158 lines
17 KiB
XML
158 lines
17 KiB
XML
<document id="C0574D76DA4F0B9AA536E8142D3C2036" ID-CLB-Dataset="276483" ID-DOI="10.1111/j.1096-3642.2009.00552.x" ID-GBIF-Dataset="3be82545-1797-46a0-95fc-3e0fc44c9af0" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438122" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440497297" checkinUser="valdenar" docAuthor="Zhang, Kuiyan, Yang, Ding, Ren, Dong & Shih, Chungkun" docDate="2010" docId="038296595833FF8CFCDB9413FBF05293" docLanguage="en" docName="j.1096-3642.2009.00552.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Uranorhagionidae ZHANG, YANG & REN" docType="treatment" docVersion="6" lastPageNumber="564" masterDocId="FFBBEE215832FF8DFFBB9529FFCA5766" masterDocTitle="An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)" masterLastPageNumber="572" masterPageNumber="563" pageNumber="564" updateTime="1699284570084" updateUser="ExternalLinkService">
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<mods:titleInfo id="91D9CA622FA6C509EC55CA3A3260ED3D">
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<mods:title id="556243FACA49FA4E219D83CD91710257">An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera)</mods:title>
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<mods:roleTerm id="25CE2528C9BE0502F87F71B8C40FD6CB">Author</mods:roleTerm>
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<mods:namePart id="A3EE033450D8340FB8D56F3522720EFD">Zhang, Kuiyan</mods:namePart>
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<mods:roleTerm id="024C5498D15AB4323A6E5D386DD6CDF9">Author</mods:roleTerm>
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<mods:namePart id="599F2FE3D707DBD30F38F816EDA1B75F">Yang, Ding</mods:namePart>
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</mods:name>
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<mods:name id="D6DF05726A20417A7931FB8864759BC8" type="personal">
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<mods:roleTerm id="BE490722AA133C6A2A29D0C4E8E5FEB4">Author</mods:roleTerm>
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<mods:namePart id="93E6C0FB2AC52E1FF190BDFB68A8B38C">Ren, Dong</mods:namePart>
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</mods:name>
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<mods:name id="DD6C5F760AC20E08D7D5DD1FC0C3C527" type="personal">
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<mods:roleTerm id="04B45F8545EF4D786C98628A88232BE1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="39D8DD80BF891F9AF139DBBDBF52B458">Shih, Chungkun</mods:namePart>
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<mods:typeOfResource id="046537E2A2EA99C58B451568EE2AC909">text</mods:typeOfResource>
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<mods:title id="C4736C9D65657BBD2E7F1F36A9B782A4">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="874867553E5A1EFF6BC712F93A1E96EE">
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<mods:date id="FEA33CCE87F102C5DB1B668F0CD74AEF">2010</mods:date>
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<mods:number id="4FDBBABBF11BA077FBFFA60143748BAC">2010-02-26</mods:number>
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<mods:number id="0A8C926A7AF6791E1062941E51A8AA35">158</mods:number>
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<mods:url id="72A46F45E52A44D7DC60F5C4CE610BA1">http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x</mods:url>
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<mods:classification id="B484E8CB2C47E4DAB692071401C9F1F7">journal article</mods:classification>
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<mods:identifier id="381D921EE828D7E1D69F9AC35A833F1B" type="CLB-Dataset">276483</mods:identifier>
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<mods:identifier id="062AE142B27F93A6160189B6D8785AB1" type="DOI">10.1111/j.1096-3642.2009.00552.x</mods:identifier>
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<mods:identifier id="3B5C017A182C2217CEE54B2D44F4DF9C" type="GBIF-Dataset">3be82545-1797-46a0-95fc-3e0fc44c9af0</mods:identifier>
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<mods:identifier id="0C3C076C4E0256D8571BD50D487D0C86" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="BD067DC6827A5EE62FB91976637B7563" type="Zenodo-Dep">5438122</mods:identifier>
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<treatment id="038296595833FF8CFCDB9413FBF05293" ID-DOI="http://doi.org/10.5281/zenodo.5491002" ID-GBIF-Taxon="217238216" ID-Zenodo-Dep="5491002" LSID="urn:lsid:plazi:treatment:038296595833FF8CFCDB9413FBF05293" httpUri="http://treatment.plazi.org/id/038296595833FF8CFCDB9413FBF05293" lastPageNumber="564" pageId="1" pageNumber="564">
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<subSubSection id="C33174C45833FF8CFCDB9413FB0C5609" pageId="1" pageNumber="564" type="nomenclature">
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<paragraph id="8B94274F5833FF8CFCDB9413FB0C5609" blockId="1.[864,1403,314,367]" pageId="1" pageNumber="564">
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<heading id="D0DC90235833FF8CFCDB9413FB0C5609" centered="true" fontSize="9" level="2" pageId="1" pageNumber="564" reason="4">
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FAMILY
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<taxonomicName id="4C2B5CCC5833FF8CFC7E9414FB88560B" authority="ZHANG, YANG & REN" authorityName="ZHANG, YANG & REN" class="Insecta" family="Uranorhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family">
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<emphasis id="B95FFB5D5833FF8CFC7E9414FB705628" bold="true" box="[965,1210,317,334]" pageId="1" pageNumber="564">URANORHAGIONIDAE</emphasis>
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ZHANG, YANG & REN
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</taxonomicName>
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<emphasis id="B95FFB5D5833FF8CFBF79474FB0C5609" bold="true" box="[1100,1222,346,367]" pageId="1" pageNumber="564">FAM. NOV.</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33174C45833FF8CFC8194A8FBF05293" pageId="1" pageNumber="564" type="materials_examined">
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<paragraph id="8B94274F5833FF8CFC8194A8FB0756F0" blockId="1.[826,1229,384,406]" box="[826,1229,384,406]" pageId="1" pageNumber="564">
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<emphasis id="B95FFB5D5833FF8CFC8194A8FB9656F3" box="[826,1116,384,406]" italics="true" pageId="1" pageNumber="564">
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<typeStatus id="549099ED5833FF8CFC8194A8FCA456F0" box="[826,878,385,406]" pageId="1" pageNumber="564">Type</typeStatus>
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genus:
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<taxonomicName id="4C2B5CCC5833FF8CFC7194A9FB9656F3" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[970,1116,384,405]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus" status="gen. nov.">Uranorhagio</taxonomicName>
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</emphasis>
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<taxonomicNameLabel id="A26C46265833FF8CFBDD94A8FB0756F0" box="[1126,1229,385,406]" pageId="1" pageNumber="564" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B94274F5833FF8CFC819486FBE65591" blockId="1.[826,1441,431,759]" pageId="1" pageNumber="564">
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<emphasis id="B95FFB5D5833FF8CFC819486FC7B56A2" box="[826,945,431,452]" italics="true" pageId="1" pageNumber="564">Diagnosis:</emphasis>
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Body stout. Head round. Mesonotum slightly convex. Hind tibia with spur(s), empodia pulvilliform. Wing slender relative to body. Veins thick and strong. Vein Sc ending slightly beyond the middle of the wing; veins
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<collectionCode id="ED3ABF8A5833FF8CFBB79703FBD45559" box="[1036,1054,554,575]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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<subScript id="17AF250A5833FF8CFBA5971AFBEC5559" attach="left" box="[1054,1062,563,575]" fontSize="5" pageId="1" pageNumber="564">1</subScript>
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and Rs
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<subScript id="17AF250A5833FF8CFBC6971AFB4F5559" attach="left" box="[1149,1157,563,575]" fontSize="5" pageId="1" pageNumber="564">1</subScript>
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long, vein
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<collectionCode id="ED3ABF8A5833FF8CFABF9703FADC5559" box="[1284,1302,554,575]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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<subScript id="17AF250A5833FF8CFAAD971AFAE45559" attach="left" box="[1302,1326,563,575]" fontSize="5" pageId="1" pageNumber="564">2+3</subScript>
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distinctly and strongly bent upwards at base, vein
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<collectionCode id="ED3ABF8A5833FF8CFAA99761FAEE553B" box="[1298,1316,584,605]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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<subScript id="17AF250A5833FF8CFA9F977BFAE65538" attach="left" box="[1316,1324,594,606]" fontSize="5" pageId="1" pageNumber="564">5</subScript>
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ending at (or beyond) wing apex; veins M
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<subScript id="17AF250A5833FF8CFB179759FB7E551A" attach="left" box="[1196,1204,624,636]" fontSize="5" pageId="1" pageNumber="564">1</subScript>
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and M
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<subScript id="17AF250A5833FF8CFAB19759FAD8551A" attach="left" box="[1290,1298,624,636]" fontSize="5" pageId="1" pageNumber="564">2</subScript>
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bifurcating, distal of crossvein m–m; cell sc wide open; cell r
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<subScript id="17AF250A5833FF8CFA2297A6FA6B55FD" attach="left" box="[1433,1441,655,667]" fontSize="5" pageId="1" pageNumber="564">4</subScript>
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slender; five posterior cells present and wide open, cell cup distinctly open; anal lobe small and narrow, alula unconspicuous.
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</paragraph>
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<paragraph id="8B94274F5833FF8CFC819639FC3C5422" blockId="1.[826,1441,784,836]" pageId="1" pageNumber="564">
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<emphasis id="B95FFB5D5833FF8CFC819639FB345443" box="[826,1278,784,805]" italics="true" pageId="1" pageNumber="564">Stratigraphic and geographic range:</emphasis>
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The Middle Jurassic,
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<collectingCountry id="F33C67DF5833FF8CFC119607FC395422" box="[938,1011,814,836]" name="China" pageId="1" pageNumber="564">China</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B94274F5833FF8CFC819674FBF05293" blockId="1.[826,1441,861,1525]" pageId="1" pageNumber="564">
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<emphasis id="B95FFB5D5833FF8CFC819674FC6C5414" box="[826,934,861,882]" italics="true" pageId="1" pageNumber="564">Remarks:</emphasis>
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<taxonomicName id="4C2B5CCC5833FF8CFC169674FBB75415" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[941,1149,861,883]" class="Insecta" family="Uranorhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family" status="fam. nov.">Uranorhagionidae</taxonomicName>
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<taxonomicNameLabel id="A26C46265833FF8CFB3F9674FB215414" box="[1156,1259,861,883]" pageId="1" pageNumber="564" rank="family">fam. nov.</taxonomicNameLabel>
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is considered to be a member of the lower Orthorrhapha, because of the stout body, the presence of tibia spur(s), and pulvilliform empodia. Contrasting with other families of lower Orthorrhapha, this new family is somewhat similar to the
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<taxonomicName id="4C2B5CCC5833FF8CFCDD96DFFC3F536A" box="[870,1013,1014,1036]" class="Insecta" family="Rhagionidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family">Rhagionidae</taxonomicName>
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family. However, we cannot put these specimens in any known families, because of their special and unique wing venation. Therefore, we erect a new family in superfamily
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<taxonomicName id="4C2B5CCC5833FF8CFB38917BFACC530E" box="[1155,1286,1106,1128]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Saxifragales" pageId="1" pageNumber="564" phylum="Tracheophyta" rank="superFamily" superFamily="Tabanoidea">Tabanoidea</taxonomicName>
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for them. We can differentiate
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<taxonomicName id="4C2B5CCC5833FF8CFBB59159FB1453E0" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[1038,1246,1136,1158]" class="Insecta" family="Uranorhagionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family" status="fam. nov.">Uranorhagionidae</taxonomicName>
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<taxonomicNameLabel id="A26C46265833FF8CFB569159FA9753E0" box="[1261,1373,1136,1158]" pageId="1" pageNumber="564" rank="family">fam. nov.</taxonomicNameLabel>
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from
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<taxonomicName id="4C2B5CCC5833FF8CFC8191A6FC0353C3" box="[826,969,1167,1189]" class="Insecta" family="Rhagionidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family">Rhagionidae</taxonomicName>
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by the following characters: veins Sc and
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<collectionCode id="ED3ABF8A5833FF8CFC819187FC8653A5" box="[826,844,1198,1219]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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<subScript id="17AF250A5833FF8CFCF79191FC9E53A2" attach="left" box="[844,852,1208,1220]" fontSize="5" pageId="1" pageNumber="564">1</subScript>
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long; vein
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<collectionCode id="ED3ABF8A5833FF8CFC749187FC2B53A5" box="[975,993,1198,1219]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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<subScript id="17AF250A5833FF8CFC5B9191FC3353A2" attach="left" box="[992,1017,1208,1220]" fontSize="5" pageId="1" pageNumber="564">2+3</subScript>
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fornical at base; vein CuA
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<subScript id="17AF250A5833FF8CFA989191FAE153A2" attach="left" box="[1315,1323,1208,1220]" fontSize="5" pageId="1" pageNumber="564">2</subScript>
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short; cell d extending to near the wing margin (in
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<taxonomicName id="4C2B5CCC5833FF8CFAB491E5FA6B5387" authority="Zhang & Yang & Ren & Shih, 2010" authorityName="Zhang & Yang & Ren & Shih" authorityYear="2010" box="[1295,1441,1228,1249]" class="Insecta" family="Uranorhagionidae" genus="Uranorhagio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B95FFB5D5833FF8CFAB491E5FA6B5387" box="[1295,1441,1228,1249]" italics="true" pageId="1" pageNumber="564">Uranorhagio</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C46265833FF8CFC8191C2FC635266" box="[826,937,1259,1280]" pageId="1" pageNumber="564" rank="genus">gen. nov.</taxonomicNameLabel>
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); hind femur swollen at apex; and nine abdominal segments. Also, this family is superficially similar to the
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<taxonomicName id="4C2B5CCC5833FF8CFC5C9001FB065258" box="[999,1228,1320,1342]" class="Insecta" family="Rhagionemestriidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="family">Rhagionemestriidae</taxonomicName>
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family (especially for
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<taxonomicName id="4C2B5CCC5833FF8CFCD8906FFC29523D" authorityName="Mostovski & Martinez-Delclos" authorityYear="2000" box="[867,995,1350,1371]" class="Insecta" family="Rhagionemestriidae" genus="Iberomosca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FFB5D5833FF8CFCD8906FFC29523D" box="[867,995,1350,1371]" italics="true" pageId="1" pageNumber="564">Iberomosca</emphasis>
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</taxonomicName>
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). But, we can easily separate the new family from
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<taxonomicName id="4C2B5CCC5833FF8CFC7E904CFB8F521C" authorityName="Mostovski & Martinez-Delclos" authorityYear="2000" box="[965,1093,1381,1402]" class="Insecta" family="Rhagionemestriidae" genus="Iberomosca" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="564" phylum="Arthropoda" rank="genus">
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<emphasis id="B95FFB5D5833FF8CFC7E904CFB8F521C" box="[965,1093,1381,1402]" italics="true" pageId="1" pageNumber="564">Iberomosca</emphasis>
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</taxonomicName>
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by the following features: vein Rs
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<subScript id="17AF250A5833FF8CFCEC90A7FC9552FC" attach="left" box="[855,863,1422,1434]" fontSize="5" pageId="1" pageNumber="564">1</subScript>
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extends much longer than vein Rs
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<subScript id="17AF250A5833FF8CFAB890A7FAC152FC" attach="left" box="[1283,1291,1422,1434]" fontSize="5" pageId="1" pageNumber="564">2</subScript>
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; vein
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<collectionCode id="ED3ABF8A5833FF8CFAE290ADFAA152FF" box="[1369,1387,1412,1433]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="564">R</collectionCode>
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||
<subScript id="17AF250A5833FF8CFAD190A7FA4952FC" attach="left" box="[1386,1411,1422,1434]" fontSize="5" pageId="1" pageNumber="564">2+3</subScript>
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is fornical at the base; cell br is longer than cell bm; the cell cup is open at the wing margin; and there is no distinct diagonal vein.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |