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<document id="1C690F9ACF84B851F7D85993E7AABAC2" ID-CLB-Dataset="276483" ID-DOI="10.1111/j.1096-3642.2009.00552.x" ID-GBIF-Dataset="3be82545-1797-46a0-95fc-3e0fc44c9af0" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440497297" checkinUser="valdenar" docAuthor="Zhang, Kuiyan, Yang, Ding, Ren, Dong &amp; Shih, Chungkun" docDate="2010" docId="038296595837FF8BFCEA919EFABE51F6" docLanguage="en" docName="j.1096-3642.2009.00552.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Strenorhagio asymmetricus Zhang &amp; Yang &amp; Ren &amp; Shih 2010" docType="treatment" docVersion="6" lastPageNumber="569" masterDocId="FFBBEE215832FF8DFFBB9529FFCA5766" masterDocTitle="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)" masterLastPageNumber="572" masterPageNumber="563" pageNumber="568" updateTime="1699284570084" updateUser="ExternalLinkService">
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<mods:title id="3E1651030BF9D5C153485E1A78AF16AF">An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)</mods:title>
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<mods:name id="79E94F27701DFDF5B8C2B393A95B45BE" type="personal">
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<mods:namePart id="9F70E02B11E76B72BAC41F74E88CEF9B">Zhang, Kuiyan</mods:namePart>
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<mods:namePart id="E87E64F91EAF310201D2E5FA869E3591">Yang, Ding</mods:namePart>
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<mods:namePart id="9A9FE78D524C047F2033F726FD49F3C1">Ren, Dong</mods:namePart>
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<mods:namePart id="6A6B8C1787F671BEC10A3C00EA025A06">Shih, Chungkun</mods:namePart>
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<mods:title id="61F674F1F73673090741EAD93D829E2B">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="5F7F16098038A8E20901691E4022651B">2010</mods:date>
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<mods:number id="4C11B9BBC5F5CEC72AE996059CF94A3B">2010-02-26</mods:number>
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<mods:identifier id="FE7246B870885113DFA1C2756379C10A" type="CLB-Dataset">276483</mods:identifier>
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<treatment id="038296595837FF8BFCEA919EFABE51F6" ID-DOI="http://doi.org/10.5281/zenodo.5491006" ID-GBIF-Taxon="217238220" ID-Zenodo-Dep="5491006" LSID="urn:lsid:plazi:treatment:038296595837FF8BFCEA919EFABE51F6" httpUri="http://treatment.plazi.org/id/038296595837FF8BFCEA919EFABE51F6" lastPageId="6" lastPageNumber="569" pageId="5" pageNumber="568">
<subSubSection id="C33174C45837FF88FCEA919EFAFA538B" pageId="5" pageNumber="568" type="nomenclature">
<paragraph id="8B94274F5837FF88FCEA919EFAFA538B" blockId="5.[849,1418,1207,1261]" pageId="5" pageNumber="568">
<heading id="D0DC90235837FF88FCEA919EFAFA538B" centered="true" fontSize="9" level="2" pageId="5" pageNumber="568" reason="4">
<taxonomicName id="4C2B5CCC5837FF88FCEA919EFB0353AA" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[849,1225,1207,1228]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="569" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5837FF88FCEA919EFB0353AA" bold="true" box="[849,1225,1207,1228]" italics="true" pageId="5" pageNumber="568">STRENORHAGIO ASYMMETRICUS</emphasis>
</taxonomicName>
ZHANG, YANG &amp; REN
<emphasis id="B95FFB5D5837FF88FC5991F3FB7A538A" bold="true" box="[994,1200,1239,1260]" pageId="5" pageNumber="568">GEN. ET SP. NOV.</emphasis>
(
<figureCitation id="13103BCA5837FF88FB7991F2FAC1538B" box="[1218,1291,1239,1261]" captionStart="Figure 5" captionStartId="6.[144,223,781,800]" captionTargetBox="[143,1424,198,751]" captionTargetId="figure-416@6.[723,1424,198,751]" captionTargetPageId="6" captionText="Figure 5. Strenorhagio asymmetricus sp. nov. Holotype, body with wings in dorsal view. A, photograph. B, camera lucida drawing." figureDoi="http://doi.org/10.5281/zenodo.5438132" httpUri="https://zenodo.org/record/5438132/files/figure.png" pageId="5" pageNumber="568">FIGS 5</figureCitation>
,
<figureCitation id="13103BCA5837FF88FAA191FEFAE2538B" box="[1306,1320,1239,1261]" captionStart="Figure 6" captionStartId="6.[144,223,1280,1299]" captionTargetBox="[144,763,877,1251]" captionTargetId="figure-438@6.[144,763,877,1251]" captionTargetPageId="6" captionText="Figure 6. Strenorhagio asymmetricus sp. nov. Paratype, photograph of body with a wing in lateral view." figureDoi="http://doi.org/10.5281/zenodo.5438134" httpUri="https://zenodo.org/record/5438134/files/figure.png" pageId="5" pageNumber="568">6</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33174C45837FF8BFC8191D4FABE51F6" lastPageId="6" lastPageNumber="569" pageId="5" pageNumber="568" type="etymology">
<paragraph id="8B94274F5837FF88FC8191D4FAE75237" blockId="5.[826,1441,1277,1361]" pageId="5" pageNumber="568">
<emphasis id="B95FFB5D5837FF88FC8191D4FC725274" box="[826,952,1277,1298]" italics="true" pageId="5" pageNumber="568">Etymology:</emphasis>
The specific name refers to the Greek ‘
<taxonomicName id="4C2B5CCC5837FF88FC849034FC165254" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[831,988,1309,1330]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="568" phylum="Arthropoda" rank="species" species="asymmetricus">
<emphasis id="B95FFB5D5837FF88FC849034FC165254" box="[831,988,1309,1330]" italics="true" pageId="5" pageNumber="568">asymmetricus</emphasis>
</taxonomicName>
’ (asymmetric), because of the asymmetric position of crossvein r–m in two wings.
</paragraph>
<paragraph id="8B94274F5837FF88FC8190A3FA9A51F2" blockId="5.[826,1441,1417,1684]" pageId="5" pageNumber="568">
<emphasis id="B95FFB5D5837FF88FC8190A3FC7B52F9" box="[826,945,1418,1439]" italics="true" pageId="5" pageNumber="568">Diagnosis:</emphasis>
Hind femur swollen at apical part; each tibia with a spur at apex. Crossvein r–m slightly proximal to beginning of vein R
<subScript id="17AF250A5837FF88FB0D90F8FB0452BB" attach="left" box="[1206,1230,1489,1501]" fontSize="5" pageId="5" pageNumber="568">
2+
<quantity id="4CD38AAA5837FF88FB7D90F8FB0452BB" box="[1222,1230,1489,1501]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="5" pageNumber="568" unit="in" value="3.0">3</quantity>
</subScript>
in left wing, and nearly coalesced with vein R
<subScript id="17AF250A5837FF88FB3490C6FB6D529D" attach="left" box="[1167,1191,1519,1531]" fontSize="5" pageId="5" pageNumber="568">2+3</subScript>
at vein Rs (vein Rs
<subScript id="17AF250A5837FF88FA2290C6FA6B529D" attach="left" box="[1433,1441,1519,1531]" fontSize="5" pageId="5" pageNumber="568">2</subScript>
nearly disappearing) in right wing; vein CuA
<subScript id="17AF250A5837FF88FAFA9327FA83517C" attach="left" box="[1345,1353,1550,1562]" fontSize="5" pageId="5" pageNumber="568">1</subScript>
arising from cell bm, crossvein m–cu long; basal trunk of veins M
<subScript id="17AF250A5837FF88FC2F9362FC565131" attach="left" box="[916,924,1611,1623]" fontSize="5" pageId="5" pageNumber="568">1</subScript>
and M
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shorter than crossvein m–m; mouths of cells sc and r
<subScript id="17AF250A5837FF88FBBA9343FBC35110" attach="left" box="[1025,1033,1642,1654]" fontSize="5" pageId="5" pageNumber="568">1</subScript>
wide, relative widths about 4: 3. Segments I–VI with black stripe at each end.
</paragraph>
<paragraph id="8B94274F5837FF88FC8193E4FB395067" blockId="5.[826,1441,1741,1793]" pageId="5" pageNumber="568">
<emphasis id="B95FFB5D5837FF88FC8193E4FC6E5184" box="[826,932,1741,1762]" italics="true" pageId="5" pageNumber="568">
<typeStatus id="549099ED5837FF88FC8193E4FC545184" box="[826,926,1741,1762]" pageId="5" pageNumber="568" type="holotype">Holotype</typeStatus>
:
</emphasis>
CNU-DIB-NN2007020, an almost complete adult body with wings in dorsal view.
</paragraph>
<paragraph id="8B94274F5837FF88FC819212FACF5009" blockId="5.[826,1441,1850,1903]" pageId="5" pageNumber="568">
<emphasis id="B95FFB5D5837FF88FC819212FC6D5036" box="[826,935,1851,1872]" italics="true" pageId="5" pageNumber="568">
<typeStatus id="549099ED5837FF88FC819212FC6A5036" box="[826,928,1851,1872]" pageId="5" pageNumber="568" type="paratype">Paratype</typeStatus>
:
</emphasis>
CNU-DIB-NN2007021, an almost complete adult body with a wing in lateral view.
</paragraph>
<caption id="DF5477C75834FF8BFF2B9624FEFC545B" ID-DOI="http://doi.org/10.5281/zenodo.5438132" ID-Zenodo-Dep="5438132" httpUri="https://zenodo.org/record/5438132/files/figure.png" pageId="6" pageNumber="569" startId="6.[144,223,781,800]" targetBox="[143,1424,198,751]" targetPageId="6">
<paragraph id="8B94274F5834FF8BFF2B9624FEFC545B" blockId="6.[144,1421,781,829]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFF2B9624FD605446" bold="true" box="[144,682,781,800]" pageId="6" pageNumber="569">
Figure 5.
<taxonomicName id="4C2B5CCC5834FF8BFEBF9624FD825446" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[260,584,781,800]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5834FF8BFEBF9624FD825446" bold="true" box="[260,584,781,800]" italics="true" pageId="6" pageNumber="569">Strenorhagio asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265834FF8BFDEA9624FD605446" box="[593,682,781,800]" pageId="6" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Holotype, body with wings in dorsal view. A, photograph. B, camera lucida drawing.
</paragraph>
</caption>
<caption id="DF5477C75834FF8BFF2B9029FD3D5257" ID-DOI="http://doi.org/10.5281/zenodo.5438134" ID-Zenodo-Dep="5438134" httpUri="https://zenodo.org/record/5438134/files/figure.png" pageId="6" pageNumber="569" startId="6.[144,223,1280,1299]" targetBox="[144,763,877,1251]" targetPageId="6">
<paragraph id="8B94274F5834FF8BFF2B9029FD3D5257" blockId="6.[144,759,1280,1329]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFF2B9029FD3D5272" bold="true" box="[144,759,1280,1300]" pageId="6" pageNumber="569">
Figure 6.
<taxonomicName id="4C2B5CCC5834FF8BFEBF9029FDA85272" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[260,610,1280,1300]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5834FF8BFEBF9029FDA85272" bold="true" box="[260,610,1280,1300]" italics="true" pageId="6" pageNumber="569">Strenorhagio asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265834FF8BFD3F9028FD3D5272" box="[644,759,1281,1300]" pageId="6" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype, photograph of body with a wing in lateral view.
</paragraph>
</caption>
<paragraph id="8B94274F5834FF8BFF2B9040FEB552BC" blockId="6.[144,759,1385,1498]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFF2B9040FE7C5218" box="[144,438,1385,1406]" italics="true" pageId="6" pageNumber="569">Type locality and horizon:</emphasis>
Daohugou Village, Shantou Township, Ningcheng County,
<collectingRegion id="49EFE9AD5834FF8BFE5790A1FD5552FB" box="[492,671,1415,1437]" country="China" name="Nei Mongol" pageId="6" pageNumber="569">Inner Mongolia</collectingRegion>
,
<collectingCountry id="F33C67DF5834FF8BFD1190AEFD3852FB" box="[682,754,1415,1437]" name="China" pageId="6" pageNumber="569">China</collectingCountry>
; Jiulongshan Formation, the Middle Jurassic (Aalenian–Bajocian).
</paragraph>
<paragraph id="8B94274F5834FF8BFF2B9320FE2F515A" blockId="6.[144,759,1544,1903]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFF2B9320FED25178" box="[144,280,1545,1566]" italics="true" pageId="6" pageNumber="569">Description:</emphasis>
Body length,
<quantity id="4CD38AAA5834FF8BFE659321FD885178" box="[478,578,1544,1566]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="569" unit="mm" value="13.5">13.5 mm</quantity>
; wing length,
<quantity id="4CD38AAA5834FF8BFF2B930EFF2D515A" box="[144,231,1575,1597]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="569" unit="mm" value="8.5">8.5 mm</quantity>
; wing width,
<quantity id="4CD38AAA5834FF8BFE33930EFE2A515A" box="[392,480,1575,1596]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="6" pageNumber="569" unit="mm" value="3.1">3.1 mm</quantity>
.
</paragraph>
<paragraph id="8B94274F5834FF8BFF13936CFD4951FF" blockId="6.[144,759,1544,1903]" pageId="6" pageNumber="569">Body stout. Head round, slightly narrower than thorax. Eyes large, bare; ommatidia clearly visible, with upper area composed of larger facets.</paragraph>
<paragraph id="8B94274F5834FF8BFF13938BFEDC5074" blockId="6.[144,759,1544,1903]" pageId="6" pageNumber="569">Legs pubescent. Hind femur swollen at apical part; each tibia with a spur at apex; first tarsomere distinctly much longer and wider than following tarsomeres.</paragraph>
<paragraph id="8B94274F5834FF8BFF139235FAA7520B" blockId="6.[144,759,1544,1903]" lastBlockId="6.[806,1421,878,1482]" pageId="6" pageNumber="569">
Wing hyaline; veins thick and strong. Vein Sc ending slightly beyond middle of wing. Veins R
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and Rs
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long; vein R
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distinctly and strongly bent upwards at base; vein R
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ending slightly beyond wing apex. Crossvein r–m at basal third of cell d; proximal to beginning of vein R
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2+
<quantity id="4CD38AAA5834FF8BFB89969FFBF054A4" box="[1074,1082,950,962]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="6" pageNumber="569" unit="in" value="3.0">3</quantity>
</subScript>
in left wing, nearly coalesced with vein R
<subScript id="17AF250A5834FF8BFC0996FDFC005486" attach="left" box="[946,970,980,992]" fontSize="5" pageId="6" pageNumber="569">2+3</subScript>
at vein Rs (with vein Rs
<subScript id="17AF250A5834FF8BFB4796FDFACE5486" attach="left" box="[1276,1284,980,992]" fontSize="5" pageId="6" pageNumber="569">2</subScript>
nearly disappearing) in right wing; vein Rs
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shorter than vein Rs
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. All branches of vein M long, veins M
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and M
<subScript id="17AF250A5834FF8BFA3E9138FA47537B" attach="left" box="[1413,1421,1041,1053]" fontSize="5" pageId="6" pageNumber="569">2</subScript>
bifurcating distal to crossvein m–m, basal trunk of veins M
<subScript id="17AF250A5834FF8BFC3E9166FC47533D" attach="left" box="[901,909,1103,1115]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
and M
<subScript id="17AF250A5834FF8BFC5D9166FC24533D" attach="left" box="[998,1006,1103,1115]" fontSize="5" pageId="6" pageNumber="569">2</subScript>
shorter than crossvein m–m. Vein CuA
<subScript id="17AF250A5834FF8BFCE29144FCAB531F" attach="left" box="[857,865,1133,1145]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
arising from cell bm, crossvein m–cu long; vein CuP present. Mouths of cells sc and r
<subScript id="17AF250A5834FF8BFB6591A5FB2C53FE" attach="left" box="[1246,1254,1164,1176]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
wide, relative widths 4: 3; mouths of cells r
<subScript id="17AF250A5834FF8BFB259183FB7C53D0" attach="left" box="[1182,1206,1194,1206]" fontSize="5" pageId="6" pageNumber="569">2+3</subScript>
and r
<subScript id="17AF250A5834FF8BFAB19183FAD853D0" attach="left" box="[1290,1298,1194,1206]" fontSize="5" pageId="6" pageNumber="569">4</subScript>
distinctly narrow. Cell bm about as wide as cell br; cell d slender, far from wing margin. Five posterior cells present, and wide open; mouths of cell m
<subScript id="17AF250A5834FF8BFA99902FFAE05274" attach="left" box="[1314,1322,1286,1298]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
slightly narrower than that of cell m
<subScript id="17AF250A5834FF8BFBC7900CFB4E5257" attach="left" box="[1148,1156,1317,1329]" fontSize="5" pageId="6" pageNumber="569">2</subScript>
; mouth of cell m
<subScript id="17AF250A5834FF8BFAEA900CFA935257" attach="left" box="[1361,1369,1317,1329]" fontSize="5" pageId="6" pageNumber="569">3</subScript>
distinctly narrower than that of cell m
<subScript id="17AF250A5834FF8BFB6E906AFB175229" attach="left" box="[1237,1245,1347,1359]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
, about half as wide as that of cell cua
<subScript id="17AF250A5834FF8BFB82904BFB8B5208" attach="left" box="[1081,1089,1378,1390]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
. Cell cup distinctly open.
</paragraph>
<paragraph id="8B94274F5834FF8BFC84905FFC5F52AC" blockId="6.[806,1421,878,1482]" pageId="6" pageNumber="569">Abdomen cylindrical, robust, and pubescent. Nine segments visible; segments I–VI with black stripe at each end.</paragraph>
<paragraph id="8B94274F5834FF8BFC9D9328FABE51F6" blockId="6.[806,1421,1537,1681]" pageId="6" pageNumber="569">
<emphasis id="B95FFB5D5834FF8BFC9D9328FC585170" box="[806,914,1537,1558]" italics="true" pageId="6" pageNumber="569">Remarks:</emphasis>
Because the position of crossvein r–m is asymmetric in the two wings, the crossvein m–cu is longer, the cell r
<subScript id="17AF250A5834FF8BFC4F9361FC365132" attach="left" box="[1012,1020,1608,1620]" fontSize="5" pageId="6" pageNumber="569">1</subScript>
is narrower, and segments I–VI have black stripe at each end, we can easily separate
<taxonomicName id="4C2B5CCC5834FF8BFC9D9352FC2E51F7" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[806,996,1659,1681]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="asymmetricus" status="sp. nov.">
<emphasis id="B95FFB5D5834FF8BFC9D9352FC2E51F7" box="[806,996,1659,1681]" italics="true" pageId="6" pageNumber="569">S. asymmetricus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265834FF8BFC559352FB8C51F6" box="[1006,1094,1659,1680]" pageId="6" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
from
<taxonomicName id="4C2B5CCC5834FF8BFB359352FAD951F6" authority="Zhang &amp; Yang &amp; Ren &amp; Shih, 2010" authorityName="Zhang &amp; Yang &amp; Ren &amp; Shih" authorityYear="2010" box="[1166,1299,1659,1680]" class="Insecta" family="Uranorhagionidae" genus="Strenorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="569" phylum="Arthropoda" rank="species" species="grimaldi" status="sp. nov.">
<emphasis id="B95FFB5D5834FF8BFB359352FAD951F6" box="[1166,1299,1659,1680]" italics="true" pageId="6" pageNumber="569">S. grimaldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C46265834FF8BFAA79352FABE51F6" box="[1308,1396,1659,1680]" pageId="6" pageNumber="569" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>
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