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<document id="87896ED20BE47BCF70A226C83B4700EB" ID-CLB-Dataset="31177" ID-DOI="10.11646/zootaxa.4363.4.1" ID-GBIF-Dataset="a141fa76-cc88-4901-944a-306171e41413" ID-ISSN="1175-5326" ID-Zenodo-Dep="1114473" ID-ZooBank="86DE1B94-63AE-4ABF-B28A-0ECEA22D2F10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513152258038" checkinUser="plazi" docAuthor="Mcfadden, Catherine S. &amp; Van Ofwegen, Leen P." docDate="2017" docId="038A07766C1C2862FF245D06FE334B22" docLanguage="en" docName="zootaxa.4363.4.1.pdf" docOrigin="Zootaxa 4363 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptophyton fustis Mcfadden &amp; Van Ofwegen 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="475" masterDocId="FFB37F0E6C0A287AFFB35C76FF964E2E" masterDocTitle="Revisionary systematics of the endemic soft coral fauna (Octocorallia: Alcyonacea: Alcyoniina) of the Agulhas Bioregion, South Africa" masterLastPageNumber="488" masterPageNumber="451" pageNumber="473" updateTime="1698509335316" updateUser="ExternalLinkService">
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<mods:title id="A1449175A677A7ADFF88268BCDD138EA">Revisionary systematics of the endemic soft coral fauna (Octocorallia: Alcyonacea: Alcyoniina) of the Agulhas Bioregion, South Africa</mods:title>
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<mods:namePart id="FF9A208EBEDD10349ACEFC95589162F8">Mcfadden, Catherine S.</mods:namePart>
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<mods:namePart id="60C92E986B8D3BCBD72C5183260E3ED8">Van Ofwegen, Leen P.</mods:namePart>
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<mods:title id="F5EF6394B2D9A00F5F8443A7C7140717">Zootaxa</mods:title>
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<mods:date id="78E683D3CDF196AE6B1CA4D2BD5D9D85">2017</mods:date>
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<treatment id="038A07766C1C2862FF245D06FE334B22" ID-DOI="http://doi.org/10.5281/zenodo.5613950" ID-GBIF-Taxon="137679934" ID-Zenodo-Dep="5613950" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22" httpUri="http://treatment.plazi.org/id/038A07766C1C2862FF245D06FE334B22" lastPageId="24" lastPageNumber="475" pageId="22" pageNumber="473">
<subSubSection id="C339E5EB6C1C286CFF245D06FEC74F85" pageId="22" pageNumber="473" type="nomenclature">
<paragraph id="8B9CB6606C1C286CFF245D06FE564FA4" blockId="22.[151,448,368,427]" box="[151,448,368,394]" pageId="22" pageNumber="473">
<heading id="D0D4010C6C1C286CFF245D06FE564FA4" bold="true" box="[151,448,368,394]" fontSize="11" level="1" pageId="22" pageNumber="473" reason="1">
<emphasis id="B9576A726C1C286CFF245D06FE564FA4" bold="true" box="[151,448,368,394]" pageId="22" pageNumber="473">
<taxonomicName id="4C23CDE36C1C286CFF245D06FEE04FA4" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[151,374,368,394]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="22" pageNumber="473" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C1C286CFF245D06FEE04FA4" bold="true" box="[151,374,368,394]" italics="true" pageId="22" pageNumber="473">Leptophyton fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C1C286CFECD5D06FE564FA4" box="[382,448,368,394]" pageId="22" pageNumber="473" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9CB6606C1C286CFF245DE4FEC74F85" blockId="22.[151,448,368,427]" box="[151,337,402,427]" pageId="22" pageNumber="473">
<figureCitation id="1318AAE56C1C286CFF245DE4FF654F85" box="[151,243,402,427]" captionStart="FIGURE 5" captionStartId="10.[151,250,1819,1841]" captionTargetBox="[232,1366,195,1796]" captionTargetId="figure@10.[232,1366,195,1796]" captionTargetPageId="10" captionText="FIGURE 5. a, Lampophyton spinatum n. sp. paratype, RMNH Coel. 40199 (SAF195), colony photographed in situ; b, Alcyonium dolium n. sp. holotype, RMNH Coel.40204 (SAF292); c, Leptophyton benayahui SAF289 photographed in situ; d, Leptophyton fustis n. sp., holotype, RMNH Coel. 40211 (SAF087), colony photographed in situ (T. fallax n. comb. purple morph visible in lower right corner); e, Circularius wilsoni n. comb. RMNH Coel. 40207 (SAF448), colonies growing on mussel shell photographed in situ; f, Tenerodus pollex n. gen., n. sp. holotype, RMNH Coel. 40219 (SAF420), colonies photographed in situ; g, Tenerodus fallax, n. comb. RMNH Coel. 40217 (SAF383), pink color morph; h, Tenerodus fallax, n. comb. SAF015, purple and yellow color morphs." httpUri="https://zenodo.org/record/1114515/files/figure.png" pageId="22" pageNumber="473">Figs. 5d</figureCitation>
,
<figureCitation id="1318AAE56C1C286CFF4C5DE5FEBC4F85" box="[255,298,403,427]" captionStart="FIGURE 13" captionStartId="22.[151,250,1937,1959]" captionTargetBox="[181,1409,695,1911]" captionTargetId="figure@22.[181,1409,695,1911]" captionTargetPageId="22" captionText="FIGURE 13. a, Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087); b, Tenerodus fallax RMNH Coel. 40215 (SAF067), attached to a polychaete tube; c, Tenerodus fallax RMNH Coel. 40217 (SAF383), growing on a mussel shell; d, Tenerodus pollex n. sp. holotype, RMNH Coel.40219 (SAF420)." httpUri="https://zenodo.org/record/1114559/files/figure.png" pageId="22" pageNumber="473">13a</figureCitation>
,
<figureCitation id="1318AAE56C1C286CFE865DE5FEC74F85" box="[309,337,403,427]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="22" pageNumber="473">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C339E5EB6C1C286CFF245DACFE384CAD" pageId="22" pageNumber="473" type="materials_examined">
<paragraph id="8B9CB6606C1C286CFF245DACFE384CAD" blockId="22.[151,1436,474,643]" pageId="22" pageNumber="473">
<materialsCitation id="3B4BBC3D6C1C286CFF245DACFBE04C39" ID-GBIF-Occurrence="1705513526" collectingDate="2008-03-11" collectionCode="RMNH" collectorName="C. S. McFadden" country="South Africa" county="Material" elevation="18" latitude="-33.98495" location="Algoa Bay" longLatPrecision="1" longitude="25.8629" municipality="Coel." pageId="22" pageNumber="473" specimenCount="1" stateProvince="Eastern Cape" typeStatus="holotype">
<emphasis id="B9576A726C1C286CFF245DACFEE84FDD" bold="true" box="[151,382,474,499]" pageId="22" pageNumber="473">
<collectingCounty id="62FDCEEC6C1C286CFF245DACFF694FDD" box="[151,255,474,499]" pageId="22" pageNumber="473">Material</collectingCounty>
examined
</emphasis>
.
<typeStatus id="549808C26C1C286CFE215DACFE694FDD" box="[402,511,474,499]" pageId="22" pageNumber="473" type="holotype">
<emphasis id="B9576A726C1C286CFE215DACFE694FDD" bold="true" box="[402,511,474,499]" pageId="22" pageNumber="473">Holotype</emphasis>
</typeStatus>
.
<collectionCode id="ED322EA56C1C286CFDA15DADFDF04FDC" box="[530,614,475,498]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
<collectingMunicipality id="6BF82C1A6C1C286CFDC15DACFD394FDD" box="[626,687,474,499]" pageId="22" pageNumber="473">Coel.</collectingMunicipality>
40211 (SAF087),
<collectingCountry id="F334F6F06C1C286CFC3C5DACFBB14FDD" box="[911,1063,474,499]" name="South Africa" pageId="22" pageNumber="473">South Africa</collectingCountry>
,
<collectingRegion id="49E778826C1C286CFB845DADFB454FDD" box="[1079,1235,475,499]" country="South Africa" name="Eastern Cape" pageId="22" pageNumber="473">
<collectingRegion id="49E778826C1C286CFB845DADFB1D4FDC" box="[1079,1163,475,498]" country="American Samoa" name="Eastern" pageId="22" pageNumber="473">Eastern</collectingRegion>
Cape
</collectingRegion>
,
<location id="8EFCE0BB6C1C286CFB575DACFAF54FDD" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22:8EFCE0BB6C1C286CFB575DACFAF54FDD" box="[1252,1379,474,499]" country="South Africa" county="Material" latitude="-33.98495" longLatPrecision="1" longitude="25.8629" municipality="Coel." name="Algoa Bay" pageId="22" pageNumber="473" stateProvince="Eastern Cape">Algoa Bay</location>
,
<location id="8EFCE0BB6C1C286CFAC75DACFF764C39" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22:8EFCE0BB6C1C286CFAC75DACFF764C39" country="South Africa" county="Material" latitude="-33.98495" longLatPrecision="1" longitude="25.8629" municipality="Coel." name="Riy Banks" pageId="22" pageNumber="473" stateProvince="Eastern Cape">Riy Banks</location>
,
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,
<geoCoordinate id="EE17D0A76C1C286CFE305D88FD844C39" box="[387,530,510,535]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="1" value="25.8629">25º51.774E</geoCoordinate>
, depth
<elevation id="000E51536C1C286CFDD15D88FD514C38" box="[610,711,510,535]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="22" pageNumber="473" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">
<quantity id="4CDB1B856C1C286CFDD15D88FD514C38" box="[610,711,510,535]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="22" pageNumber="473" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
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, coll.
<collectorName id="26D6D3B66C1C286CFCB85D89FC2B4C39" box="[779,957,510,535]" pageId="22" pageNumber="473">C.S. McFadden</collectorName>
,
<collectingDate id="EFD969486C1C286CFC7B5D89FBE04C39" box="[968,1142,510,535]" pageId="22" pageNumber="473" value="2008-03-11">
<date id="FF9D90A06C1C286CFC7B5D89FBE04C39" box="[968,1142,510,535]" pageId="22" pageNumber="473" value="2008-03-11">11 March 2008</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B4BBC3D6C1C286CFBCC5D88FECE4C15" ID-GBIF-Occurrence="1705513529" collectionCode="RMNH" country="South Africa" location="Coel." pageId="22" pageNumber="473" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<typeStatus id="549808C26C1C286CFBCC5D88FB614C39" box="[1151,1271,510,535]" pageId="22" pageNumber="473" type="paratype">
<emphasis id="B9576A726C1C286CFBCC5D88FB614C39" bold="true" box="[1151,1271,510,535]" pageId="22" pageNumber="473">Paratypes</emphasis>
</typeStatus>
.
<collectionCode id="ED322EA56C1C286CFAB65D89FACF4C38" box="[1285,1369,511,534]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
<location id="8EFCE0BB6C1C286CFAD35D88FA0A4C39" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22:8EFCE0BB6C1C286CFAD35D88FA0A4C39" box="[1376,1436,510,535]" country="South Africa" name="Coel." pageId="22" pageNumber="473" stateProvince="Eastern Cape">Coel.</location>
40213 (SAF086)
</materialsCitation>
,
<materialsCitation id="3B4BBC3D6C1C286CFED75E55FC464C15" ID-GBIF-Occurrence="1705513532" box="[356,976,546,571]" collectingDate="2008-03-11" collectionCode="RMNH" collectorName="C. S. McFadden" country="South Africa" county="Material" elevation="18" latitude="-33.98495" location="Coel." longLatPrecision="1" longitude="25.8629" municipality="Coel." pageId="22" pageNumber="473" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<collectionCode id="ED322EA56C1C286CFED75E55FE2E4C14" box="[356,440,547,570]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="22" pageNumber="473">RMNH</collectionCode>
<location id="8EFCE0BB6C1C286CFE735E54FE6B4C15" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22:8EFCE0BB6C1C286CFE735E54FE6B4C15" box="[448,509,546,571]" name="Coel." pageId="22" pageNumber="473">Coel.</location>
40212 (SAF088), same data as holotype
</materialsCitation>
.
<materialsCitation id="3B4BBC3D6C1C286CFC685E54FE3D4CAD" ID-GBIF-Occurrence="1705513534" collectingDate="2000-03-22" collectionCode="RMNH" collectorName="Grootbank &amp; J. Starmer" country="South Africa" county="Material" elevation="9" latitude="-33.98448" location="Plettenberg Bay" longLatPrecision="1" longitude="25.864017" municipality="Other" pageId="22" pageNumber="473" specimenCount="1" stateProvince="Eastern Cape" typeStatus="paratype">
<emphasis id="B9576A726C1C286CFC685E54FB014C15" bold="true" box="[987,1175,546,571]" pageId="22" pageNumber="473">
<collectingMunicipality id="6BF82C1A6C1C286CFC685E54FBB54C15" box="[987,1059,546,571]" pageId="22" pageNumber="473">Other</collectingMunicipality>
material.
</emphasis>
SAF099, same data as holotype except
<geoCoordinate id="EE17D0A76C1C286CFEE15E30FE494C71" box="[338,479,582,607]" direction="south" orientation="latitude" pageId="22" pageNumber="473" precision="1" value="-33.98448">33º59.069S</geoCoordinate>
,
<geoCoordinate id="EE17D0A76C1C286CFE5F5E30FDE44C71" box="[492,626,582,607]" direction="east" orientation="longitude" pageId="22" pageNumber="473" precision="1" value="25.864017">25º51.841E</geoCoordinate>
. UF2435,
<collectingCountry id="F334F6F06C1C286CFD5B5E30FCEA4C71" box="[744,892,582,607]" name="South Africa" pageId="22" pageNumber="473">South Africa</collectingCountry>
,
<location id="8EFCE0BB6C1C286CFC3B5E30FBAB4C71" LSID="urn:lsid:plazi:treatment:038A07766C1C2862FF245D06FE334B22:8EFCE0BB6C1C286CFC3B5E30FBAB4C71" box="[904,1085,582,607]" country="South Africa" latitude="-33.98448" longLatPrecision="1" longitude="25.864017" municipality="Other" name="Plettenberg Bay" pageId="22" pageNumber="473">Plettenberg Bay</location>
,
<collectorName id="26D6D3B66C1C286CFBF95E30FB534C71" box="[1098,1221,582,607]" pageId="22" pageNumber="473">Grootbank</collectorName>
, depth
<elevation id="000E51536C1C286CFAA55E31FAD24C70" box="[1302,1348,583,607]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="473" unit="m" value="9.0">
<quantity id="4CDB1B856C1C286CFAA55E31FAD24C70" box="[1302,1348,583,607]" metricMagnitude="0" metricUnit="m" metricValue="9.0" pageId="22" pageNumber="473" unit="m" value="9.0">9 m</quantity>
</elevation>
, coll.
<collectorName id="26D6D3B66C1C286CFA395E31FF674CAD" pageId="22" pageNumber="473">J. Starmer</collectorName>
,
<collectingDate id="EFD969486C1C286CFF4F5E1DFE3D4CAD" box="[252,427,618,643]" pageId="22" pageNumber="473" value="2000-03-22">
<date id="FF9D90A06C1C286CFF4F5E1DFE3D4CAD" box="[252,427,618,643]" pageId="22" pageNumber="473" value="2000-03-22">22 March 2000</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF5CE6E86C1C286CFF245BE7FCB849CB" httpUri="https://zenodo.org/record/1114559/files/figure.png" pageId="22" pageNumber="473" targetBox="[181,1409,695,1911]" targetPageId="22">
<paragraph id="8B9CB6606C1C286CFF245BE7FCB849CB" blockId="22.[151,1436,1937,2021]" pageId="22" pageNumber="473">
<emphasis id="B9576A726C1C286CFF245BE7FEB74989" bold="true" box="[151,289,1937,1959]" pageId="22" pageNumber="473">FIGURE 13.</emphasis>
a,
<taxonomicName id="4C23CDE36C1C286CFEF05BE7FE694989" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[323,511,1937,1959]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="22" pageNumber="473" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C1C286CFEF05BE7FE694989" box="[323,511,1937,1959]" italics="true" pageId="22" pageNumber="473">Leptophyton fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C1C286CFDB45BE7FDD74989" box="[519,577,1937,1959]" pageId="22" pageNumber="473" rank="species">
<emphasis id="B9576A726C1C286CFDB45BE7FDD74989" bold="true" box="[519,577,1937,1959]" pageId="22" pageNumber="473">n. sp.</emphasis>
</taxonomicNameLabel>
holotype, RMNH Coel. 40211 (SAF087); b
<taxonomicName id="4C23CDE36C1C286CFBBF5BE7FB554989" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[1036,1219,1937,1959]" pageId="22" pageNumber="473" rank="subSpecies" subSpecies="fallax">
,
<emphasis id="B9576A726C1C286CFBAA5BE7FB554989" box="[1049,1219,1937,1959]" italics="true" pageId="22" pageNumber="473">Tenerodus fallax</emphasis>
</taxonomicName>
RMNH Coel. 40215 (SAF067), attached to a polychaete tube; c
<taxonomicName id="4C23CDE36C1C286CFDE45BC7FC9949E8" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[599,783,1968,1990]" pageId="22" pageNumber="473" rank="subSpecies" subSpecies="fallax">
,
<emphasis id="B9576A726C1C286CFDD75BC6FC9949E8" box="[612,783,1968,1990]" italics="true" pageId="22" pageNumber="473">Tenerodus fallax</emphasis>
</taxonomicName>
RMNH Coel. 40217 (SAF383), growing on a mussel shell; d,
<taxonomicName id="4C23CDE36C1C286CFF245BB9FED549CB" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[151,323,1999,2021]" class="Anthozoa" family="Paralcyoniidae" genus="Tenerodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="22" pageNumber="473" phylum="Cnidaria" rank="species" species="pollex" status="sp. nov.">
<emphasis id="B9576A726C1C286CFF245BB9FED549CB" box="[151,323,1999,2021]" italics="true" pageId="22" pageNumber="473">Tenerodus pollex</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C1C286CFEFA5BB9FE1449CB" box="[329,386,1999,2021]" pageId="22" pageNumber="473" rank="species">
<emphasis id="B9576A726C1C286CFEFA5BB9FE1449CB" bold="true" box="[329,386,1999,2021]" pageId="22" pageNumber="473">n. sp.</emphasis>
</taxonomicNameLabel>
holotype, RMNH Coel. 40219 (SAF420).
</paragraph>
</caption>
<caption id="DF5CE6E86C1D286DFF245BCEFC2649C2" httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="23" pageNumber="474" targetBox="[169,1417,193,1955]" targetPageId="23">
<paragraph id="8B9CB6606C1D286DFF245BCEFC2649C2" blockId="23.[151,1436,1976,2028]" pageId="23" pageNumber="474">
<emphasis id="B9576A726C1D286DFF245BCEFEB649E0" bold="true" box="[151,288,1976,1998]" pageId="23" pageNumber="474">FIGURE 14.</emphasis>
<taxonomicName id="4C23CDE36C1D286DFE9B5BCEFE7549E0" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[296,483,1976,1998]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="23" pageNumber="474" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C1D286DFE9B5BCEFE7549E0" box="[296,483,1976,1998]" italics="true" pageId="23" pageNumber="474">Leptophyton fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C1D286DFE585BCEFDB349E0" box="[491,549,1976,1998]" pageId="23" pageNumber="474" rank="species">
<emphasis id="B9576A726C1D286DFE585BCEFDB349E0" bold="true" box="[491,549,1976,1998]" pageId="23" pageNumber="474">n. sp.</emphasis>
</taxonomicNameLabel>
holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk.
</paragraph>
</caption>
<subSubSection id="C339E5EB6C122862FF745CE1FA814CDE" pageId="24" pageNumber="475" type="description">
<paragraph id="8B9CB6606C122862FF745CE1FE824F82" blockId="24.[151,1437,151,1293]" pageId="24" pageNumber="475">
<emphasis id="B9576A726C122862FF745CE1FEC74E9E" bold="true" box="[199,337,151,176]" pageId="24" pageNumber="475">Description</emphasis>
. The
<typeStatus id="549808C26C122862FE2C5CE1FD974E9E" box="[415,513,151,176]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
is an erect lobate colony
<quantity id="4CDB1B856C122862FCFB5CE1FC0D4E81" box="[840,923,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="24" pageNumber="475" unit="cm" value="5.7">5.7 cm</quantity>
in height, with a poorly demarcated stalk approximately
<quantity id="4CDB1B856C122862FEF15CCAFE064EFA" box="[322,400,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="24" pageNumber="475" unit="cm" value="2.5">2.5 cm</quantity>
long and 1.0 cm in diameter (
<figureCitation id="1318AAE56C122862FD415CCAFCC24EFA" box="[754,852,188,213]" captionStart="FIGURE 13" captionStartId="22.[151,250,1937,1959]" captionTargetBox="[181,1409,695,1911]" captionTargetId="figure@22.[181,1409,695,1911]" captionTargetPageId="22" captionText="FIGURE 13. a, Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087); b, Tenerodus fallax RMNH Coel. 40215 (SAF067), attached to a polychaete tube; c, Tenerodus fallax RMNH Coel. 40217 (SAF383), growing on a mussel shell; d, Tenerodus pollex n. sp. holotype, RMNH Coel.40219 (SAF420)." httpUri="https://zenodo.org/record/1114559/files/figure.png" pageId="24" pageNumber="475">Fig. 13a</figureCitation>
). It is a fragment of a larger colony (
<figureCitation id="1318AAE56C122862FAB75CCAFACC4EFB" box="[1284,1370,188,213]" captionStart="FIGURE 5" captionStartId="10.[151,250,1819,1841]" captionTargetBox="[232,1366,195,1796]" captionTargetId="figure@10.[232,1366,195,1796]" captionTargetPageId="10" captionText="FIGURE 5. a, Lampophyton spinatum n. sp. paratype, RMNH Coel. 40199 (SAF195), colony photographed in situ; b, Alcyonium dolium n. sp. holotype, RMNH Coel.40204 (SAF292); c, Leptophyton benayahui SAF289 photographed in situ; d, Leptophyton fustis n. sp., holotype, RMNH Coel. 40211 (SAF087), colony photographed in situ (T. fallax n. comb. purple morph visible in lower right corner); e, Circularius wilsoni n. comb. RMNH Coel. 40207 (SAF448), colonies growing on mussel shell photographed in situ; f, Tenerodus pollex n. gen., n. sp. holotype, RMNH Coel. 40219 (SAF420), colonies photographed in situ; g, Tenerodus fallax, n. comb. RMNH Coel. 40217 (SAF383), pink color morph; h, Tenerodus fallax, n. comb. SAF015, purple and yellow color morphs." httpUri="https://zenodo.org/record/1114515/files/figure.png" pageId="24" pageNumber="475">Fig. 5d</figureCitation>
). The surface of the stalk is slightly rough, with numerous longitudinal creases and bumps formed by denser aggregations of sclerites. Four elongate, primary lobes,
<quantity id="4CDB1B856C122862FDDE5D72FD7C4F32" box="[621,746,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="2.7" metricValueMin="0.5" pageId="24" pageNumber="475" unit="cm" value="1.6" valueMax="2.7" valueMin="0.5">0.52.7 cm</quantity>
in length and 0.51.0 cm in diameter, arise from the stalk; the larger lobes branch to give rise to a few smaller lobes. The polyps are distributed uniformly over the surfaces of the lobes. Polyps are retractile, although in the
<typeStatus id="549808C26C122862FD395D3AFD7A4F4B" box="[650,748,332,357]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
a majority were preserved with the tentacles extended or in various stages of incomplete retraction. The colony has a flabby consistency, and the surface tissue is thin and easily torn.
</paragraph>
<paragraph id="8B9CB6606C122862FF745DC1FC544C13" blockId="24.[151,1437,151,1293]" pageId="24" pageNumber="475">
The polyps have small spindles and rods,
<quantity id="4CDB1B856C122862FD2C5DC1FCD54FE1" box="[671,835,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="24" pageNumber="475" unit="mm" value="0.115" valueMax="0.16" valueMin="0.07">0.070.16 mm</quantity>
long, in the tentacles (
<figureCitation id="1318AAE56C122862FBF75DC1FB334FFE" box="[1092,1189,439,464]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="24" pageNumber="475">Fig. 14a</figureCitation>
). Around the bases of the polyps and in the surface of the polyparium and stalk are radiates,
<quantity id="4CDB1B856C122862FC7B5DAAFBFB4FDA" box="[968,1133,476,501]" metricMagnitude="-4" metricUnit="m" metricValue="1.15" metricValueMax="1.5" metricValueMin="0.8" pageId="24" pageNumber="475" unit="mm" value="0.11499999999999999" valueMax="0.15" valueMin="0.08">0.080.15 mm</quantity>
long (
<figureCitation id="1318AAE56C122862FB0A5DAAFA8B4FDB" box="[1209,1309,476,501]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="24" pageNumber="475">Fig. 14b</figureCitation>
), and stout club-like sclerites,
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long (
<figureCitation id="1318AAE56C122862FDE85D89FD2B4C36" box="[603,701,511,536]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="24" pageNumber="475">Fig. 14c</figureCitation>
). The surface of the stalk base has predominantly small radiates,
<quantity id="4CDB1B856C122862FF245E52FEAC4C12" box="[151,314,548,573]" metricMagnitude="-5" metricUnit="m" metricValue="7" metricValueMax="9.0" metricValueMin="5.0" pageId="24" pageNumber="475" unit="mm" value="0.07" valueMax="0.09" valueMin="0.05">0.050.09 mm</quantity>
long (
<figureCitation id="1318AAE56C122862FE325E52FE754C13" box="[385,483,548,573]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="24" pageNumber="475">Fig. 14d</figureCitation>
). The interior of the colony lacks sclerites.
</paragraph>
<paragraph id="8B9CB6606C122862FF745E31FD804CE2" blockId="24.[151,1437,151,1293]" pageId="24" pageNumber="475">
All three
<typeStatus id="549808C26C122862FE865E3EFE364C4E" box="[309,416,584,608]" pageId="24" pageNumber="475" type="paratype">paratypes</typeStatus>
are smaller colonies or portions of colonies,
<quantity id="4CDB1B856C122862FC1A5E31FC614C71" box="[937,1015,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="24" pageNumber="475" unit="cm" value="2.5">2.5 cm</quantity>
tall with the stalk constituting about half of the total height. They have from 13 primary lobes, each subdivided into a few secondary lobes. In all other respects they resemble the
<typeStatus id="549808C26C122862FE755EF9FDBA4C86" box="[454,556,655,680]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
. Spherical gametes,
<quantity id="4CDB1B856C122862FCA25EF9FC224C89" box="[785,948,655,680]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.5" metricValueMin="2.5" pageId="24" pageNumber="475" unit="mm" value="0.4" valueMax="0.55" valueMin="0.25">0.250.55 mm</quantity>
in diameter, can be seen within the stalk of RMNH Coel. 40212 and SAF099.
</paragraph>
<paragraph id="8B9CB6606C122862FF745EA1FA814CDE" blockId="24.[151,1437,151,1293]" box="[199,1303,727,752]" pageId="24" pageNumber="475">
<emphasis id="B9576A726C122862FF745EA1FE9C4CDE" bold="true" box="[199,266,727,752]" pageId="24" pageNumber="475">Color</emphasis>
. In life, pale orange to salmon-pink (
<figureCitation id="1318AAE56C122862FD195EA1FD684CDE" box="[682,766,727,752]" captionStart="FIGURE 5" captionStartId="10.[151,250,1819,1841]" captionTargetBox="[232,1366,195,1796]" captionTargetId="figure@10.[232,1366,195,1796]" captionTargetPageId="10" captionText="FIGURE 5. a, Lampophyton spinatum n. sp. paratype, RMNH Coel. 40199 (SAF195), colony photographed in situ; b, Alcyonium dolium n. sp. holotype, RMNH Coel.40204 (SAF292); c, Leptophyton benayahui SAF289 photographed in situ; d, Leptophyton fustis n. sp., holotype, RMNH Coel. 40211 (SAF087), colony photographed in situ (T. fallax n. comb. purple morph visible in lower right corner); e, Circularius wilsoni n. comb. RMNH Coel. 40207 (SAF448), colonies growing on mussel shell photographed in situ; f, Tenerodus pollex n. gen., n. sp. holotype, RMNH Coel. 40219 (SAF420), colonies photographed in situ; g, Tenerodus fallax, n. comb. RMNH Coel. 40217 (SAF383), pink color morph; h, Tenerodus fallax, n. comb. SAF015, purple and yellow color morphs." httpUri="https://zenodo.org/record/1114515/files/figure.png" pageId="24" pageNumber="475">Fig. 5d</figureCitation>
). White or cream in alcohol. Sclerites colorless.
</paragraph>
</subSubSection>
<subSubSection id="C339E5EB6C122862FF745E8AFE644D16" pageId="24" pageNumber="475" type="etymology">
<paragraph id="8B9CB6606C122862FF745E8AFE644D16" blockId="24.[151,1437,151,1293]" pageId="24" pageNumber="475">
<emphasis id="B9576A726C122862FF745E8AFED04D3B" bold="true" box="[199,326,764,789]" pageId="24" pageNumber="475">Etymology</emphasis>
. From the Latin
<taxonomicName id="4C23CDE36C122862FDA95E8AFDC34D3A" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[538,597,764,788]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis">
<emphasis id="B9576A726C122862FDA95E8AFDC34D3A" box="[538,597,764,788]" italics="true" pageId="24" pageNumber="475">fustis</emphasis>
</taxonomicName>
, a club or cudgel, in reference to the stout, club-shaped sclerites that surround the base of the polyp.
</paragraph>
</subSubSection>
<subSubSection id="C339E5EB6C122862FF745F32FE334B22" pageId="24" pageNumber="475" type="discussion">
<paragraph id="8B9CB6606C122862FF745F32FE334B22" blockId="24.[151,1437,151,1293]" pageId="24" pageNumber="475">
<emphasis id="B9576A726C122862FF745F32FEA24D73" bold="true" box="[199,308,836,861]" pageId="24" pageNumber="475">Remarks</emphasis>
. Although
<taxonomicName id="4C23CDE36C122862FE0B5F33FD824D72" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[440,532,836,860]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C122862FE0B5F33FD824D72" box="[440,532,836,860]" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FDAE5F32FDC94D73" box="[541,607,836,861]" pageId="24" pageNumber="475" rank="species">
<emphasis id="B9576A726C122862FDAE5F32FDC94D73" bold="true" box="[541,607,836,861]" pageId="24" pageNumber="475">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C23CDE36C122862FD2F5F33FCA74D73" authorityName=", Ofwegen &amp; Schleyer" authorityYear="1997" box="[668,817,836,861]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="benayahui">
<emphasis id="B9576A726C122862FD2F5F33FCA74D73" box="[668,817,836,861]" italics="true" pageId="24" pageNumber="475">L. benayahui</emphasis>
</taxonomicName>
share a similar colony growth form, weakly armored polyps, and both lack sclerites in the colony interior, the form of the sclerites differs greatly between the two congeners (
<figureCitation id="1318AAE56C122862FEAB5FFAFEE34D8A" box="[280,373,908,933]" captionStart="FIGURE 12" captionStartId="21.[151,250,1775,1797]" captionTargetBox="[151,1436,587,1753]" captionTargetId="figure@21.[151,1436,587,1753]" captionTargetPageId="21" captionText="FIGURE 12. Leptophyton benayahui, RMNH Coel.40209 (SAF333). a, tentaclesclerites; b, sclerites from proximal region of polyps and colony surface; c, sclerites from base of colony." httpUri="https://zenodo.org/record/1114557/files/figure.png" pageId="24" pageNumber="475">Figs. 12</figureCitation>
,
<figureCitation id="1318AAE56C122862FE325FFAFE364D8A" box="[385,416,908,932]" captionStart="FIGURE 14" captionStartId="23.[151,250,1976,1998]" captionTargetBox="[169,1417,193,1955]" captionTargetId="figure@23.[169,1417,193,1955]" captionTargetPageId="23" captionText="FIGURE 14. Leptophyton fustis n. sp. holotype, RMNH Coel. 40211 (SAF087). a, tentacle sclerites; b, radiates from colony surface; c, club-like forms from colony surface; d, sclerites from base of stalk." httpUri="https://zenodo.org/record/1114561/files/figure.png" pageId="24" pageNumber="475">14</figureCitation>
). At present, their geographic ranges are not known to overlap, with
<taxonomicName id="4C23CDE36C122862FB175FFBFAA04D8B" authorityName=", Ofwegen &amp; Schleyer" authorityYear="1997" box="[1188,1334,908,933]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="benayahui">
<emphasis id="B9576A726C122862FB175FFBFAA04D8B" box="[1188,1334,908,933]" italics="true" pageId="24" pageNumber="475">L. benayahui</emphasis>
</taxonomicName>
found in the subtropical regions of
<collectingRegion id="49E778826C122862FE775FD9FDE24DE6" box="[452,628,943,968]" country="South Africa" name="KwaZulu-Natal" pageId="24" pageNumber="475">KwaZulu-Natal</collectingRegion>
and
<taxonomicName id="4C23CDE36C122862FD1D5FC7FC9F4DE6" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[686,777,944,968]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C122862FD1D5FC7FC9F4DE6" box="[686,777,944,968]" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FCA15FD9FCC54DE6" box="[786,851,943,968]" pageId="24" pageNumber="475" rank="species">
<emphasis id="B9576A726C122862FCA15FD9FCC54DE6" bold="true" box="[786,851,943,968]" pageId="24" pageNumber="475">n. sp.</emphasis>
</taxonomicNameLabel>
in the cold waters of the
<collectingRegion id="49E778826C122862FBCE5FC6FA824DE6" box="[1149,1300,944,968]" country="South Africa" name="Eastern Cape" pageId="24" pageNumber="475">
<collectingRegion id="49E778826C122862FBCE5FC6FB474DE9" box="[1149,1233,944,967]" country="American Samoa" name="Eastern" pageId="24" pageNumber="475">Eastern</collectingRegion>
Cape
</collectingRegion>
(Algoa and Plettenberg Bays). Superficially, expanded colonies of
<taxonomicName id="4C23CDE36C122862FCB25FA3FCCC4DC2" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[769,858,980,1004]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C122862FCB25FA3FCCC4DC2" box="[769,858,980,1004]" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FCD15FA2FC344DC3" box="[866,930,980,1005]" pageId="24" pageNumber="475" rank="species">
<emphasis id="B9576A726C122862FCD15FA2FC344DC3" bold="true" box="[866,930,980,1005]" pageId="24" pageNumber="475">n. sp.</emphasis>
</taxonomicNameLabel>
resemble the pink color morph of
<taxonomicName id="4C23CDE36C122862FA9A5FA2FF414A3E" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" baseAuthorityName="Luttschwager" baseAuthorityYear="1922" class="Anthozoa" family="Paralcyoniidae" genus="Tenerodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fallax" status="comb. nov.">
<emphasis id="B9576A726C122862FA9A5FA2FF414A3E" italics="true" pageId="24" pageNumber="475">Tenerodus fallax</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FF6E5F8EFEA84A3E" box="[221,318,1015,1040]" pageId="24" pageNumber="475">n. comb.</taxonomicNameLabel>
(compare
<figureCitation id="1318AAE56C122862FE075F81FDF64A3E" box="[436,608,1015,1040]" captionStart="FIGURE 5" captionStartId="10.[151,250,1819,1841]" captionTargetBox="[232,1366,195,1796]" captionTargetId="figure@10.[232,1366,195,1796]" captionTargetPageId="10" captionText="FIGURE 5. a, Lampophyton spinatum n. sp. paratype, RMNH Coel. 40199 (SAF195), colony photographed in situ; b, Alcyonium dolium n. sp. holotype, RMNH Coel.40204 (SAF292); c, Leptophyton benayahui SAF289 photographed in situ; d, Leptophyton fustis n. sp., holotype, RMNH Coel. 40211 (SAF087), colony photographed in situ (T. fallax n. comb. purple morph visible in lower right corner); e, Circularius wilsoni n. comb. RMNH Coel. 40207 (SAF448), colonies growing on mussel shell photographed in situ; f, Tenerodus pollex n. gen., n. sp. holotype, RMNH Coel. 40219 (SAF420), colonies photographed in situ; g, Tenerodus fallax, n. comb. RMNH Coel. 40217 (SAF383), pink color morph; h, Tenerodus fallax, n. comb. SAF015, purple and yellow color morphs." httpUri="https://zenodo.org/record/1114515/files/figure.png" pageId="24" pageNumber="475">
Figs. 5d and
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) with which it may co-occur. The form and arrangement of sclerites in the two species differ greatly, however. The heavily armored polyps and surface of
<taxonomicName id="4C23CDE36C122862FB84586BFB034A1B" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" baseAuthorityName="Luttschwager" baseAuthorityYear="1922" box="[1079,1173,1052,1077]" class="Anthozoa" family="Paralcyoniidae" genus="Tenerodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fallax">
<emphasis id="B9576A726C122862FB84586BFB034A1B" box="[1079,1173,1052,1077]" italics="true" pageId="24" pageNumber="475">T. fallax</emphasis>
</taxonomicName>
give the contracted or preserved colony a firm consistency. In contrast,
<taxonomicName id="4C23CDE36C122862FD685837FCAE4A76" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[731,824,1088,1112]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C122862FD685837FCAE4A76" box="[731,824,1088,1112]" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FCF75849FC114A76" box="[836,903,1087,1112]" pageId="24" pageNumber="475" rank="species">
<emphasis id="B9576A726C122862FCF75849FC114A76" bold="true" box="[836,903,1087,1112]" pageId="24" pageNumber="475">n. sp.</emphasis>
</taxonomicNameLabel>
lacks sclerites in the colony interior and has smaller, more sparsely arranged sclerites in the colony surface, giving the colonies a weak, flabby consistency.
<taxonomicName id="4C23CDE36C122862FA365813FF454A8E" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis">
<emphasis id="B9576A726C122862FA365813FF454A8E" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C23CDE36C122862FDB958FEFD5B4A8E" baseAuthorityName="Verseveldt &amp; Williams" baseAuthorityYear="1988" box="[522,717,1159,1184]" class="Anthozoa" family="Alcyoniidae" genus="Gersemia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="liltvedi">
<emphasis id="B9576A726C122862FDB958FEFD5B4A8E" box="[522,717,1159,1184]" italics="true" pageId="24" pageNumber="475">Gersemia liltvedi</emphasis>
</taxonomicName>
by its fully retractile polyps that are distributed over the entire surface of the branched lobes rather than being restricted to terminal catkins.
<taxonomicName id="4C23CDE36C122862FB9358DBFB204AEB" authorityName=", Ofwegen &amp; Schleyer" authorityYear="1997" box="[1056,1206,1196,1221]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="benayahui">
<emphasis id="B9576A726C122862FB9358DBFB204AEB" box="[1056,1206,1196,1221]" italics="true" pageId="24" pageNumber="475">L. benayahui</emphasis>
</taxonomicName>
and
<taxonomicName id="4C23CDE36C122862FB4058DBFAD94AEA" authority="Mcfadden &amp; Van Ofwegen, 2017" authorityName="Mcfadden &amp; Van Ofwegen" authorityYear="2017" box="[1267,1359,1196,1220]" class="Anthozoa" family="Nephtheidae" genus="Leptophyton" higherTaxonomySource="GBIF" kingdom="Animalia" order="Alcyonacea" pageId="24" pageNumber="475" phylum="Cnidaria" rank="species" species="fustis" status="sp. nov.">
<emphasis id="B9576A726C122862FB4058DBFAD94AEA" box="[1267,1359,1196,1220]" italics="true" pageId="24" pageNumber="475">L. fustis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264D7096C122862FAE958DAFA0A4AEB" box="[1370,1436,1196,1221]" pageId="24" pageNumber="475" rank="species">
<emphasis id="B9576A726C122862FAE958DAFA0A4AEB" bold="true" box="[1370,1436,1196,1221]" pageId="24" pageNumber="475">n. sp.</emphasis>
</taxonomicNameLabel>
differed at all of the loci we sequenced, with pairwise genetic distances of 2.6% at 28S rDNA, 5.46.0% at
<emphasis id="B9576A726C122862FA8E58A6FAF84AC9" box="[1341,1390,1232,1255]" italics="true" pageId="24" pageNumber="475">COI</emphasis>
and 2.12.5% at 16S rDNA.
</paragraph>
</subSubSection>
</treatment>
</document>