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<document id="E4B7D781C76A9F9B8C53597FB53F733B" ID-DOI="10.11646/zootaxa.4093.3.1" ID-GBIF-Dataset="85f537f5-faf4-4a89-839b-8f56feadb0e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="264106" ID-ZooBank="565068DE-7042-483D-99FD-50BD2FA86BC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458719684729" checkinUser="plazi" docAuthor="Pollock, Darren A." docDate="2016" docId="038A6816711A433E2EDCF9D333E5FBAB" docLanguage="en" docName="zootaxa.4093.3.1.pdf.imf" docOrigin="Zootaxa 4093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eurypinae" docType="key" docVersion="9" lastPageNumber="303" masterDocId="FFB3106E711B433C2E4BFF88367EFFDC" masterDocTitle="Revision of Thisiomorphus Pic (Coleoptera: Mycteridae: Eurypinae) with descriptions of eleven new species from Central and South America and a key to genera of Neotropical Eurypinae" masterLastPageNumber="322" masterPageNumber="301" pageNumber="302" updateTime="1698185977904" updateUser="plazi">
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<mods:title id="17FE9F3F4A625D8C56CF17A0B826B2AC">Revision of Thisiomorphus Pic (Coleoptera: Mycteridae: Eurypinae) with descriptions of eleven new species from Central and South America and a key to genera of Neotropical Eurypinae</mods:title>
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<mods:name id="00FC6D718B5E3951A00A8342FA0DAE38" type="personal">
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<mods:namePart id="49B5A5B85CC3E829D94EE93D100529F0">Pollock, Darren A.</mods:namePart>
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<mods:date id="B7B1B69C3F639EA7AF3112807DDA1121">2016</mods:date>
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<treatment id="038A6816711A433E2EDCF9D333E5FBAB" ID-DOI="http://doi.org/10.5281/zenodo.6063826" ID-GBIF-Taxon="119257835" ID-Zenodo-Dep="6063826" LSID="urn:lsid:plazi:treatment:038A6816711A433E2EDCF9D333E5FBAB" httpUri="http://treatment.plazi.org/id/038A6816711A433E2EDCF9D333E5FBAB" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<subSubSection id="C3398A8B711A433D2EDCF9D33229F9A9" box="[151,1111,1627,1653]" pageId="1" pageNumber="302" type="nomenclature">
<paragraph id="8B9CD900711A433D2EDCF9D33229F9A9" blockId="1.[151,1111,1627,1653]" box="[151,1111,1627,1653]" pageId="1" pageNumber="302">
<heading id="D0D46E6C711A433D2EDCF9D33229F9A9" bold="true" box="[151,1111,1627,1653]" fontSize="11" level="1" pageId="1" pageNumber="302" reason="1">
<emphasis id="B9570512711A433D2EDCF9D33229F9A9" bold="true" box="[151,1111,1627,1653]" pageId="1" pageNumber="302">
Key to adults of described Neotropical (including Chile) genera of
<taxonomicName id="4C23A283711A433D2D98F9D33229F9A9" box="[979,1111,1627,1653]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3398A8B711A433E2EDCF92933E5FBAB" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302" type="key">
<keyStep id="30D7C1A5711A433D2EDCF92933E5F92D" pageId="1" pageNumber="302">
<paragraph id="8B9CD900711A433D2EDCF92933E5F90F" blockId="1.[151,1436,1697,2011]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF92933E5F90F" pageId="1" pageNumber="302">1 Head with distinct, shallowly impressed ocular grooves, narrowly separated from eyes, traceable from frontal canthus to poste- rior extent of eye...................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B9CD900711A433D2EDCF95433E5F92D" blockId="1.[151,1436,1697,2011]" box="[151,1435,1756,1777]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF95433E5F92D" box="[151,1435,1756,1777]" pageId="1" pageNumber="302">- Head without such an ocular groove....................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A5711A433D2EDCF97133E2F8BA" pageId="1" pageNumber="302">
<paragraph id="8B9CD900711A433D2EDCF97133E2F8F7" blockId="1.[151,1436,1697,2011]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF97133E2F8F7" pageId="1" pageNumber="302">
2 Procoxal cavities closed; dorsal surface without conspicuous setae; elytral shape generally conical, widest posteriorly (
<figureCitation id="1318C585711A433D2B04F97133EBF8D2" box="[1359,1429,1785,1806]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="1" pageNumber="302">Fig. 21</figureCitation>
).....................................................................
<taxonomicName id="4C23A283711A433D2A6BF89E32B3F8F7" authority="Gray" authorityName="Gray" box="[1056,1229,1814,1835]" class="Insecta" family="Mycteridae" genus="Stilpnonotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B9570512711A433D2A6BF89E32E9F8F7" box="[1056,1175,1814,1835]" italics="true" pageId="1" pageNumber="302">Stilpnonotus</emphasis>
Gray
</taxonomicName>
(7 described species)
</keyLead>
</paragraph>
<paragraph id="8B9CD900711A433D2EDCF8BC33E2F8BA" blockId="1.[151,1436,1697,2011]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF8BC33E2F8BA" pageId="1" pageNumber="302">
- Procoxal cavities narrowly open; dorsal surface with conspicuous setae (although short in some species); elytral shape gener- ally parallel-sided (
<figureCitation id="1318C585711A433D2FD1F8D9379EF8BA" box="[410,480,1873,1894]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="1" pageNumber="302">Fig. 22</figureCitation>
)........................................
<taxonomicName id="4C23A283711A433D2D85F8D932BFF8BA" authority="Champion" authorityName="Champion" box="[974,1217,1873,1894]" class="Insecta" family="Mycteridae" genus="Conomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="B9570512711A433D2D85F8D93228F8BA" box="[974,1110,1873,1894]" italics="true" pageId="1" pageNumber="302">Conomorphus</emphasis>
Champion
</taxonomicName>
(10 described species)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A5711A433D2EDCF8E633E5F862" pageId="1" pageNumber="302">
<paragraph id="8B9CD900711A433D2EDCF8E633E5F85F" blockId="1.[151,1436,1697,2011]" box="[151,1435,1902,1923]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF8E633E5F85F" box="[151,1435,1902,1923]" pageId="1" pageNumber="302">
3 Eye with conspicuous intrafacetal setae (e.g.,
<figureCitation id="1318C585711A433D2CDCF8E634A2F85F" box="[663,732,1902,1923]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="1" pageNumber="302">Fig. 32</figureCitation>
)......................................................... 4
</keyLead>
</paragraph>
<paragraph id="8B9CD900711A433D2EDCF80433E5F862" blockId="1.[151,1436,1697,2011]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF80433E5F862" pageId="1" pageNumber="302">- Eye either without intrafacetal setae, or setae so short they are not noticeable using standard light microscopy, definitely not conspicuous.......................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A5711A433E2EDCF84E33E5FF70" lastPageId="2" lastPageNumber="303" pageId="1" pageNumber="302">
<paragraph id="8B9CD900711A433D2EDCF84E33E5F807" blockId="1.[151,1436,1697,2011]" box="[151,1435,1990,2011]" pageId="1" pageNumber="302">
<keyLead id="30D27A35711A433D2EDCF84E33E5F807" box="[151,1435,1990,2011]" pageId="1" pageNumber="302">
4. Head with distinct, deep frontal furrows (e.g.,
<figureCitation id="1318C585711A433D2CD6F84E3574F807" box="[669,778,1990,2011]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="1" pageNumber="302">Figs 32–33</figureCitation>
)..................................................... 5
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFF1F33E5FF70" blockId="2.[151,1436,151,1143]" box="[151,1435,151,172]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFF1F33E5FF70" box="[151,1435,151,172]" pageId="2" pageNumber="303">- Head without deep frontal furrows........................................................................ 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFF3D33EBFE9D" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFF3D33E7FF27" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFF3D33E7FF27" pageId="2" pageNumber="303">
5 Mandibles bluntly unidentate apically; eyes small, widely separated (ocular index&gt; 60); antennomeres elongate, filiform to subserrate; dorsal body surface with light and dark setae, producing a mottled appearance (
<figureCitation id="1318C5857119433E2A0CFF5A32EEFF3B" box="[1095,1168,210,231]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="2" pageNumber="303">Fig. 23</figureCitation>
)...........................................................................................
<taxonomicName id="4C23A2837119433E2A5EFF7A32B8FEDB" authority="Solier" authorityName="Solier" box="[1045,1222,242,263]" class="Insecta" family="Mycteridae" genus="Loboglossa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2A5EFF7A32F9FEDB" box="[1045,1159,242,263]" italics="true" pageId="2" pageNumber="303">Loboglossa</emphasis>
Solier
</taxonomicName>
(6 described species)1
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFE8733EBFE9D" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFE8733EBFE9D" pageId="2" pageNumber="303">- Mandibles bidentate apically; eyes larger, more narrowly separated (ocular index &lt;60); antennomeres variable; dorsal body surface either uniform in color, or with longitudinally oriented color pattern, not mottled............................ 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFEC233E2FE6B" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFEC233E2FEA0" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFEC233E2FEA0" pageId="2" pageNumber="303">
6 Penultimate tarsomere very wide, wider than length of distal tarsomere (
<figureCitation id="1318C5857119433E2DD0FEC2359DFE83" box="[923,995,330,351]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 32</figureCitation>
); frontal furrows relatively straight, subparallel; ommatidia coarse..............................................
<taxonomicName id="4C23A2837119433E2DAFFEEF320DFEA0" authority="Kirby" authorityName="Kirby" box="[996,1139,359,380]" class="Insecta" family="Mycteridae" genus="Eurypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2DAFFEEF324AFEA0" box="[996,1076,359,380]" italics="true" pageId="2" pageNumber="303">Eurypus</emphasis>
Kirby
</taxonomicName>
(6 described species) (
<figureCitation id="1318C5857119433E2B04FEEF33EBFEA0" box="[1359,1429,359,380]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="2" pageNumber="303">Fig. 24</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFE0C33E2FE6B" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFE0C33E2FE6B" pageId="2" pageNumber="303">
- Penultimate tarsomere wide, but width less than length of distal tarsomere; frontal furrows somewhat arcuate, distinctly convergent anteriorly (
<figureCitation id="1318C5857119433E2FD3FE2A379EFE6B" box="[408,480,418,439]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 33</figureCitation>
); ommatidia fine......................
<emphasis id="B95705127119433E2DC6FE2A3274FE6B" box="[909,1034,418,439]" italics="true" pageId="2" pageNumber="303">Cladobradus</emphasis>
Pic (1 species:
<taxonomicName id="4C23A2837119433E2AEAFE2A3345FE6B" authority="Pic" authorityName="Pic" box="[1185,1339,418,439]" class="Insecta" family="Mycteridae" genus="Conomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="atritarsis">
<emphasis id="B95705127119433E2AEAFE2A32CFFE6B" box="[1185,1201,418,439]" italics="true" pageId="2" pageNumber="303">C</emphasis>
.
<emphasis id="B95705127119433E2AF6FE2A336BFE6B" box="[1213,1301,418,439]" italics="true" pageId="2" pageNumber="303">atritarsis</emphasis>
Pic
</taxonomicName>
) (
<figureCitation id="1318C5857119433E2B04FE2A33EBFE6B" box="[1359,1429,418,439]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="2" pageNumber="303">Fig. 25</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFE3733E5FDD3" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFE3733E5FE2D" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFE3733E5FE2D" pageId="2" pageNumber="303">
<specimenCount id="9D2512897119433E2EDCFE37376CFE08" box="[151,274,447,468]" pageId="2" pageNumber="303" type="male">7 Male</specimenCount>
sex patch consisting of setae on both ventrites 2 and 3; wing rudimentary, not extended to elytral apex................................................................
<taxonomicName id="4C23A2837119433E2D78FE543246FE2D" authority="Champion" authorityName="Champion" box="[819,1080,476,497]" class="Insecta" family="Mycteridae" genus="Conomorphinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2D78FE5435B0FE2D" box="[819,974,476,497]" italics="true" pageId="2" pageNumber="303">Conomorphinus</emphasis>
Champion
</taxonomicName>
(1 species:
<taxonomicName id="4C23A2837119433E2AE1FE5433E8FE2D" authority="Champion" authorityName="Champion" box="[1194,1430,476,497]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="bolivianus">
<emphasis id="B95705127119433E2AE1FE5432C4FE2D" box="[1194,1210,476,497]" italics="true" pageId="2" pageNumber="303">C</emphasis>
.
<emphasis id="B95705127119433E2A8DFE543354FE2D" box="[1222,1322,476,497]" italics="true" pageId="2" pageNumber="303">bolivianus</emphasis>
Champion
</taxonomicName>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFE7233E5FDD3" blockId="2.[151,1436,151,1143]" box="[151,1435,506,527]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFE7233E5FDD3" box="[151,1435,506,527]" pageId="2" pageNumber="303">
- Male sex patch restricted to
<date id="FF9DFFC07119433E2FBBFE72346EFDD3" box="[496,528,506,527]" pageId="2" pageNumber="303">V2</date>
; wing fully developed, extended to elytral apex.................................... 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFD9F33E2FD58" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFD9F33E5FD95" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFD9F33E5FD95" pageId="2" pageNumber="303">
<specimenCount id="9D2512897119433E2EDCFD9F376CFDF0" box="[151,274,535,556]" pageId="2" pageNumber="303" type="male">8 Male</specimenCount>
sex patch consisting of distinctly raised, rounded “platform” with either no setae, or setae short and dense; known only from Chile.............................................
<taxonomicName id="4C23A2837119433E2D2BFDBC32EEFD95" authority="Fairmaire &amp; Germain" authorityName="Fairmaire &amp; Germain" box="[864,1168,564,585]" class="Insecta" family="Mycteridae" genus="Batobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2D2BFDBC35C4FD95" box="[864,954,564,585]" italics="true" pageId="2" pageNumber="303">Batobius</emphasis>
Fairmaire &amp; Germain
</taxonomicName>
(4 described spp.) (
<figureCitation id="1318C5857119433E2B05FDBC33EAFD95" box="[1358,1428,564,585]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 27</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFDDA33E2FD58" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFDDA33E2FD58" pageId="2" pageNumber="303">
- Male sex patch consisting of relatively long setae, borne on raised medial section of ventrite 2; Known only from Brazil.............................
<taxonomicName id="4C23A2837119433E2C43FDE7348BFD58" authority="Pollock" authorityName="Pollock" box="[520,757,623,644]" class="Insecta" family="Mycteridae" genus="Brasilaccoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2C43FDE734DBFD58" box="[520,677,623,644]" italics="true" pageId="2" pageNumber="303">Brasilaccoderus</emphasis>
Pollock
</taxonomicName>
(2 species:
<taxonomicName id="4C23A2837119433E2D2DFDE73244FD58" authority="Champion" authorityName="Champion" box="[870,1082,623,644]" class="Insecta" family="Mycteridae" genus="Brasilaccoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="B95705127119433E2D2DFDE7350BFD58" box="[870,885,623,644]" italics="true" pageId="2" pageNumber="303">B</emphasis>
.
<emphasis id="B95705127119433E2DCAFDE735BFFD58" box="[897,961,623,644]" italics="true" pageId="2" pageNumber="303">scaber</emphasis>
(Champion)
</taxonomicName>
and
<taxonomicName id="4C23A2837119433E2A22FDE73342FD58" authority="Pollock" authorityName="Pollock" box="[1129,1340,623,644]" class="Insecta" family="Mycteridae" genus="Brasilaccoderus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="plaumanni">
<emphasis id="B95705127119433E2A22FDE73206FD58" box="[1129,1144,623,644]" italics="true" pageId="2" pageNumber="303">B</emphasis>
.
<emphasis id="B95705127119433E2ACFFDE73294FD58" box="[1156,1258,623,644]" italics="true" pageId="2" pageNumber="303">plaumanni</emphasis>
Pollock
</taxonomicName>
) (
<figureCitation id="1318C5857119433E2B04FDE733EBFD58" box="[1359,1429,623,644]" captionStart="FIGURES 21 – 26" captionStartId="8.[151,264,1387,1409]" captionTargetBox="[218,1376,170,1355]" captionTargetId="figure@8.[196,1385,153,1365]" captionTargetPageId="8" captionText="FIGURES 21 – 26. Dorsal habitus of representative Neotropical Eurypinae. 21. Stilpnonotus sp.; 22. Conomorphus sp.; 23. Loboglossa variipennis Solier; 24. Eurypus sp.; 25. Cladobradus atritarsis Pic, holotype, TL = 12 mm; 26. Brasilaccoderus plaumanni Pollock." httpUri="https://zenodo.org/record/264110/files/figure.png" pageId="2" pageNumber="303">Fig. 26</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFD0433E5FD25" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFD0433E5FD63" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFD0433E5FD63" pageId="2" pageNumber="303">
9 Head distinctly rounded; eyes large, width across eyes equal to or greater than width of pronotum; pronotum with lateral carina present, at least in basal half; male sex patch on
<date id="FF9DFFC07119433E2CE4FD2234ABFD63" box="[687,725,682,703]" pageId="2" pageNumber="303">V3.</date>
......................................................... 10
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFD4F33E5FD25" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFD4F33E5FD25" pageId="2" pageNumber="303">
- Head not distinctly rounded; eyes usually smaller, width across eyes less than or equal to width of pronotum; pronotum with lateral carina absent, or at most vaguely suggested only, in basal half; male sex patch on
<date id="FF9DFFC07119433E2A79FD6C321EFD25" box="[1074,1120,740,761]" pageId="2" pageNumber="303">V2.</date>
........................ 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFC8A33E5FCB3" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFC8A33E2FCE8" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFC8A33E2FCE8" pageId="2" pageNumber="303">
10 TL of most specimens&gt;
<quantity id="4CDB74E57119433E2F9BFC8A3462FCCB" box="[464,540,770,791]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="303" unit="mm" value="3.5">3.5 mm</quantity>
; apical maxillary palpomere cultriform; apical labial palpomere ovate with truncate apex...........................................
<taxonomicName id="4C23A2837119433E2CF5FC9735F0FCE8" authority="Champion" authorityName="Champion" box="[702,910,799,820]" class="Insecta" family="Mycteridae" genus="Cleodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2CF5FC97355DFCE8" box="[702,803,799,820]" italics="true" pageId="2" pageNumber="303">Cleodaeus</emphasis>
Champion
</taxonomicName>
(1 described species:
<taxonomicName id="4C23A2837119433E2A2AFC973345FCE8" authority="Champion" authorityName="Champion" box="[1121,1339,799,820]" class="Insecta" family="Mycteridae" genus="Cleodaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="rugiceps">
<emphasis id="B95705127119433E2A2AFC97320FFCE8" box="[1121,1137,799,820]" italics="true" pageId="2" pageNumber="303">C</emphasis>
.
<emphasis id="B95705127119433E2A36FC9732AEFCE8" box="[1149,1232,799,820]" italics="true" pageId="2" pageNumber="303">rugiceps</emphasis>
Champion
</taxonomicName>
) (
<figureCitation id="1318C5857119433E2B04FC9733EBFCE8" box="[1359,1429,799,820]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 30</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFCB433E5FCB3" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFCB433E5FCB3" pageId="2" pageNumber="303">
- TL of most specimens &lt;
<quantity id="4CDB74E57119433E2F9BFCB43462FC8D" box="[464,540,828,849]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="303" unit="mm" value="3.5">3.5 mm</quantity>
; apical maxillary palpomere securiform; apical labial palpomere narrowed distally, pyriform...........................................................
<taxonomicName id="4C23A2837119433E2DECFCD23219FCB3" authority="Champion" authorityName="Champion" box="[935,1127,858,879]" class="Insecta" family="Mycteridae" genus="Physcius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2DECFCD23582FCB3" box="[935,1020,858,879]" italics="true" pageId="2" pageNumber="303">Physcius</emphasis>
Champion
</taxonomicName>
(22 described species) (
<figureCitation id="1318C5857119433E2B04FCD233EBFCB3" box="[1359,1429,858,879]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 31</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFCFF33E5FC38" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFCFF33E2FC76" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFCFF33E2FC76" pageId="2" pageNumber="303">
11 Eyes flat and not protuberant, facets coarse, width of single eye about half length; frontal furrows absent; male sex patch absent........................................................
<taxonomicName id="4C23A2837119433E2D89FC1D320DFC76" authority="Champion" authorityName="Champion" box="[962,1139,917,938]" class="Insecta" family="Mycteridae" genus="Thisias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2D89FC1D3279FC76" box="[962,1031,917,938]" italics="true" pageId="2" pageNumber="303">Thisias</emphasis>
Champion
</taxonomicName>
(5 described species) (
<figureCitation id="1318C5857119433E2B04FC1D33EBFC76" box="[1359,1429,917,938]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 28</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFC3A33E5FC38" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFC3A33E5FC38" pageId="2" pageNumber="303">- Eyes more protuberant, facets fine, width of single eye distinctly greater than half length; frontal furrows at least suggested, often rather deep; male sex patch present, on ventrite 2....................................................... 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30D7C1A57119433E2EDCFC6533E5FBAB" pageId="2" pageNumber="303">
<paragraph id="8B9CD9007119433E2EDCFC6533E2FBC3" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFC6533E2FBC3" pageId="2" pageNumber="303">
12 Head with frons slightly to distinctly sunken about level of antennal insertions; pronotal disc with pair of distinct, rounded depressions...................................................
<taxonomicName id="4C23A2837119433E2DFFFB82320DFBC3" authority="Pic" authorityName="Pic" box="[948,1139,1034,1055]" class="Insecta" family="Mycteridae" genus="Physiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2DFFFB823233FBC3" box="[948,1101,1034,1055]" italics="true" pageId="2" pageNumber="303">Physiomorphus</emphasis>
Pic
</taxonomicName>
(8 described species) (
<figureCitation id="1318C5857119433E2B04FB8233EBFBC3" box="[1359,1429,1034,1055]" captionStart="FIGURES 27 – 33" captionStartId="9.[151,264,1635,1657]" captionTargetBox="[167,1412,206,1593]" captionTargetId="figure@9.[151,1436,193,1614]" captionTargetPageId="9" captionText="FIGURES 27 – 33. Representative Neotropical Eurypinae. 27 – 31. Dorsal habitus. Batobius sp.; 28. Thisias? marmoratus Champion, TL = 8.6 mm; 29. Physiomorphus subcostulatus Pollock; 30. Cleodaeus rugiceps Champion, TL = 5.1 mm; 31. Physcius sp., TL = 3.0 mm. 32 – 33. Head detail. 32. Eurypus sp.; 33. Cladobradus atritarsis Pic, holotype." httpUri="https://zenodo.org/record/264111/files/figure.png" pageId="2" pageNumber="303">Fig. 29</figureCitation>
)
</keyLead>
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFBAF33E5FBAB" blockId="2.[151,1436,151,1143]" pageId="2" pageNumber="303">
<keyLead id="30D27A357119433E2EDCFBAF33E5FBAB" pageId="2" pageNumber="303">
- Head with frons flat, not sunken; pronotal disc uniformly convex, without pair of distinct, rounded depressions (in some specimens – notably
<taxonomicName id="4C23A2837119433E2FCBFBCD3478FB86" box="[384,518,1093,1114]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B95705127119433E2FCBFBCD37F2FB86" box="[384,396,1093,1114]" italics="true" pageId="2" pageNumber="303">T</emphasis>
.
<emphasis id="B95705127119433E2FDCFBCD3478FB86" box="[407,518,1093,1114]" italics="true" pageId="2" pageNumber="303">brasiliensis</emphasis>
</taxonomicName>
– with slight indentations, but not distinct, rounded depressions)....................................................................................................
<taxonomicName id="4C23A2837119433E2A39FBEA3358FBAB" authority="Pic" authorityName="Pic" box="[1138,1318,1122,1143]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2A39FBEA337CFBAB" box="[1138,1282,1122,1143]" italics="true" pageId="2" pageNumber="303">Thisiomorphus</emphasis>
Pic
</taxonomicName>
(
<figureCitation id="1318C5857119433E2B78FBEA33EAFBAB" box="[1331,1428,1122,1143]" captionStart-0="FIGURES 1 – 6" captionStart-1="FIGURES 7 – 12" captionStartId-0="5.[151,264,1662,1684]" captionStartId-1="6.[151,264,1598,1620]" captionTargetBox-0="[212,1374,202,1600]" captionTargetBox-1="[248,1353,186,1561]" captionTargetId-0="figure@5.[203,1384,193,1641]" captionTargetId-1="figure@6.[226,1360,181,1577]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1 – 6. Dorsal habitus of Thisiomorphus species. 1. T. andrewsi, n. sp., male holotype; 2. T. bolivianus, n. sp., female holotype; 3. T. brasiliensis, n. sp., male holotype; 4. T. caeruleus, n. sp., female; 5. T. convergens, n. sp., female holotype; 6. T. davidsoni, n. sp., female holotype." captionText-1="FIGURES 7 – 12. Dorsal habitus of Thisiomorphus species. 7. T. curticornis, n. sp., male holotype; 8. T. festivus, n. sp., male holotype; 9. T. inaequalis, n. sp., male holotype; 10. T. latus Pic, holotype; 11. T. osaensis, n. sp., female; 12. T. solisi, n. sp., male holotype." httpUri-0="https://zenodo.org/record/264107/files/figure.png" httpUri-1="https://zenodo.org/record/264108/files/figure.png" pageId="2" pageNumber="303">Figs 1–12</figureCitation>
)
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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