301 lines
37 KiB
XML
301 lines
37 KiB
XML
<document id="AD3E7F1F7C73F86CE48792ECB0B6C9FF" ID-DOI="10.11646/zootaxa.5029.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5454137" ID-ZooBank="4EB4DF61-5DA9-4021-A6D6-00142C31B5E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630655531429" checkinUser="plazi" docAuthor="Schwarzhans, Werner W. & Møller, Peter R." docDate="2021" docId="0389CB1C9A1F2958FF005C8FFF175FED" docLanguage="en" docName="zootaxa.5029.1.1.pdf" docOrigin="Zootaxa 5029 (1)" docSource="http://zoobank.org/4eb4df61-5da9-4021-a6d6-00142c31b5e5" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Porogadus turgidus Schwarzhans & Møller 2021, n. sp." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="FFB0B3649A322968FF975A12FFDD5F38" masterDocTitle="Revision of the ‘ dragon-head’ cusk eels of the genus Porogadus (Teleostei: Ophidiidae), with description of eight new species and one new genus" masterLastPageNumber="96" masterPageNumber="1" pageNumber="46" updateTime="1699281781800" updateUser="plazi">
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<mods:title id="75F3DE6A1070CC70E25E846270CF1156">Revision of the ‘ dragon-head’ cusk eels of the genus Porogadus (Teleostei: Ophidiidae), with description of eight new species and one new genus</mods:title>
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<mods:namePart id="2C90466AFDFD2627B2E186AD6958701E">Schwarzhans, Werner W.</mods:namePart>
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<mods:namePart id="20440D7C67B3A1669D5E419EE59445CD">Møller, Peter R.</mods:namePart>
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<mods:title id="CF626B73C953F6A02DE6EA00174F0A85">Zootaxa</mods:title>
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<mods:part id="0C51075DD17923F60B54A1C44000594F">
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<mods:date id="D5044A0861E2711A22B9017C32693211">2021</mods:date>
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<mods:number id="7B001997F1C9DE5E373CD6C7819496C8">2021-09-03</mods:number>
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<mods:number id="76540F7C47540308D331475464A702CA">5029</mods:number>
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<mods:identifier id="EBB1AA7FA2A4822482E77F76451B073E" type="DOI">10.11646/zootaxa.5029.1.1</mods:identifier>
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<treatment id="0389CB1C9A1F2958FF005C8FFF175FED" ID-DOI="http://doi.org/10.5281/zenodo.5495391" ID-Zenodo-Dep="5495391" LSID="urn:lsid:plazi:treatment:0389CB1C9A1F2958FF005C8FFF175FED" httpUri="http://treatment.plazi.org/id/0389CB1C9A1F2958FF005C8FFF175FED" lastPageId="48" lastPageNumber="49" pageId="45" pageNumber="46">
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<subSubSection id="C33A29819A1F2945FF005C8FFE0D598F" box="[151,464,1693,1720]" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="8B9F7A0A9A1F2945FF005C8FFE0D598F" blockId="45.[151,464,1693,1755]" box="[151,464,1693,1720]" pageId="45" pageNumber="46">
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<heading id="D0D7CD669A1F2945FF005C8FFE0D598F" bold="true" box="[151,464,1693,1720]" fontSize="11" level="2" pageId="45" pageNumber="46" reason="4">
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<emphasis id="B954A6189A1F2945FF005C8FFE0D598F" bold="true" box="[151,464,1693,1720]" pageId="45" pageNumber="46">
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<taxonomicName id="4C2001899A1F2945FF005C8FFE5B5980" authority="Schwarzhans & Møller, 2021" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[151,390,1693,1720]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="45" pageNumber="46" phylum="Chordata" rank="species" species="turgidus" status="sp. nov.">
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<emphasis id="B954A6189A1F2945FF005C8FFE5B5980" bold="true" box="[151,390,1693,1720]" italics="true" pageId="45" pageNumber="46">Porogadus turgidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2671B639A1F2945FE1A5C8FFE0D598F" box="[397,464,1693,1719]" pageId="45" pageNumber="46" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C33A29819A1F2945FF005CD3FE6859E3" box="[151,437,1729,1755]" pageId="45" pageNumber="46" type="description">
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<paragraph id="8B9F7A0A9A1F2945FF005CD3FE6859E3" blockId="45.[151,464,1693,1755]" box="[151,437,1729,1755]" pageId="45" pageNumber="46">
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<figureCitation id="131B668F9A1F2945FF005CD3FF2959E3" box="[151,244,1729,1755]" captionStart="FIGURE 20" captionStartId="47.[151,250,1939,1963]" captionTargetBox="[151,1436,311,1904]" captionTargetId="figure-64@47.[151,1436,309,1916]" captionTargetPageId="47" captionText="FIGURE 20. Porogadus turgidus n. sp.; 20A–F, BMNH 1995.11.22.6, holotype, 295 mm SL, 20°55’N, 31°11’W, 4500–4610 m, (A–B photographs, C–D head drawings, E otolith inner face, F otolith ventral view); 20G–H, BMNH 1995.11.22.5, paratype, 282 mm SL, 20°55’N, 31°11’W, 4500–4610 m, otolith (G inner face, H ventral view)." figureDoi="http://doi.org/10.5281/zenodo.5454177" httpUri="https://zenodo.org/record/5454177/files/figure.png" pageId="45" pageNumber="46">Figs. 20</figureCitation>
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,
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<figureCitation id="131B668F9A1F2945FF685CD3FEC359E3" box="[255,286,1729,1755]" captionStart="FIGURE 44" captionStartId="82.[151,250,1887,1911]" captionTargetBox="[151,1436,1183,1857]" captionTargetId="figure-169@82.[151,1436,1178,1863]" captionTargetPageId="82" captionText="FIGURE 44. Geographic distribution of the species of the Porogadus miles group." figureDoi="http://doi.org/10.5281/zenodo.5454225" httpUri="https://zenodo.org/record/5454225/files/figure.png" pageId="45" pageNumber="46">44</figureCitation>
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,
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<figureCitation id="131B668F9A1F2945FEBE5CD3FE9559E3" box="[297,328,1729,1755]" captionStart="FIGURE 49" captionStartId="88.[151,250,1943,1967]" captionTargetBox="[178,1409,194,1915]" captionTargetId="figure-26@88.[175,1412,181,1919]" captionTargetPageId="88" captionText="FIGURE 49. Depth versus distance from shelf plots for specimens of the Porogadus miles group (upper panel) and the Porogadus gracilis group (lower panel)." figureDoi="http://doi.org/10.5281/zenodo.5454235" httpUri="https://zenodo.org/record/5454235/files/figure.png" pageId="45" pageNumber="46">49</figureCitation>
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, Tab. 1–7
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A29819A1F2945FF005D1BFD9658A8" pageId="45" pageNumber="46" type="materials_examined">
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||
<paragraph id="8B9F7A0A9A1F2945FF005D1BFD9658A8" blockId="45.[151,1437,1801,2044]" pageId="45" pageNumber="46">
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||
<emphasis id="B954A6189A1F2945FF005D1BFEA1581B" bold="true" box="[151,380,1801,1827]" pageId="45" pageNumber="46">Material examined</emphasis>
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(
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<materialsCitation id="3B4870579A1F2945FE195D1BFD9658A8" collectingDate="1993-10-16" collectionCode="R" collectorName="S. Discovery Cruise & N. Merrett & N. Merrett." elevation="4555" latitude="20.916666" longLatPrecision="1266" longitude="-31.183332" pageId="45" pageNumber="46" specimenCode="BMNH 1995.11" specimenCount="2" typeStatus="holotype">
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<specimenCount id="9D26B1839A1F2945FE195D1BFDFC581C" box="[398,545,1801,1828]" pageId="45" pageNumber="46" type="generic">2 specimens</specimenCount>
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||
):
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||
<typeStatus id="549BC4A89A1F2945FDA25D1BFD40581B" box="[565,669,1801,1827]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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<specimenCode id="DB86D2719A1F2945FD305D18FCBC581C" box="[679,865,1801,1828]" collectionCode="BMNH" pageId="45" pageNumber="46">BMNH 1995.11</specimenCode>
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.22.6,
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<quantity id="4CD8D7EF9A1F2945FC305D1BFBDB581C" box="[935,1030,1801,1828]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" pageId="45" pageNumber="46" unit="mm" value="295.0">295 mm</quantity>
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SL,
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<geoCoordinate id="EE141CCD9A1F2945FBD75D1BFB7C581C" box="[1088,1185,1801,1828]" degrees="20" direction="north" minutes="55" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="20.916666">20°55’N</geoCoordinate>
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||
<geoCoordinate id="EE141CCD9A1F2945FB3D5D1BFACD581B" box="[1194,1296,1801,1827]" degrees="31" direction="west" minutes="11" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-31.183332">31°11’W</geoCoordinate>
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||
,
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||
<quantity id="4CD8D7EF9A1F2945FA895D1BFF725870" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.61" metricValueMin="4.5" pageId="45" pageNumber="46" unit="m" value="4555.0" valueMax="4610.0" valueMin="4500.0">
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||
<elevation id="000D9D399A1F2945FA895D1BFF725870" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.61" metricValueMin="4.5" pageId="45" pageNumber="46" unit="m" value="4555.0" valueMax="4610.0" valueMin="4500.0">4500–4610 m</elevation>
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</quantity>
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,
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<collectionCode id="ED31E2CF9A1F2945FF2B5D3FFF0E587F" box="[188,211,1837,1863]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="45" pageNumber="46">R</collectionCode>
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||
.
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||
<collectionCode id="ED31E2CF9A1F2945FF405D3FFF33587F" box="[215,238,1837,1863]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="45" pageNumber="46">R</collectionCode>
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||
.
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<collectorName id="26D51FDC9A1F2945FF655D3FFE0F587F" box="[242,466,1837,1863]" pageId="45" pageNumber="46">S. Discovery Cruise</collectorName>
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semi-balloon otter trawl,
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<date id="FF9E5CCA9A1F2945FD6E5D3FFC66587F" box="[761,955,1837,1863]" pageId="45" pageNumber="46" value="1993-10-16">
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||
<collectingDate id="EFDAA5229A1F2945FD6E5D3FFC66587F" box="[761,955,1837,1863]" pageId="45" pageNumber="46" value="1993-10-16">16 October 1993</collectingDate>
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||
</date>
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||
, collected by
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<collectorName id="26D51FDC9A1F2945FBCF5D3CFB095870" box="[1112,1236,1838,1864]" pageId="45" pageNumber="46">N. Merrett</collectorName>
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;
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<typeStatus id="549BC4A89A1F2945FB775D3CFA9D5870" box="[1248,1344,1838,1864]" pageId="45" pageNumber="46" type="paratype">paratype</typeStatus>
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<specimenCode id="DB86D2719A1F2945FADF5D3CFF285854" collectionCode="BMNH" pageId="45" pageNumber="46">BMNH 1995.11</specimenCode>
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.22.5,
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<quantity id="4CD8D7EF9A1F2945FEA05D43FE495854" box="[311,404,1873,1900]" metricMagnitude="-1" metricUnit="m" metricValue="2.82" pageId="45" pageNumber="46" unit="mm" value="282.0">282 mm</quantity>
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SL,
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<geoCoordinate id="EE141CCD9A1F2945FE5E5D43FDF75854" box="[457,554,1873,1900]" degrees="20" direction="north" minutes="55" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="20.916666">20°55’N</geoCoordinate>
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<geoCoordinate id="EE141CCD9A1F2945FDA65D43FD4A5853" box="[561,663,1873,1899]" degrees="31" direction="west" minutes="11" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="-31.183332">31°11’W</geoCoordinate>
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,
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||
<quantity id="4CD8D7EF9A1F2945FD355D43FCE25854" box="[674,831,1873,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.61" metricValueMin="4.5" pageId="45" pageNumber="46" unit="m" value="4555.0" valueMax="4610.0" valueMin="4500.0">
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||
<elevation id="000D9D399A1F2945FD355D43FCE25854" box="[674,831,1873,1900]" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.61" metricValueMin="4.5" pageId="45" pageNumber="46" unit="m" value="4555.0" valueMax="4610.0" valueMin="4500.0">4500–4610 m</elevation>
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</quantity>
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,
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<collectionCode id="ED31E2CF9A1F2945FCDC5D43FCBF5853" box="[843,866,1873,1899]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="45" pageNumber="46">R</collectionCode>
|
||
.
|
||
<collectionCode id="ED31E2CF9A1F2945FCF15D43FCA05853" box="[870,893,1873,1899]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="45" pageNumber="46">R</collectionCode>
|
||
.
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||
<collectorName id="26D51FDC9A1F2945FC175D43FB835853" box="[896,1118,1873,1899]" pageId="45" pageNumber="46">S. Discovery Cruise</collectorName>
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semi-balloon otter trawl,
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||
<date id="FF9E5CCA9A1F2945FA175D43FEEE58B7" pageId="45" pageNumber="46" value="1993-10-16">
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<collectingDate id="EFDAA5229A1F2945FA175D43FEEE58B7" pageId="45" pageNumber="46" value="1993-10-16">16 October 1993</collectingDate>
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||
</date>
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||
, collected by
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<collectorName id="26D51FDC9A1F2945FE5A5D64FD9658A8" box="[461,587,1910,1936]" pageId="45" pageNumber="46">N. Merrett.</collectorName>
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="C33A29819A1F2946FF505D8BFD305E25" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="diagnosis">
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<paragraph id="8B9F7A0A9A1F2946FF505D8BFD305E25" blockId="45.[151,1437,1801,2044]" lastBlockId="46.[151,1437,151,2013]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
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<emphasis id="B954A6189A1F2945FF505D8BFE9F588B" bold="true" box="[199,322,1945,1971]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
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Precaudal vertebrae 17–18; long gill rakers on first gill arch 11–15; HL:HD 1.82–1.85; all head spines strong, present on ethmoidal, lacrimal, prefrontal, interorbital, supraorbital, sphenotic, 5th infraorbital, supratemporal, inner and outer posttemporal, inner preopercular rim, absent along outer preopercular rim; opercular spine sharp, strong, extruding; additional (3rd) postorbital pore, pores on top of head in front of nape; large opercular flap without ridges; lower lateral line pores until beginning of anal fin 18–21; vomer with broad dentition patch (4–7 rows of teeth); palatines with broad dentition patch (6 rows of teeth); otolith with clearly separated ostial and caudal colliculi; OL:OH = 1.6; OL:TCL = 2.45–2.85.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A29819A1C2946FF505B34FCE45915" pageId="46" pageNumber="47" type="description">
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<paragraph id="8B9F7A0A9A1C2946FF505B34FD365EE9" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
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<emphasis id="B954A6189A1C2946FF505B34FE845E78" bold="true" box="[199,345,294,320]" pageId="46" pageNumber="47">Description.</emphasis>
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Meristics: precaudal vertebrae 17 (17–18), 1 (1–2) last vertebrae without ribs; pectoral-fin rays 16 (16–18); D/V = 4 (4–5); D/A = 26 (24–26); V/A = 18 (18). Gill rakers in
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<typeStatus id="549BC4A89A1C2946FC5E5B59FBF65E5D" box="[969,1067,331,357]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
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on lower first gill arch with nine plate-shaped rakers, followed by a series of 11 long rakers. The lower four of those intercept by single plate shaped rakers. Upper gill raker with a series of three plate shaped and three slightly longer rakers intercepted, followed by two plate shaped rakers; total long rakers 11 (11–15).
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</paragraph>
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<paragraph id="8B9F7A0A9A1C2946FF505BC9FD215DBD" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">Morphometrics: in % of SL: HL 16.0 (15.3–16.0); maximal HD 9.0 (8.0–9.0); HD through center of eye 6.5 (5.2–6.5); bony interorbital width 2.6 (2.6–3.0); snout length 5.4 (5.0–5.4); upper jaw length 9.3 (9.2–9.3); predorsal 15.7 (15.7); preanal 30.5 (29.6–30.5); prepelvic 13.8 (13.0–13.8); prepectoral 16.6 (16.1–16.6); pectoral length 9.5 (9.5–9.7). Relations: HL:HD = 1.82 (1.82–1.85); HL to snout length 2.95 (2.95–3.07); preanal to predorsal 1.95 (1.88–1.95); predorsal to prepectoral 0.94 (0.94–0.97).</paragraph>
|
||
<paragraph id="8B9F7A0A9A1C2946FF50589DFEB35B61" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
|
||
Slender fish with long tapering tail and long, pointed snout. Maximal size of fishes investigated
|
||
<quantity id="4CD8D7EF9A1C2946FA99589CFAB05D91" box="[1294,1389,654,681]" metricMagnitude="-1" metricUnit="m" metricValue="2.95" pageId="46" pageNumber="47" unit="mm" value="295.0">295 mm</quantity>
|
||
SL. Head long and slender, with flat dorsal profile, with strong spines as follows: ethmoidal (1), lacrimal (2), prefrontal (1), interorbital (3), supraorbital (1), sphenotic (2–3), 5th infraorbital (1), supratemporal (1), inner and outer posttemporal (3–5), inner preopercular rim (3–4), no spines along outer preopercular rim. Opercle with sharp, strong, extruding spine. Eye small located in strongly asymmetric orbit. Maxilla extending far beyond eye, strongly widened posteriorly and with distinct supramaxilla. Infra-/postorbital and mandibular-preopercular pores wide, postorbital with 3 pores, the 3rd uppermost one being small, head top with 3 pores on occiput in front of nape and cavernous system extending between squamation of occiput and sphenotic and supraorbital spines. Head squamation on opercle, cheeks, occiput, frontal, around eyes, and possibly on rear of maxilla (indications of scale-pockets). Opercle with three distinct large neuromasts behind preopercular edge; opercular flap large, without radial ridges. All three lateral line rows reasonably well visible. Upper lateral line row with 8–9 pores; lower lateral line row with 18–21 pores until beginning of anal fin; central lateral line row long, lower and central lateral line rows fading behind beginning of anal fin.
|
||
</paragraph>
|
||
<paragraph id="8B9F7A0A9A1C2946FF505E71FAD65BD1" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">Dentition. All teeth tiny and cone-shaped. Vomer with a broad dentition patch with 4–7 rows of teeth anteriorly; palatines with broad dentition patch with 6 rows of teeth throughout. Premaxilla tooth patches not fused anteriorly; ca. 7 teeth rows anteriorly and 3–4 rows posteriorly. Dentary tooth patches fused anteriorly; ca. 6 teeth rows anteriorly and 2–3 rows posteriorly. Median basibranchial tooth patch short, sometimes anteriorly widened.</paragraph>
|
||
<paragraph id="8B9F7A0A9A1C2946FF505EE1FC085ADD" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">
|
||
Otolith morphology (n = 2). Size up to
|
||
<quantity id="4CD8D7EF9A1C2946FD165EE1FD055A35" box="[641,728,1267,1293]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="46" pageNumber="47" unit="mm" value="4.9">4.9 mm</quantity>
|
||
in length (
|
||
<typeStatus id="549BC4A89A1C2946FCC65EE1FC645A36" box="[849,953,1267,1294]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
); OL:OH = 1.6; OH:OT = 2.2–2.6. Moderately thick and moderately elongate otolith, anteriorly broadly rounded, posteriorly slightly expanded and somewhat tapering. Dorsal rim with marked, broadly rounded predorsal lobe, ventral rim regularly curved. All rims smooth. Inner face slightly convex, smooth, with very short, centrally positioned sulcus; OL:TCL = 2.45–2.85. Sulcus with shallow, clearly separated ostial and caudal colliculi; ostial colliculum nearly twice as long and wide as caudal colliculum; OCL:CCL = 1.85–1.90. Dorsal field with indistinct depression; ventral field smooth, without ventral furrow. Outer face smooth, with mild umbo opposite to ostium on inner face.
|
||
</paragraph>
|
||
<paragraph id="8B9F7A0A9A1C2946FF505FFDFCE45915" blockId="46.[151,1437,151,2013]" pageId="46" pageNumber="47">Coloration. Live coloration not known. Color of preserved specimens medium brown; belly and frontal part of head top slightly darker than body, opercle distinctly darker.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A29819A1C2947FF505C24FED75F89" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="discussion">
|
||
<paragraph id="8B9F7A0A9A1C2947FF505C24FED75F89" blockId="46.[151,1437,151,2013]" lastBlockId="47.[151,1436,150,285]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
|
||
<emphasis id="B954A6189A1C2946FF505C24FE915968" bold="true" box="[199,332,1590,1616]" pageId="46" pageNumber="47">Discussion.</emphasis>
|
||
<taxonomicName id="4C2001899A1C2946FEC15C25FDE85969" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[342,565,1591,1617]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="turgidus">
|
||
<emphasis id="B954A6189A1C2946FEC15C25FDE85969" box="[342,565,1591,1617]" italics="true" pageId="46" pageNumber="47">Porogadus turgidus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C2001899A1C2946FD475C25FCF45969" authorityName=", MNHN" authorityYear="1995" box="[720,809,1591,1617]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FD475C25FCF45969" box="[720,809,1591,1617]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
and the co-occurring
|
||
<taxonomicName id="4C2001899A1C2946FBBF5C25FB385969" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[1064,1253,1591,1617]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1C2946FBBF5C25FB385969" box="[1064,1253,1591,1617]" italics="true" pageId="46" pageNumber="47">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2001899A1C2946FAB75C25FA485969" box="[1312,1429,1591,1617]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FAB75C25FA485969" box="[1312,1429,1591,1617]" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
. External differences are subtle and depending on good preservation (see also discussion to
|
||
<taxonomicName id="4C2001899A1C2946FB275C49FAD4594D" authorityName=", MNHN" authorityYear="1995" box="[1200,1289,1627,1653]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FB275C49FAD4594D" box="[1200,1289,1627,1653]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C2001899A1C2946FAB55C49FF2E59A1" authorityName="Schwarzhans & Møller" authorityYear="2021" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="turgidus">
|
||
<emphasis id="B954A6189A1C2946FAB55C49FF2E59A1" italics="true" pageId="46" pageNumber="47">Porogadus turgidus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C2001899A1C2946FE1C5C6DFE3F59A1" authorityName=", MNHN" authorityYear="1995" box="[395,482,1663,1689]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FE1C5C6DFE3F59A1" box="[395,482,1663,1689]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2001899A1C2946FD8C5C6DFD5359A1" box="[539,654,1663,1689]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FD8C5C6DFD5359A1" box="[539,654,1663,1689]" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
in the lesser pores of the lower lateral line until the beginning of the anal fin of 18–21 (vs 22–27). Its predorsal length is the shortest in this group (<16 % of SL vs 17–20 %). From
|
||
<taxonomicName id="4C2001899A1C2946FA1F5CB1FEEB59D9" authorityName="Schwarzhans & Møller" authorityYear="2021" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1C2946FA1F5CB1FEEB59D9" italics="true" pageId="46" pageNumber="47">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
it differs in the broader vomer (4–7 rows of teeth vs 1–3, rarely 4). It shares with
|
||
<taxonomicName id="4C2001899A1C2946FB2D5CD5FAA959D9" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[1210,1396,1735,1761]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1C2946FB2D5CD5FAA959D9" box="[1210,1396,1735,1761]" italics="true" pageId="46" pageNumber="47">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
the lack of spines along the outer preopercular rim (vs present in
|
||
<taxonomicName id="4C2001899A1C2946FCDD5CF9FC60583D" box="[842,957,1771,1797]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FCDD5CF9FC60583D" box="[842,957,1771,1797]" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2001899A1C2946FC635CF9FB97583D" authorityName=", MNHN" authorityYear="1995" box="[1012,1098,1771,1797]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FC635CF9FB97583D" box="[1012,1098,1771,1797]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
). It may further differ from
|
||
<taxonomicName id="4C2001899A1C2946FA1F5CF9FF335811" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FA1F5CF9FF335811" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
in the presence of scales on the rear part of the maxilla (scale-pockets; vs no scales on the maxilla). Another useful character to distinguish these four species is the otolith, when preserved. The otoliths of
|
||
<taxonomicName id="4C2001899A1C2946FB3A5D21FAF95875" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[1197,1316,1843,1869]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="turgidus">
|
||
<emphasis id="B954A6189A1C2946FB3A5D21FAF95875" box="[1197,1316,1843,1869]" italics="true" pageId="46" pageNumber="47">P. turgidus</emphasis>
|
||
</taxonomicName>
|
||
share with those of
|
||
<taxonomicName id="4C2001899A1C2946FF615D45FE6D5849" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[246,432,1879,1905]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1C2946FF615D45FE6D5849" box="[246,432,1879,1905]" italics="true" pageId="46" pageNumber="47">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
the compressed shape (OL:OH = 1.05–1.6 vs 1.8–2.05 for
|
||
<taxonomicName id="4C2001899A1C2946FBD55D45FB695849" box="[1090,1204,1879,1905]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FBD55D45FB695849" box="[1090,1204,1879,1905]" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
). Also, most otoliths of
|
||
<taxonomicName id="4C2001899A1C2946FF225D69FED658AD" authorityName=", MNHN" authorityYear="1995" box="[181,267,1915,1941]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FF225D69FED658AD" box="[181,267,1915,1941]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
are more elongate (OL:OH = 1.7–2.0, rarely 1.55–1.65, see above). With the otoliths of
|
||
<taxonomicName id="4C2001899A1C2946FB735D69FAE758AD" authorityName=", MNHN" authorityYear="1995" box="[1252,1338,1915,1941]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FB735D69FAE758AD" box="[1252,1338,1915,1941]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
it shares the clearly separated colliculi (vs a single, joined colliculum in
|
||
<taxonomicName id="4C2001899A1C2946FCC85D8DFBC75881" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[863,1050,1951,1977]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1C2946FCC85D8DFBC75881" box="[863,1050,1951,1977]" italics="true" pageId="46" pageNumber="47">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2001899A1C2946FBC65D8DFB195881" box="[1105,1220,1951,1977]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1C2946FBC65D8DFB195881" box="[1105,1220,1951,1977]" italics="true" pageId="46" pageNumber="47">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
). The otoliths of
|
||
<taxonomicName id="4C2001899A1C2946FA1F5D8DFF2E58E5" authorityName="Schwarzhans & Møller" authorityYear="2021" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="turgidus">
|
||
<emphasis id="B954A6189A1C2946FA1F5D8DFF2E58E5" italics="true" pageId="46" pageNumber="47">P. turgidus</emphasis>
|
||
</taxonomicName>
|
||
differ from those of
|
||
<taxonomicName id="4C2001899A1C2946FE755DD1FDE758E5" authorityName=", MNHN" authorityYear="1995" box="[482,570,1987,2013]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="miles">
|
||
<emphasis id="B954A6189A1C2946FE755DD1FDE758E5" box="[482,570,1987,2013]" italics="true" pageId="46" pageNumber="47">P. miles</emphasis>
|
||
</taxonomicName>
|
||
in the very short sulcus as expressed in the ration OL:TCL of 2.45–2.85 (vs 1.7–2.05).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A29819A1D2947FF505AA8FE6E5E25" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="8B9F7A0A9A1D2947FF505AA8FE6E5E25" blockId="47.[151,1436,150,285]" pageId="47" pageNumber="48">
|
||
<emphasis id="B954A6189A1D2947FF505AA8FEBC5FEC" bold="true" box="[199,353,186,212]" pageId="47" pageNumber="48">Distribution.</emphasis>
|
||
<taxonomicName id="4C2001899A1D2947FEFF5AA9FD9B5FED" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[360,582,187,213]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="turgidus">
|
||
<emphasis id="B954A6189A1D2947FEFF5AA9FD9B5FED" box="[360,582,187,213]" italics="true" pageId="47" pageNumber="48">Porogadus turgidus</emphasis>
|
||
</taxonomicName>
|
||
is one of three deepwater species of the genus occurring in the
|
||
<collectingCountry id="F3373A9A9A1D2947FA8D5AA9FA415FED" box="[1306,1436,187,213]" name="Cape Verde" pageId="47" pageNumber="48">Cape Verde</collectingCountry>
|
||
Basin of the northeastern Atlantic (the other two being
|
||
<taxonomicName id="4C2001899A1D2947FD615ACDFC725FC1" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[758,943,223,249]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="caboverdensis">
|
||
<emphasis id="B954A6189A1D2947FD615ACDFC725FC1" box="[758,943,223,249]" italics="true" pageId="47" pageNumber="48">P. caboverdensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2001899A1D2947FC735ACDFB885FC1" box="[996,1109,223,249]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="mendax">
|
||
<emphasis id="B954A6189A1D2947FC735ACDFB885FC1" box="[996,1109,223,249]" italics="true" pageId="47" pageNumber="48">P. mendax</emphasis>
|
||
</taxonomicName>
|
||
). It has been caught at depths between
|
||
<quantity id="4CD8D7EF9A1D2947FF6C5B10FE725E25" box="[251,431,258,285]" metricMagnitude="3" metricUnit="m" metricValue="4.555" metricValueMax="4.61" metricValueMin="4.5" pageId="47" pageNumber="48" unit="m" value="4555.0" valueMax="4610.0" valueMin="4500.0">4500 to 4610 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F2A829A1D2947FF005D81FBDD58CB" ID-DOI="http://doi.org/10.5281/zenodo.5454177" ID-Zenodo-Dep="5454177" httpUri="https://zenodo.org/record/5454177/files/figure.png" pageId="47" pageNumber="48" startId="47.[151,250,1939,1963]" targetBox="[151,1436,311,1904]" targetPageId="47">
|
||
<paragraph id="8B9F7A0A9A1D2947FF005D81FBDD58CB" blockId="47.[151,1437,1939,2035]" pageId="47" pageNumber="48">
|
||
<emphasis id="B954A6189A1D2947FF005D81FEFD5893" bold="true" box="[151,288,1939,1963]" pageId="47" pageNumber="48">FIGURE 20.</emphasis>
|
||
<taxonomicName id="4C2001899A1D2947FEB05D81FE325893" authority="Schwarzhans & Møller, 2021" authorityName="Schwarzhans & Møller" authorityYear="2021" box="[295,495,1939,1963]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="47" pageNumber="48" phylum="Chordata" rank="species" species="turgidus" status="sp. nov.">
|
||
<emphasis id="B954A6189A1D2947FEB05D81FE325893" box="[295,495,1939,1963]" italics="true" pageId="47" pageNumber="48">Porogadus turgidus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954A6189A1D2947FE625D81FDF25893" bold="true" box="[501,559,1939,1963]" pageId="47" pageNumber="48">
|
||
<taxonomicNameLabel id="A2671B639A1D2947FE625D81FDF25893" box="[501,559,1939,1963]" pageId="47" pageNumber="48" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
;
|
||
<emphasis id="B954A6189A1D2947FDAB5D81FD595893" bold="true" box="[572,644,1939,1963]" pageId="47" pageNumber="48">20A–F</emphasis>
|
||
, BMNH 1995.11.22.6, holotype, 295 mm SL, 20°55’N, 31°11’W, 4500–4610 m, (A–B photographs, C–D head drawings, E otolith inner face, F otolith ventral view);
|
||
<emphasis id="B954A6189A1D2947FB925DA5FB8F58F7" bold="true" box="[1029,1106,1975,1999]" pageId="47" pageNumber="48">20G–H</emphasis>
|
||
, BMNH 1995.11.22.5, paratype, 282 mm SL, 20°55’N, 31°11’W, 4500–4610 m, otolith (G inner face, H ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A29819A022958FF505A85FF175FED" pageId="48" pageNumber="49" type="etymology">
|
||
<paragraph id="8B9F7A0A9A022958FF505A85FF175FED" blockId="48.[151,1436,151,213]" pageId="48" pageNumber="49">
|
||
<emphasis id="B954A6189A022958FF505A85FE935F89" bold="true" box="[199,334,151,177]" pageId="48" pageNumber="49">Etymology.</emphasis>
|
||
From
|
||
<taxonomicName id="4C2001899A022958FE0A5A85FD8D5F8A" authority="(Latin)" baseAuthorityName="Latin" box="[413,592,151,178]" class="Actinopterygii" family="Ophidiidae" genus="Porogadus" kingdom="Animalia" order="Ophidiiformes" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="turgidus">turgidus (Latin)</taxonomicName>
|
||
= inflated, referring to the large otolith size compared to its small sulcus size.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |