FBB8FBC3" box="[1009,1035,1126,1151]" italics="true" pageId="7" pageNumber="8">sb</emphasis>
present, dental row reduced to 47 teeth. Lyrifissures of pedipalps as shown in 
<figureCitation id="13048094777B4246FD47FB0BFCACFB18" box="[665,799,1162,1188]" captionStart="FIGURES 13 – 17" captionStartId="9.[151,269,1880,1904]" captionTargetBox="[205,1386,203,1839]" captionTargetId="figure@9.[191,1389,194,1859]" captionTargetPageId="9" captionText="FIGURES 13 – 17. Titanobochica magna gen. nov., sp. nov., male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15 – 17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus." httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="7" pageNumber="8">Figs 13–14</figureCitation>
.
</paragraph>
<paragraph id="8B809C11777B4246FF1BFB31FE3AFAE3" blockId="7.[151,1437,152,1935]" pageId="7" pageNumber="8">
Legs smooth, tarsi not swollen basally, claws smooth and apically blunt, subterminal setae apically feathery. Leg I (
<figureCitation id="13048094777B4246FE86FB54FE11FB53" box="[344,418,1237,1263]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Fig. 8</figureCitation>
): metatarsus extremely short and apparently fused with tarsus, together composing a rigid article. Leg IV (
<figureCitation id="13048094777B4246FE51FB7BFDBBFAA8" box="[399,520,1274,1300]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Figs 9–10</figureCitation>
, 
<figureCitation id="13048094777B4246FDCDFB7BFD87FAA8" box="[531,564,1274,1300]" captionStart="FIGURES 28 – 31" captionStartId="12.[151,270,1797,1821]" captionTargetBox="[151,1436,194,1773]" captionTargetId="figure@12.[151,1436,194,1775]" captionTargetPageId="12" captionText="FIGURES 28 – 31. Titanobochica magna gen. nov., sp. nov., male paratype from Gruta do Vale Telheiro. SEM images. 28. fingers of the left chela, dorsal view, A: glandular pore magnified; 29. tip of the chelal movable finger, with venedens, A: magnification of a tooth; 30. right chelal hand, partial dorsal view; 31. leg IV, detail of claws and subterminal feathery setae. Scale bars (mm): 0.009 (Fig. 28 A), 0.01 (Fig. 29 A), 0.075 (Figs 29, 31), 0.75 (Fig. 30), 1.00 (Fig. 28)." httpUri="https://zenodo.org/record/199340/files/figure.png" pageId="7" pageNumber="8">31</figureCitation>
): femur(F)/patella(P) suture strongly oblique, femur evidently shorter than ¼ length of combined segments, according to Muchmore’s formula (1998): F/F+P: 0.15 (0.14–0.16); articles without tactile setae.
</paragraph>
<paragraph id="8B809C11777B4246FF1BFAEBFBB2F833" blockId="7.[151,1437,152,1935]" pageId="7" pageNumber="8">
<emphasis id="B94B4003777B4246FF1BFAEBFED2FA38" bold="true" box="[197,353,1386,1412]" pageId="7" pageNumber="8">Tritonymph.</emphasis>
Carapacal chaetotaxy: 31–39 setae, formula (total number of setae in each row followed by number of lateral setae in parentheses) 4–6:17–19(6–7):4–6:5–8. Coxal area: apex of the pedipalp with 2 setae, pedipalpal coxa 10–14 setae, pedal coxa I: 13–15, II: 11–16, III: 9–14, IV: 12–16. Opisthosomal tergites I-X chaetotaxy (totals, followed by number of median setae in parentheses): 4–6: 6: 6: 6–8: 6–9(1): 6–8(1): 6– 8(1): 6–8(1): 6–7: 4; sternite II: 4–5 setae, III-X (discal/posterior): 5/11–12: 6–8/7–8: 3–4/9–12: 3–5/8–12:2– 5/8–11: 2–4/8–10: 1–3/6–8: 6, segment XI: 6–8 setae, anal cone with 2 tergal and 2 sternal setae, stigmata sternite III: 6–7 setae, IV: 4–5. Cheliceral palm with 4 setae, some specimens with 4 setae on one chelicera and 5 on other, subgaleal seta on movable finger 0.79–0.80 from base; galea short and simple, length: 
<quantity id="4CC731F4777B4246FAB6F9F1FF7BF913" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="8" unit="mm" value="0.04">0.04 mm</quantity>
; fixed finger with 8–9 teeth, the distal one larger than the rest; movable finger with 7–12 teeth, the distal one very large and sublateral; rallum with 4 externally serrate blades, with the same shape as in adults; serrula exterior with 30–34 blades, serrula interior with 25–31 blades. Pedipalp: chelal fixed finger about as long as femur; trichobothrium 
<emphasis id="B94B4003777B4246FE76F887FE77F8A3" box="[424,452,1798,1823]" italics="true" pageId="7" pageNumber="8">ist</emphasis>
proximal of 
<emphasis id="B94B4003777B4246FDBFF887FD31F8A3" box="[609,642,1798,1823]" italics="true" pageId="7" pageNumber="8">est</emphasis>
; fixed finger with 93–100 teeth; movable finger with 64–68 teeth, some basal ones without dental canal. Trichobothria 
<emphasis id="B94B4003777B4246FCC6F8AAFC88F8F8" box="[792,827,1835,1860]" italics="true" pageId="7" pageNumber="8">isb</emphasis>
and 
<emphasis id="B94B4003777B4246FCA9F8AAFC21F8F8" box="[887,914,1835,1860]" italics="true" pageId="7" pageNumber="8">sb</emphasis>
lacking. Legs: tarsi of all four legs basally swollen, maximum width of leg I at 0.18–0.22 of total length from base, leg IV (
<figureCitation id="13048094777B4246FB7BF8D1FAB0F8D6" box="[1189,1283,1872,1898]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="7" pageNumber="8">Fig. 12</figureCitation>
) 0.29–0.32; metatarsus leg I apparently fused with the tarsus; leg IV: F/F+P 0.16–0.17.
</paragraph>
<caption id="DF40CC9977744249FF49F8B7FD85F852" httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="8" pageNumber="9" targetBox="[232,1360,200,1810]" targetPageId="8">
<paragraph id="8B809C1177744249FF49F8B7FD85F852" blockId="8.[151,1437,1846,2030]" pageId="8" pageNumber="9">
<emphasis id="B94B400377744249FF49F8B7FEEBF8F2" bold="true" box="[151,344,1846,1870]" pageId="8" pageNumber="9">FIGURES 3–12.</emphasis>
<taxonomicName id="4C3FE79277744249FE81F8B6FDE1F8F2" box="[351,594,1847,1870]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B400377744249FE81F8B6FDE1F8F2" box="[351,594,1847,1870]" italics="true" pageId="8" pageNumber="9">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B400377744249FD84F8B7FD70F8F2" bold="true" box="[602,707,1846,1870]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="A278FD7877744249FD84F8B7FD70F8F2" box="[602,707,1846,1870]" pageId="8" pageNumber="9" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="B94B400377744249FD0CF8B7FC9FF8F2" bold="true" box="[722,812,1846,1870]" pageId="8" pageNumber="9">sp. nov.</emphasis>
, male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: 
<emphasis id="B94B400377744249FE81F856FEC9F852" box="[351,378,2007,2030]" italics="true" pageId="8" pageNumber="9">ga</emphasis>
: cheliceral galea.
</paragraph>
</caption>
<caption id="DF40CC9977754248FF49F8D9FF4BF853" httpUri="https://zenodo.org/record/199337/files/figure.png" pageId="9" pageNumber="10" targetBox="[205,1386,203,1839]" targetPageId="9">
<paragraph id="8B809C1177754248FF49F8D9FF4BF853" blockId="9.[151,1437,1880,2031]" pageId="9" pageNumber="10">
<emphasis id="B94B400377754248FF49F8D9FEEEF8CC" bold="true" box="[151,349,1880,1904]" pageId="9" pageNumber="10">FIGURES 13–17.</emphasis>
<taxonomicName id="4C3FE79277754248FEBDF8D9FDF9F8D3" box="[355,586,1880,1903]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B400377754248FEBDF8D9FDF9F8D3" box="[355,586,1880,1903]" italics="true" pageId="9" pageNumber="10">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B400377754248FD8FF8D9FD06F8CC" bold="true" box="[593,693,1880,1904]" pageId="9" pageNumber="10">
<taxonomicNameLabel id="A278FD7877754248FD8FF8D9FD06F8CC" box="[593,693,1880,1904]" pageId="9" pageNumber="10" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, 
<emphasis id="B94B400377754248FD1CF8D9FCA4F8CC" bold="true" box="[706,791,1880,1904]" pageId="9" pageNumber="10">sp. nov.</emphasis>
, male holotype. 13. left pedipalp, dorsal view; 14. left chela, antiaxial view, some teeth magnified; 15. tip of fixed finger of left chela, ventral view; 16. tip of fixed finger of left chela, antiaxial view; 17. tip of movable finger of left chela, antiaxial view. Scale bars (mm): 0.20 (Figs 15–17), 1.00 (Figs 13, 14). Abbreviations: trichobothrial terminology following Chamberlin (1931); cs: chemosensory setae; nr: nodus ramosus.
</paragraph>
</caption>
<caption id="DF40CC997776424BFF49F81FFB84F849" httpUri="https://zenodo.org/record/199338/files/figure.png" pageId="10" pageNumber="11" targetBox="[210,1361,194,1922]" targetPageId="10">
<paragraph id="8B809C117776424BFF49F81FFB84F849" blockId="10.[151,1436,1950,2038]" pageId="10" pageNumber="11">
<emphasis id="B94B40037776424BFF49F81FFEEDF80A" bold="true" box="[151,350,1950,1974]" pageId="10" pageNumber="11">FIGURES 18–23.</emphasis>
<taxonomicName id="4C3FE7927776424BFEBBF81EFDFDF80A" box="[357,590,1951,1974]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B40037776424BFEBBF81EFDFDF80A" box="[357,590,1951,1974]" italics="true" pageId="10" pageNumber="11">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B40037776424BFD8AF81FFCAFF80A" bold="true" box="[596,796,1950,1974]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A278FD787776424BFD8AF81FFD09F80A" box="[596,698,1950,1974]" pageId="10" pageNumber="11" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, male holotype. 18. coxae of pedipalp and leg I; 19. genital sternites, ventral; 20. left stigma of sternite III; 21. left stigma of sternite IV; 22. sternites II 
<emphasis id="B94B40037776424BFB42F83FFB1AF86A" bold="true" box="[1180,1193,1982,2006]" pageId="10" pageNumber="11">–</emphasis>
VII, ventral view; 23. tergites I 
<emphasis id="B94B40037776424BFF2AF85FFEB2F84A" bold="true" box="[244,257,2014,2038]" pageId="10" pageNumber="11">–</emphasis>
VII, dorsal view. Scale bars (mm): 0.20 (Figs 19 
<emphasis id="B94B40037776424BFCDBF85FFCA1F84A" bold="true" box="[773,786,2014,2038]" pageId="10" pageNumber="11">–</emphasis>
21), 0.50 (Figs 18, 22, 23).
</paragraph>
</caption>
<caption id="DF40CC997777424AFF49F8FEFC73F86B" httpUri="https://zenodo.org/record/199339/files/figure.png" pageId="11" pageNumber="12" targetBox="[176,1411,194,1896]" targetPageId="11">
<paragraph id="8B809C117777424AFF49F8FEFC73F86B" blockId="11.[151,1436,1919,2007]" pageId="11" pageNumber="12">
<emphasis id="B94B40037777424AFF49F8FEFEECF82B" bold="true" box="[151,351,1919,1943]" pageId="11" pageNumber="12">FIGURES 24–27.</emphasis>
<taxonomicName id="4C3FE7927777424AFEB8F801FDFCF82B" box="[358,591,1920,1943]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B40037777424AFEB8F801FDFCF82B" box="[358,591,1920,1943]" italics="true" pageId="11" pageNumber="12">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B40037777424AFD88F8FEFCACF82B" bold="true" box="[598,799,1919,1943]" pageId="11" pageNumber="12">
<taxonomicNameLabel id="A278FD787777424AFD88F8FEFD0EF82B" box="[598,701,1919,1943]" pageId="11" pageNumber="12" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, male paratype from Gruta do Vale Telheiro. SEM images. 24. carapace; 25. opisthosoma, tergite VII partially teratological; 26. chelicerae; 27. distal portion of fingers of left chelicera. Scale bars (mm): 0.12 (Fig. 27), 0.30 (Fig 26), 1.00 (Figs 24, 25).
</paragraph>
</caption>
<caption id="DF40CC997770424DFF49F884FF48F820" httpUri="https://zenodo.org/record/199340/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1436,194,1773]" targetPageId="12">
<paragraph id="8B809C117770424DFF49F884FF48F820" blockId="12.[151,1437,1797,1948]" pageId="12" pageNumber="13">
<emphasis id="B94B40037770424DFF49F884FEECF8A1" bold="true" box="[151,351,1797,1821]" pageId="12" pageNumber="13">FIGURES 28–31.</emphasis>
<taxonomicName id="4C3FE7927770424DFEB8F884FDFCF8A0" box="[358,591,1797,1820]" class="Arachnida" family="Bochicidae" genus="Titanobochica" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="magna" status="gen. nov.">
<emphasis id="B94B40037770424DFEB8F884FDFCF8A0" box="[358,591,1797,1820]" italics="true" pageId="12" pageNumber="13">Titanobochica magna</emphasis>
</taxonomicName>
<emphasis id="B94B40037770424DFD88F884FCACF8A1" bold="true" box="[598,799,1797,1821]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A278FD787770424DFD88F884FD0EF8A1" box="[598,701,1797,1821]" pageId="12" pageNumber="13" rank="genus">gen. nov.</taxonomicNameLabel>
, sp. nov.
</emphasis>
, male paratype from Gruta do Vale Telheiro. SEM images. 28. fingers of the left chela, dorsal view, A: glandular pore magnified; 29. tip of the chelal movable finger, with venedens, A: magnification of a tooth; 30. right chelal hand, partial dorsal view; 31. leg IV, detail of claws and subterminal feathery setae. Scale bars (mm): 0.009 (Fig. 28A), 0.01 (Fig. 29A), 0.075 (Figs 29, 31), 0.75 (Fig. 30), 1.00 (Fig. 28).
</paragraph>
</caption>
<paragraph id="8B809C117771424CFF1BFF16FCB0FDCE" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
<emphasis id="B94B40037771424CFF1BFF16FEC2FF0D" bold="true" box="[197,369,151,177]" pageId="13" pageNumber="14">Deutonymph.</emphasis>
Carapacal chaetotaxy: 25–27 setae, formula (total number of setae in each row, followed by number of lateral setae in parentheses) 4:13–14(3–4):4:4–5. Coxal area: pedipalpal apex with 2 setae; pedipalpal coxa 7–8 setae, pedal coxa I: 8–10, II: 7–8, III: 6–8, IV: 8. Tergites I-X chaetotaxy: 4–5: 4: 6: 6: 6: 6: 6: 6: 6: 4–6, without median setae; sternite II without setae, sternites III–X uniseriate, some with median subdiscal setae: 6–7: 2/5–6: 1–2/8–10: 2/8–10: 2/8: 2/8: 0–1/7: 6, segment XI: 6–7 setae, anal cone with 2+2 setae, stigmata sternite III: 4–5 setae, IV: 3–4. Cheliceral palm with 4 setae, subgaleal seta on movable finger 0.74–0.75; galea short and simple, length 
<quantity id="4CC731F47771424CFD45FEF9FCE3FE2E" box="[667,848,376,402]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="13" pageNumber="14" unit="mm" value="0.03" valueMax="0.04" valueMin="0.02">0.02–0.04 mm</quantity>
; fixed finger with 7–10 teeth, distal tooth large; movable finger with 8–10 teeth, distal tooth large and sublateral; rallum with 3 unilaterally serrate blades; serrula exterior with 27 blades, serrula interior with 22. Pedipalp: chelal fixed finger longer than the femur; trichobothrium 
<emphasis id="B94B40037771424CFE8BFE69FEC1FDBD" box="[341,370,488,513]" italics="true" pageId="13" pageNumber="14">ist</emphasis>
proximal of 
<emphasis id="B94B40037771424CFDCCFE69FD87FDBD" box="[530,564,488,513]" italics="true" pageId="13" pageNumber="14">est</emphasis>
; fixed finger with 83–84 teeth; movable with 55–58 teeth, basal 12–17 flattened, all with dental canal. Trichobothria 
<emphasis id="B94B40037771424CFD62FD8CFD57FD9A" box="[700,740,525,550]" italics="true" pageId="13" pageNumber="14">esb</emphasis>
, 
<emphasis id="B94B40037771424CFD2AFD8CFCA4FD9A" box="[756,791,525,550]" italics="true" pageId="13" pageNumber="14">isb</emphasis>
, 
<emphasis id="B94B40037771424CFCF8FD8CFCF2FD9A" box="[806,833,525,550]" italics="true" pageId="13" pageNumber="14">sb</emphasis>
and 
<emphasis id="B94B40037771424CFCA2FD8CFC23FD9A" box="[892,912,525,550]" italics="true" pageId="13" pageNumber="14">st</emphasis>
lacking. Tarsi of legs basally swollen, leg I maximum width 0.20 from base, leg IV 0.29; suture between metatarsus and tarsus leg I almost indistinguishable (
<figureCitation id="130480947771424CFEB1FDD9FE76FDCE" box="[367,453,600,626]" captionStart="FIGURES 3 – 12" captionStartId="8.[151,273,1846,1870]" captionTargetBox="[232,1360,200,1810]" captionTargetId="figure@8.[221,1369,197,1822]" captionTargetPageId="8" captionText="FIGURES 3 – 12. Titanobochica magna gen. nov., sp. nov., male holotype, unless otherwise noted. 3. carapace; 4. anterior margin of carapace; 5. right chelicera; 6. distal portion of fingers of right chelicera; 7. cheliceral rallum; 8. left leg I, lateral view; 9. left leg IV, lateral view; 10. distal portion of tarsus and apotele of left leg IV, lateral view. Deutonymph paratype: 11. metatarsus and tarsus of right leg I, lateral view; 12. tarsus of left leg IV, lateral view, tritonymph paratype. Scale bars (mm): 0.10 (Fig. 7), 0.20 (Figs 6, 10); 0.25 (Fig. 5), 0.50 (Figs 3, 4, 11, 12), 1.00 (Figs 8, 9). Abbreviations: ga: cheliceral galea." httpUri="https://zenodo.org/record/199336/files/figure.png" pageId="13" pageNumber="14">Fig. 11</figureCitation>
); leg IV F/F+P: 0.19–0.20.
</paragraph>
</subSubSection>
<subSubSection id="C325CF9A7771424CFF1BFDFCFC4EFA8B" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8B809C117771424CFF1BFDFCFE51FD00" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
<emphasis id="B94B40037771424CFF1BFDFCFEF2FD2B" bold="true" box="[197,321,637,663]" pageId="13" pageNumber="14">Remarks.</emphasis>
No significant differences are found in the form and length of the cheliceral galea between males, females and nymphs.
</paragraph>
<paragraph id="8B809C117771424CFF1BFD49FE28FC90" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
The development of the chelal femur length and slenderness relative to the other pedipalpal articles is progressive from the nymphal to adult stages, as shown in the ratios of the femur, femur/fixed finger and chela/femur (
<tableCitation id="C6BDA9AA7771424CFEECFC93FE3EFC90" box="[306,397,786,812]" captionStart="TABLE 2" captionStartId="5.[151,240,375,399]" captionText="TABLE 2. Titanobochica magna gen. nov., sp. nov.: measurements and ratios. M: male; F: female; T: tritonymph., D:" pageId="13" pageNumber="14">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8B809C117771424CFF1BFCB9FC80FCCB" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
Heterodentate marginal teeth of fixed chelal finger have been previously reported in 
<taxonomicName id="4C3FE7927771424CFB3BFCB9FADCFCEE" box="[1253,1391,824,850]" class="Arachnida" family="Bochicidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Bochicidae</taxonomicName>
for 
<taxonomicName id="4C3FE7927771424CFF49FCDCFC9CFCCB" authority="Chamberlin, 1947 (Chamberlin 1947)" authorityName="Chamberlin, 1947 (Chamberlin" authorityYear="1947" box="[151,815,861,887]" class="Arachnida" family="Bochicidae" genus="Vachonium" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="B94B40037771424CFF49FCDCFEDAFCCA" box="[151,361,861,886]" italics="true" pageId="13" pageNumber="14">Vachonium boneti</emphasis>
<bibRefCitation id="EFAEE1E07771424CFEAFFCDCFDF6FCCB" author="Chamberlin" box="[369,581,861,887]" pageId="13" pageNumber="14" refString="Chamberlin, J. C. (1947) The Vachoniidae - a new family of false scorpions represented by two new species from caves in Yucatan (Arachnida, Chelonethida, Neobisioidea). Bulletin of the University of Utah, Biological Series, 10 (4), 1 - 15." type="journal article" year="1947">Chamberlin, 1947</bibRefCitation>
(
<bibRefCitation id="EFAEE1E07771424CFD8AFCDCFC95FCCB" author="Chamberlin" box="[596,806,861,887]" pageId="13" pageNumber="14" refString="Chamberlin, J. C. (1947) The Vachoniidae - a new family of false scorpions represented by two new species from caves in Yucatan (Arachnida, Chelonethida, Neobisioidea). Bulletin of the University of Utah, Biological Series, 10 (4), 1 - 15." type="journal article" year="1947">Chamberlin 1947</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8B809C117771424CFF1BFC03FE50FC5B" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
Presence of chemosensory setae on the palm of the chela was reported for 
<taxonomicName id="4C3FE7927771424CFBE4FC03FB0DFC20" box="[1082,1214,898,924]" class="Arachnida" family="Bochicidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Bochicidae</taxonomicName>
by 
<bibRefCitation id="EFAEE1E07771424CFB32FC03FA2BFC20" author="Judson" box="[1260,1432,898,924]" pageId="13" pageNumber="14" refString="Judson, M. L. I. (2007) A new and endangered pseudoscorpion of the genus Lagynochthonius (Arachnida, Chelonethi, Chthoniidae) from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini. Zootaxa, 1627, 53 - 68." type="journal article" year="2007">Judson (2007)</bibRefCitation>
, who assumed the chemosensory function of these setae because they are smaller, thinner and more strongly inclined than ordinary setae.
</paragraph>
<paragraph id="8B809C117771424CFF1BFC73FC4EFA8B" blockId="13.[151,1437,151,1335]" pageId="13" pageNumber="14">
Tritonymphs and deutonymphs show tarsi swollen basally, which does not occur in adults. Basally swollen tarsi have been described for nymphs and, exceptionally, for adults of some genera of the family 
<taxonomicName id="4C3FE7927771424CFF49FBBCFEA8FBEB" box="[151,283,1085,1111]" class="Arachnida" family="Syarinidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Syarinidae</taxonomicName>
: 
<taxonomicName id="4C3FE7927771424CFEF8FBBCFD56FBEB" authority="Beier (Vachon 1954)" authorityName="Beier (Vachon" authorityYear="1954" box="[294,741,1085,1111]" class="Arachnida" family="Syarinidae" genus="Pseudoblothrus" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFEF8FBBCFE52FBEA" box="[294,481,1085,1110]" italics="true" pageId="13" pageNumber="14">Pseudoblothrus</emphasis>
Beier (
<bibRefCitation id="EFAEE1E07771424CFDE4FBBCFD6FFBEB" author="Vachon" box="[570,732,1085,1111]" pageId="13" pageNumber="14" refString="Vachon, M. (1954) Remarques morphologiques et anatomiques sur les Pseudoscorpions (Arachnides) appartenant au genre Pseudoblothrus (Beier) (Fam. Syarinidae J. C. C.) (a propos de la description de P. strinatii n. sp., des cavernes de Suisse). Bulletin du Museum National d'Histoire Naturelle, Paris, (2) 26, 212 - 219." type="journal article" year="1954">Vachon 1954</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C3FE7927771424CFD2CFBBCFC53FBEB" authority="Beier" authorityName="Beier" box="[754,992,1085,1111]" class="Arachnida" family="Syarinidae" genus="Hadoblothrus" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFD2CFBBCFC24FBEA" box="[754,919,1085,1110]" italics="true" pageId="13" pageNumber="14">Hadoblothrus</emphasis>
Beier
</taxonomicName>
and 
<taxonomicName id="4C3FE7927771424CFBC5FBBCFF53FBC0" authority="Beier (Mahnert 1980)" authorityName="Beier (Mahnert" authorityYear="1980" class="Arachnida" family="Syarinidae" genus="Microcreagrella" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFBC5FBBCFB6FFBEA" box="[1051,1244,1085,1110]" italics="true" pageId="13" pageNumber="14">Microcreagrella</emphasis>
Beier (
<bibRefCitation id="EFAEE1E07771424CFAEBFBBCFF6BFBC0" author="Mahnert" pageId="13" pageNumber="14" refString="Mahnert, V. (1980) Pseudoskorpione (Arachnida) aus Hohlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Grotte d'Italia, (4 a) 8, 21 - 38." type="journal article" year="1980">Mahnert 1980</bibRefCitation>
)
</taxonomicName>
. In the family 
<taxonomicName id="4C3FE7927771424CFE4FFBE3FDA4FBC0" box="[401,535,1122,1148]" class="Arachnida" family="Bochicidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="family">Bochicidae</taxonomicName>
this characteristic has also been reported for nymphal stages of the genera 
<taxonomicName id="4C3FE7927771424CFF49FB09FE4DFB1E" authority="Muchmore, 1972" authorityName="Muchmore" authorityYear="1972" box="[151,510,1160,1186]" class="Arachnida" family="Bochicidae" genus="Mexobisium" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFF49FB09FE94FB1D" box="[151,295,1160,1185]" italics="true" pageId="13" pageNumber="14">Mexobisium</emphasis>
<bibRefCitation id="EFAEE1E07771424CFEF1FB09FE4DFB1E" author="Muchmore" box="[303,510,1160,1186]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1972) New diplosphyronid pseudoscorpions, mainly cavernicolous, from Mexico (Arachnida, Pseudoscorpionida). Transactions of the American Microscopical Society, 91, 261 - 276." type="journal article" year="1972">Muchmore, 1972</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C3FE7927771424CFDD5FB09FD74FB1D" box="[523,711,1160,1185]" class="Arachnida" family="Bochicidae" genus="Paravachonium" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFDD5FB09FD74FB1D" box="[523,711,1160,1185]" italics="true" pageId="13" pageNumber="14">Paravachonium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3FE7927771424CFCDDFB09FC35FB1D" box="[771,902,1160,1185]" class="Arachnida" family="Bochicidae" genus="Vachonium" kingdom="Animalia" order="Pseudoscorpiones" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B94B40037771424CFCDDFB09FC35FB1D" box="[771,902,1160,1185]" italics="true" pageId="13" pageNumber="14">Vachonium</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFAEE1E07771424CFC48FB09FBDEFB1E" author="Muchmore" box="[918,1133,1160,1186]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1973 a) The pseudoscorpion genus Mexobisium in middle America (Arachnida, Pseudoscorpionida). Bulletin of the Association for Mexican Cave Studies, 5, 63 - 72." type="journal article" year="1973" yearSuffix="a">Muchmore 1973a</bibRefCitation>
, 
<bibRefCitation id="EFAEE1E07771424CFBA7FB09FB74FB1E" author="Muchmore" box="[1145,1223,1160,1186]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1973 b) New and little known pseudoscorpions, mainly from caves in Mexico (Arachnida, Pseudoscorpionida). Bulletin of the Association for Mexican Cave Studies, 5, 47 - 62." type="journal article" year="1973" yearSuffix="b">1973b</bibRefCitation>
, 
<bibRefCitation id="EFAEE1E07771424CFB0DFB09FAACFB1E" author="Muchmore" box="[1235,1311,1160,1186]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1982 c) Some new species of pseudoscorpions from caves in Mexico (Arachnida, Pseudoscorpionida). Bulletin for the Association of Mexican Cave Studies, 8, 63 - 78." type="journal article" year="1982" yearSuffix="c">1982c</bibRefCitation>
), to which 
<emphasis id="B94B40037771424CFF49FB2CFEF0FB7A" box="[151,323,1197,1222]" italics="true" pageId="13" pageNumber="14">Titanobochica</emphasis>
is now added; apparently in correlation with this characteristic, the nymphal stages of these bochicid genera also usually show fused metatarsi and tarsi of legs I and II (
<bibRefCitation id="EFAEE1E07771424CFBD2FB53FB6CFB50" author="Muchmore" box="[1036,1247,1234,1260]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1973 a) The pseudoscorpion genus Mexobisium in middle America (Arachnida, Pseudoscorpionida). Bulletin of the Association for Mexican Cave Studies, 5, 63 - 72." type="journal article" year="1973" yearSuffix="a">Muchmore 1973a</bibRefCitation>
, 
<bibRefCitation id="EFAEE1E07771424CFB35FB53FA84FB50" author="Muchmore" box="[1259,1335,1234,1260]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1973 b) New and little known pseudoscorpions, mainly from caves in Mexico (Arachnida, Pseudoscorpionida). Bulletin of the Association for Mexican Cave Studies, 5, 47 - 62." type="journal article" year="1973" yearSuffix="b">1973b</bibRefCitation>
, 
<bibRefCitation id="EFAEE1E07771424CFA9DFB53FA3CFB50" author="Muchmore" box="[1347,1423,1234,1260]" pageId="13" pageNumber="14" refString="Muchmore, W. B. (1982 c) Some new species of pseudoscorpions from caves in Mexico (Arachnida, Pseudoscorpionida). Bulletin for the Association of Mexican Cave Studies, 8, 63 - 78." type="journal article" year="1982" yearSuffix="c">1982c</bibRefCitation>
). 
<emphasis id="B94B40037771424CFF49FB79FE8CFAAD" box="[151,319,1272,1297]" italics="true" pageId="13" pageNumber="14">Titanobochica</emphasis>
shows a rigid suture between metatarsi and tarsi of legs I–II in adults and tritonymphs, whereas in deutonymphs they have a very indistinct, almost unrecognizable suture.
</paragraph>
</subSubSection>
</treatment>
</document>