315 lines
39 KiB
XML
315 lines
39 KiB
XML
<document id="D03A95BCC1A41AFC5CD8F0BC900C1FAB" ID-CLB-Dataset="27673" ID-DOI="10.11646/zootaxa.4545.4.5" ID-GBIF-Dataset="847bdbc0-1d3a-4c4b-b4db-b72c57f29048" ID-ISSN="1175-5326" ID-Zenodo-Dep="2618303" ID-ZooBank="A95821A9-9E6D-4914-B02A-EA1753B9587D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553929485073" checkinUser="plazi" docAuthor="Ermilov, Sergey G. & Khaustov, Alexander A." docDate="2019" docId="038287D33366FFD51EA2594DFBE4FA6D" docLanguage="en" docName="zootaxa.4545.2.5.pdf" docOrigin="Zootaxa 4545 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pergalumna pietertheroni Ermilov & Khaustov 2019, sp. nov." docType="treatment" docVersion="9" lastPageNumber="534" masterDocId="FFBBFFAB3364FFD61E355F45FFE6FFC7" masterDocTitle="New Galumnoidea (Acari, Oribatida) from Tanzania" masterLastPageNumber="547" masterPageNumber="531" pageNumber="533" updateTime="1698715666792" updateUser="ExternalLinkService">
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<mods:mods id="5221E947FF2DF1BF6FCCC5E42609CBC1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5BE02320D3550BD549DDEEB6C8609E8C">
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<mods:title id="385DF84CBCB0CC301E2EBB5E065123EE">New Galumnoidea (Acari, Oribatida) from Tanzania</mods:title>
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<mods:namePart id="BFB55A6F943F4E6BD31C5450EF2DDD48">Ermilov, Sergey G.</mods:namePart>
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<mods:namePart id="8EFE92A474DB6924B2B95C6366299D08">Khaustov, Alexander A.</mods:namePart>
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<mods:typeOfResource id="0BB784CB75FFE35BEB0CB49C752B3A08">text</mods:typeOfResource>
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<mods:title id="27BBF6C45B542E625E9E363753C7D56B">Zootaxa</mods:title>
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<mods:part id="C1B155A9F9E65A76722D4DD89B175336">
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<mods:date id="2FCC5662E71F54379F9F09D79591B21A">2019</mods:date>
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<mods:number id="B3A10499830C3BF159B3D234856D548C">2019-01-21</mods:number>
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<mods:number id="8AC52CFDCDBC8CEC8F15164D49FC5EBB">4545</mods:number>
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<mods:number id="C736004BE1B485950B739108EABE3FB5">4</mods:number>
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<mods:classification id="C1866BBB0801203D303D8F2887F51FB3">journal article</mods:classification>
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<mods:identifier id="89CC49764D4833D86A60838968426CA5" type="DOI">10.11646/zootaxa.4545.4.5</mods:identifier>
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<treatment id="038287D33366FFD51EA2594DFBE4FA6D" ID-DOI="http://doi.org/10.5281/zenodo.5930123" ID-GBIF-Taxon="154530266" ID-Zenodo-Dep="5930123" LSID="urn:lsid:plazi:treatment:038287D33366FFD51EA2594DFBE4FA6D" httpUri="http://treatment.plazi.org/id/038287D33366FFD51EA2594DFBE4FA6D" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
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<subSubSection id="C331654E3366FFD41EA2594DFDD3F9E4" box="[151,565,1544,1571]" pageId="2" pageNumber="533" type="nomenclature">
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<paragraph id="8B9436C53366FFD41EA2594DFDD3F9E4" blockId="2.[151,565,1544,1605]" box="[151,565,1544,1571]" pageId="2" pageNumber="533">
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<heading id="D0DC81A93366FFD41EA2594DFDD3F9E4" bold="true" box="[151,565,1544,1571]" fontSize="11" level="1" pageId="2" pageNumber="533" reason="1">
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<emphasis id="B95FEAD73366FFD41EA2594DFDD3F9E4" bold="true" box="[151,565,1544,1571]" pageId="2" pageNumber="533">
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<taxonomicName id="4C2B4D463366FFD41EA2594DFE36F9E5" ID-CoL="8TFLJ" authority="Ermilov & Khaustov, 2019" authorityName="Ermilov & Khaustov" authorityYear="2019" box="[151,464,1544,1570]" class="Arachnida" family="Galumnidae" genus="Pergalumna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="2" pageNumber="533" phylum="Arthropoda" rank="species" species="pietertheroni" status="sp. nov.">
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<emphasis id="B95FEAD73366FFD41EA2594DFE36F9E5" bold="true" box="[151,464,1544,1570]" italics="true" pageId="2" pageNumber="533">Pergalumna pietertheroni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26C57AC3366FFD41FE2594CFDD3F9E4" box="[471,565,1545,1571]" pageId="2" pageNumber="533" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C331654E3366FFD41EA25969FE9DF983" box="[151,379,1580,1605]" pageId="2" pageNumber="533" type="description">
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<paragraph id="8B9436C53366FFD41EA25969FE9DF983" blockId="2.[151,565,1544,1605]" box="[151,379,1580,1605]" pageId="2" pageNumber="533">
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(
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<figureCitation id="13102A403366FFD41EAA5969FEC6F982" box="[159,288,1580,1605]" captionStart-0="FIGURES 14–18" captionStart-1="FIGURES 19–21" captionStart-2="FIGURES 22–25" captionStartId-0="7.[151,264,1925,1948]" captionStartId-1="8.[151,264,1750,1773]" captionStartId-2="9.[151,264,1949,1972]" captionTargetBox-0="[190,1417,197,1824]" captionTargetBox-1="[218,1339,296,1625]" captionTargetBox-2="[210,1404,228,1877]" captionTargetId-0="figure@7.[174,1419,193,1954]" captionTargetId-1="figure@8.[150,1436,225,1709]" captionTargetId-2="figure@9.[150,1436,193,1928]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." captionText-1="FIGURES 19–21. Pergalumna pietertheroni sp. nov., adult: 19—anterior part of body, lateral view (gnathosoma, pteromorph and legs not shown); 20—posterior part of body, lateral view; 21—posterior view. Scale bar 50 µm." captionText-2="FIGURES 22–25. Pergalumna pietertheroni sp. nov., adult: 22—leg I, right, antiaxial view; 23—trochanter, femur and genu of leg II, right, ventroantiaxial view; 24—trochanter, femur and genu of leg III, left, antiaxial view; 25—leg IV, left, antiaxial view. Scale bar 20 µm." httpUri-0="https://zenodo.org/record/2618311/files/figure.png" httpUri-1="https://zenodo.org/record/2618313/files/figure.png" httpUri-2="https://zenodo.org/record/2618315/files/figure.png" pageId="2" pageNumber="533">Figs 14–25</figureCitation>
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,
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<figureCitation id="13102A403366FFD41F1E5969FE94F983" box="[299,370,1580,1604]" captionStart="FIGURES 38–45" captionStartId="13.[151,264,1836,1859]" captionTargetBox="[150,1436,193,1806]" captionTargetId="figure@13.[150,1436,193,1806]" captionTargetPageId="13" captionText="FIGURES 38–45. Galumnoidea, adults, microscope images: 38—Pergalumna hugocoetzeeae sp. nov., prodorsal sculpturing and rostrum, dorsoanterior view; 39—Pergalumna hugocoetzeeae sp. nov., dorsosejugal suture; 40—Pergalumna hugocoetzeeae sp. nov., dorsophragma and sejugal porose area; 41—Pergalumna pietertheroni sp. nov., granulate sculpturing in anogenital region; 42—Pergalumna pietertheroni sp. nov., dorsophragma and sejugal porose area; 43—Galumnella paraokinawana sp. nov., foveolate ornamentation in epimeral region; 44—Galumnella paraokinawana sp. nov., reticulate ornamentation between genital and anal apertures; 45—Galumnella paraokinawana sp. nov., claw of leg tarsus II." httpUri="https://zenodo.org/record/2618323/files/figure.png" pageId="2" pageNumber="533">41, 42</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C331654E3366FFD41EA25931FED2F8DA" pageId="2" pageNumber="533" type="diagnosis">
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<paragraph id="8B9436C53366FFD41EA25931FED2F8DA" blockId="2.[151,1436,1652,2037]" pageId="2" pageNumber="533">
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<emphasis id="B95FEAD73366FFD41EA25931FEF7F94A" bold="true" box="[151,273,1652,1677]" pageId="2" pageNumber="533">Diagnosis.</emphasis>
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Body size: 315–348 × 215–249. Body surface microgranulate. Rostral, lamellar and interlamellar setae short, thin, smooth,
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<emphasis id="B95FEAD73366FFD41FB559DCFE71F977" box="[384,407,1689,1712]" italics="true" pageId="2" pageNumber="533">ro</emphasis>
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longest. Bothridial setae long, setiform, smooth. Dorsosejugal suture present. With three pairs of rounded porose areas,
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<emphasis id="B95FEAD73366FFD41FCF59F9FDFFF913" box="[506,537,1724,1748]" italics="true" pageId="2" pageNumber="533">A1</emphasis>
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absent. Porose areas
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<emphasis id="B95FEAD73366FFD41D2659F9FCD4F914" box="[787,818,1724,1747]" italics="true" pageId="2" pageNumber="533">Aa</emphasis>
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located between
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<emphasis id="B95FEAD73366FFD41DC959F9FBF4F912" box="[1020,1042,1724,1749]" italics="true" pageId="2" pageNumber="533">la</emphasis>
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and
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<emphasis id="B95FEAD73366FFD41A7859F9FB8FF912" box="[1101,1129,1724,1749]" italics="true" pageId="2" pageNumber="533">lm</emphasis>
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, closer to
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<emphasis id="B95FEAD73366FFD41AD059F9FB1DF912" box="[1253,1275,1724,1749]" italics="true" pageId="2" pageNumber="533">la</emphasis>
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. Median pore absent. Epimeral setal formula: 3–1–3–3. Epimeral and anogenital setae short, thin, smooth. Postanal porose area elongate oval.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331654E3366FFD51EF25862FD05FBA1" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533" type="description">
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<paragraph id="8B9436C53366FFD41EF25862FAC3F8A2" blockId="2.[151,1436,1652,2037]" pageId="2" pageNumber="533">
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<emphasis id="B95FEAD73366FFD41EF25862FEBFF887" bold="true" box="[199,345,1831,1856]" pageId="2" pageNumber="533">Description.</emphasis>
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<emphasis id="B95FEAD73366FFD41F56586CFDE3F887" box="[355,517,1833,1856]" italics="true" pageId="2" pageNumber="533">Measurements</emphasis>
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. Body length: 332 (
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<typeStatus id="549088673366FFD41CC45862FCBFF887" box="[753,857,1831,1856]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
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), 315–348 (five
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<typeStatus id="549088673366FFD41A2E586DFB6AF887" box="[1051,1164,1832,1856]" pageId="2" pageNumber="533" type="paratype">paratypes</typeStatus>
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); notogaster width: 232 (
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<typeStatus id="549088673366FFD41EAA5809FEE1F8A2" box="[159,263,1868,1893]" pageId="2" pageNumber="533" type="holotype">holotype</typeStatus>
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), 215–249 (five
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<typeStatus id="549088673366FFD41F8B5809FDC8F8A3" box="[446,558,1868,1892]" pageId="2" pageNumber="533" type="paratype">paratypes</typeStatus>
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). Females larger than males: 348 × 249 versus 315–332 × 215–232.
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</paragraph>
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<paragraph id="8B9436C53366FFD41EF25834FC4EF86B" blockId="2.[151,1436,1652,2037]" pageId="2" pageNumber="533">
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<emphasis id="B95FEAD73366FFD41EF25834FEA3F84F" box="[199,325,1905,1928]" italics="true" pageId="2" pageNumber="533">Integument</emphasis>
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(
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<figureCitation id="13102A403366FFD41F61582AFE40F84F" box="[340,422,1903,1928]" captionStart="FIGURES 38–45" captionStartId="13.[151,264,1836,1859]" captionTargetBox="[150,1436,193,1806]" captionTargetId="figure@13.[150,1436,193,1806]" captionTargetPageId="13" captionText="FIGURES 38–45. Galumnoidea, adults, microscope images: 38—Pergalumna hugocoetzeeae sp. nov., prodorsal sculpturing and rostrum, dorsoanterior view; 39—Pergalumna hugocoetzeeae sp. nov., dorsosejugal suture; 40—Pergalumna hugocoetzeeae sp. nov., dorsophragma and sejugal porose area; 41—Pergalumna pietertheroni sp. nov., granulate sculpturing in anogenital region; 42—Pergalumna pietertheroni sp. nov., dorsophragma and sejugal porose area; 43—Galumnella paraokinawana sp. nov., foveolate ornamentation in epimeral region; 44—Galumnella paraokinawana sp. nov., reticulate ornamentation between genital and anal apertures; 45—Galumnella paraokinawana sp. nov., claw of leg tarsus II." httpUri="https://zenodo.org/record/2618323/files/figure.png" pageId="2" pageNumber="533">Fig. 41</figureCitation>
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). Body color pale yellowish. Body surface (including subcapitular mentum and genital and anal plates) densely microgranulate (diameter of granules less than 1).
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</paragraph>
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<paragraph id="8B9436C53366FFD51EF258F2FDDAFEDA" blockId="2.[151,1436,1652,2037]" lastBlockId="3.[151,1437,151,1451]" lastPageId="3" lastPageNumber="534" pageId="2" pageNumber="533">
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<emphasis id="B95FEAD73366FFD41EF258F2FEA2F817" box="[199,324,1975,2000]" italics="true" pageId="2" pageNumber="533">Prodorsum</emphasis>
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(
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<figureCitation id="13102A403366FFD41F6258F2FE06F817" box="[343,480,1975,2000]" captionStart="FIGURES 14–18" captionStartId="7.[151,264,1925,1948]" captionTargetBox="[190,1417,197,1824]" captionTargetId="figure@7.[174,1419,193,1954]" captionTargetPageId="7" captionText="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." httpUri="https://zenodo.org/record/2618311/files/figure.png" pageId="2" pageNumber="533">Figs 14, 15</figureCitation>
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,
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<figureCitation id="13102A403366FFD41FC558FDFDE8F817" box="[496,526,1976,2000]" captionStart="FIGURES 19–21" captionStartId="8.[151,264,1750,1773]" captionTargetBox="[218,1339,296,1625]" captionTargetId="figure@8.[150,1436,225,1709]" captionTargetPageId="8" captionText="FIGURES 19–21. Pergalumna pietertheroni sp. nov., adult: 19—anterior part of body, lateral view (gnathosoma, pteromorph and legs not shown); 20—posterior part of body, lateral view; 21—posterior view. Scale bar 50 µm." httpUri="https://zenodo.org/record/2618313/files/figure.png" pageId="2" pageNumber="533">19</figureCitation>
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,
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<figureCitation id="13102A403366FFD41C2A58FDFDD9F817" box="[543,575,1976,2000]" captionStart="FIGURES 38–45" captionStartId="13.[151,264,1836,1859]" captionTargetBox="[150,1436,193,1806]" captionTargetId="figure@13.[150,1436,193,1806]" captionTargetPageId="13" captionText="FIGURES 38–45. Galumnoidea, adults, microscope images: 38—Pergalumna hugocoetzeeae sp. nov., prodorsal sculpturing and rostrum, dorsoanterior view; 39—Pergalumna hugocoetzeeae sp. nov., dorsosejugal suture; 40—Pergalumna hugocoetzeeae sp. nov., dorsophragma and sejugal porose area; 41—Pergalumna pietertheroni sp. nov., granulate sculpturing in anogenital region; 42—Pergalumna pietertheroni sp. nov., dorsophragma and sejugal porose area; 43—Galumnella paraokinawana sp. nov., foveolate ornamentation in epimeral region; 44—Galumnella paraokinawana sp. nov., reticulate ornamentation between genital and anal apertures; 45—Galumnella paraokinawana sp. nov., claw of leg tarsus II." httpUri="https://zenodo.org/record/2618323/files/figure.png" pageId="2" pageNumber="533">42</figureCitation>
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). Rostrum rounded. Lamellar and sublamellar lines thin, parallel, curving backwards at ventral ends. Prodorsal leg niches and lateral ridges of prodorsum well developed. Rostral (20–28), lamellar (8) and interlamellar (12) setae setiform, thin, smooth. Insertions of lamellar setae clearly distanced from lamellar lines. Bothridial setae (90–102) setiform, smooth. Exobothridial setae and their alveoli absent. Sejugal porose areas (12–16 × 4) elongate oval, transversely oriented, located posterolateral to interlamellar setae. Dorsophragmata distinctly elongated.
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</paragraph>
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<paragraph id="8B9436C53367FFD51EF25E6CFDE6FE11" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
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<emphasis id="B95FEAD73367FFD51EF25E6CFEA2FE87" box="[199,324,297,320]" italics="true" pageId="3" pageNumber="534">Notogaster</emphasis>
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(
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<figureCitation id="13102A403367FFD51F675E62FE4FFE87" box="[338,425,295,320]" captionStart="FIGURES 14–18" captionStartId="7.[151,264,1925,1948]" captionTargetBox="[190,1417,197,1824]" captionTargetId="figure@7.[174,1419,193,1954]" captionTargetPageId="7" captionText="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." httpUri="https://zenodo.org/record/2618311/files/figure.png" pageId="3" pageNumber="534">Figs 14</figureCitation>
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,
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<figureCitation id="13102A403367FFD51F815E6DFE18FE87" box="[436,510,296,320]" captionStart="FIGURES 19–21" captionStartId="8.[151,264,1750,1773]" captionTargetBox="[218,1339,296,1625]" captionTargetId="figure@8.[150,1436,225,1709]" captionTargetPageId="8" captionText="FIGURES 19–21. Pergalumna pietertheroni sp. nov., adult: 19—anterior part of body, lateral view (gnathosoma, pteromorph and legs not shown); 20—posterior part of body, lateral view; 21—posterior view. Scale bar 50 µm." httpUri="https://zenodo.org/record/2618313/files/figure.png" pageId="3" pageNumber="534">19–21</figureCitation>
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). Dorsosejugal suture present, simple. With 10 pairs of setal alveoli and three pairs of rounded porose areas (
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<emphasis id="B95FEAD73367FFD51F845E09FE36FEA4" box="[433,464,332,355]" italics="true" pageId="3" pageNumber="534">Aa</emphasis>
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, 8–10;
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<emphasis id="B95FEAD73367FFD51C115E09FDA5FEA3" box="[548,579,332,356]" italics="true" pageId="3" pageNumber="534">A2</emphasis>
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, 6–8;
|
||
<emphasis id="B95FEAD73367FFD51CBC5E09FD4EFEA3" box="[649,680,332,356]" italics="true" pageId="3" pageNumber="534">A3</emphasis>
|
||
, 4–6),
|
||
<emphasis id="B95FEAD73367FFD51CC35E09FCF3FEA3" box="[758,789,332,356]" italics="true" pageId="3" pageNumber="534">A1</emphasis>
|
||
absent. Porose areas
|
||
<emphasis id="B95FEAD73367FFD51A315E09FBC2FEA4" box="[1028,1060,332,355]" italics="true" pageId="3" pageNumber="534">Aa</emphasis>
|
||
located between
|
||
<emphasis id="B95FEAD73367FFD51AD15E09FB1CFEA2" box="[1252,1274,332,357]" italics="true" pageId="3" pageNumber="534">la</emphasis>
|
||
and
|
||
<emphasis id="B95FEAD73367FFD51B1A5E09FAADFEA2" box="[1327,1355,332,357]" italics="true" pageId="3" pageNumber="534">lm</emphasis>
|
||
, closer to
|
||
<emphasis id="B95FEAD73367FFD51E865E2AFF2FFE4F" box="[179,201,367,392]" italics="true" pageId="3" pageNumber="534">la</emphasis>
|
||
. Median pore absent. All lyrifissures distinct,
|
||
<emphasis id="B95FEAD73367FFD51CFB5E34FD0CFE4F" box="[718,746,369,392]" italics="true" pageId="3" pageNumber="534">im</emphasis>
|
||
located anterolateral to
|
||
<emphasis id="B95FEAD73367FFD51DCD5E35FBF1FE4F" box="[1016,1047,368,392]" italics="true" pageId="3" pageNumber="534">A1</emphasis>
|
||
,
|
||
<emphasis id="B95FEAD73367FFD51A105E34FBDDFE4F" box="[1061,1083,369,392]" italics="true" pageId="3" pageNumber="534">ip</emphasis>
|
||
between
|
||
<emphasis id="B95FEAD73367FFD51A905E34FB55FE4F" box="[1189,1203,369,392]" italics="true" pageId="3" pageNumber="534">p</emphasis>
|
||
<subScript id="17AF34803367FFD51A865E3BFB5DFE4B" attach="left" box="[1203,1211,382,396]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
and
|
||
<emphasis id="B95FEAD73367FFD51AC45E34FB19FE4F" box="[1265,1279,369,392]" italics="true" pageId="3" pageNumber="534">p</emphasis>
|
||
<subScript id="17AF34803367FFD51ACA5E3BFAE1FE4B" attach="left" box="[1279,1287,382,396]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
and closer to
|
||
<emphasis id="B95FEAD73367FFD51EA25EDDFF43FE68" box="[151,165,408,431]" italics="true" pageId="3" pageNumber="534">p</emphasis>
|
||
<subScript id="17AF34803367FFD51E905EE0FF4BFE74" attach="left" box="[165,173,421,435]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
,
|
||
<emphasis id="B95FEAD73367FFD51E885ED3FF35FE68" box="[189,211,406,431]" italics="true" pageId="3" pageNumber="534">ih</emphasis>
|
||
anterior to
|
||
<emphasis id="B95FEAD73367FFD51F695EDDFE8CFE68" box="[348,362,408,431]" italics="true" pageId="3" pageNumber="534">p</emphasis>
|
||
<subScript id="17AF34803367FFD51F5F5EE0FE94FE74" attach="left" box="[362,370,421,435]" fontSize="6" pageId="3" pageNumber="534">3</subScript>
|
||
,
|
||
<emphasis id="B95FEAD73367FFD51FB75EDDFE45FE68" box="[386,419,408,431]" italics="true" pageId="3" pageNumber="534">ips</emphasis>
|
||
posterior to
|
||
<emphasis id="B95FEAD73367FFD51C0D5EDDFDA0FE68" box="[568,582,408,431]" italics="true" pageId="3" pageNumber="534">p</emphasis>
|
||
<subScript id="17AF34803367FFD51C735EE0FDA8FE74" attach="left" box="[582,590,421,435]" fontSize="6" pageId="3" pageNumber="534">3</subScript>
|
||
. Opisthonotal gland openings located lateral to
|
||
<emphasis id="B95FEAD73367FFD51A425ED2FB70FE68" box="[1143,1174,407,431]" italics="true" pageId="3" pageNumber="534">A2</emphasis>
|
||
. Circumgastric sigillar band present, but poorly visible.
|
||
</paragraph>
|
||
<paragraph id="8B9436C53367FFD51EF25EA4FA9AFDA1" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25EA4FEB0FE3D" box="[199,342,481,506]" italics="true" pageId="3" pageNumber="534">Gnathosoma</emphasis>
|
||
(
|
||
<figureCitation id="13102A403367FFD51F5F5EA4FE15FE3D" box="[362,499,481,506]" captionStart="FIGURES 14–18" captionStartId="7.[151,264,1925,1948]" captionTargetBox="[190,1417,197,1824]" captionTargetId="figure@7.[174,1419,193,1954]" captionTargetPageId="7" captionText="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." httpUri="https://zenodo.org/record/2618311/files/figure.png" pageId="3" pageNumber="534">Figs 16–18</figureCitation>
|
||
). Subcapitulum size: 106–110 × 90–94. Subcapitular setae setiform, smooth,
|
||
<emphasis id="B95FEAD73367FFD51BBB5EA7FA7AFE3E" box="[1422,1436,482,505]" italics="true" pageId="3" pageNumber="534">a</emphasis>
|
||
(20–28) longer than
|
||
<emphasis id="B95FEAD73367FFD51FB25D43FE7DFDDA" box="[391,411,518,541]" italics="true" pageId="3" pageNumber="534">m</emphasis>
|
||
(12) and
|
||
<emphasis id="B95FEAD73367FFD51C275D40FDC6FDD9" box="[530,544,517,542]" italics="true" pageId="3" pageNumber="534">h</emphasis>
|
||
(4–6);
|
||
<emphasis id="B95FEAD73367FFD51C4F5D43FD6EFDDA" box="[634,648,518,541]" italics="true" pageId="3" pageNumber="534">a</emphasis>
|
||
thickest,
|
||
<emphasis id="B95FEAD73367FFD51CC95D40FCECFDD9" box="[764,778,517,542]" italics="true" pageId="3" pageNumber="534">h</emphasis>
|
||
thinnest. Adoral setae (10–12) setiform, heavily barbed. Length of palps 73–77. Postpalpal setae (6) spiniform, smooth. Length of chelicerae 127–131. Cheliceral setae setiform, barbed,
|
||
<emphasis id="B95FEAD73367FFD51F695D08FE63FDA1" box="[348,389,589,614]" italics="true" pageId="3" pageNumber="534">cha</emphasis>
|
||
(41–45) longer than
|
||
<emphasis id="B95FEAD73367FFD51C445D08FD7CFDA1" box="[625,666,589,614]" italics="true" pageId="3" pageNumber="534">chb</emphasis>
|
||
(24–26). Trägårdh’s organ of chelicerae long, elongate triangular.
|
||
</paragraph>
|
||
<paragraph id="8B9436C53367FFD51EF25D34FC7EFD31" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25D34FD46FD4E" box="[199,672,625,650]" italics="true" pageId="3" pageNumber="534">Epimeral and lateral podosomal regions</emphasis>
|
||
(
|
||
<figureCitation id="13102A403367FFD51C865D34FCE8FD4D" box="[691,782,625,650]" captionStart="FIGURES 14–18" captionStartId="7.[151,264,1925,1948]" captionTargetBox="[190,1417,197,1824]" captionTargetId="figure@7.[174,1419,193,1954]" captionTargetPageId="7" captionText="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." httpUri="https://zenodo.org/record/2618311/files/figure.png" pageId="3" pageNumber="534">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation id="13102A403367FFD51D2B5D37FCDBFD4D" box="[798,829,626,650]" captionStart="FIGURES 19–21" captionStartId="8.[151,264,1750,1773]" captionTargetBox="[218,1339,296,1625]" captionTargetId="figure@8.[150,1436,225,1709]" captionTargetPageId="8" captionText="FIGURES 19–21. Pergalumna pietertheroni sp. nov., adult: 19—anterior part of body, lateral view (gnathosoma, pteromorph and legs not shown); 20—posterior part of body, lateral view; 21—posterior view. Scale bar 50 µm." httpUri="https://zenodo.org/record/2618313/files/figure.png" pageId="3" pageNumber="534">19</figureCitation>
|
||
). Anterior tectum of epimere I smooth. Pedotecta I broadly rounded, pedotecta II triangular in ventral view. Discidia triangular. Epimeral setal formula: 3–1–3–3. Epimeral setae (4–6) setiform, thin, smooth. Circumpedal carinae (slightly developed in mediodistal part) of medium length, directed to insertions of setae
|
||
<emphasis id="B95FEAD73367FFD51CA95D98FD5EFD31" box="[668,696,733,758]" italics="true" pageId="3" pageNumber="534">3b</emphasis>
|
||
, but reaching them.
|
||
</paragraph>
|
||
<paragraph id="8B9436C53367FFD51EF25C44FBE1FC6A" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25C44FE74FCDE" box="[199,402,769,794]" italics="true" pageId="3" pageNumber="534">Anogenital region</emphasis>
|
||
(
|
||
<figureCitation id="13102A403367FFD51F945C44FE1FFCDD" box="[417,505,769,794]" captionStart="FIGURES 14–18" captionStartId="7.[151,264,1925,1948]" captionTargetBox="[190,1417,197,1824]" captionTargetId="figure@7.[174,1419,193,1954]" captionTargetPageId="7" captionText="FIGURES 14–18. Pergalumna pietertheroni sp. nov., adult: 14—dorsal view; 15—ventral view (gnathosoma, legs and right pteromorph not shown); 16—subcapitulum, ventral view; 17—palp, left, paraxial view; 18—chelicera, left, paraxial view. Scale bar 50 µm (14, 15), 20 µm (16–18)." httpUri="https://zenodo.org/record/2618311/files/figure.png" pageId="3" pageNumber="534">Figs 15</figureCitation>
|
||
,
|
||
<figureCitation id="13102A403367FFD51C335C47FDA8FCDD" box="[518,590,770,794]" captionStart="FIGURES 19–21" captionStartId="8.[151,264,1750,1773]" captionTargetBox="[218,1339,296,1625]" captionTargetId="figure@8.[150,1436,225,1709]" captionTargetPageId="8" captionText="FIGURES 19–21. Pergalumna pietertheroni sp. nov., adult: 19—anterior part of body, lateral view (gnathosoma, pteromorph and legs not shown); 20—posterior part of body, lateral view; 21—posterior view. Scale bar 50 µm." httpUri="https://zenodo.org/record/2618313/files/figure.png" pageId="3" pageNumber="534">19–21</figureCitation>
|
||
,
|
||
<figureCitation id="13102A403367FFD51C6E5C47FD9CFCDD" box="[603,634,770,794]" captionStart="FIGURES 38–45" captionStartId="13.[151,264,1836,1859]" captionTargetBox="[150,1436,193,1806]" captionTargetId="figure@13.[150,1436,193,1806]" captionTargetPageId="13" captionText="FIGURES 38–45. Galumnoidea, adults, microscope images: 38—Pergalumna hugocoetzeeae sp. nov., prodorsal sculpturing and rostrum, dorsoanterior view; 39—Pergalumna hugocoetzeeae sp. nov., dorsosejugal suture; 40—Pergalumna hugocoetzeeae sp. nov., dorsophragma and sejugal porose area; 41—Pergalumna pietertheroni sp. nov., granulate sculpturing in anogenital region; 42—Pergalumna pietertheroni sp. nov., dorsophragma and sejugal porose area; 43—Galumnella paraokinawana sp. nov., foveolate ornamentation in epimeral region; 44—Galumnella paraokinawana sp. nov., reticulate ornamentation between genital and anal apertures; 45—Galumnella paraokinawana sp. nov., claw of leg tarsus II." httpUri="https://zenodo.org/record/2618323/files/figure.png" pageId="3" pageNumber="534">41</figureCitation>
|
||
). Six pairs of genital, one pair of aggenital, two pairs of anal and three pairs of adanal setae similar in length (4–6), setiform, thin, smooth. Anterior edge of genital plates with two setae. Aggenital setae equally distanced from genital and anal plates. Adanal lyrifissures located close and parallel to anal plates. Adanal setae
|
||
<emphasis id="B95FEAD73367FFD51FB55C28FE7AFC41" box="[384,412,877,902]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51FA95C39FE42FC4D" attach="left" box="[412,420,892,906]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
posterior,
|
||
<emphasis id="B95FEAD73367FFD51C285C28FDDFFC41" box="[541,569,877,902]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51C0C5C39FDA7FC4D" attach="left" box="[569,577,892,906]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
posterolateral,
|
||
<emphasis id="B95FEAD73367FFD51CC55C28FCEAFC41" box="[752,780,877,902]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51D395C39FCF2FC4D" attach="left" box="[780,788,892,906]" fontSize="6" pageId="3" pageNumber="534">3</subScript>
|
||
lateral to anal aperture;
|
||
<emphasis id="B95FEAD73367FFD51A1F5C28FBA0FC41" box="[1066,1094,877,902]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51A735C39FBA8FC4D" attach="left" box="[1094,1102,892,906]" fontSize="6" pageId="3" pageNumber="534">3</subScript>
|
||
anterolateral to
|
||
<emphasis id="B95FEAD73367FFD51B325C28FACDFC41" box="[1287,1323,877,902]" italics="true" pageId="3" pageNumber="534">iad</emphasis>
|
||
. Distance
|
||
<emphasis id="B95FEAD73367FFD51EA25CD1FF55FC6A" box="[151,179,916,941]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51E865CE6FF5DFC76" attach="left" box="[179,187,931,945]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
–
|
||
<emphasis id="B95FEAD73367FFD51EFD5CD1FF02FC6A" box="[200,228,916,941]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51ED15CE6FF0AFC76" attach="left" box="[228,236,931,945]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
shorter than
|
||
<emphasis id="B95FEAD73367FFD51F4A5CD1FE7DFC6A" box="[383,411,916,941]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51FAE5CE6FE45FC76" attach="left" box="[411,419,931,945]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
–
|
||
<emphasis id="B95FEAD73367FFD51F845CD1FE2BFC6A" box="[433,461,916,941]" italics="true" pageId="3" pageNumber="534">ad</emphasis>
|
||
<subScript id="17AF34803367FFD51FF85CE6FE33FC76" attach="left" box="[461,469,931,945]" fontSize="6" pageId="3" pageNumber="534">3</subScript>
|
||
. Postanal porose area elongate oval (8–12 × 4–6).
|
||
</paragraph>
|
||
<paragraph id="8B9436C53367FFD51EF25CF9FD05FBA1" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25CF9FF1AFC14" box="[199,252,956,979]" italics="true" pageId="3" pageNumber="534">Legs</emphasis>
|
||
(
|
||
<figureCitation id="13102A403367FFD51F3E5CFEFE76FC13" box="[267,400,955,980]" captionStart="FIGURES 22–25" captionStartId="9.[151,264,1949,1972]" captionTargetBox="[210,1404,228,1877]" captionTargetId="figure@9.[150,1436,193,1928]" captionTargetPageId="9" captionText="FIGURES 22–25. Pergalumna pietertheroni sp. nov., adult: 22—leg I, right, antiaxial view; 23—trochanter, femur and genu of leg II, right, ventroantiaxial view; 24—trochanter, femur and genu of leg III, left, antiaxial view; 25—leg IV, left, antiaxial view. Scale bar 20 µm." httpUri="https://zenodo.org/record/2618315/files/figure.png" pageId="3" pageNumber="534">Figs 22–25</figureCitation>
|
||
). Median claw with basal tooth, distinctly thicker than laterals, all slightly barbed on dorsal side. Porose areas on all femora and on trochanters III, IV poorly visible. Formulas of leg setation and solenidia: I (
|
||
<date id="FF9510053367FFD51EAA5B46FEC8FBDC" box="[159,302,1027,1051]" pageId="3" pageNumber="534" value="1920-03-01" valueMax="1920-04-04" valueMin="1920-03-01">1–4–3–4–20</date>
|
||
) [1–2–2], II (
|
||
<date id="FF9510053367FFD51FDC5B46FD9EFBDB" box="[489,632,1027,1052]" pageId="3" pageNumber="534" value="2015-03-01" valueMax="2015-04-04" valueMin="2015-03-01">1–4–3–4–15</date>
|
||
) [1–1–2], III (
|
||
<date id="FF9510053367FFD51D085B46FC2DFBDB" box="[829,971,1027,1052]" pageId="3" pageNumber="534" value="2015-01-01" valueMax="2015-03-02" valueMin="2015-01-01">1–2–1–3–15</date>
|
||
) [1–1–0], IV (
|
||
<date id="FF9510053367FFD51AA75B46FAC6FBDC" box="[1170,1312,1027,1051]" pageId="3" pageNumber="534" value="2012-02-01" valueMax="2012-03-02" valueMin="2012-02-01">1–2–2–3–12</date>
|
||
) [0–1–0]; homologies of setae and solenidia indicated in
|
||
<tableCitation id="C6A9037E3367FFD51C9C5B62FD1BFBF8" box="[681,765,1063,1088]" captionStart="TABLE 1" captionStartId="14.[151,239,917,940]" captionText="TABLE 1. Leg setation and solenidia of adult Pergalumna hugocoetzeeae sp. nov., Pergalumna pietertheroni sp. nov. and Galumnella paraokinawana sp. nov." pageId="3" pageNumber="534">Table 1</tableCitation>
|
||
. Famulus on tarsi I inserted anterior to solenidia ω
|
||
<subScript id="17AF34803367FFD51B0E5B73FAA5FB83" attach="left" box="[1339,1347,1078,1092]" fontSize="6" pageId="3" pageNumber="534">1</subScript>
|
||
and ω
|
||
<subScript id="17AF34803367FFD51BB85B73FA73FB83" attach="left" box="[1421,1429,1078,1092]" fontSize="6" pageId="3" pageNumber="534">2</subScript>
|
||
. Solenidion on tibiae IV inserted in their middle part.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331654E3367FFD51EF25B37FD95FB69" pageId="3" pageNumber="534" type="materials_examined">
|
||
<paragraph id="8B9436C53367FFD51EF25B37FD95FB69" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25B37FE63FB4C" bold="true" box="[199,389,1138,1163]" pageId="3" pageNumber="534">Type deposition</emphasis>
|
||
. The
|
||
<typeStatus id="549088673367FFD51FFD5B37FDCCFB4C" box="[456,554,1138,1163]" pageId="3" pageNumber="534" type="holotype">holotype</typeStatus>
|
||
and two
|
||
<typeStatus id="549088673367FFD51CA05B36FCE6FB4C" box="[661,768,1139,1163]" pageId="3" pageNumber="534" type="paratype">paratypes</typeStatus>
|
||
are deposited in the collection of SMNH.
|
||
<materialsCitation id="3B433C983367FFD51AD25B37FD88FB69" ID-GBIF-Occurrence="2147525286" collectionCode="TSUMZ" pageId="3" pageNumber="534" specimenCount="1" typeStatus="paratype">
|
||
Three
|
||
<typeStatus id="549088673367FFD51B055B36FA7AFB4C" box="[1328,1436,1139,1163]" pageId="3" pageNumber="534" type="paratype">paratypes</typeStatus>
|
||
are deposited in the collection of
|
||
<collectionCode id="ED3AAE003367FFD51C395BD3FD88FB69" box="[524,622,1174,1198]" pageId="3" pageNumber="534">TSUMZ</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331654E3367FFD51EF25BFFFCB0FB31" pageId="3" pageNumber="534" type="etymology">
|
||
<paragraph id="8B9436C53367FFD51EF25BFFFCB0FB31" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25BFFFEA8FB14" bold="true" box="[199,334,1210,1235]" pageId="3" pageNumber="534">Etymology.</emphasis>
|
||
The specific name is dedicated to our friend and colleague, well-known acarologist, Dr. Pieter Theron (North-West University, Potchefstroom,
|
||
<collectingCountry id="F33C76553367FFD51C805B98FCACFB31" box="[693,842,1245,1270]" name="South Africa" pageId="3" pageNumber="534">South Africa</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C331654E3367FFD51EF25A47FBE4FA6D" pageId="3" pageNumber="534" type="discussion">
|
||
<paragraph id="8B9436C53367FFD51EF25A47FBE4FA6D" blockId="3.[151,1437,151,1451]" pageId="3" pageNumber="534">
|
||
<emphasis id="B95FEAD73367FFD51EF25A47FEDDFADC" bold="true" box="[199,315,1282,1307]" pageId="3" pageNumber="534">Remarks.</emphasis>
|
||
In having dorsosejugal suture, three pairs of rounded notogastral porose areas, setiform bothridial setae and short rostral, lamellar and interlamellar setae,
|
||
<taxonomicName id="4C2B4D463367FFD51D305A60FBC0FAF9" authority="Ermilov & Khaustov, 2019" authorityName="Ermilov & Khaustov" authorityYear="2019" box="[773,1062,1317,1342]" class="Arachnida" family="Galumnidae" genus="Pergalumna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="pietertheroni" status="sp. nov.">
|
||
<emphasis id="B95FEAD73367FFD51D305A60FBC0FAF9" box="[773,1062,1317,1342]" italics="true" pageId="3" pageNumber="534">Pergalumna pietertheroni</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B95FEAD73367FFD51A195A60FB60FAF9" bold="true" box="[1068,1158,1317,1342]" pageId="3" pageNumber="534">
|
||
<taxonomicNameLabel id="A26C57AC3367FFD51A195A60FB60FAF9" box="[1068,1158,1317,1342]" pageId="3" pageNumber="534" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically most similar to
|
||
<taxonomicName id="4C2B4D463367FFD51F3D5A0FFD5AFAA5" authority="Mahunka, 1984" authorityName="Mahunka" authorityYear="1984" box="[264,700,1354,1379]" class="Arachnida" family="Galumnidae" genus="Pergalumna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="3" pageNumber="534" phylum="Arthropoda" rank="species" species="tanzanica">
|
||
<emphasis id="B95FEAD73367FFD51F3D5A0FFDE0FAA5" box="[264,518,1354,1379]" italics="true" pageId="3" pageNumber="534">Pergalumna tanzanica</emphasis>
|
||
Mahunka, 1984
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F33C76553367FFD51D345A0FFC83FAA4" box="[769,869,1354,1379]" name="Tanzania" pageId="3" pageNumber="534">Tanzania</collectingCountry>
|
||
(see
|
||
<bibRefCitation id="EFBA4B343367FFD51D955A0FFBB1FAA4" author="Mahunka, S." box="[928,1111,1354,1379]" pageId="3" pageNumber="534" pagination="87 - 136" refId="ref7918" refString="Mahunka, S. (1984 b) Oribatids of the Eastern Part of the Ethiopian Region (Acari). V. Acta Zoologica Hungarica, 30 (1 - 2), 87 - 136." type="journal article" year="1984">
|
||
Mahunka 1984
|
||
<emphasis id="B95FEAD73367FFD51A7C5A0FFBB1FAA4" box="[1097,1111,1354,1379]" italics="true" pageId="3" pageNumber="534">b</emphasis>
|
||
</bibRefCitation>
|
||
), but clearly differs from the latter species by the morphology of bothridial setae (smooth versus heavily, unilaterally ciliate) and position of porose areas
|
||
<emphasis id="B95FEAD73367FFD51F1C5AD6FEAEFA6D" box="[297,328,1427,1450]" italics="true" pageId="3" pageNumber="534">Aa</emphasis>
|
||
(located closer to
|
||
<emphasis id="B95FEAD73367FFD51C225AD7FDCBFA6C" box="[535,557,1426,1451]" italics="true" pageId="3" pageNumber="534">la</emphasis>
|
||
versus equally distanced from
|
||
<emphasis id="B95FEAD73367FFD51DBF5AD7FC46FA6C" box="[906,928,1426,1451]" italics="true" pageId="3" pageNumber="534">la</emphasis>
|
||
and
|
||
<emphasis id="B95FEAD73367FFD51DE35AD7FC14FA6C" box="[982,1010,1426,1451]" italics="true" pageId="3" pageNumber="534">lm</emphasis>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |