1096 lines
178 KiB
XML
1096 lines
178 KiB
XML
<document id="2EC969952C7289CFBFFC622038629502" ID-CLB-Dataset="26520" ID-DOI="10.11646/zootaxa.4614.3.6" ID-GBIF-Dataset="fd229573-d281-4c7a-a9cc-86e8cf041c60" ID-ISSN="1175-5326" ID-Zenodo-Dep="3243076" ID-ZooBank="2A3C63DE-4C89-4433-80F8-FD5FFC250DDA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1560229891125" checkinUser="plazi" docAuthor="Ng, Paul Y. C. & Ng, Peter K. L." docDate="2019" docId="0394AC519C78FF88748FFF40FDC7FBC2" docLanguage="en" docName="zootaxa.4614.3.6.pdf" docOrigin="Zootaxa 4614 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma spectrum Ng & Ng 2019, n. sp." docType="treatment" docVersion="9" lastPageNumber="539" masterDocId="FFADD4299C79FF827418FFD9FFA1FFD1" masterDocTitle="Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo" masterLastPageNumber="540" masterPageNumber="529" pageNumber="530" updateTime="1698734479808" updateUser="ExternalLinkService">
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<mods:title id="F991B97140C7F9C4256FA6A8F6C66777">Geosesarma spectrum, a new species of semiterrestrial vampire crab (Crustacea: Decapoda: Brachyura: Sesarmidae) from Brunei Darussalam, Borneo</mods:title>
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<mods:namePart id="CD703F0FAB43E305349D6F5099AA7E80">Ng, Paul Y. C.</mods:namePart>
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<mods:namePart id="1AB0F856AC789845ADED42F269582199">Ng, Peter K. L.</mods:namePart>
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<mods:title id="B17AB1489B097438D30151E5E1E25408">Zootaxa</mods:title>
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<mods:date id="BAC059BD6AE0C03332E366C3319788D2">2019</mods:date>
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<paragraph id="8B821D479C78FF83748FFF40FE4AFF65" blockId="1.[151,491,153,216]" box="[151,491,153,180]" pageId="1" pageNumber="530">
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<heading id="D0CAAA2B9C78FF83748FFF40FE4AFF65" bold="true" box="[151,491,153,180]" fontSize="11" level="1" pageId="1" pageNumber="530" reason="1">
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<emphasis id="B949C1559C78FF83748FFF40FE4AFF65" bold="true" box="[151,491,153,180]" pageId="1" pageNumber="530">
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<taxonomicName id="4C3D66C49C78FF83748FFF40FE03FF65" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[151,418,153,180]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="530" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
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<emphasis id="B949C1559C78FF83748FFF40FE03FF65" bold="true" box="[151,418,153,180]" italics="true" pageId="1" pageNumber="530">Geosesarma spectrum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27A7C2E9C78FF8375B1FF43FE4AFF65" box="[425,491,154,180]" pageId="1" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B821D479C78FF83748FFF64FEF9FF09" blockId="1.[151,491,153,216]" box="[151,344,189,216]" pageId="1" pageNumber="530">
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(
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<figureCitation id="130601C29C78FF837487FF64FEAAFF06" box="[159,267,189,215]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1901,1926]" captionStartId-1="2.[151,250,1650,1675]" captionStartId-2="3.[151,250,1937,1962]" captionTargetBox-1="[187,1400,668,1630]" captionTargetBox-2="[188,1399,885,1917]" captionTargetId-0="figure@1.[187,1400,808,1881]" captionTargetId-1="figure@2.[187,1400,668,1630]" captionTargetId-2="figure@3.[188,1399,885,1917]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." captionText-1="FIGURE 2. Geosesarma spectrum n. sp., colour in life. A, B, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740); C, paratype male (7.9 × 7.6 mm) (ZRC 2018.0741); D, paratype female (10.2 × 9.8 mm) (ZRC 2018.0741). All from Brunei." captionText-2="FIGURE 3. Geosesarma spectrum n. sp., colour in life. A, female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B–D, female (11.2 × 10.7 mm) (ZRC 2018.0741), Brunei.A, overall habitus; B, ovigerous female with late-stage eggs; C, female with newly hatched juvenile crabs on right side of carapace; D, female with juvenile crabs on right side of carapace." httpUri-0="https://zenodo.org/record/3404937/files/figure.png" httpUri-1="https://zenodo.org/record/3243078/files/figure.png" httpUri-2="https://zenodo.org/record/3243080/files/figure.png" pageId="1" pageNumber="530">Figs. 1–3</figureCitation>
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||
,
|
||
<figureCitation id="130601C29C78FF83750FFF64FEF1FF09" box="[279,336,189,216]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStart-4="FIGURE 9" captionStart-5="FIGURE 10" captionStartId-0="5.[151,250,1057,1082]" captionStartId-1="5.[151,250,1933,1958]" captionStartId-2="6.[151,250,1587,1612]" captionStartId-3="7.[151,250,1864,1889]" captionStartId-4="8.[151,250,1811,1836]" captionStartId-5="9.[151,250,1593,1618]" captionTargetBox-0="[151,1436,181,1037]" captionTargetBox-1="[151,1436,181,1567]" captionTargetBox-2="[151,1436,181,1567]" captionTargetBox-3="[177,1411,921,1845]" captionTargetBox-4="[247,1340,198,1775]" captionTargetBox-5="[151,1436,597,1574]" captionTargetId-0="figure@5.[151,1436,181,1037]" captionTargetId-1="figure@5.[151,1436,1248,1914]" captionTargetId-2="figure@6.[151,1436,181,1567]" captionTargetId-3="figure@7.[177,1411,921,1845]" captionTargetId-4="figure@8.[228,1359,181,1791]" captionTargetId-5="figure@9.[151,1436,597,1574]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionText-0="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." captionText-1="FIGURE 6. A, C, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, D, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak. A, B, frontal view of cephalothorax; C, D, inner view of left chela." captionText-2="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." captionText-3="FIGURE 8. A, B, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; C, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; D, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; E, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, male thoracic sternum and pleon; B–E, male pleonal somites 3–6 and telson." captionText-4="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." captionText-5="FIGURE 10. A, C, Geosesarma spectrum n. sp., paratype ovigerous female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B, D, G. katibas Ng, 1995, female (12.3 x 11.8 mm) (ZRC 2016.0296), Sarawak. A, B, female thoracic sternum and pleon; C, D, female sternopleonal cavity and vulvae." httpUri-0="https://zenodo.org/record/3404939/files/figure.png" httpUri-1="https://zenodo.org/record/3404941/files/figure.png" httpUri-2="https://zenodo.org/record/3243084/files/figure.png" httpUri-3="https://zenodo.org/record/3243086/files/figure.png" httpUri-4="https://zenodo.org/record/3243088/files/figure.png" httpUri-5="https://zenodo.org/record/3243090/files/figure.png" pageId="1" pageNumber="530">5–10</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C78FF83748FFEDBFAA3FECB" box="[151,1282,257,282]" pageId="1" pageNumber="530" type="reference_group">
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<paragraph id="8B821D479C78FF83748FFEDBFAA3FECB" blockId="1.[151,1282,257,282]" box="[151,1282,257,282]" pageId="1" pageNumber="530">
|
||
<treatmentCitationGroup id="AB2D3A699C78FF83748FFEDBFAA3FECB" box="[151,1282,257,282]" pageId="1" pageNumber="530">
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||
<taxonomicName id="4C3D66C49C78FF83748FFEDBFDC8FECB" authority="- Choy, 1991: 129" authorityName="- Choy" authorityPageNumber="129" authorityYear="1991" box="[151,617,257,282]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="530" phylum="Arthropoda" rank="species" species="gracillima">
|
||
<emphasis id="B949C1559C78FF83748FFEDBFEB7FEC8" box="[151,278,258,281]" italics="true" pageId="1" pageNumber="530">Geosesarma</emphasis>
|
||
cf.
|
||
<emphasis id="B949C1559C78FF837525FEDBFE06FECB" box="[317,423,258,282]" italics="true" pageId="1" pageNumber="530">gracillima</emphasis>
|
||
—
|
||
<treatmentCitation id="0A9C3B569C78FF8375D8FED8FDC8FECB" author="Choy, S. C." box="[448,617,257,282]" page="129" pageId="1" pageNumber="530" year="1991">
|
||
<bibRefCitation id="EFAC60B69C78FF8375D8FED8FDC8FECB" author="Choy, S. C." box="[448,617,257,282]" pageId="1" pageNumber="530" pagination="117 - 153" refId="ref4838" refString="Choy, S. C. (1991) The crustacean fauna of negara Brunei Darussalam. Brunei Museum Journal, 7 (3), 117 - 153." type="journal article" year="1991">Choy, 1991: 129</bibRefCitation>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
, 144 (not
|
||
<taxonomicName id="4C3D66C49C78FF8376D7FEDBFB5AFECB" ID-CoL="84HS9" authority="De Man, 1902" authorityName="De Man" authorityYear="1902" box="[719,1275,258,282]" class="Malacostraca" family="Sesarmidae" genus="Sesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="530" phylum="Arthropoda" rank="subSpecies" species="ocypoda" subGenus="Sesarma" subSpecies="gracillima">
|
||
<emphasis id="B949C1559C78FF8376D7FEDBFC87FEC8" box="[719,806,258,281]" italics="true" pageId="1" pageNumber="530">Sesarma</emphasis>
|
||
(
|
||
<emphasis id="B949C1559C78FF83772EFEDBFC2CFEC8" box="[822,909,258,281]" italics="true" pageId="1" pageNumber="530">Sesarma</emphasis>
|
||
)
|
||
<emphasis id="B949C1559C78FF837784FEDBFBC3FECB" box="[924,1122,258,282]" italics="true" pageId="1" pageNumber="530">ocypoda gracillima</emphasis>
|
||
De
|
||
<bibRefCitation id="EFAC60B69C78FF837093FEDBFB5AFECB" author="Man, J. G. De" box="[1163,1275,258,282]" pageId="1" pageNumber="530" pagination="467 - 929" refId="ref5276" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kukenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25, 467 - 929., pls. 19 - 27. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">Man, 1902</bibRefCitation>
|
||
</taxonomicName>
|
||
)
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
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<paragraph id="8B821D479C78FF80748FFE9EFDA5FE7E" blockId="1.[151,1437,327,786]" lastBlockId="2.[151,1437,152,647]" lastPageId="2" lastPageNumber="531" pageId="1" pageNumber="530">
|
||
<subSubSection id="C3274ECC9C78FF83748FFE9EFC29FCC3" pageId="1" pageNumber="530" type="materials_examined">
|
||
<materialsCitation id="3B55171A9C78FF83748FFE9EFC29FCC3" ID-GBIF-Occurrence="2397873773" pageId="1" pageNumber="530">
|
||
<emphasis id="B949C1559C78FF83748FFE9EFE20FEB3" bold="true" box="[151,385,327,354]" pageId="1" pageNumber="530">Material examined.</emphasis>
|
||
Holotype: male (13.6 × 13.5 mm) (ZRC 2018.0740), forest trails around Ulu Ulu resort, Ulu Temburong National Park, 4°33’21.2”N, 115°9’16.1”E, Brunei Darussalam, coll. P.Y.C. Ng, September 2018. Para- types: 2 males (7.9 × 7.6 mm, 12.2 × 12.1 mm), 2 females (15.1 × 14.3 mm, 10.2 × 9.8 mm), 1 ex-ovigerous female with 10 juvenile crabs (11.2 × 10.7 mm) (ZRC 2018.0741), same data as holotype. Others: 1 young male (ZRC 2018.0744), on leaf of sapling, waist height, trail to West Ridge, Belalong, Brunei, coll. I. Das, 27 March 1992; 1 male (10.8 × 10.7 mm) (ZRC 2018.0745), Kuala Belalong Field Studies Centre, Brunei, coll. S. Choy, 10 February 1991; 2 males (10.8 × 10.7 mm, 10.2 × 9.8 mm), 2 females (10.5 × 10.0 mm, 10.2 × 9.9 mm) (ZRC 2017.1215), in pitfall trap, West Ridge, Kuala Belalong Field Studies Centre, Brunei, coll. J. K. Charles, 7 February 1993; 2 males (larger 10.8 × 10.5 mm), 1 female (ZRC 2011.1023), station 19, near Pagon House, on forest floor, Kuala Belalong Field Studies Centre, Brunei, coll. Samahan, 15 February 1993; 1 young male (5.7 × 5.4 mm) (ZRC 2018.0746), in
|
||
<emphasis id="B949C1559C78FF8374ADFD69FE12FD1B" box="[181,435,688,714]" italics="true" pageId="1" pageNumber="530">Nepenthes bicalcarata</emphasis>
|
||
(Hooker, 1893: Nepentheceae), Seria, Brunei, coll. C. Clarke, 29 August 1989; 1 young male (ZRC 2016.0135), Belait peat swamp, Sungei Mendaram, Brunei, coll. I. Kwan, 18 February 2013; 1 juvenile female (ZRC 2018.0747), Lamunin, Brunei, coll. 30 October 1989.
|
||
</materialsCitation>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C7BFF8074DFFF41FDA5FE7E" pageId="2" pageNumber="531" type="materials_examined">
|
||
<emphasis id="B949C1559C7BFF8074DFFF41FE76FF62" bold="true" box="[199,471,152,179]" pageId="2" pageNumber="531">Comparative material.</emphasis>
|
||
<taxonomicName id="4C3D66C49C7BFF8075C5FF40FCE6FF62" authority=", Ng, 1995 a" authorityName=", Ng" authorityYear="1995" box="[477,839,152,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7BFF8075C5FF40FD60FF63" box="[477,705,152,179]" italics="true" pageId="2" pageNumber="531">Geosesarma katibas</emphasis>
|
||
,
|
||
<bibRefCitation id="EFAC60B69C7BFF8076D6FF40FCE6FF62" author="Ng, P. K. L." box="[718,839,152,179]" pageId="2" pageNumber="531" pagination="57 - 72" refId="ref5556" refString="Ng, P. K. L. (1995 a) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995a</bibRefCitation>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="5486A3E59C7BFF807749FF41FC12FF63" box="[849,947,152,178]" pageId="2" pageNumber="531" type="holotype">holotype</typeStatus>
|
||
male (13.5 ×
|
||
<quantity id="4CC5B0A29C7BFF807056FF41FB13FF62" box="[1102,1202,152,179]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="2" pageNumber="531" unit="mm" value="13.5">13.5 mm</quantity>
|
||
) (
|
||
<collectionCode id="ED2C85829C7BFF8070D0FF40FAA1FF62" box="[1224,1280,153,179]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC</collectionCode>
|
||
1996.01834), among dead leaves, quadrat about
|
||
<quantity id="4CC5B0A29C7BFF807603FF65FDF5FF06" box="[539,596,188,215]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="531" unit="m" value="50.0">50 m</quantity>
|
||
from Sungei Katibas, east bank, about
|
||
<quantity id="4CC5B0A29C7BFF807016FF64FBF4FF06" box="[1038,1109,189,215]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="531" unit="m" value="100.0">100 m</quantity>
|
||
from mouth of Melinau, Ulu Katibas, Song District,
|
||
<collectingCountry id="F32A5DD79C7BFF807585FF39FDA1FF2B" box="[413,512,224,250]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF807614FF39FDD4FF2B" box="[524,629,224,250]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll. R.B. Stuebing,
|
||
<date id="FF833B879C7BFF80777CFF39FBB3FF2A" box="[868,1042,224,251]" pageId="2" pageNumber="531" value="1994-03-25">25 March 1994</date>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF807006FF38FDD1FECF" ID-GBIF-Occurrence="2252314109" collectionCode="ZRC" pageId="2" pageNumber="531" specimenCode="ZRC 1995.0274" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D3BD6CE9C7BFF807006FF38FB71FF2B" box="[1054,1232,224,251]" pageId="2" pageNumber="531" type="male" typeStatus="paratype">1 paratype male</specimenCount>
|
||
(13.2 ×
|
||
<quantity id="4CC5B0A29C7BFF807137FF38FA35FF2A" box="[1327,1428,225,251]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" pageId="2" pageNumber="531" unit="mm" value="13.1">13.1 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB9BB53C9C7BFF807487FEDCFEFAFECE" box="[159,347,260,287]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 1995.0274</specimenCode>
|
||
), same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF807664FEDCFF73FEB6" ID-GBIF-Occurrence="2252314094" collectingDate="2013-01-02" collectionCode="ZRC" collectorName="Sungei Boeng & Sungei Julau & Sarikei & J. Grinang" country="Malaysia" location="Sarawak" pageId="2" pageNumber="531" specimenCode="ZRC 2016.0296" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
|
||
<specimenCount id="9D3BD6CE9C7BFF807664FEDCFD75FECF" box="[636,724,260,287]" pageId="2" pageNumber="531" type="male">3 males</specimenCount>
|
||
(largest 12.2 ×
|
||
<quantity id="4CC5B0A29C7BFF807794FEDCFC54FECE" box="[908,1013,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="2" pageNumber="531" unit="mm" value="11.9">11.9 mm</quantity>
|
||
),
|
||
<specimenCount id="9D3BD6CE9C7BFF807011FEDCFBD6FECF" box="[1033,1143,260,287]" pageId="2" pageNumber="531" type="female">2 females</specimenCount>
|
||
(largest 12.3 ×
|
||
<quantity id="4CC5B0A29C7BFF807137FEDCFA35FECE" box="[1327,1428,261,287]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="531" unit="mm" value="11.8">11.8 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB9BB53C9C7BFF807487FEF0FEFBFE92" box="[159,346,296,323]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2016.0296</specimenCode>
|
||
),
|
||
<collectorName id="26C878919C7BFF807573FEF1FDADFE92" box="[363,524,296,323]" pageId="2" pageNumber="531">Sungei Boeng</collectorName>
|
||
, tributary of
|
||
<collectorName id="26C878919C7BFF807683FEF1FC8FFE93" box="[667,814,296,322]" pageId="2" pageNumber="531">Sungei Julau</collectorName>
|
||
,
|
||
<collectorName id="26C878919C7BFF807720FEF1FC2BFE93" box="[824,906,296,322]" pageId="2" pageNumber="531">Sarikei</collectorName>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80778DFEF1FC59FE93" box="[917,1016,296,322]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80701BFEF1FBCDFE93" box="[1027,1132,296,322]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26C878919C7BFF8070B7FEF0FA87FE93" box="[1199,1318,296,323]" pageId="2" pageNumber="531">J. Grinang</collectorName>
|
||
,
|
||
<collectingDate id="EFC7C26F9C7BFF807129FEF0FF73FEB6" pageId="2" pageNumber="531" value="2013-01-02">
|
||
<date id="FF833B879C7BFF807129FEF0FF73FEB6" pageId="2" pageNumber="531" value="2013-01-02">2 January 2013</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF8074C7FE94FA20FEB6" ID-GBIF-Occurrence="2252314100" box="[223,1409,332,359]" collectingDate="1994-03-22" collectionCode="ZRC" collectorName="Sungei Katibas" country="Malaysia" location="Sarawak" pageId="2" pageNumber="531" specimenCode="ZRC 2018.0748" specimenCount="1" typeStatus="paratype">
|
||
1 young male (
|
||
<specimenCode id="DB9BB53C9C7BFF807593FE94FDE6FEB6" box="[395,583,333,359]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2018.0748</specimenCode>
|
||
), station Q99, East bank,
|
||
<collectorName id="26C878919C7BFF807774FE95FBBBFEB7" box="[876,1050,332,358]" pageId="2" pageNumber="531">Sungei Katibas</collectorName>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80703FFE95FB2BFEB7" box="[1063,1162,332,358]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
, coll.
|
||
<collectingDate id="EFC7C26F9C7BFF8070C8FE94FA20FEB6" box="[1232,1409,332,359]" pageId="2" pageNumber="531" value="1994-03-22">
|
||
<date id="FF833B879C7BFF8070C8FE94FA20FEB6" box="[1232,1409,332,359]" pageId="2" pageNumber="531" value="1994-03-22">22 March 1994</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF807196FE94FDA0FE7E" ID-GBIF-Occurrence="2252314096" collectingDate="2007-04-14" collectionCode="ZRC" collectorName="Samarahan & Bintulu Division & T. M. Leong" country="Malaysia" pageId="2" pageNumber="531" specimenCode="ZRC 2007.0257" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
|
||
<specimenCount id="9D3BD6CE9C7BFF807196FE94FF6CFE5B" pageId="2" pageNumber="531" type="male">1 male</specimenCount>
|
||
(carapace crushed, ca.
|
||
<quantity id="4CC5B0A29C7BFF8075D5FEA8FDBAFE5A" box="[461,539,369,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="2" pageNumber="531" unit="mm" value="14.0">14 mm</quantity>
|
||
carapace width) (
|
||
<specimenCode id="DB9BB53C9C7BFF8076C6FEA8FC39FE5A" box="[734,920,368,395]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2007.0257</specimenCode>
|
||
), on road from
|
||
<collectorName id="26C878919C7BFF807026FEA9FB1BFE5B" box="[1086,1210,368,394]" pageId="2" pageNumber="531">Samarahan</collectorName>
|
||
to
|
||
<collectorName id="26C878919C7BFF8070C3FEA9FA39FE5B" box="[1243,1432,368,394]" pageId="2" pageNumber="531">Bintulu Division</collectorName>
|
||
, coll.
|
||
<collectorName id="26C878919C7BFF8074D7FE4CFEF7FE7E" box="[207,342,405,431]" pageId="2" pageNumber="531">T.M. Leong</collectorName>
|
||
,
|
||
<collectingDate id="EFC7C26F9C7BFF807579FE4CFDA0FE7E" box="[353,513,404,431]" pageId="2" pageNumber="531" value="2007-04-14">
|
||
<date id="FF833B879C7BFF807579FE4CFDA0FE7E" box="[353,513,404,431]" pageId="2" pageNumber="531" value="2007-04-14">14 April 2007</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</subSubSection>
|
||
</paragraph>
|
||
<caption id="DF424DCF9C78FF83748FF8B4FCEBF87B" httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="1" pageNumber="530" targetBox="[187,1400,808,1881]" targetPageId="1">
|
||
<paragraph id="8B821D479C78FF83748FF8B4FCEBF87B" pageId="1" pageNumber="530">
|
||
<emphasis id="B949C1559C78FF83748FF8B4FEB2F857" bold="true" box="[151,275,1901,1926]" pageId="1" pageNumber="530">FIGURE 1.</emphasis>
|
||
<emphasis id="B949C1559C78FF837501F8B7FE5AF857" box="[281,507,1902,1926]" italics="true" pageId="1" pageNumber="530">Geosesarma spectrum</emphasis>
|
||
<emphasis id="B949C1559C78FF83761AF8B7FD9AF857" bold="true" box="[514,571,1902,1926]" pageId="1" pageNumber="530">n. sp.</emphasis>
|
||
, colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C7BFF8074DFFE60FDA5FD56" blockId="2.[151,1437,152,647]" pageId="2" pageNumber="531">
|
||
<taxonomicName id="4C3D66C49C7BFF8074DFFE60FD0BFE02" authority="(De Man, 1902)" authorityName="De Man" authorityYear="1902" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[199,682,440,467]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7BFF8074DFFE60FE44FE03" box="[199,485,440,467]" italics="true" pageId="2" pageNumber="531">Geosesarma gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7BFF807639FE60FD03FE02" author="Man, J. G. De" box="[545,674,441,467]" pageId="2" pageNumber="531" pagination="467 - 929" refId="ref5276" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kukenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25, 467 - 929., pls. 19 - 27. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">Man, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
:
|
||
<specimenCount id="9D3BD6CE9C7BFF8076AFFE61FCAEFE03" box="[695,783,440,466]" pageId="2" pageNumber="531" type="male">6 males</specimenCount>
|
||
(largest 12.1 ×
|
||
<quantity id="4CC5B0A29C7BFF8077D0FE60FB90FE02" box="[968,1073,441,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="2" pageNumber="531" unit="mm" value="11.8">11.8 mm</quantity>
|
||
),
|
||
<specimenCount id="9D3BD6CE9C7BFF80705EFE60FB15FE03" box="[1094,1204,440,467]" pageId="2" pageNumber="531" type="female">7 females</specimenCount>
|
||
(largest 11.1 ×
|
||
<quantity id="4CC5B0A29C7BFF807173FE60FF66FE26" metricMagnitude="-2" metricUnit="m" metricValue="1.04" pageId="2" pageNumber="531" unit="mm" value="10.4">10.4 mm</quantity>
|
||
), 1 ovigerous female (12.0 ×
|
||
<quantity id="4CC5B0A29C7BFF80760CFE05FDD9FE26" box="[532,632,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="2" pageNumber="531" unit="mm" value="11.6">11.6 mm</quantity>
|
||
) (egg size
|
||
<quantity id="4CC5B0A29C7BFF8076EDFE04FCEAFE26" box="[757,843,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="531" unit="mm" value="1.2">1.2 mm</quantity>
|
||
in diameter after preservation) (ZRC 2007.262), besides trail in Glen Forest, arboreal, Lambir Hills National Park,
|
||
<collectingCountry id="F32A5DD79C7BFF80777BFDD9FC67FDCB" box="[867,966,512,538]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF8077C9FDD9FB9BFDCB" box="[977,1082,512,538]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll. T.M. Leong
|
||
<emphasis id="B949C1559C7BFF807111FDD8FAE0FDCB" box="[1289,1345,512,538]" italics="true" pageId="2" pageNumber="531">et al.</emphasis>
|
||
, 22, 23,
|
||
<date id="FF833B879C7BFF80748FFDFDFE95FDEE" box="[151,308,548,575]" pageId="2" pageNumber="531" value="2007-04-25">25 April 2007</date>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF807527FDFCFF5BFDB3" ID-GBIF-Occurrence="2252314090" collectionCode="ZRC" country="Malaysia" location="Lambir Hills National Park" pageId="2" pageNumber="531" specimenCode="ZRC 2011.1031" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D3BD6CE9C7BFF807527FDFCFE01FDEF" box="[319,416,548,575]" pageId="2" pageNumber="531" type="female">1 female</specimenCount>
|
||
(
|
||
<specimenCode id="DB9BB53C9C7BFF8075B6FDFCFDC6FDEE" box="[430,615,549,575]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2011.1031</specimenCode>
|
||
), on leaf, far from water, near termite mound,
|
||
<location id="8EE24B9C9C7BFF80707EFDFDFA39FDEF" LSID="urn:lsid:plazi:treatment:0394AC519C78FF88748FFF40FDC7FBC2:8EE24B9C9C7BFF80707EFDFDFA39FDEF" box="[1126,1432,548,574]" country="Malaysia" name="Lambir Hills National Park" pageId="2" pageNumber="531">Lambir Hills National Park</location>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80748FFD91FF5BFDB3" box="[151,250,584,610]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B55171A9C7BFF80751FFD91FE5EFD56" ID-GBIF-Occurrence="2252314083" collectingDate="2001-02-19" collectionCode="ZRC" collectorName="H. S. Yong" country="Malaysia" location="Niah National Park" pageId="2" pageNumber="531" specimenCode="ZRC 2011.1030" specimenCount="1" specimenCount-male="1">
|
||
coll.
|
||
<collectorName id="26C878919C7BFF807559FD90FE18FDB2" box="[321,441,585,611]" pageId="2" pageNumber="531">H.S. Yong</collectorName>
|
||
, 1995;
|
||
<specimenCount id="9D3BD6CE9C7BFF807617FD90FDFAFDB3" box="[527,603,584,611]" pageId="2" pageNumber="531" type="male">1 male</specimenCount>
|
||
(13.0 ×
|
||
<quantity id="4CC5B0A29C7BFF8076D8FD90FC86FDB2" box="[704,807,585,611]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="2" pageNumber="531" unit="mm" value="12.8">12.8 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB9BB53C9C7BFF807727FD90FC5AFDB2" box="[831,1019,585,611]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2011.1030</specimenCode>
|
||
),
|
||
<location id="8EE24B9C9C7BFF807017FD91FB4FFDB3" LSID="urn:lsid:plazi:treatment:0394AC519C78FF88748FFF40FDC7FBC2:8EE24B9C9C7BFF807017FD91FB4FFDB3" box="[1039,1262,584,610]" country="Malaysia" name="Niah National Park" pageId="2" pageNumber="531">Niah National Park</location>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF8070E3FD91FAFFFDB3" box="[1275,1374,584,610]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
, coll. local rangers,
|
||
<collectingDate id="EFC7C26F9C7BFF80752DFDB4FE5EFD56" box="[309,511,620,647]" pageId="2" pageNumber="531" value="2001-02-19">
|
||
<date id="FF833B879C7BFF80752DFDB4FE5EFD56" box="[309,511,620,647]" pageId="2" pageNumber="531" value="2001-02-19">19 February 2001</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF424DCF9C7BFF80748FF9ABFAACF97E" httpUri="https://zenodo.org/record/3243078/files/figure.png" pageId="2" pageNumber="531" startId="2.[151,250,1650,1675]" targetBox="[187,1400,668,1630]" targetPageId="2">
|
||
<paragraph id="8B821D479C7BFF80748FF9ABFAACF97E" blockId="2.[151,1436,1650,1711]" pageId="2" pageNumber="531">
|
||
<emphasis id="B949C1559C7BFF80748FF9ABFEB2F95A" bold="true" box="[151,275,1650,1675]" pageId="2" pageNumber="531">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C3D66C49C7BFF807502F9AAFE5DF95B" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[282,508,1651,1674]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7BFF807502F9AAFE5DF95B" box="[282,508,1651,1674]" italics="true" pageId="2" pageNumber="531">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7BFF80761BF9ABFD9DF95B" box="[515,572,1650,1674]" pageId="2" pageNumber="531" rank="species">
|
||
<emphasis id="B949C1559C7BFF80761BF9ABFD9DF95B" bold="true" box="[515,572,1650,1674]" pageId="2" pageNumber="531">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, colour in life. A, B, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740); C, paratype male (7.9 × 7.6 mm) (ZRC 2018.0741); D, paratype female (10.2 × 9.8 mm) (ZRC 2018.0741). All from Brunei.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C7BFF8074DFF904FAFFF856" blockId="2.[151,1437,1756,2035]" pageId="2" pageNumber="531">
|
||
<treatmentCitationGroup id="AB2D3A699C7BFF8074DFF904FAFFF856" infered="true" pageId="2" pageNumber="531">
|
||
<taxonomicName id="4C3D66C49C7BFF8074DFF904FD26F926" ID-CoL="3FS8V" authority="Ng & Grinang, 2004: 1" authorityName="Ng & Grinang" authorityPageNumber="1" authorityYear="2004" box="[199,647,1757,1783]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="531" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B949C1559C7BFF8074DFF904FE22F926" box="[199,387,1757,1783]" italics="true" pageId="2" pageNumber="531">Geosesarma bau</emphasis>
|
||
<treatmentCitation id="0A9C3B569C7BFF807590F904FD26F926" author="Ng, P. K. L. & Grinang, J." box="[392,647,1757,1783]" page="1" pageId="2" pageNumber="531" year="2004">
|
||
<bibRefCitation id="EFAC60B69C7BFF807590F904FD26F926" author="Ng, P. K. L. & Grinang, J." box="[392,647,1757,1783]" pageId="2" pageNumber="531" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">
|
||
Ng & Grinang, 2004:
|
||
<specimenCount id="9D3BD6CE9C7BFF807661F904FD26F926" box="[633,647,1757,1783]" pageId="2" pageNumber="531" type="female">1</specimenCount>
|
||
</bibRefCitation>
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
female (ZRC 2018.0750), in cup of
|
||
<taxonomicName id="4C3D66C49C7BFF80700FF904FA3DF926" authority="(Jack, 1835)" authorityName="Jack" authorityYear="1835" baseAuthorityName="Jack" baseAuthorityYear="1835" box="[1047,1436,1756,1783]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="2" pageNumber="531" phylum="Tracheophyta" rank="species" species="ampullaria">
|
||
<emphasis id="B949C1559C7BFF80700FF904FAACF926" box="[1047,1293,1757,1783]" italics="true" pageId="2" pageNumber="531">Nepenthes ampullaria</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC60B69C7BFF807103F904FA35F926" author="Jack, W" box="[1307,1428,1756,1783]" pageId="2" pageNumber="531" pagination="253 - 272" refId="ref5161" refString="Jack, W (1835) Description on Malayan Plants. Companion to the Botanical Magazine, 1, 121 - 157, 253 - 272." type="journal article" year="1835">Jack, 1835</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
[Nepentheceae], Mount Santubong,
|
||
<collectingCountry id="F32A5DD79C7BFF807628F8D8FD32F8CA" box="[560,659,1793,1819]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF8076B8F8D8FCA8F8CA" box="[672,777,1793,1819]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll. W. Denzer,
|
||
<date id="FF833B879C7BFF8077CEF8D8FBD6F8CA" box="[982,1143,1793,1819]" pageId="2" pageNumber="531" value="1992-04-10">10 April 1992</date>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF80709CF8D8FD6DF8B2" ID-GBIF-Occurrence="2252314104" collectingDate="1995-09-05" collectionCode="ZRC" collectorName="Kuching & Serian & Kuap & P. K. L. Ng" country="Malaysia" latitude="1.4191389" location="Sarawak" longLatPrecision="1" longitude="110.34803" pageId="2" pageNumber="531" specimenCode="ZRC 2002.0228" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="9D3BD6CE9C7BFF80709CF8D8FB46F8CA" box="[1156,1255,1793,1819]" pageId="2" pageNumber="531" type="female">1 female</specimenCount>
|
||
(9.6 ×
|
||
<quantity id="4CC5B0A29C7BFF807124F8D8FA35F8CA" box="[1340,1428,1793,1819]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="531" unit="mm" value="9.3">9.3 mm</quantity>
|
||
) (
|
||
<specimenCode id="DB9BB53C9C7BFF807487F8FCFEFBF8EE" box="[159,346,1829,1855]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC 2002.0228</specimenCode>
|
||
), road from
|
||
<collectorName id="26C878919C7BFF8075FBF8FCFDE2F8EE" box="[483,579,1829,1855]" pageId="2" pageNumber="531">Kuching</collectorName>
|
||
to
|
||
<collectorName id="26C878919C7BFF807670F8FCFD13F8EE" box="[616,690,1829,1855]" pageId="2" pageNumber="531">Serian</collectorName>
|
||
, 10th mile rock, into
|
||
<collectorName id="26C878919C7BFF8077BBF8FCFC43F8EE" box="[931,994,1829,1855]" pageId="2" pageNumber="531">Kuap</collectorName>
|
||
,
|
||
<geoCoordinate id="EE097B809C7BFF8077F6F8FDFBD6F8EE" box="[1006,1143,1828,1855]" degrees="1" direction="north" minutes="25" orientation="latitude" pageId="2" pageNumber="531" precision="1" seconds="8.9" value="1.4191389">1°25’8.9” N</geoCoordinate>
|
||
<geoCoordinate id="EE097B809C7BFF807066F8FDFA88F8EE" box="[1150,1321,1828,1855]" degrees="110" direction="east" minutes="20" orientation="longitude" pageId="2" pageNumber="531" precision="1" seconds="52.9" value="110.34803">110°20’52.9”E</geoCoordinate>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80712DF8FCFA39F8EE" box="[1333,1432,1829,1855]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80748FF890FEA1F8B2" box="[151,256,1865,1891]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll.
|
||
<collectorName id="26C878919C7BFF80755BF890FE13F8B2" box="[323,434,1865,1891]" pageId="2" pageNumber="531">P.K.L. Ng</collectorName>
|
||
<emphasis id="B949C1559C7BFF8075A1F890FE50F8B2" box="[441,497,1865,1891]" italics="true" pageId="2" pageNumber="531">et al.</emphasis>
|
||
,
|
||
<collectingDate id="EFC7C26F9C7BFF8075E6F891FD6DF8B2" box="[510,716,1864,1891]" pageId="2" pageNumber="531" value="1995-09-05">
|
||
<date id="FF833B879C7BFF8075E6F891FD6DF8B2" box="[510,716,1864,1891]" pageId="2" pageNumber="531" value="1995-09-05">5 September 1995</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B55171A9C7BFF8076CFF890FAF8F856" ID-GBIF-Occurrence="2252314085" collectionCode="ZRC" country="Malaysia" county="Sarikei" location="River Chuat" municipality="Sibu" pageId="2" pageNumber="531" specimenCount="1">
|
||
1 ovigerous female (11.2 ×
|
||
<quantity id="4CC5B0A29C7BFF807011F890FBCCF8B2" box="[1033,1133,1865,1891]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="2" pageNumber="531" unit="mm" value="10.9">10.9 mm</quantity>
|
||
) (egg size about 1.0 mm in diameter, eggs wrinkled) (
|
||
<collectionCode id="ED2C85829C7BFF8075A4F8B4FE55F856" box="[444,500,1901,1927]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="531">ZRC</collectionCode>
|
||
1965.7.29.5),
|
||
<location id="8EE24B9C9C7BFF80768FF8B4FC82F856" LSID="urn:lsid:plazi:treatment:0394AC519C78FF88748FFF40FDC7FBC2:8EE24B9C9C7BFF80768FF8B4FC82F856" box="[663,803,1901,1927]" country="Malaysia" county="Sarikei" municipality="Sibu" name="River Chuat" pageId="2" pageNumber="531">River Chuat</location>
|
||
,
|
||
<collectingMunicipality id="6BE6873D9C7BFF807735F8B4FCC2F856" box="[813,867,1901,1927]" pageId="2" pageNumber="531">Sibu</collectingMunicipality>
|
||
, near
|
||
<collectingCounty id="62E365CB9C7BFF8077BEF8B4FC59F856" box="[934,1016,1901,1927]" pageId="2" pageNumber="531">Sarikei</collectingCounty>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80701BF8B4FBC7F856" box="[1027,1126,1901,1927]" name="Malaysia" pageId="2" pageNumber="531">Sarawak</collectingCountry>
|
||
,
|
||
<collectingCountry id="F32A5DD79C7BFF80706AF8B4FB7AF856" box="[1138,1243,1901,1927]" name="Malaysia" pageId="2" pageNumber="531">Malaysia</collectingCountry>
|
||
, coll. 1948
|
||
</materialsCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<subSubSection id="C3274ECC9C7BFF8174DFF849FBE0FD1E" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531" type="diagnosis">
|
||
<paragraph id="8B821D479C7BFF8174DFF849FCE0FDEE" blockId="2.[151,1437,1756,2035]" lastBlockId="3.[151,1437,152,863]" lastPageId="3" lastPageNumber="532" pageId="2" pageNumber="531">
|
||
<emphasis id="B949C1559C7BFF8074DFF849FE3FF87A" bold="true" box="[199,414,1936,1963]" pageId="2" pageNumber="531">Diagnosis of male.</emphasis>
|
||
Carapace quadrate, marginally wider than long or subequal, width to length ratio 1.01–1.06, lateral margins parallel, gently diverging towards posterior carapace margin (
|
||
<figureCitation id="130601C29C7BFF807011F86CFBCFF81E" box="[1033,1134,1973,1999]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="2" pageNumber="531">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7BFF807064F86DFB03F81E" box="[1148,1186,1972,1999]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="2" pageNumber="531">5A</figureCitation>
|
||
); dorsal surfaces with well-defined regions, anterior half with numerous distinct tubercles of varying sizes, posterior half with smaller, lower tubercles (
|
||
<figureCitation id="130601C29C7AFF81754EFF40FE1AFF62" box="[342,443,153,179]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF8175D1FF41FE4EFF62" box="[457,495,152,179]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="3" pageNumber="532">5A</figureCitation>
|
||
); frontal margin distinctly deflexed, frontal lobes broad, with truncated margins in dorsal view, separated by shallow median concavity; postfrontal and postorbital cristae sharp, distinct (
|
||
<figureCitation id="130601C29C7AFF81711EFF64FAC8FF06" box="[1286,1385,189,215]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF81716BFF65FA39FF06" box="[1395,1432,188,215]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="3" pageNumber="532">5A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF81748FFF38FF1CFF2D" box="[151,189,225,252]" captionStart="FIGURE 6" captionStartId="5.[151,250,1933,1958]" captionTargetId="figure@5.[151,1436,1248,1914]" captionTargetPageId="5" captionText="FIGURE 6. A, C, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, D, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak. A, B, frontal view of cephalothorax; C, D, inner view of left chela." httpUri="https://zenodo.org/record/3404941/files/figure.png" pageId="3" pageNumber="532">6A</figureCitation>
|
||
); external orbital angle triangular, directed obliquely anteriorly, clearly extending past lateral carapace margins, separated from first epibranchial tooth by distinct cleft; first epibranchial tooth low, blunt, second epibranchial tooth indistinct, separated from first by shallow notch (
|
||
<figureCitation id="130601C29C7AFF8176A1FEF0FCBDFE92" box="[697,796,297,323]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF817730FEF1FCEFFE92" box="[808,846,296,323]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="3" pageNumber="532">5A</figureCitation>
|
||
); merus of third maxilliped rounded; exopod slender, flagellum absent (
|
||
<figureCitation id="130601C29C7AFF817579FE94FE1AFEB6" box="[353,443,333,359]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="3" pageNumber="532">Fig. 9A</figureCitation>
|
||
); outer surfaces of palm of chela covered with small rounded granules; dorsal margin of dactylus with 11 sharp, anteriorly directed tubercles (
|
||
<figureCitation id="130601C29C7AFF8176F7FEA8FCF0FE5A" box="[751,849,369,395]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1C</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF817747FEA8FC20FE5A" box="[863,897,369,395]" captionStart="FIGURE 6" captionStartId="5.[151,250,1933,1958]" captionTargetId="figure@5.[151,1436,1248,1914]" captionTargetPageId="5" captionText="FIGURE 6. A, C, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, D, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak. A, B, frontal view of cephalothorax; C, D, inner view of left chela." httpUri="https://zenodo.org/record/3404941/files/figure.png" pageId="3" pageNumber="532">6C</figureCitation>
|
||
, 7A); ambulatory legs with long, slender merus with sharp subdistal spine on dorsal margin, surface weakly rugose, propodus slender, relatively long (
|
||
<figureCitation id="130601C29C7AFF81712AFE4CFA39FE7E" box="[1330,1432,405,431]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1A</figureCitation>
|
||
, 7G, H); pleon triangular; somite 6 broadest, with lateral margins gently convex; telson triangular, longer than broad, lateral margins gently convex (
|
||
<figureCitation id="130601C29C7AFF8175EFFE04FDFBFE26" box="[503,602,477,503]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="3" pageNumber="532">Figs. 1B</figureCitation>
|
||
, 8A, B); G1 stout, proximal, distal part bent at angle of ca. 45º from longitudinal axis (
|
||
<figureCitation id="130601C29C7AFF817558FDD8FE04FDCA" box="[320,421,513,539]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="3" pageNumber="532">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF8175AAFDD8FE61FDCA" box="[434,448,513,539]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="3" pageNumber="532">9</figureCitation>
|
||
B–E), distal part elongate in lateral view, spatuliform from marginal view, with distinct median cleft on tip when viewed ventrally (
|
||
<figureCitation id="130601C29C7AFF817667FDFCFD42FDEE" box="[639,739,549,575]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="3" pageNumber="532">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF8176F7FDFCFC95FDEE" box="[751,820,549,575]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="3" pageNumber="532">9D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B821D479C7AFF8174DFFD90FBE0FD1E" blockId="3.[151,1437,152,863]" pageId="3" pageNumber="532">
|
||
<emphasis id="B949C1559C7AFF8174DFFD90FE91FDB2" bold="true" box="[199,304,585,611]" pageId="3" pageNumber="532">Females.</emphasis>
|
||
Females resemble the males in all non-sexual characters, with chelae equal, relatively more slender, less inflated, tubercles on outer surface of palm and dorsal margin of dactylus smaller, less distinct (
|
||
<figureCitation id="130601C29C7AFF817108FDB4FA39FD56" box="[1296,1432,621,647]" captionStart="FIGURE 3" captionStartId="3.[151,250,1937,1962]" captionTargetBox="[188,1399,885,1917]" captionTargetId="figure@3.[188,1399,885,1917]" captionTargetPageId="3" captionText="FIGURE 3. Geosesarma spectrum n. sp., colour in life. A, female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B–D, female (11.2 × 10.7 mm) (ZRC 2018.0741), Brunei.A, overall habitus; B, ovigerous female with late-stage eggs; C, female with newly hatched juvenile crabs on right side of carapace; D, female with juvenile crabs on right side of carapace." httpUri="https://zenodo.org/record/3243080/files/figure.png" pageId="3" pageNumber="532">Figs. 3A, B</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7AFF81748FFD49FF1DFD7A" box="[151,188,656,683]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="3" pageNumber="532">5C</figureCitation>
|
||
). Pleon ovate, covering most of thoracic sternum (
|
||
<figureCitation id="130601C29C7AFF8176E1FD48FCC2FD7A" box="[761,867,657,683]" captionStart="FIGURE 10" captionStartId="9.[151,250,1593,1618]" captionTargetBox="[151,1436,597,1574]" captionTargetId="figure@9.[151,1436,597,1574]" captionTargetPageId="9" captionText="FIGURE 10. A, C, Geosesarma spectrum n. sp., paratype ovigerous female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B, D, G. katibas Ng, 1995, female (12.3 x 11.8 mm) (ZRC 2016.0296), Sarawak. A, B, female thoracic sternum and pleon; C, D, female sternopleonal cavity and vulvae." httpUri="https://zenodo.org/record/3243090/files/figure.png" pageId="3" pageNumber="532">Fig. 10A</figureCitation>
|
||
); vulvae positioned on anterior margin of thoracic sternite 6, with a round, stiff operculum, without sternal opercular cover (
|
||
<figureCitation id="130601C29C7AFF8177D6FD6CFB94FD1E" box="[974,1077,693,719]" captionStart="FIGURE 10" captionStartId="9.[151,250,1593,1618]" captionTargetBox="[151,1436,597,1574]" captionTargetId="figure@9.[151,1436,597,1574]" captionTargetPageId="9" captionText="FIGURE 10. A, C, Geosesarma spectrum n. sp., paratype ovigerous female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B, D, G. katibas Ng, 1995, female (12.3 x 11.8 mm) (ZRC 2016.0296), Sarawak. A, B, female thoracic sternum and pleon; C, D, female sternopleonal cavity and vulvae." httpUri="https://zenodo.org/record/3243090/files/figure.png" pageId="3" pageNumber="532">Fig. 10C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C7AFF8674DFFD01FC0AFAAA" lastPageId="4" lastPageNumber="533" pageId="3" pageNumber="532" type="description">
|
||
<paragraph id="8B821D479C7AFF8174DFFD01FE7DFC8E" blockId="3.[151,1437,152,863]" pageId="3" pageNumber="532">
|
||
<emphasis id="B949C1559C7AFF8174DFFD01FE9EFD22" bold="true" box="[199,319,728,755]" pageId="3" pageNumber="532">Variation.</emphasis>
|
||
The G1s of a smaller
|
||
<typeStatus id="5486A3E59C7AFF817628FD00FD31FD22" box="[560,656,729,755]" pageId="3" pageNumber="532" type="paratype">paratype</typeStatus>
|
||
male (7.9 ×
|
||
<quantity id="4CC5B0A29C7AFF817702FD00FCD1FD22" box="[794,880,729,755]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="3" pageNumber="532" unit="mm" value="7.6">7.6 mm</quantity>
|
||
) agree well with that of the
|
||
<typeStatus id="5486A3E59C7AFF8170BAFD00FAA9FD22" box="[1186,1288,729,755]" pageId="3" pageNumber="532" type="holotype">holotype</typeStatus>
|
||
. The external orbital angle is more slender, relatively shorter, blunter and does not extend past the lateral margins of the carapace, and the tubercles on the anterior half of the carapace are also less pronounced (
|
||
<figureCitation id="130601C29C7AFF817011FCF8FBC0FCEA" box="[1033,1121,800,827]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="3" pageNumber="532">Fig. 5B</figureCitation>
|
||
). The chelae resemble those of the female detailed above.
|
||
</paragraph>
|
||
<caption id="DF424DCF9C7AFF81748FF848FB19F820" httpUri="https://zenodo.org/record/3243080/files/figure.png" pageId="3" pageNumber="532" startId="3.[151,250,1937,1962]" targetBox="[188,1399,885,1917]" targetPageId="3">
|
||
<paragraph id="8B821D479C7AFF81748FF848FB19F820" blockId="3.[151,1436,1937,2034]" pageId="3" pageNumber="532">
|
||
<emphasis id="B949C1559C7AFF81748FF848FEB5F87B" bold="true" box="[151,276,1937,1962]" pageId="3" pageNumber="532">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C3D66C49C7AFF817503F84BFE5FF878" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[283,510,1938,1961]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="532" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7AFF817503F84BFE5FF878" box="[283,510,1938,1961]" italics="true" pageId="3" pageNumber="532">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7AFF81761DF84BFD9EF87B" box="[517,575,1938,1962]" pageId="3" pageNumber="532" rank="species">
|
||
<emphasis id="B949C1559C7AFF81761DF84BFD9EF87B" bold="true" box="[517,575,1938,1962]" pageId="3" pageNumber="532">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, colour in life. A, female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B–D, female (11.2 × 10.7 mm) (ZRC 2018.0741), Brunei. A, overall habitus; B, ovigerous female with late-stage eggs; C, female with newly hatched juvenile crabs on right side of carapace; D, female with juvenile crabs on right side of carapace.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF424DCF9C7DFF86748FFC6BFC81FBE6" httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="4" pageNumber="533" startId="4.[151,250,946,971]" targetBox="[187,1400,181,927]" targetPageId="4">
|
||
<paragraph id="8B821D479C7DFF86748FFC6BFC81FBE6" blockId="4.[151,1437,946,1079]" pageId="4" pageNumber="533">
|
||
<emphasis id="B949C1559C7DFF86748FFC6BFEB5FC1A" bold="true" box="[151,276,946,971]" pageId="4" pageNumber="533">FIGURE 4.</emphasis>
|
||
A,
|
||
<taxonomicName id="4C3D66C49C7DFF867522FC6AFDD2FC1A" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" box="[314,627,946,971]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7DFF867522FC6AFDA8FC1A" box="[314,521,947,971]" italics="true" pageId="4" pageNumber="533">Geosesarma katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B,
|
||
<taxonomicName id="4C3D66C49C7DFF86719CFC6AFEE6FC3E" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7DFF86719CFC6AFF41FC3E" italics="true" pageId="4" pageNumber="533">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C,
|
||
<taxonomicName id="4C3D66C49C7DFF86704BFC0EFA39FC3E" authority="(De Man, 1892)" authorityName="De Man" authorityYear="1892" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[1107,1432,983,1007]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7DFF86704BFC0EFB4FFC3E" box="[1107,1262,983,1007]" italics="true" pageId="4" pageNumber="533">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7DFF867138FC0EFA30FC3E" author="Man, J. G. De" box="[1312,1425,983,1007]" pageId="4" pageNumber="533" pagination="225 - 264" refId="ref5188" refString="Man, J. G. De (1892) Carcinological studies in the Leyden Museum. No. 6. Notes from the Leyden Museum, 14, 225 - 264., pls. 7 - 10." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male, Lambir Hills National Park, Sarawak (specimen not collected); D,
|
||
<taxonomicName id="4C3D66C49C7DFF867769FC22FB12FBC2" authority="(De Man, 1892)" authorityName="De Man" authorityYear="1892" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[881,1203,1019,1043]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7DFF867769FC22FBAAFBC2" box="[881,1035,1019,1043]" italics="true" pageId="4" pageNumber="533">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7DFF867024FC22FB0DFBC2" author="Man, J. G. De" box="[1084,1196,1019,1043]" pageId="4" pageNumber="533" pagination="225 - 264" refId="ref5188" refString="Man, J. G. De (1892) Carcinological studies in the Leyden Museum. No. 6. Notes from the Leyden Museum, 14, 225 - 264., pls. 7 - 10." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C7DFF8674DFFBBDFC0AFAAA" blockId="4.[151,1437,1124,2016]" pageId="4" pageNumber="533">
|
||
<emphasis id="B949C1559C7DFF8674DFFBBDFE33FBAE" bold="true" box="[199,402,1124,1151]" pageId="4" pageNumber="533">Live colouration.</emphasis>
|
||
In live specimens, the anterior half of the carapace is a deep, rusty, reddish-brown; the posterior half of the carapace and ambulatory meri being pale purple, while the carpi, propodi and dactyli are pale orange (
|
||
<figureCitation id="130601C29C7DFF867487FB74FEA4FB16" box="[159,261,1197,1223]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="4" pageNumber="533">Figs. 1A</figureCitation>
|
||
, 2A, 3A). The carpus, merus and palm of the chelipeds are bright purple in adult males, slightly duller in females and smaller specimens, with the fingers bright orange (
|
||
<figureCitation id="130601C29C7DFF867790FB08FBB1FB3A" box="[904,1040,1233,1259]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="4" pageNumber="533">Figs. 1A, C</figureCitation>
|
||
, 2B, 3B). The posterior half of the ambulatory legs are covered in white spots, which are faint to absent in large adult males but distinct in juveniles and all females (
|
||
<figureCitation id="130601C29C7DFF867555FAC0FE71FAE5" box="[333,464,1305,1332]" captionStart="FIGURE 2" captionStartId="2.[151,250,1650,1675]" captionTargetBox="[187,1400,668,1630]" captionTargetId="figure@2.[187,1400,668,1630]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma spectrum n. sp., colour in life. A, B, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740); C, paratype male (7.9 × 7.6 mm) (ZRC 2018.0741); D, paratype female (10.2 × 9.8 mm) (ZRC 2018.0741). All from Brunei." httpUri="https://zenodo.org/record/3243078/files/figure.png" pageId="4" pageNumber="533">Figs. 2C, D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7DFF8675C3FAC0FD81FAE2" box="[475,544,1305,1331]" captionStart="FIGURE 4" captionStartId="4.[151,250,946,971]" captionTargetBox="[187,1400,181,927]" captionTargetId="figure@4.[187,1400,181,927]" captionTargetPageId="4" captionText="FIGURE 4. A, Geosesarma katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B, G. katibas Ng, 1995, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C, G. gracillimum (De Man, 1892), male, Lambir Hills National Park, Sarawak (specimen not collected); D, G. gracillimum (De Man, 1892), female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected)." httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="4" pageNumber="533">4A, C</figureCitation>
|
||
). The white spots sometimes extend to the posterior half of the carapace in small specimens (
|
||
<figureCitation id="130601C29C7DFF867503FAE4FE00FA89" box="[283,417,1341,1368]" captionStart="FIGURE 2" captionStartId="2.[151,250,1650,1675]" captionTargetBox="[187,1400,668,1630]" captionTargetId="figure@2.[187,1400,668,1630]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma spectrum n. sp., colour in life. A, B, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740); C, paratype male (7.9 × 7.6 mm) (ZRC 2018.0741); D, paratype female (10.2 × 9.8 mm) (ZRC 2018.0741). All from Brunei." httpUri="https://zenodo.org/record/3243078/files/figure.png" pageId="4" pageNumber="533">Figs. 2C, D</figureCitation>
|
||
). The ventral surfaces are a pale yellow with hints of purple, particularly on the telson (
|
||
<figureCitation id="130601C29C7DFF86716DFAE4FF18FAAA" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="4" pageNumber="533">Fig 1B</figureCitation>
|
||
). The eyes are olive green, and the eggs are orangish red (
|
||
<figureCitation id="130601C29C7DFF86775DFAB8FC3CFAAA" box="[837,925,1377,1403]" captionStart="FIGURE 4" captionStartId="4.[151,250,946,971]" captionTargetBox="[187,1400,181,927]" captionTargetId="figure@4.[187,1400,181,927]" captionTargetPageId="4" captionText="FIGURE 4. A, Geosesarma katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B, G. katibas Ng, 1995, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C, G. gracillimum (De Man, 1892), male, Lambir Hills National Park, Sarawak (specimen not collected); D, G. gracillimum (De Man, 1892), female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected)." httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="4" pageNumber="533">Fig. 4B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C7DFF8674DFFA5FFE54FA12" pageId="4" pageNumber="533" type="etymology">
|
||
<paragraph id="8B821D479C7DFF8674DFFA5FFE54FA12" blockId="4.[151,1437,1124,2016]" pageId="4" pageNumber="533">
|
||
<emphasis id="B949C1559C7DFF8674DFFA5FFEEFFA71" bold="true" box="[199,334,1414,1440]" pageId="4" pageNumber="533">Etymology.</emphasis>
|
||
From the Latin for a colorful image, alluding to the striking colour of the species in life. The name is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C7DFF8474DFFA17FF44F865" lastPageId="6" lastPageNumber="535" pageId="4" pageNumber="533" type="discussion">
|
||
<paragraph id="8B821D479C7DFF8674DFFA17FF48F982" blockId="4.[151,1437,1124,2016]" pageId="4" pageNumber="533">
|
||
<emphasis id="B949C1559C7DFF8674DFFA17FE9AFA39" bold="true" box="[199,315,1486,1512]" pageId="4" pageNumber="533">Remarks.</emphasis>
|
||
In the structure of the G1, carapace features and colouration in life,
|
||
<taxonomicName id="4C3D66C49C7DFF867059FA17FA9AFA36" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[1089,1339,1486,1512]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7DFF867059FA17FA9AFA36" box="[1089,1339,1486,1512]" italics="true" pageId="4" pageNumber="533">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7DFF86715AFA17FA23FA39" box="[1346,1410,1486,1512]" pageId="4" pageNumber="533" rank="species">
|
||
<emphasis id="B949C1559C7DFF86715AFA17FA23FA39" bold="true" box="[1346,1410,1486,1512]" pageId="4" pageNumber="533">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is closest to
|
||
<taxonomicName id="4C3D66C49C7DFF86751DFA2BFE55F9DA" authority="Ng, 1995 a" authorityName=", Ng" authorityYear="1995" box="[261,500,1520,1548]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7DFF86751DFA2BFED9F9DA" box="[261,376,1521,1548]" italics="true" pageId="4" pageNumber="533">G. katibas</emphasis>
|
||
<bibRefCitation id="EFAC60B69C7DFF867566FA2BFE55F9DA" author="Ng, P. K. L." box="[382,500,1520,1548]" pageId="4" pageNumber="533" pagination="57 - 72" refId="ref5556" refString="Ng, P. K. L. (1995 a) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995a</bibRefCitation>
|
||
</taxonomicName>
|
||
, from central
|
||
<collectingCountry id="F32A5DD79C7DFF867695FA28FD50F9DA" box="[653,753,1521,1547]" name="Malaysia" pageId="4" pageNumber="533">Sarawak</collectingCountry>
|
||
. It can, however, be distinguished from the latter by the shape of the external orbital angle and epibranchial teeth, length of the ambulatory leg propodi, as well as the structure of the G1.
|
||
</paragraph>
|
||
<paragraph id="8B821D479C7DFF8674DFF987FC63F831" blockId="4.[151,1437,1124,2016]" pageId="4" pageNumber="533">
|
||
In
|
||
<taxonomicName id="4C3D66C49C7DFF8674FDF987FECCF9A6" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[229,365,1630,1656]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7DFF8674FDF987FECCF9A6" box="[229,365,1630,1656]" italics="true" pageId="4" pageNumber="533">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7DFF86756BF987FE11F9A9" box="[371,432,1630,1656]" pageId="4" pageNumber="533" rank="species">
|
||
<emphasis id="B949C1559C7DFF86756BF987FE11F9A9" bold="true" box="[371,432,1630,1656]" pageId="4" pageNumber="533">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, the external orbital angle is typically marginally narrower, and the cleft separating it from the first epibranchial tooth deeper (
|
||
<figureCitation id="130601C29C7DFF86763FF958FD25F94A" box="[551,644,1665,1691]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="4" pageNumber="533">Figs.1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7DFF867689F959FD3CF94A" box="[657,669,1664,1691]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="4" pageNumber="533">5</figureCitation>
|
||
A–C) (vs. marginally broader external orbital angle, shallower cleft separating external orbital angle from the first epibranchial tooth,
|
||
<figureCitation id="130601C29C7DFF867769F97CFC53F911" box="[881,1010,1700,1728]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="4" pageNumber="533">Figs. 5D, E</figureCitation>
|
||
). The propodus of the ambulatory legs are longer (
|
||
<figureCitation id="130601C29C7DFF867501F910FEDFF932" box="[281,382,1737,1763]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="4" pageNumber="533">Figs. 1A</figureCitation>
|
||
, 7G, H) (vs. comparatively shorter in similar sized specimens,
|
||
<figureCitation id="130601C29C7DFF867052F910FB1CF935" box="[1098,1213,1737,1764]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="4" pageNumber="533">
|
||
Figs.
|
||
<date id="FF833B879C7DFF867092F913FB02F935" box="[1162,1187,1738,1764]" pageId="4" pageNumber="533">7I</date>
|
||
, J
|
||
</figureCitation>
|
||
). The proximal and distal parts of the G1 are less strongly bent when viewed laterally, and the tip of the distal part has a distinct median cleft, clearly visible from the ventral view (
|
||
<figureCitation id="130601C29C7DFF867662F8C8FD7CF8FD" box="[634,733,1809,1836]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="4" pageNumber="533">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7DFF8676F0F8C8FD54F8FA" box="[744,757,1809,1835]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="4" pageNumber="533">9</figureCitation>
|
||
B–E) (vs. proximal and distal parts of G1 more strongly bent when viewed laterally, distinct median cleft at tip absent,
|
||
<figureCitation id="130601C29C7DFF86770DF8ECFCD5F881" box="[789,884,1845,1872]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="4" pageNumber="533">Figs. 7E</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7DFF867767F8ECFC67F881" box="[895,966,1845,1872]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="4" pageNumber="533">9G, H</figureCitation>
|
||
).
|
||
<taxonomicName id="4C3D66C49C7DFF8677C1F8EFFB1DF89E" authorityName=", Ng" authorityYear="1995" box="[985,1212,1845,1872]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7DFF8677C1F8EFFB1DF89E" box="[985,1212,1845,1872]" italics="true" pageId="4" pageNumber="533">Geosesarma katibas</emphasis>
|
||
</taxonomicName>
|
||
was described from Ulu Katibas basin, Song District,
|
||
<collectingCountry id="F32A5DD79C7DFF867613F880FDCFF8A2" box="[523,622,1881,1907]" name="Malaysia" pageId="4" pageNumber="533">Sarawak</collectingCountry>
|
||
, with more recent material collected from Kanowit basin, Sarikei district, ca.
|
||
<quantity id="4CC5B0A29C7DFF8674D8F8A5FEA9F846" box="[192,264,1916,1943]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="533" unit="km" value="50.0">50 km</quantity>
|
||
away, two major downstream tributaries of the Rajang river. Given the limited distributional ranges of many
|
||
<emphasis id="B949C1559C7DFF8674C4F87BFECEF86D" box="[220,367,1954,1980]" italics="true" pageId="4" pageNumber="533">
|
||
<taxonomicName id="4C3D66C49C7DFF8674C4F87BFECAF86D" authorityName="De Man" authorityYear="1892" box="[220,363,1954,1980]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="genus">Geosesarma</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
it can be assumed that the distribution does not overlap with
|
||
<taxonomicName id="4C3D66C49C7DFF867007F87BFB09F86A" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[1055,1192,1954,1980]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="533" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7DFF867007F87BFB09F86A" box="[1055,1192,1954,1980]" italics="true" pageId="4" pageNumber="533">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7DFF8670B7F87BFB4FF86D" box="[1199,1262,1954,1980]" pageId="4" pageNumber="533" rank="species">
|
||
<emphasis id="B949C1559C7DFF8670B7F87BFB4FF86D" bold="true" box="[1199,1262,1954,1980]" pageId="4" pageNumber="533">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, which have so far only been collected in several localities in
|
||
<collectingCountry id="F32A5DD79C7DFF867682F81CFD48F80E" box="[666,745,1989,2015]" name="Brunei" pageId="4" pageNumber="533">Brunei</collectingCountry>
|
||
, over
|
||
<quantity id="4CC5B0A29C7DFF867735F81CFCDAF80E" box="[813,891,1989,2015]" metricMagnitude="5" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="533" unit="km" value="400.0">400km</quantity>
|
||
away.
|
||
</paragraph>
|
||
<caption id="DF424DCF9C7CFF87748FFBF8FD40FB1B" httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="5" pageNumber="534" startId="5.[151,250,1057,1082]" targetBox="[151,1436,181,1037]" targetPageId="5">
|
||
<paragraph id="8B821D479C7CFF87748FFBF8FD40FB1B" pageId="5" pageNumber="534">
|
||
<emphasis id="B949C1559C7CFF87748FFBF8FEB4FBEB" bold="true" box="[151,277,1057,1082]" pageId="5" pageNumber="534">FIGURE 5.</emphasis>
|
||
Dorsal view of carapace. A,
|
||
<emphasis id="B949C1559C7CFF877652FBFBFC8FFBE8" box="[586,814,1058,1081]" italics="true" pageId="5" pageNumber="534">Geosesarma spectrum</emphasis>
|
||
<emphasis id="B949C1559C7CFF87772FFBFBFCD2FBEB" bold="true" box="[823,883,1058,1082]" pageId="5" pageNumber="534">n. sp.</emphasis>
|
||
, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B,
|
||
<emphasis id="B949C1559C7CFF877512FB9FFE29FB8C" box="[266,392,1094,1117]" italics="true" pageId="5" pageNumber="534">G. spectrum</emphasis>
|
||
<emphasis id="B949C1559C7CFF877597FB9FFE6BFB8F" bold="true" box="[399,458,1094,1118]" pageId="5" pageNumber="534">n. sp.</emphasis>
|
||
, paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C,
|
||
<emphasis id="B949C1559C7CFF87704BFB9FFB71FB8C" box="[1107,1232,1094,1117]" italics="true" pageId="5" pageNumber="534">G. spectrum</emphasis>
|
||
<emphasis id="B949C1559C7CFF8770C0FB9FFAB2FB8F" bold="true" box="[1240,1299,1094,1118]" pageId="5" pageNumber="534">n. sp.</emphasis>
|
||
, paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D,
|
||
<emphasis id="B949C1559C7CFF8776B5FBB3FCB4FB50" box="[685,789,1129,1153]" italics="true" pageId="5" pageNumber="534">G. katibas</emphasis>
|
||
Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E,
|
||
<emphasis id="B949C1559C7CFF877503FB57FE25FB74" box="[283,388,1165,1189]" italics="true" pageId="5" pageNumber="534">G. katibas</emphasis>
|
||
Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F,
|
||
<emphasis id="B949C1559C7CFF87708DFB57FA93FB74" box="[1173,1330,1165,1189]" italics="true" pageId="5" pageNumber="534">G. gracillimum</emphasis>
|
||
(De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF424DCF9C7CFF87748FF854FF75F83F" httpUri="https://zenodo.org/record/3404941/files/figure.png" pageId="5" pageNumber="534" targetBox="[151,1436,1248,1914]" targetPageId="5">
|
||
<paragraph id="8B821D479C7CFF87748FF854FF75F83F" pageId="5" pageNumber="534">
|
||
<emphasis id="B949C1559C7CFF87748FF854FEB5F877" bold="true" box="[151,276,1933,1958]" pageId="5" pageNumber="534">FIGURE 6.</emphasis>
|
||
A, C,
|
||
<emphasis id="B949C1559C7CFF877541F856FD9DF877" box="[345,572,1935,1958]" italics="true" pageId="5" pageNumber="534">Geosesarma spectrum</emphasis>
|
||
<emphasis id="B949C1559C7CFF87765BF857FDDCF877" bold="true" box="[579,637,1934,1958]" pageId="5" pageNumber="534">n. sp.</emphasis>
|
||
, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, D,
|
||
<emphasis id="B949C1559C7CFF87712BF856FA3DF877" box="[1331,1436,1934,1958]" italics="true" pageId="5" pageNumber="534">G. katibas</emphasis>
|
||
Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak. A, B, frontal view of cephalothorax; C, D, inner view of left chela.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF424DCF9C7FFF84748FF9EAFCA6F90A" httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="6" pageNumber="535" startId="6.[151,250,1587,1612]" targetBox="[151,1436,181,1567]" targetPageId="6">
|
||
<paragraph id="8B821D479C7FFF84748FF9EAFCA6F90A" blockId="6.[151,1437,1587,1756]" pageId="6" pageNumber="535">
|
||
<emphasis id="B949C1559C7FFF84748FF9EAFEB2F99A" bold="true" box="[151,275,1587,1612]" pageId="6" pageNumber="535">FIGURE 7.</emphasis>
|
||
A, D, G, H,
|
||
<taxonomicName id="4C3D66C49C7FFF84758AF9EDFDD5F99A" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[402,628,1588,1611]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7FFF84758AF9EDFDD5F99A" box="[402,628,1588,1611]" italics="true" pageId="6" pageNumber="535">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7FFF847662F9EAFD12F99A" box="[634,691,1587,1611]" pageId="6" pageNumber="535" rank="species">
|
||
<emphasis id="B949C1559C7FFF847662F9EAFD12F99A" bold="true" box="[634,691,1587,1611]" pageId="6" pageNumber="535">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J,
|
||
<taxonomicName id="4C3D66C49C7FFF84719CF9EDFEE7F9A1" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7FFF84719CF9EDFF41F9BE" italics="true" pageId="6" pageNumber="535">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L,
|
||
<taxonomicName id="4C3D66C49C7FFF8477D7F981FAB3F9BE" authority="(De Man, 1892)" authorityName="De Man" authorityYear="1892" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[975,1298,1623,1647]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7FFF8477D7F981FBC8F9BE" box="[975,1129,1623,1647]" italics="true" pageId="6" pageNumber="535">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7FFF847083F981FAAAF9BE" author="Man, J. G. De" box="[1179,1291,1623,1647]" pageId="6" pageNumber="535" pagination="225 - 264" refId="ref5188" refString="Man, J. G. De (1892) Carcinological studies in the Leyden Museum. No. 6. Notes from the Leyden Museum, 14, 225 - 264., pls. 7 - 10." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C7FFF8474DFF8D3FF44F865" blockId="6.[151,1437,1802,2044]" pageId="6" pageNumber="535">
|
||
Among the other known Bornean species of
|
||
<taxonomicName id="4C3D66C49C7FFF8476A1F8D3FCE4F8F5" authorityName="De Man" authorityYear="1892" box="[697,837,1802,1828]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949C1559C7FFF8476A1F8D3FCE4F8F5" box="[697,837,1802,1828]" italics="true" pageId="6" pageNumber="535">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
, the general structure of the G1, absence of flagellum on third maxilliped, and presence of a row of distinct tubercles on the dorsal margin of the cheliped fingers, ally
|
||
<taxonomicName id="4C3D66C49C7FFF84748FF88BFE30F8BD" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[151,401,1874,1900]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7FFF84748FF88BFE30F8BD" box="[151,401,1874,1900]" italics="true" pageId="6" pageNumber="535">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7FFF847580F88BFE76F8BD" box="[408,471,1874,1900]" pageId="6" pageNumber="535" rank="species">
|
||
<emphasis id="B949C1559C7FFF847580F88BFE76F8BD" bold="true" box="[408,471,1874,1900]" pageId="6" pageNumber="535">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
with
|
||
<taxonomicName id="4C3D66C49C7FFF847600F88BFCF6F8BD" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" box="[536,855,1874,1900]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B949C1559C7FFF847600F88BFDC5F8BD" box="[536,612,1874,1900]" italics="true" pageId="6" pageNumber="535">G. bau</emphasis>
|
||
<bibRefCitation id="EFAC60B69C7FFF847674F88BFCF6F8BD" author="Ng, P. K. L. & Grinang, J." box="[620,855,1874,1900]" pageId="6" pageNumber="535" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C7FFF84777AF88BFB68F8BD" authority="(De Man, 1902)" authorityName="De Man" authorityYear="1902" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[866,1225,1874,1900]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7FFF84777AF88BFBAFF8BD" box="[866,1038,1874,1900]" italics="true" pageId="6" pageNumber="535">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7FFF84705EF88BFB60F8BD" author="Man, J. G. De" box="[1094,1217,1874,1900]" pageId="6" pageNumber="535" pagination="467 - 929" refId="ref5276" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kukenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25, 467 - 929., pls. 19 - 27. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">Man, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D66C49C7FFF847118F88BFE9BF841" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="larsi">
|
||
<emphasis id="B949C1559C7FFF847118F88BFAF4F8BD" box="[1280,1365,1874,1900]" italics="true" pageId="6" pageNumber="535">G. larsi</emphasis>
|
||
<bibRefCitation id="EFAC60B69C7FFF847145F88BFE9BF841" author="Ng, P. K. L. & Grinang, J." pageId="6" pageNumber="535" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
(from
|
||
<collectingCountry id="F32A5DD79C7FFF847592F8AFFE4DF840" box="[394,492,1910,1937]" name="Malaysia" pageId="6" pageNumber="535">Sarawak</collectingCountry>
|
||
) as well as
|
||
<taxonomicName id="4C3D66C49C7FFF84766CF8AFFC26F841" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[628,903,1909,1936]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B949C1559C7FFF84766CF8AFFCB2F841" box="[628,787,1910,1936]" italics="true" pageId="6" pageNumber="535">G. ambawang</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D66C49C7FFF8477DCF8AFFB68F841" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[964,1225,1909,1936]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="pontianak">
|
||
<emphasis id="B949C1559C7FFF8477DCF8AFFBF9F841" box="[964,1112,1910,1936]" italics="true" pageId="6" pageNumber="535">G. pontianak</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
(from Kalimantan Barat).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C7FFF8B74DFF867FB37FE92" lastPageId="9" lastPageNumber="538" pageId="6" pageNumber="535" type="reference_group">
|
||
<paragraph id="8B821D479C7FFF8574DFF867FEC4FF2A" blockId="6.[151,1437,1802,2044]" lastBlockId="7.[151,1437,152,899]" lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535">
|
||
<taxonomicName id="4C3D66C49C7FFF8474DFF867FE47F809" ID-CoL="3FS99" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[199,486,1982,2008]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="535" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7FFF8474DFF867FE47F809" box="[199,486,1982,2008]" italics="true" pageId="6" pageNumber="535">Geosesarma gracillimum</emphasis>
|
||
</taxonomicName>
|
||
was originally described from Baram,
|
||
<collectingCountry id="F32A5DD79C7FFF8477B3F867FBAFF809" box="[939,1038,1982,2008]" name="Malaysia" pageId="6" pageNumber="535">Sarawak</collectingCountry>
|
||
, and has been reported from other parts of northern
|
||
<collectingCountry id="F32A5DD79C7FFF847540F83BFE19F82D" box="[344,440,2018,2044]" name="Malaysia" pageId="6" pageNumber="535">Sarawak</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAC60B69C7FFF8475DFF83BFDCAF82D" author="Holthuis, L. B." box="[455,619,2018,2044]" pageId="6" pageNumber="535" pagination="3 - 47" refId="ref5094" refString="Holthuis, L. B. (1979) Cavernicolous and terrestrial decapod crustacea from Northern Sarawak, Borneo. Zoologische Verhandelingen, 171, 3 - 47, figs. 1 - 9, pls. 1 - 8." type="journal article" year="1979">Holthuis 1979</bibRefCitation>
|
||
; Ng 2015). The species has previously been recorded from
|
||
<collectingCountry id="F32A5DD79C7FFF84711BF83BFAEEF82D" box="[1283,1359,2018,2044]" name="Brunei" pageId="6" pageNumber="535">Brunei</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFAC60B69C7FFF857146F83BFF73FF62" author="Choy, S. C." lastPageId="7" lastPageNumber="536" pageId="6" pageNumber="535" pagination="117 - 153" refId="ref4838" refString="Choy, S. C. (1991) The crustacean fauna of negara Brunei Darussalam. Brunei Museum Journal, 7 (3), 117 - 153." type="journal article" year="1991">Choy 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC60B69C7EFF8574C5FF40FE60FF62" author="Ng, P. K. L. & Grinang, J." box="[221,449,153,179]" pageId="7" pageNumber="536" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
; Ng 2015). However, a more detailed examination of the Bruneian
|
||
<taxonomicName id="4C3D66C49C7EFF8570B3FF40FAF7FF62" ID-CoL="3FS99" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[1195,1366,153,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7EFF8570B3FF40FAF7FF62" box="[1195,1366,153,179]" italics="true" pageId="7" pageNumber="536">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
material shows that they all belong instead to
|
||
<taxonomicName id="4C3D66C49C7EFF857674FF64FD57FF06" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[620,758,189,215]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7EFF857674FF64FD57FF06" box="[620,758,189,215]" italics="true" pageId="7" pageNumber="536">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7EFF8576E6FF64FC9FFF06" box="[766,830,189,215]" pageId="7" pageNumber="536" rank="species">
|
||
<emphasis id="B949C1559C7EFF8576E6FF64FC9FFF06" bold="true" box="[766,830,189,215]" pageId="7" pageNumber="536">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, thus restricting the distribution of
|
||
<taxonomicName id="4C3D66C49C7EFF8570CAFF64FADFFF06" ID-CoL="3FS99" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[1234,1406,189,215]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7EFF8570CAFF64FADFFF06" box="[1234,1406,189,215]" italics="true" pageId="7" pageNumber="536">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
to northern
|
||
<collectingCountry id="F32A5DD79C7EFF8574E5FF38FEC0FF2A" box="[253,353,225,251]" name="Malaysia" pageId="7" pageNumber="536">Sarawak</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B821D479C7EFF8574DFFEDCFF47FDCA" blockId="7.[151,1437,152,899]" pageId="7" pageNumber="536">
|
||
<taxonomicName id="4C3D66C49C7EFF8574DFFEDCFE62FECE" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[199,451,261,287]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7EFF8574DFFEDCFE62FECE" box="[199,451,261,287]" italics="true" pageId="7" pageNumber="536">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7EFF8575D3FEDCFDADFECE" box="[459,524,261,287]" pageId="7" pageNumber="536" rank="species">
|
||
<emphasis id="B949C1559C7EFF8575D3FEDCFDADFECE" bold="true" box="[459,524,261,287]" pageId="7" pageNumber="536">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3D66C49C7EFF8576BDFEDCFCF3FECE" ID-CoL="3FS99" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[677,850,261,287]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7EFF8576BDFEDCFCF3FECE" box="[677,850,261,287]" italics="true" pageId="7" pageNumber="536">G. gracillimum</emphasis>
|
||
</taxonomicName>
|
||
in having a relatively more narrow external orbital tooth that is directed obliquely anteriorly, with the outer margin distinctly convex, extending clearly past the lateral carapace margins (
|
||
<figureCitation id="130601C29C7EFF857575FE94FE1AFEB6" box="[365,443,332,359]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="7" pageNumber="536">Figs. 5</figureCitation>
|
||
A–C) (vs. a broader external orbital tooth that is directed obliquely outwards, with the outer margin straight to mildly convex, extending only slightly past lateral carapace margins,
|
||
<figureCitation id="130601C29C7EFF8570BCFEA8FB59FE5A" box="[1188,1272,368,395]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="7" pageNumber="536">Fig. 5F</figureCitation>
|
||
); the distinctly more slender ambulatory meri (
|
||
<figureCitation id="130601C29C7EFF8575EDFE4CFDCDFE7E" box="[501,620,405,431]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Fig. 7G, H</figureCitation>
|
||
) (vs. proportionately broader ambulatory meri,
|
||
<figureCitation id="130601C29C7EFF857066FE4CFB52FE7E" box="[1150,1267,405,431]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Fig. 7K, L</figureCitation>
|
||
); and a slightly shorter and relatively stockier G1 with a distinct median cleft on the tip of distal part (
|
||
<figureCitation id="130601C29C7EFF857041FE60FB1CFE02" box="[1113,1213,441,467]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7EFF8570D1FE60FB77FE02" box="[1225,1238,441,467]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="7" pageNumber="536">9</figureCitation>
|
||
B–E) (vs. proportionately longer, slenderer G1 with a shallower and less distinct median cleft on the tip of the distal part,
|
||
<figureCitation id="130601C29C7EFF857123FE04FA39FE26" box="[1339,1432,477,503]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Figs. 7F</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7EFF85748FFDD8FF04FDCA" box="[151,165,513,539]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="7" pageNumber="536">9</figureCitation>
|
||
I–M).
|
||
</paragraph>
|
||
<paragraph id="8B821D479C7EFF8574DFFDFCFC67FD1E" blockId="7.[151,1437,152,899]" pageId="7" pageNumber="536">
|
||
<taxonomicName id="4C3D66C49C7EFF8574DFFDFCFE63FDEE" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[199,450,549,575]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7EFF8574DFFDFCFE63FDEE" box="[199,450,549,575]" italics="true" pageId="7" pageNumber="536">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7EFF8575D1FDFCFDA9FDEE" box="[457,520,549,575]" pageId="7" pageNumber="536" rank="species">
|
||
<emphasis id="B949C1559C7EFF8575D1FDFCFDA9FDEE" bold="true" box="[457,520,549,575]" pageId="7" pageNumber="536">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be distinguished from
|
||
<taxonomicName id="4C3D66C49C7EFF857722FDFCFC31FDEE" ID-CoL="97ec6452-b6f7-4951-b4f8-f18236a0b515" authorityName="Ng & Grinang" authorityYear="2018" box="[826,912,549,575]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="larsi">
|
||
<emphasis id="B949C1559C7EFF857722FDFCFC31FDEE" box="[826,912,549,575]" italics="true" pageId="7" pageNumber="536">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
by possessing distinctly more numerous granules on the carapace surface (
|
||
<figureCitation id="130601C29C7EFF8575FAFD90FDE7FDB2" box="[482,582,585,611]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="7" pageNumber="536">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7EFF85764AFD91FDFFFDB2" box="[594,606,584,611]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="7" pageNumber="536">5</figureCitation>
|
||
A–C) (vs. carapace surface with fewer granules,
|
||
<bibRefCitation id="EFAC60B69C7EFF85709DFD90FACAFDB2" author="Ng, P. K. L. & Grinang, J." box="[1157,1387,585,611]" pageId="7" pageNumber="536" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang 2018</bibRefCitation>
|
||
: fig. 2A-C); the relatively shorter ambulatory meri and propodi (
|
||
<figureCitation id="130601C29C7EFF857734FDB4FC2EFD56" box="[812,911,621,647]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="7" pageNumber="536">Figs. 1A</figureCitation>
|
||
, 7G, H) (vs. proportionately longer ambulatory meri and propodi,
|
||
<bibRefCitation id="EFAC60B69C7EFF85757DFD48FDE8FD7A" author="Ng, P. K. L. & Grinang, J." box="[357,585,657,683]" pageId="7" pageNumber="536" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang 2018</bibRefCitation>
|
||
: figs 2A, B, 3D–F); and a distinctly shorter and stouter G1 (
|
||
<figureCitation id="130601C29C7EFF8570FCFD48FAE9FD7A" box="[1252,1352,657,683]" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7EFF85714BFD48FAC1FD7A" box="[1363,1376,657,683]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="7" pageNumber="536">9</figureCitation>
|
||
B–E) (vs. relatively longer, more slender G1;
|
||
<bibRefCitation id="EFAC60B69C7EFF85764EFD6CFC9AFD1E" author="Ng, P. K. L. & Grinang, J." box="[598,827,693,719]" pageId="7" pageNumber="536" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang 2018</bibRefCitation>
|
||
: fig. 5B–G).
|
||
</paragraph>
|
||
<paragraph id="8B821D479C7EFF8574DFFD00FB01FC52" blockId="7.[151,1437,152,899]" pageId="7" pageNumber="536">
|
||
<taxonomicName id="4C3D66C49C7EFF8574DFFD00FE61FD22" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[199,448,729,755]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7EFF8574DFFD00FE61FD22" box="[199,448,729,755]" italics="true" pageId="7" pageNumber="536">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7EFF8575DDFD00FDA3FD22" box="[453,514,729,755]" pageId="7" pageNumber="536" rank="species">
|
||
<emphasis id="B949C1559C7EFF8575DDFD00FDA3FD22" bold="true" box="[453,514,729,755]" pageId="7" pageNumber="536">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C3D66C49C7EFF85768AFD00FD7CFD22" ID-CoL="3FS8V" authorityName="Ng & Grinang" authorityYear="2004" box="[658,733,729,755]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B949C1559C7EFF85768AFD00FD7CFD22" box="[658,733,729,755]" italics="true" pageId="7" pageNumber="536">G. bau</emphasis>
|
||
</taxonomicName>
|
||
in having the external orbital tooth longer and directed obliquely anteriorly (
|
||
<figureCitation id="130601C29C7EFF857537FD24FEDBFCC6" box="[303,378,764,791]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="7" pageNumber="536">Figs. 5</figureCitation>
|
||
A–C) (vs. external orbital tooth proportionately shorter and directed obliquely outwards,
|
||
<bibRefCitation id="EFAC60B69C7EFF857146FD24FE94FCEA" author="Ng, P. K. L. & Grinang, J." pageId="7" pageNumber="536" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 9A, B); relatively shorter ambulatory legs (
|
||
<figureCitation id="130601C29C7EFF857755FCF8FC11FCEA" box="[845,944,801,827]" captionStart="FIGURE 1" captionStartId="1.[151,250,1901,1926]" captionTargetId="figure@1.[187,1400,808,1881]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." httpUri="https://zenodo.org/record/3404937/files/figure.png" pageId="7" pageNumber="536">Figs. 1A</figureCitation>
|
||
, 7G, H) (vs. proportionately longer ambulatory legs,
|
||
<bibRefCitation id="EFAC60B69C7EFF85751DFC9CFE46FC8E" author="Ng, P. K. L. & Grinang, J." box="[261,487,837,863]" pageId="7" pageNumber="536" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: figs. 8A, B); and a relatively stouter G1 with the distal part more strongly bent (
|
||
<figureCitation id="130601C29C7EFF85717CFC9CFF1AFC52" captionStart="FIGURE 7" captionStartId="6.[151,250,1587,1612]" captionTargetBox="[151,1436,181,1567]" captionTargetId="figure@6.[151,1436,181,1567]" captionTargetPageId="6" captionText="FIGURE 7. A, D, G, H, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, E, I, J, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; C, F, K, L, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A–C, B, outer view of dactylus of left chela; D–F, ventral view of left G1; G, I, left third ambulatory leg; H, J, left fourth ambulatory leg; K, right third ambulatory leg (transposed for comparative purposes); L, right fourth ambulatory leg (transposed for comparative purposes)." httpUri="https://zenodo.org/record/3243084/files/figure.png" pageId="7" pageNumber="536">Figs. 7D</figureCitation>
|
||
,
|
||
<figureCitation id="130601C29C7EFF8574DFFCB0FF75FC52" box="[199,212,873,899]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="7" pageNumber="536">9</figureCitation>
|
||
B–E) (vs. more slender G1 with distal part less bent,
|
||
<bibRefCitation id="EFAC60B69C7EFF857730FCB0FBADFC52" author="Ng, P. K. L. & Grinang, J." box="[808,1036,873,899]" pageId="7" pageNumber="536" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: figs. 9D–F).
|
||
</paragraph>
|
||
<caption id="DF424DCF9C7EFF85748FF891FE93F820" httpUri="https://zenodo.org/record/3243086/files/figure.png" pageId="7" pageNumber="536" startId="7.[151,250,1864,1889]" targetBox="[177,1411,921,1845]" targetPageId="7">
|
||
<paragraph id="8B821D479C7EFF85748FF891FE93F820" blockId="7.[151,1436,1864,2033]" pageId="7" pageNumber="536">
|
||
<emphasis id="B949C1559C7EFF85748FF891FEB2F8B0" bold="true" box="[151,275,1864,1889]" pageId="7" pageNumber="536">FIGURE 8.</emphasis>
|
||
A, B,
|
||
<taxonomicName id="4C3D66C49C7EFF85754EF890FD99F8B1" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[342,568,1865,1888]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C7EFF85754EF890FD99F8B1" box="[342,568,1865,1888]" italics="true" pageId="7" pageNumber="536">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C7EFF857627F890FDD9F8B0" box="[575,632,1865,1889]" pageId="7" pageNumber="536" rank="species">
|
||
<emphasis id="B949C1559C7EFF857627F890FDD9F8B0" bold="true" box="[575,632,1865,1889]" pageId="7" pageNumber="536">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; C,
|
||
<taxonomicName id="4C3D66C49C7EFF857110F890FF6DF854" ID-CoL="8YX9B" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7EFF857110F890FAD1F8B0" box="[1288,1392,1865,1889]" italics="true" pageId="7" pageNumber="536">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; D,
|
||
<taxonomicName id="4C3D66C49C7EFF857761F8B4FBE8F854" ID-CoL="8YX9B" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" box="[889,1097,1900,1925]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7EFF857761F8B4FC43F854" box="[889,994,1901,1925]" italics="true" pageId="7" pageNumber="536">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; E,
|
||
<taxonomicName id="4C3D66C49C7EFF8575CFF848FD05F878" ID-CoL="8YX9B" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" box="[471,676,1936,1961]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C7EFF8575CFF848FD9FF878" box="[471,574,1937,1961]" italics="true" pageId="7" pageNumber="536">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; F,
|
||
<taxonomicName id="4C3D66C49C7EFF8570CEF848FEAEF81C" ID-CoL="3FS99" authority="(De Man, 1892)" authorityName="De Man" authorityYear="1892" baseAuthorityName="De Man" baseAuthorityYear="1892" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="536" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C7EFF8570CEF848FAD0F878" box="[1238,1393,1937,1961]" italics="true" pageId="7" pageNumber="536">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C7EFF85748FF86CFEA9F81C" author="Man, J. G. De" box="[151,264,1973,1997]" pageId="7" pageNumber="536" pagination="225 - 264" refId="ref5188" refString="Man, J. G. De (1892) Carcinological studies in the Leyden Museum. No. 6. Notes from the Leyden Museum, 14, 225 - 264., pls. 7 - 10." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, male thoracic sternum and pleon; B–E, male pleonal somites 3–6 and telson.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF424DCF9C71FF8A748FF8CAFF67F80E" httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="8" pageNumber="537" startId="8.[151,250,1811,1836]" targetBox="[247,1340,198,1775]" targetPageId="8">
|
||
<paragraph id="8B821D479C71FF8A748FF8CAFF67F80E" blockId="8.[151,1437,1811,2015]" pageId="8" pageNumber="537">
|
||
<emphasis id="B949C1559C71FF8A748FF8CAFEB5F8FD" bold="true" box="[151,276,1811,1836]" pageId="8" pageNumber="537">FIGURE 9.</emphasis>
|
||
A–F,
|
||
<taxonomicName id="4C3D66C49C71FF8A754BF8CDFD97F8FA" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[339,566,1812,1835]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C71FF8A754BF8CDFD97F8FA" box="[339,566,1812,1835]" italics="true" pageId="8" pageNumber="537">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B949C1559C71FF8A7626F8CAFDD9F8FA" bold="true" box="[574,632,1811,1835]" pageId="8" pageNumber="537">
|
||
<taxonomicNameLabel id="A27A7C2E9C71FF8A7626F8CAFDD9F8FA" box="[574,632,1811,1835]" pageId="8" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H,
|
||
<taxonomicName id="4C3D66C49C71FF8A712BF8CDFF58F881" ID-CoL="8YX9B" authority="Ng, 1995" authorityName=", Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C71FF8A712BF8CDFA3DF8FA" box="[1331,1436,1811,1835]" italics="true" pageId="8" pageNumber="537">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M,
|
||
<taxonomicName id="4C3D66C49C71FF8A7742F8E1FB05F89E" ID-CoL="3FS99" authority="(De Man, 1892)" authorityName="De Man" authorityYear="1892" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[858,1188,1847,1871]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="537" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C71FF8A7742F8E1FC57F89E" box="[858,1014,1847,1871]" italics="true" pageId="8" pageNumber="537">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C71FF8A7032F8E1FB3CF89E" author="Man, J. G. De" box="[1066,1181,1847,1871]" pageId="8" pageNumber="537" pagination="225 - 264" refId="ref5188" refString="Man, J. G. De (1892) Carcinological studies in the Leyden Museum. No. 6. Notes from the Leyden Museum, 14, 225 - 264., pls. 7 - 10." type="journal article" year="1892">Man, 1892</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C70FF8B74DFFF40FB37FE92" blockId="9.[151,1437,152,575]" pageId="9" pageNumber="538">
|
||
<taxonomicName id="4C3D66C49C70FF8B74DFFF40FE62FF62" ID-CoL="84FTR" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[199,451,153,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C70FF8B74DFFF40FE62FF62" box="[199,451,153,179]" italics="true" pageId="9" pageNumber="538">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C70FF8B75D4FF40FDADFF62" box="[460,524,153,179]" pageId="9" pageNumber="538" rank="species">
|
||
<emphasis id="B949C1559C70FF8B75D4FF40FDADFF62" bold="true" box="[460,524,153,179]" pageId="9" pageNumber="538">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
can be separated from
|
||
<taxonomicName id="4C3D66C49C70FF8B7703FF40FB90FF62" ID-CoL="3FS8P" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[795,1073,152,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B949C1559C70FF8B7703FF40FC1BFF62" box="[795,954,153,179]" italics="true" pageId="9" pageNumber="538">G. ambawang</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D66C49C70FF8B7077FF40FAD8FF62" ID-CoL="6KBJ5" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[1135,1401,152,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="pontianak">
|
||
<emphasis id="B949C1559C70FF8B7077FF40FAA2FF62" box="[1135,1283,153,179]" italics="true" pageId="9" pageNumber="538">G. pontianak</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
, in having the anterior carapace regions more granular (
|
||
<figureCitation id="130601C29C70FF8B76EBFF64FCE1FF06" box="[755,832,188,215]" captionStart="FIGURE 5" captionStartId="5.[151,250,1057,1082]" captionTargetBox="[151,1436,181,1037]" captionTargetId="figure@5.[151,1436,181,1037]" captionText="FIGURE 5. Dorsal view of carapace. A, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; B, G. spectrum n. sp., paratype male (7.9 × 7.6 mm), (ZRC 2018.0741), Brunei; C, G. spectrum n. sp., paratype fe- male (10.2 × 9.8 mm) (ZRC 2018.0741), Brunei; D, G. katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Sarawak; E, G. katibas Ng, 1995, paratype male (13.2 × 13.1 mm) (ZRC 1995.0274), Sarawak; F, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak." httpUri="https://zenodo.org/record/3404939/files/figure.png" pageId="9" pageNumber="538">Figs. 5</figureCitation>
|
||
A–C) (vs. anterior carapace region less granular, Ng 2015: figs. 8A, B, 11A, B); the merus of the third maxilliped is shorter than the ischium (
|
||
<figureCitation id="130601C29C70FF8B7073FF38FB63FF2A" box="[1131,1218,225,251]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="9" pageNumber="538">Fig. 9A</figureCitation>
|
||
) (vs. merus of third maxilliped equal or subequal to ischium, Ng 2015: figs. 9A, 13A); and the absence of a distinct subdistal hump on the G1 (
|
||
<figureCitation id="130601C29C70FF8B74EAFEF0FE92FE92" box="[242,307,297,323]" captionStart="FIGURE 9" captionStartId="8.[151,250,1811,1836]" captionTargetBox="[247,1340,198,1775]" captionTargetId="figure@8.[228,1359,181,1791]" captionTargetPageId="8" captionText="FIGURE 9. A–F, Geosesarma spectrum n. sp., holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei; G, H, G. katibas Ng, 1995, male (12.2 × 11.9 mm) (ZRC 2016.0296), Sarawak; I–M, G. gracillimum (De Man, 1892), male (12.1 × 11.8 mm) (ZRC 2007.262), Sarawak. A, left third maxilliped; B, left G1 (dorsal view); C, left G1 (ventral view); D, G, K, distal part of left G1 (dorsal view); E, H, distal part of left G1 (ventral view); F, left G2 (dorsal view); I, J left G1 (ventral views at different angles); L, M, distal part of left G1 (ventral views at different angles). Scales: A–C, F, I, J = 1.0 mm; D, E, G, H, K–M = 0.5 mm." httpUri="https://zenodo.org/record/3243088/files/figure.png" pageId="9" pageNumber="538">Fig. 9</figureCitation>
|
||
B–E) (vs. subdistal hump present on G1, Ng 2015: figs. 9D–I, 13D–H, J–M).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3274ECC9C70FF8874DFFE94FDC7FBC2" lastPageId="10" lastPageNumber="539" pageId="9" pageNumber="538" type="description">
|
||
<paragraph id="8B821D479C70FF8B74DFFE94FE15FDEE" blockId="9.[151,1437,152,575]" pageId="9" pageNumber="538">
|
||
The live colour of adult males of
|
||
<taxonomicName id="4C3D66C49C70FF8B7652FE94FD74FEB6" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[586,725,333,359]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C70FF8B7652FE94FD74FEB6" box="[586,725,333,359]" italics="true" pageId="9" pageNumber="538">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C70FF8B76C6FE94FCBFFEB6" box="[734,798,333,359]" pageId="9" pageNumber="538" rank="species">
|
||
<emphasis id="B949C1559C70FF8B76C6FE94FCBFFEB6" bold="true" box="[734,798,333,359]" pageId="9" pageNumber="538">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is almost identical to that of
|
||
<taxonomicName id="4C3D66C49C70FF8B706AFE94FB46FEB6" authorityName=", Ng" authorityYear="1995" box="[1138,1255,333,359]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C70FF8B706AFE94FB46FEB6" box="[1138,1255,333,359]" italics="true" pageId="9" pageNumber="538">G. katibas</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="130601C29C70FF8B70E0FE94FAC4FEB6" box="[1272,1381,333,359]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="1.[151,250,1901,1926]" captionStartId-1="2.[151,250,1650,1675]" captionStartId-2="3.[151,250,1937,1962]" captionTargetBox-1="[187,1400,668,1630]" captionTargetBox-2="[188,1399,885,1917]" captionTargetId-0="figure@1.[187,1400,808,1881]" captionTargetId-1="figure@2.[187,1400,668,1630]" captionTargetId-2="figure@3.[188,1399,885,1917]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="FIGURE 1. Geosesarma spectrum n. sp., colour in life, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740), Brunei. A, overall habitus; B, thoracic sternum and pleon; C, outer view of right chela." captionText-1="FIGURE 2. Geosesarma spectrum n. sp., colour in life. A, B, holotype male (13.6 × 13.5 mm) (ZRC 2018.0740); C, paratype male (7.9 × 7.6 mm) (ZRC 2018.0741); D, paratype female (10.2 × 9.8 mm) (ZRC 2018.0741). All from Brunei." captionText-2="FIGURE 3. Geosesarma spectrum n. sp., colour in life. A, female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B–D, female (11.2 × 10.7 mm) (ZRC 2018.0741), Brunei.A, overall habitus; B, ovigerous female with late-stage eggs; C, female with newly hatched juvenile crabs on right side of carapace; D, female with juvenile crabs on right side of carapace." httpUri-0="https://zenodo.org/record/3404937/files/figure.png" httpUri-1="https://zenodo.org/record/3243078/files/figure.png" httpUri-2="https://zenodo.org/record/3243080/files/figure.png" pageId="9" pageNumber="538">Figs. 1–3</figureCitation>
|
||
, 4A, B), and is diagnostic for these two species. Compared to
|
||
<taxonomicName id="4C3D66C49C70FF8B7704FEA8FC07FE5A" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[796,934,369,395]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C70FF8B7704FEA8FC07FE5A" box="[796,934,369,395]" italics="true" pageId="9" pageNumber="538">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C70FF8B77B6FEA8FC4CFE5A" box="[942,1005,369,395]" pageId="9" pageNumber="538" rank="species">
|
||
<emphasis id="B949C1559C70FF8B77B6FEA8FC4CFE5A" bold="true" box="[942,1005,369,395]" pageId="9" pageNumber="538">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B77E4FEA8FBF3FE5A" authorityName="Ng & Grinang" authorityYear="2018" box="[1020,1106,369,395]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="larsi">
|
||
<emphasis id="B949C1559C70FF8B77E4FEA8FBF3FE5A" box="[1020,1106,369,395]" italics="true" pageId="9" pageNumber="538">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
is a deep purplish-red on the anterior half of the carapace, ambulatory meri and chelipeds (
|
||
<bibRefCitation id="EFAC60B69C70FF8B7752FE4CFB91FE7E" author="Ng, P. K. L. & Grinang, J." box="[842,1072,405,431]" pageId="9" pageNumber="538" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang 2018</bibRefCitation>
|
||
: fig. 1).
|
||
<taxonomicName id="4C3D66C49C70FF8B7096FE4CFA3DFE7E" authorityName="Ng" authorityYear="2015" box="[1166,1436,405,431]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B949C1559C70FF8B7096FE4CFA3DFE7E" box="[1166,1436,405,431]" italics="true" pageId="9" pageNumber="538">Geosesarma ambawang</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D66C49C70FF8B74DFFE60FEFBFE02" authorityName="Ng" authorityYear="2015" box="[199,346,441,467]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="pontianak">
|
||
<emphasis id="B949C1559C70FF8B74DFFE60FEFBFE02" box="[199,346,441,467]" italics="true" pageId="9" pageNumber="538">G. pontianak</emphasis>
|
||
</taxonomicName>
|
||
are a reddish-brown on the anterior half of the carapace and ambulatory legs, with the chelipeds a uniform bright red (Ng 2015: fig. 10).
|
||
<taxonomicName id="4C3D66C49C70FF8B767AFE04FCD6FE26" box="[610,887,477,503]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="gracilimum">
|
||
<emphasis id="B949C1559C70FF8B767AFE04FCD6FE26" box="[610,887,477,503]" italics="true" pageId="9" pageNumber="538">Geosesarma gracilimum</emphasis>
|
||
</taxonomicName>
|
||
is a uniform deep red to orange (
|
||
<figureCitation id="130601C29C70FF8B70EBFE04FACCFE26" box="[1267,1389,477,503]" captionStart="FIGURE 4" captionStartId="4.[151,250,946,971]" captionTargetBox="[187,1400,181,927]" captionTargetId="figure@4.[187,1400,181,927]" captionTargetPageId="4" captionText="FIGURE 4. A, Geosesarma katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B, G. katibas Ng, 1995, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C, G. gracillimum (De Man, 1892), male, Lambir Hills National Park, Sarawak (specimen not collected); D, G. gracillimum (De Man, 1892), female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected)." httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="9" pageNumber="538">Fig. 4C, D</figureCitation>
|
||
) including the chelae, whereas
|
||
<taxonomicName id="4C3D66C49C70FF8B75CFFDD8FD82FDCA" authorityName="Ng & Grinang" authorityYear="2004" box="[471,547,513,539]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B949C1559C70FF8B75CFFDD8FD82FDCA" box="[471,547,513,539]" italics="true" pageId="9" pageNumber="538">G. bau</emphasis>
|
||
</taxonomicName>
|
||
have an orange to red carapace, with the chelae and legs brown (Ng 2015: 54, J. Grinang, pers. comm.).
|
||
</paragraph>
|
||
<caption id="DF424DCF9C70FF8B748FF9E0FD8AF94B" httpUri="https://zenodo.org/record/3243090/files/figure.png" pageId="9" pageNumber="538" startId="9.[151,250,1593,1618]" targetBox="[151,1436,597,1574]" targetPageId="9">
|
||
<paragraph id="8B821D479C70FF8B748FF9E0FD8AF94B" blockId="9.[151,1437,1593,1690]" pageId="9" pageNumber="538">
|
||
<emphasis id="B949C1559C70FF8B748FF9E0FE80F983" bold="true" box="[151,289,1593,1618]" pageId="9" pageNumber="538">FIGURE 10.</emphasis>
|
||
A, C,
|
||
<taxonomicName id="4C3D66C49C70FF8B757FF9E3FDEAF983" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[359,587,1594,1618]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C70FF8B757FF9E3FDEAF983" box="[359,587,1594,1618]" italics="true" pageId="9" pageNumber="538">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C70FF8B764AF9E3FD2DF983" box="[594,652,1594,1618]" pageId="9" pageNumber="538" rank="species">
|
||
<emphasis id="B949C1559C70FF8B764AF9E3FD2DF983" bold="true" box="[594,652,1594,1618]" pageId="9" pageNumber="538">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype ovigerous female (15.1 × 14.3 mm) (ZRC 2018.0741), Brunei; B, D,
|
||
<taxonomicName id="4C3D66C49C70FF8B74AFF987FE26F9A7" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" box="[183,391,1629,1654]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C70FF8B74AFF987FEBEF9A7" box="[183,287,1630,1654]" italics="true" pageId="9" pageNumber="538">G. katibas</emphasis>
|
||
Ng, 1995
|
||
</taxonomicName>
|
||
, female (12.3 x 11.8 mm) (ZRC 2016.0296), Sarawak. A, B, female thoracic sternum and pleon; C, D, female sternopleonal cavity and vulvae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B821D479C70FF8874DFF911FB3BFE92" blockId="9.[151,1437,1736,2015]" lastBlockId="10.[151,1437,152,1043]" lastPageId="10" lastPageNumber="539" pageId="9" pageNumber="538">
|
||
Of the 13 species of
|
||
<taxonomicName id="4C3D66C49C70FF8B75AFF910FDE2F932" authorityName="De Man" authorityYear="1892" box="[439,579,1737,1763]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949C1559C70FF8B75AFF910FDE2F932" box="[439,579,1737,1763]" italics="true" pageId="9" pageNumber="538">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
now known from Borneo (including the present new one), only three have third maxilliped exopods which possess a long flagellum:
|
||
<taxonomicName id="4C3D66C49C70FF8B773BF934FB23F8D7" authority="(De Man, 1899)" authorityName="De Man" authorityYear="1899" baseAuthorityName="De Man" baseAuthorityYear="1899" box="[803,1154,1772,1799]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="amphinome">
|
||
<emphasis id="B949C1559C70FF8B773BF934FC69F8D7" box="[803,968,1772,1799]" italics="true" pageId="9" pageNumber="538">G. amphinome</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C70FF8B7018F934FBDBF8D7" author="Man, J. G. De" box="[1024,1146,1772,1799]" pageId="9" pageNumber="538" pagination="53 - 144" refId="ref5228" refString="Man, J. G. De (1899) Zoological Results of the Dutch Scientific Expedition to Central Borneo. The Crustacea. Part II, Brachyura. Notes from the Leyden Museum, 21, 53 - 144., pls. 5 - 12." type="journal article" year="1899">Man, 1899</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(Kapuas, West Kalimantan),
|
||
<taxonomicName id="4C3D66C49C70FF8B74D7F8C8FE7EF8FB" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[207,479,1807,1835]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="pylaemenes">
|
||
<emphasis id="B949C1559C70FF8B74D7F8C8FED2F8FB" box="[207,371,1808,1835]" italics="true" pageId="9" pageNumber="538">G. pylaemenes</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
(Anjungan, West Kalimantan), and
|
||
<taxonomicName id="4C3D66C49C70FF8B7774F8C8FB66F8FA" authority="(Serene, 1968)" authorityName="Serene" authorityYear="1968" baseAuthorityName="Serene" baseAuthorityYear="1968" box="[876,1223,1808,1835]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="B949C1559C70FF8B7774F8C8FBBFF8FB" box="[876,1054,1808,1835]" italics="true" pageId="9" pageNumber="538">G. sarawakense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFAC60B69C70FF8B7033F8C9FB1EF8FA" author="Serene, R." box="[1067,1215,1808,1835]" pageId="9" pageNumber="538" pagination="1084 - 1095" refId="ref6407" refString="Serene, R. (1968) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1 - 2." type="journal article" year="1968">Serène, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(western
|
||
<collectingCountry id="F32A5DD79C70FF8B712BF8C9FA34F8FA" box="[1331,1429,1808,1835]" name="Malaysia" pageId="9" pageNumber="538">Sarawak</collectingCountry>
|
||
) (see also Ng 2015: 51). The other 10 species which are distributed all over Borneo:
|
||
<taxonomicName id="4C3D66C49C70FF8B7047F8ECFAD5F89F" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[1119,1396,1843,1871]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B949C1559C70FF8B7047F8ECFB5FF89F" box="[1119,1278,1844,1871]" italics="true" pageId="9" pageNumber="538">G. ambawang</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B7199F8ECFE2FF8A3" authority="Ng, 1995 b" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="aurantium">
|
||
<emphasis id="B949C1559C70FF8B7199F8ECFEADF8A3" italics="true" pageId="9" pageNumber="538">G. aurantium</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B750CF880FE2FF8A3" author="Ng, P. K. L." box="[276,398,1879,1907]" pageId="9" pageNumber="538" pagination="65 - 70" refId="ref5612" refString="Ng, P. K. L. (1995 b) Geosesarma aurantium, a new sesarmine land crab (Crustacea: Decapoda: Brachyura: Grapsidae) from Sabah. Malayan Nature Journal, 49, 65 - 70." type="journal article" year="1995">Ng, 1995b</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B7582F880FD7DF8A3" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" box="[410,732,1880,1907]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B949C1559C70FF8B7582F880FE46F8A3" box="[410,487,1880,1907]" italics="true" pageId="9" pageNumber="538">G. bau</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B75F7F880FD7DF8A3" author="Ng, P. K. L. & Grinang, J." box="[495,732,1880,1907]" pageId="9" pageNumber="538" pagination="299 - 325" refId="ref5833" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Sarawak Bau Limestone Biodiversity. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B76F0F880FC5CF8A3" authority="Ng, 2002" authorityName="Ng" authorityYear="2002" box="[744,1021,1880,1907]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="danumense">
|
||
<emphasis id="B949C1559C70FF8B76F0F880FC28F8A3" box="[744,905,1880,1907]" italics="true" pageId="9" pageNumber="538">G. danumense</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B7789F880FC5CF8A3" author="Ng, P. K. L." box="[913,1021,1880,1907]" pageId="9" pageNumber="538" pagination="303 - 310" refId="ref5654" refString="Ng, P. K. L. (2002) On a new species of terrestrial crab of the genus Geosesarma (Crustacea: Brachyura: Sesarmidae) from Sabah, Malaysia. Malayan Nature Journal, 56 (3), 303 - 310." type="journal article" year="2002">Ng, 2002</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B7011F880FAD5F8A3" authority="(De Man, 1902)" authorityName="De Man" authorityYear="1902" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[1033,1396,1880,1907]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B949C1559C70FF8B7011F880FB14F8A3" box="[1033,1205,1880,1907]" italics="true" pageId="9" pageNumber="538">G. gracillimum</emphasis>
|
||
(De
|
||
<bibRefCitation id="EFAC60B69C70FF8B70F7F880FACCF8A3" author="Man, J. G. De" box="[1263,1389,1880,1907]" pageId="9" pageNumber="538" pagination="467 - 929" refId="ref5276" refString="Man, J. G. De (1902) Die von Herrn Professor Kukenthal in Indischen Archipel gesammelten Dekapoden und Stomatopoden. In: W. Kukenthal, Ergebnisse einer zoologischen Forschungsreise in den Molukken und Borneo. Abhandlungen der Senckenbergischen naturforschenden Gesellschaft, 25, 467 - 929., pls. 19 - 27. https: // doi. org / 10.5962 / bhl. title. 10409" type="journal article" year="1902">Man, 1902</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B7199F880FEC9F847" authority="Ng, 1995 a" authorityName=", Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="katibas">
|
||
<emphasis id="B949C1559C70FF8B7199F880FF49F847" italics="true" pageId="9" pageNumber="538">G. katibas</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B74E8F8A4FEC9F847" author="Ng, P. K. L." box="[240,360,1915,1943]" pageId="9" pageNumber="538" pagination="57 - 72" refId="ref5556" refString="Ng, P. K. L. (1995 a) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995a</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B756CF8A4FD1FF847" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[372,702,1916,1943]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="larsi">
|
||
<emphasis id="B949C1559C70FF8B756CF8A4FE68F847" box="[372,457,1916,1943]" italics="true" pageId="9" pageNumber="538">G. larsi</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B75C9F8A4FD1FF847" author="Ng, P. K. L. & Grinang, J." box="[465,702,1916,1943]" pageId="9" pageNumber="538" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B76D2F8A4FC71F847" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[714,976,1915,1943]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="pontianak">
|
||
<emphasis id="B949C1559C70FF8B76D2F8A4FCFCF847" box="[714,861,1916,1943]" italics="true" pageId="9" pageNumber="538">G. pontianak</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C3D66C49C70FF8B77C4F8A4FB76F847" authority="Ng, 1992" authorityName="Ng" authorityYear="1992" box="[988,1239,1916,1943]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="sabanum">
|
||
<emphasis id="B949C1559C70FF8B77C4F8A4FBC2F847" box="[988,1123,1916,1943]" italics="true" pageId="9" pageNumber="538">G. sabanum</emphasis>
|
||
<bibRefCitation id="EFAC60B69C70FF8B7073F8A4FB76F847" author="Ng, P. K. L." box="[1131,1239,1916,1943]" pageId="9" pageNumber="538" pagination="210 - 213" refId="ref5497" refString="Ng, P. K. L. (1992) Geosesarma sabanus, a new sesarmine crab (Decapoda, Brachyura, Grapsidae) from the forests of Sabah, East Malaysia, Borneo. Crustaceana, 63 (2), 210 - 213. https: // doi. org / 10.1163 / 156854092 X 00596" type="journal article" year="1992">Ng, 1992</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C3D66C49C70FF8B710BF8A4FA3CF847" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[1299,1437,1917,1943]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="538" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C70FF8B710BF8A4FA3CF847" box="[1299,1437,1917,1943]" italics="true" pageId="9" pageNumber="538">G. spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C70FF8B748FF878FF77F86A" box="[151,214,1953,1979]" pageId="9" pageNumber="538" rank="species">
|
||
<emphasis id="B949C1559C70FF8B748FF878FF77F86A" bold="true" box="[151,214,1953,1979]" pageId="9" pageNumber="538">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have an exopod that completely lacks any trace of a flagellum. The reasons for this are not clear but may be associated with their habits. Those species with a well-developed flagellum on the exopod of the third maxillipeds may have more aquatic habits (it is certainly so for
|
||
<taxonomicName id="4C3D66C49C73FF8876C1FF40FCDFFF62" authorityName="Ng" authorityYear="2015" box="[729,894,153,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="pylaemenes">
|
||
<emphasis id="B949C1559C73FF8876C1FF40FCDFFF62" box="[729,894,153,179]" italics="true" pageId="10" pageNumber="539">G. pylaemenes</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C3D66C49C73FF8877ACFF40FBC9FF62" baseAuthorityName="Serene" baseAuthorityYear="1968" box="[948,1128,153,179]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="B949C1559C73FF8877ACFF40FBC9FF62" box="[948,1128,153,179]" italics="true" pageId="10" pageNumber="539">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
(see Ng 2015; unpublished data). From what is known (based mainly on available collection data), species which lack the flagellum do tend to be more terrestrial, climbing trees and can often be found far from streams (
|
||
<bibRefCitation id="EFAC60B69C73FF8877C2FF38FBEAFF2A" author="Ng, P. K. L." box="[986,1099,224,251]" pageId="10" pageNumber="539" pagination="57 - 72" refId="ref5556" refString="Ng, P. K. L. (1995 a) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995a</bibRefCitation>
|
||
, b, 2015;
|
||
<bibRefCitation id="EFAC60B69C73FF8870ADFF38FA39FF2A" author="Ebin, J. & Chung, A. Y. C." box="[1205,1432,225,251]" pageId="10" pageNumber="539" pagination="50 - 51" refId="ref5054" refString="Ebin, J. & Chung, A. Y. C. (2012) Revisiting the elegant Bornean endemic crab of Mt Silam, Sabah. Malayan Naturalist, 65 (4), 50 - 51." type="journal article" year="2012">Ebin & Chung 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC60B69C73FF88748FFEDCFEDCFECE" author="Ng, P. K. L. & Grinang, J." box="[151,381,261,287]" pageId="10" pageNumber="539" pagination="569 - 575" refId="ref5920" refString="Ng, P. K. L. & Grinang, J. (2018) A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak. Zootaxa, 4508 (4), 569 - 575. https: // doi. org / 10.11646 / zootaxa. 4508.4.5" type="journal article" year="2018">Ng & Grinang 2018</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC60B69C73FF887591FEDCFD97FECE" author="Ng, P. K. L. & Ng, P. Y. C." box="[393,566,261,287]" pageId="10" pageNumber="539" pagination="89 - 112" refId="ref6095" refString="Ng, P. K. L. & Ng, P. Y. C. (2018) The freshwater crabs of Danum Valley Conservation Area in Sabah, East Malaysia, with a description of a new species of Thelphusula Bott, 1969 (Crustacea, Brachyura, Gecarcinucidae, Potamidae, Sesarmidae). Zookeys, 760, 89 - 112. https: // doi. org / 10.3897 / zookeys. 760.24787" type="journal article" year="2018">Ng & Ng 2018</bibRefCitation>
|
||
; unpublished data). A detailed ecological study of all the species will need to be made to better understand these patterns and why the flagellum has been reduced or lost.
|
||
</paragraph>
|
||
<paragraph id="8B821D479C73FF8874DFFE95FC8FFD1E" blockId="10.[151,1437,152,1043]" pageId="10" pageNumber="539">
|
||
<emphasis id="B949C1559C73FF8874DFFE95FE26FEB6" bold="true" box="[199,391,332,359]" pageId="10" pageNumber="539">Ecological notes</emphasis>
|
||
. The recent specimens of
|
||
<taxonomicName id="4C3D66C49C73FF8876B6FE94FC09FEB6" authority="Ng & Ng, 2019" authorityName="Ng & Ng" authorityYear="2019" box="[686,936,333,359]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="spectrum" status="sp. nov.">
|
||
<emphasis id="B949C1559C73FF8876B6FE94FC09FEB6" box="[686,936,333,359]" italics="true" pageId="10" pageNumber="539">Geosesarma spectrum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27A7C2E9C73FF8877B7FE94FC4FFEB6" box="[943,1006,333,359]" pageId="10" pageNumber="539" rank="species">
|
||
<emphasis id="B949C1559C73FF8877B7FE94FC4FFEB6" bold="true" box="[943,1006,333,359]" pageId="10" pageNumber="539">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
were nocturnal, living in soil burrows on slopes, under well shaded canopy in primary rainforest within Ulu
|
||
<collectingRegion id="49F9D3A59C73FF8877ACFEA8FB94FE5A" box="[948,1077,369,395]" country="Brunei" name="Temburong" pageId="10" pageNumber="539">Temburong</collectingRegion>
|
||
National Park and several hundred metres from any stream. Adult individuals were observed at night (ca. 8 pm) at the edge of their burrows, and more individuals, including juveniles, were found foraging on leaf litter and low foliage later in the night. A single juvenile was collected in pitcher plant,
|
||
<taxonomicName id="4C3D66C49C73FF887654FE04FD60FE26" box="[588,705,477,503]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="539" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B949C1559C73FF887654FE04FD60FE26" box="[588,705,477,503]" italics="true" pageId="10" pageNumber="539">Nepenthes</emphasis>
|
||
</taxonomicName>
|
||
sp., suggesting juveniles may take refuge in phytotelmata instead of digging burrows. This is a frequently observed habit among many
|
||
<taxonomicName id="4C3D66C49C73FF8877ADFDD8FBE0FDCA" authorityName="De Man" authorityYear="1892" box="[949,1089,513,539]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949C1559C73FF8877ADFDD8FBE0FDCA" box="[949,1089,513,539]" italics="true" pageId="10" pageNumber="539">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
species (see
|
||
<bibRefCitation id="EFAC60B69C73FF8870C3FDD8FA36FDCA" author="Ng, P. K. L. & Lim, R. P." box="[1243,1431,513,539]" pageId="10" pageNumber="539" pagination="393 - 402" refId="ref6036" refString="Ng, P. K. L. & Lim, R. P. (1987) The taxonomy and biology of the nepenthiphilous sesarmine freshwater crab, Geosesarma malayanum Ng and Lim, 1986 (Crustacea, Decapoda, Brachyura, Grapsidae) from Peninsular Malaysia. Malayan Nature Journal, 41, 393 - 402." type="journal article" year="1987">Ng & Lim 1987</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC60B69C73FF88748FFDFCFF5CFDEE" author="Ng, P. K. L." box="[151,253,549,575]" pageId="10" pageNumber="539" refId="ref5449" refString="Ng, P. K. L. (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, viii + 156 pp., 63 figs., 4 colour pls." type="book" year="1988">Ng 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFAC60B69C73FF887511FDFCFE61FDEE" author="Tan, H. H. & Ng, P. K. L." box="[265,448,549,575]" pageId="10" pageNumber="539" pagination="201 - 205" refId="ref6492" refString="Tan, H. H. & Ng, P. K. L. (2008) First record in Singapore of a nepenthiphilous crab, Geosesarma perracae (Crustacea: Decapoda: Sesarmidae). Nature in Singapore, 1, 201 - 205." type="journal article" year="2008">Tan & Ng 2008</bibRefCitation>
|
||
). This has precedence, as one of the specimens (ZRC 2018.0746) was collected from a pitcher plant: “
|
||
<taxonomicName id="4C3D66C49C73FF887558FD90FE6DFDB2" authorityName="De Man" authorityYear="1892" box="[320,460,585,611]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B949C1559C73FF887558FD90FE6DFDB2" box="[320,460,585,611]" italics="true" pageId="10" pageNumber="539">Geosesarma</emphasis>
|
||
</taxonomicName>
|
||
is a wholly freshwater genus and some specimens, possibly of
|
||
<taxonomicName id="4C3D66C49C73FF887081FD90FA90FDB2" authorityName="- Choy" authorityYear="1991" box="[1177,1329,585,611]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="gracillima">
|
||
<emphasis id="B949C1559C73FF887081FD90FA90FDB2" box="[1177,1329,585,611]" italics="true" pageId="10" pageNumber="539">G. gracillima</emphasis>
|
||
</taxonomicName>
|
||
, were recently caught from the peat swamp at Seria (near the old railway). A few specimens were also collected from inside the pitcher of the pitcher plant
|
||
<taxonomicName id="4C3D66C49C73FF8875E0FD48FD57FD7A" box="[504,758,657,683]" class="Magnoliopsida" family="Nepenthaceae" genus="Nepenthes" kingdom="Plantae" order="Caryophyllales" pageId="10" pageNumber="539" phylum="Tracheophyta" rank="species" species="bicalcurata">
|
||
<emphasis id="B949C1559C73FF8875E0FD48FD57FD7A" box="[504,758,657,683]" italics="true" pageId="10" pageNumber="539">Nepenthes bicalcurata</emphasis>
|
||
</taxonomicName>
|
||
[sic]. In this respect,
|
||
<taxonomicName id="4C3D66C49C73FF8877F7FD48FB27FD7A" authorityName="- Choy" authorityYear="1991" box="[1007,1158,657,683]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="gracillima">
|
||
<emphasis id="B949C1559C73FF8877F7FD48FB27FD7A" box="[1007,1158,657,683]" italics="true" pageId="10" pageNumber="539">G. gracillima</emphasis>
|
||
</taxonomicName>
|
||
from Seria is somewhat nepenthiphilous” (
|
||
<bibRefCitation id="EFAC60B69C73FF88757DFD6CFDBDFD1E" author="Choy, S. C." box="[357,540,693,719]" pageId="10" pageNumber="539" pagination="117 - 153" refId="ref4838" refString="Choy, S. C. (1991) The crustacean fauna of negara Brunei Darussalam. Brunei Museum Journal, 7 (3), 117 - 153." type="journal article" year="1991">Choy 1991: 144</bibRefCitation>
|
||
) (see also
|
||
<bibRefCitation id="EFAC60B69C73FF88768CFD6CFC80FD1E" author="Clarke, C." box="[660,801,693,719]" pageId="10" pageNumber="539" refId="ref4868" refString="Clarke, C. (1997) Nepenthes of Borneo. Natural History Publications (Borneo), xii + 207 pp." type="book" year="1997">Clarke 1997</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B821D479C73FF8874DFFD00FDC7FBC2" blockId="10.[151,1437,152,1043]" pageId="10" pageNumber="539">
|
||
One ovigerous adult female was observed wandering on leaf litter during the day. This specimen had its left eye damaged, and was probably attacked by a predator, and evicted from its burrow. The
|
||
<specimenCount id="9D3BD6CE9C73FF887026FD25FB32FCC6" box="[1086,1171,764,791]" pageId="10" pageNumber="539" type="egg">15 eggs</specimenCount>
|
||
were eyed and believed to be close to hatching (
|
||
<figureCitation id="130601C29C73FF8875BEFCF8FE5EFCEA" box="[422,511,801,827]" captionStart="FIGURE 4" captionStartId="4.[151,250,946,971]" captionTargetBox="[187,1400,181,927]" captionTargetId="figure@4.[187,1400,181,927]" captionTargetPageId="4" captionText="FIGURE 4. A, Geosesarma katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B, G. katibas Ng, 1995, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C, G. gracillimum (De Man, 1892), male, Lambir Hills National Park, Sarawak (specimen not collected); D, G. gracillimum (De Man, 1892), female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected)." httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="10" pageNumber="539">Fig. 4B</figureCitation>
|
||
). The number of eggs were probably more originally but some were probably lost during capture. The brooding female was maintained in captivity for 10 days, during which the eggs hatched, and several juvenile crabs were found clinging to the carapace of the mother (
|
||
<figureCitation id="130601C29C73FF8877D7FCB0FBE5FC52" box="[975,1092,873,899]" captionStart="FIGURE 4" captionStartId="4.[151,250,946,971]" captionTargetBox="[187,1400,181,927]" captionTargetId="figure@4.[187,1400,181,927]" captionTargetPageId="4" captionText="FIGURE 4. A, Geosesarma katibas Ng, 1995, holotype male (13.5 × 13.5 mm) (ZRC 1996.01834), Katibas, Sarawak; B, G. katibas Ng, 1995, male, Sarikei, Sarawak (photograph J. Grinang, specimen not collected); C, G. gracillimum (De Man, 1892), male, Lambir Hills National Park, Sarawak (specimen not collected); D, G. gracillimum (De Man, 1892), female, Niah National Park, Sarawak (photograph J. Grinang, specimen not collected)." httpUri="https://zenodo.org/record/3243082/files/figure.png" pageId="10" pageNumber="539">Fig 4C, D</figureCitation>
|
||
). The juvenile crabs continued to cling to the mother for three days, leaving after completing their first molt. The present observation of the young crabs clinging to the carapace of the mother is notable, and known in several species, such as
|
||
<taxonomicName id="4C3D66C49C73FF8870DDFC68FE87FC3E" authority="Ng & Tan, 1995" authorityName="Ng & Tan" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="539" phylum="Arthropoda" rank="species" species="notophorum">
|
||
<emphasis id="B949C1559C73FF8870DDFC68FAD3FC1A" box="[1221,1394,945,971]" italics="true" pageId="10" pageNumber="539">G. notophorum</emphasis>
|
||
<bibRefCitation id="EFAC60B69C73FF887162FC68FE87FC3E" author="Ng, P. K. L. & Tan, C. G. S." pageId="10" pageNumber="539" pagination="390 - 395" refId="ref6250" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. https: // doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Ng & Tan, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Pulau Lingga,
|
||
<collectingCountry id="F32A5DD79C73FF88760BFC0CFD21FC3E" box="[531,640,981,1007]" name="Indonesia" pageId="10" pageNumber="539">Indonesia</collectingCountry>
|
||
(cf.
|
||
<bibRefCitation id="EFAC60B69C73FF8876ACFC0CFCC7FC3E" author="Ng, P. K. L. & Tan, C. G. S." box="[692,870,980,1007]" pageId="10" pageNumber="539" pagination="390 - 395" refId="ref6250" refString="Ng, P. K. L. & Tan, C. G. S. (1995) Geosesarma notophorum sp. nov. (Decapoda, Brachyura, Grapsidae, Sesarminae), a terrestrial crab from Sumatra, with novel brooding behaviour. Crustaceana, 68 (3), 390 - 395. https: // doi. org / 10.1163 / 156854095 X 00557" type="journal article" year="1995">Ng & Tan 1995</bibRefCitation>
|
||
). This is the first confirmed record of this mode of brooding behavior for a Bornean species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |