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<document id="3E50DB06E1B40E2D2EA470C5B70124BE" ID-CLB-Dataset="7224" ID-DOI="10.11646/zootaxa.4958.1.23" ID-GBIF-Dataset="5555ed0d-e35c-40fe-8752-208b91837b07" ID-ISSN="1175-5326" ID-Zenodo-Dep="4692086" ID-ZooBank="828DB95F-03D4-437B-B360-A28CDF6B409D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1618465144353" checkinUser="plazi" docAuthor="Swanson, Daniel R." docDate="2021" docId="038287A1B939E50B3AC48D1AFB96FA50" docLanguage="en" docName="zootaxa.4958.1.23.pdf" docOrigin="Zootaxa 4958 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heza nebrias Swanson 2021, sp. nov." docType="treatment" docUuid="AD56FDC8-34E4-4298-BEFA-F87E35B59835" docUuidSource="ZooBank" docVersion="8" lastPageNumber="371" masterDocId="FFBBFFD9B93DE50E3A538F40FFD4FFCD" masterDocTitle="Four new species of Heza Amyot &amp; Audinet-Serville (Heteroptera: Reduviidae Harpactorinae) from the Neotropics" masterLastPageNumber="379" masterPageNumber="366" pageNumber="370" updateTime="1698936887257" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F390F2228C51595258D73AE54733DE1E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="BC3AB7B1BE0CA4971907E6EABE508F2A">Four new species of Heza Amyot &amp; Audinet-Serville (Heteroptera: Reduviidae Harpactorinae) from the Neotropics</mods:title>
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<mods:namePart id="33C650D60E8842AFE81502BE2CF6B653">Swanson, Daniel R.</mods:namePart>
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<mods:title id="769025807728336C67168F5275E9CA9A">Zootaxa</mods:title>
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<mods:date id="83D064E88090E3D16F516FFCB1CDE937">2021</mods:date>
<mods:detail id="BD90477C60381C7D78529BDE43A652ED" type="pubDate">
<mods:number id="ACDDB48B93C08D99CE29012318961B31">2021-04-14</mods:number>
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<mods:number id="B4E286F07A35E6CFA035CF051EAD0BB4">4958</mods:number>
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<mods:classification id="B35ECBCBF742F07B9BCFB3F56D970421">journal article</mods:classification>
<mods:identifier id="9B8CF0831FA82F309BF931D13420DC3C" type="CLB-Dataset">7224</mods:identifier>
<mods:identifier id="FE0911C7DF876B247A7A3C8D54AA8092" type="DOI">10.11646/zootaxa.4958.1.23</mods:identifier>
<mods:identifier id="3DAEF32EA5B3FF73ABFB6F3E94ADE03A" type="GBIF-Dataset">5555ed0d-e35c-40fe-8752-208b91837b07</mods:identifier>
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<mods:identifier id="1E0202D8798977B059078D67981DB06C" type="Zenodo-Dep">4692086</mods:identifier>
<mods:identifier id="0E391C9AD4CD69357A6103267C7AA90D" type="ZooBank">828DB95F-03D4-437B-B360-A28CDF6B409D</mods:identifier>
</mods:mods>
<treatment id="038287A1B939E50B3AC48D1AFB96FA50" ID-DOI="http://doi.org/10.5281/zenodo.4710996" ID-GBIF-Taxon="181531341" ID-Zenodo-Dep="4710996" LSID="urn:lsid:zoobank.org:act:AD56FDC8-34E4-4298-BEFA-F87E35B59835" httpUri="http://treatment.plazi.org/id/038287A1B939E50B3AC48D1AFB96FA50" lastPageId="5" lastPageNumber="371" pageId="4" pageNumber="370">
<subSubSection id="C331653CB939E50A3AC48D1AFC1CFD5A" pageId="4" pageNumber="370" type="nomenclature">
<paragraph id="8B9436B7B939E50A3AC48D1AFE4FFDBE" blockId="4.[151,968,601,697]" box="[151,411,601,628]" pageId="4" pageNumber="370">
<heading id="D0DC81DBB939E50A3AC48D1AFE4FFDBE" bold="true" box="[151,411,601,628]" fontSize="11" level="1" pageId="4" pageNumber="370" reason="1">
<emphasis id="B95FEAA5B939E50A3AC48D1AFE4FFDBE" bold="true" box="[151,411,601,628]" pageId="4" pageNumber="370">
<taxonomicName id="4C2B4D34B939E50A3AC48D1AFEE2FDB9" authority="Swanson, 2021" authorityName="Swanson" authorityYear="2021" box="[151,310,601,628]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="nebrias" status="sp. nov.">
<emphasis id="B95FEAA5B939E50A3AC48D1AFEE2FDB9" bold="true" box="[151,310,601,628]" italics="true" pageId="4" pageNumber="370">Heza nebrias</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26C57DEB939E50A3B6E8D19FE4FFDBE" box="[317,411,601,627]" pageId="4" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9436B7B939E50A3AC48D3CFC1CFD5A" blockId="4.[151,968,601,697]" box="[151,968,636,663]" pageId="4" pageNumber="370">
<uri id="FFBA3AB5B939E50A3AC48D3CFC1CFD5A" box="[151,968,636,663]" pageId="4" pageNumber="370">
urn:lsid:zoobank.org:act:
<uuid id="FF8D0C62B939E50A3BF58D3CFC1CFD5A" box="[422,968,636,663]" pageId="4" pageNumber="370">AD56FDC8-34E4-4298-BEFA-F87E35B59835</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C331653CB939E50A3AC48DDFFE5BFD74" box="[151,399,671,697]" pageId="4" pageNumber="370" type="description">
<paragraph id="8B9436B7B939E50A3AC48DDFFE5BFD74" blockId="4.[151,968,601,697]" box="[151,399,671,697]" pageId="4" pageNumber="370">
(
<figureCitation id="13102A32B939E50A3ACC8DDFFF3BFD74" box="[159,239,671,697]" captionStart="FIGURES 1–4" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[239,1346,193,1907]" captionTargetId="figure-21@3.[229,1358,179,1917]" captionTargetPageId="3" captionText="FIGURES 1–4. New species of Heza, dorsal habitus: (1) Heza graziae sp. nov., male holotype; (2) Heza nebrias sp. nov., male holotype; (3) Heza phthinica sp. nov., male holotype; (C) Heza viridipes sp. nov., male holotype. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4692088" httpUri="https://zenodo.org/record/4692088/files/figure.png" pageId="4" pageNumber="370">Figs. 2</figureCitation>
,
<figureCitation id="13102A32B939E50A3AA88DDFFEDFFD74" box="[251,267,671,697]" captionStart="FIGURES 5–8" captionStartId="6.[151,264,1943,1967]" captionTargetBox="[205,1365,181,1919]" captionTargetId="figure-15@6.[205,1382,181,1919]" captionTargetPageId="6" captionText="FIGURES 5–8. New species of Heza, lateral habitus: (5) Heza graziae sp. nov., male holotype; (6) Heza nebrias sp. nov., male holotype; (7) Heza phthinica sp. nov., male holotype; (8) Heza viridipes sp. nov., male holotype. All show left side, except H. phthinica reflected." figureDoi="http://doi.org/10.5281/zenodo.4692092" httpUri="https://zenodo.org/record/4692092/files/figure.png" pageId="4" pageNumber="370">6</figureCitation>
,
<figureCitation id="13102A32B939E50A3B448DDFFE8AFD74" box="[279,350,671,697]" captionStart="FIGURES 9–16" captionStartId="9.[151,264,1910,1934]" captionTargetBox="[231,1356,199,1886]" captionTargetId="figure-21@9.[231,1356,199,1886]" captionTargetPageId="9" captionText="FIGURES 9–16. New species of Heza, pygophore: (9) Heza graziae sp. nov., caudal view; (10) Heza graziae sp. nov., lateral view; (11) Heza nebrias sp. nov., caudal view; (12) Heza nebrias sp. nov., lateral view; (13) Heza phthinica sp. nov., caudal view; (14) Heza phthinica sp. nov., lateral view; (15) Heza viridipes sp. nov., caudal view; (16) Heza viridipes sp. nov., lateral view." figureDoi="http://doi.org/10.5281/zenodo.4692096" httpUri="https://zenodo.org/record/4692096/files/figure.png" pageId="4" pageNumber="370">11, 12</figureCitation>
,
<figureCitation id="13102A32B939E50A3B398DDFFE53FD74" box="[362,391,671,697]" captionStart="FIGURES 17–20" captionStartId="10.[151,264,1658,1682]" captionTargetBox="[151,1436,885,1635]" captionTargetId="figure-322@10.[151,1436,885,1635]" captionTargetPageId="10" captionText="FIGURES 17–20. New species of Heza, median process of pygophore (with line drawing): (17) Heza graziae sp. nov.; (18) Heza nebrias sp. nov.; (19) Heza phthinica sp. nov.; (20) Heza viridipes sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4692100" httpUri="https://zenodo.org/record/4692100/files/figure.png" pageId="4" pageNumber="370">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C331653CB939E50A3AC48DA4FCB9FC8E" pageId="4" pageNumber="370" type="materials_examined">
<paragraph id="8B9436B7B939E50A3AC48DA4FCB9FC8E" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3AC48DA4FE91FD33" bold="true" box="[151,325,740,766]" pageId="4" pageNumber="370">Type material:</emphasis>
<materialsCitation id="3B433CEAB939E50A3B188DA4FA92FD33" ID-GBIF-Occurrence="3083764313" box="[331,1350,740,767]" collectingDate="1982-06-20" collectingDateMax="1996-04-23" collectingDateMin="1982-06-20" collectionCode="OSUC" collectorName="Roy W. Rings" country="Peru" elevation="150" location="Posada" municipality="Tingo Maria" pageId="4" pageNumber="370" specimenCode="OSUC 651120" specimenCount="2" specimenCount-male="2" stateProvince="Rondonia" typeStatus="holotype">
<typeStatus id="54908815B939E50A3B188DA4FE36FD33" box="[331,482,740,766]" pageId="4" pageNumber="370" type="holotype">HOLOTYPE</typeStatus>
,
<specimenCount id="9D2DFD3EB939E50A3BB88DA5FDD6FD32" box="[491,514,741,767]" pageId="4" pageNumber="370" type="male">♂</specimenCount>
,
<collectingCountry id="F33C7627B939E50A385F8DA5FD8CFD32" box="[524,600,741,767]" name="Peru" pageId="4" pageNumber="370">PERU</collectingCountry>
:
<collectingMunicipality id="6BF0ACCDB939E50A38338DA4FD39FD33" box="[608,749,740,766]" pageId="4" pageNumber="370">Tingo Maria</collectingMunicipality>
,
<date id="FF951077B939E50A38A58DA4FC5FFD33" box="[758,907,740,767]" pageId="4" pageNumber="370" value="1982-06-20">
<collectingDate id="EFD1E99FB939E50A38A58DA4FC5FFD33" box="[758,907,740,767]" pageId="4" pageNumber="370" value="1982-06-20">20 June 1982</collectingDate>
</date>
,
<collectorName id="26DE5361B939E50A39C78DA5FBFBFD33" box="[916,1071,740,767]" pageId="4" pageNumber="370">Roy W. Rings</collectorName>
,
<specimenCode id="DB8D9ECCB939E50A3E6A8DA4FB08FD32" box="[1081,1244,740,767]" collectionCode="OSUC" country="USA" httpUri="http://grbio.org/cool/x820-3v0f" name="Oregon State University" pageId="4" pageNumber="370">OSUC 651120</specimenCode>
(
<collectionCode id="ED3AAE72B939E50A3EBA8DA4FAEEFD33" box="[1257,1338,740,766]" country="USA" httpUri="http://grbio.org/cool/x820-3v0f" name="Oregon State University" pageId="4" pageNumber="370">OSUC</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B433CEAB939E50A3F188DA4FCB9FC8E" ID-GBIF-Occurrence="3083764305" collectingDate="1982-06-20" collectingDateMax="1996-04-23" collectingDateMin="1982-06-20" collectionCode="MSUC" collectorName="F. W. Stehr &amp; M. C. Nielsen" country="Peru" elevation="150" location="Posada" municipality="Tingo Maria" pageId="4" pageNumber="370" specimenCode="OSUC 651120" specimenCount="2" specimenCount-male="2" stateProvince="Rondonia" typeStatus="holotype">
<collectionCode id="ED3AAE72B939E50A3F188DA4FF34FCEC" pageId="4" pageNumber="370">PARATYPE</collectionCode>
,
<specimenCount id="9D2DFD3EB939E50A3AB98C47FED5FCEC" box="[234,257,775,801]" pageId="4" pageNumber="370" type="male">♂</specimenCount>
,
<collectingCountry id="F33C7627B939E50A3B5F8C47FEA2FCEC" box="[268,374,775,801]" name="Brazil" pageId="4" pageNumber="370">BRAZIL</collectingCountry>
:
<collectingRegion id="49EFF855B939E50A3B2D8C47FE3AFCEC" box="[382,494,775,801]" country="Brazil" name="Rondonia" pageId="4" pageNumber="370">Rondonia</collectingRegion>
,
<location id="8EF4606CB939E50A3BAB8C47FD9EFCEC" LSID="urn:lsid:plazi:treatment:038287A1B939E50B3AC48D1AFB96FA50:8EF4606CB939E50A3BAB8C47FD9EFCEC" box="[504,586,775,801]" country="Peru" municipality="Tingo Maria" name="Posada" pageId="4" pageNumber="370" stateProvince="Rondonia">Posada</location>
,
<collectorName id="26DE5361B939E50A38078C47FCCAFCEC" box="[596,798,775,801]" pageId="4" pageNumber="370">Rancho Grande, S</collectorName>
10-17’ -51.7” W62-52’ 7.3”, ca
<quantity id="4CD39B52B939E50A3E288C47FB15FCEC" box="[1147,1217,775,801]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="4" pageNumber="370" unit="km" value="60.0">60 km</quantity>
<collectorName id="26DE5361B939E50A3E958C47FA8BFCEC" box="[1222,1375,775,801]" pageId="4" pageNumber="370">S. Ariquemes</collectorName>
, elev.
<quantity id="4CD39B52B939E50A3AC48C69FF34FC89" box="[151,224,809,836]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="370" unit="m" value="150.0">
<elevation id="0006D184B939E50A3AC48C69FF34FC89" box="[151,224,809,836]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="370" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="FF951077B939E50A3ABF8C69FE61FC8E" box="[236,437,809,836]" pageId="4" pageNumber="370" value="1996-04-15" valueMax="1996-04-23" valueMin="1996-04-15">
<collectingDate id="EFD1E99FB939E50A3ABF8C69FE61FC8E" box="[236,437,809,836]" pageId="4" pageNumber="370" value="1996-04-15" valueMax="1996-04-23" valueMin="1996-04-15">15–23 April 1996</collectingDate>
</date>
,
<collectorName id="26DE5361B939E50A3B938C6AFDEFFC8E" box="[448,571,809,836]" pageId="4" pageNumber="370">F. W. Stehr</collectorName>
&amp;
<collectorName id="26DE5361B939E50A380C8C6AFD29FC8E" box="[607,765,809,836]" pageId="4" pageNumber="370">M. C. Nielsen</collectorName>
(
<collectionCode id="ED3AAE72B939E50A39588C69FCB5FC8E" box="[779,865,809,835]" pageId="4" pageNumber="370">MSUC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C331653CB939E50A3A948C0CFA49FC44" pageId="4" pageNumber="370" type="etymology">
<paragraph id="8B9436B7B939E50A3A948C0CFA49FC44" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948C0CFE86FCAB" bold="true" box="[199,338,844,870]" pageId="4" pageNumber="370">Etymology:</emphasis>
The specific epithet comes from the Greek adjective νεβρίας, -ου (Latinized
<taxonomicName id="4C2B4D34B939E50A3EF08C0CFB23FCAB" authorityName="Swanson" authorityYear="2021" box="[1187,1271,844,870]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="nebrias">
<emphasis id="B95FEAA5B939E50A3EF08C0CFB23FCAB" box="[1187,1271,844,870]" italics="true" pageId="4" pageNumber="370">nebrias</emphasis>
</taxonomicName>
,
<emphasis id="B95FEAA5B939E50A3F578C0CFA86FCAB" box="[1284,1362,844,870]" italics="true" pageId="4" pageNumber="370">nebrou</emphasis>
) ‘dappled like a fawn’ and references the tomentose spotting of the hemelytral base and to a lesser extent, the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C331653CB939E50A3A948CD1FD74FBFB" pageId="4" pageNumber="370" type="diagnosis">
<paragraph id="8B9436B7B939E50A3A948CD1FD74FBFB" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948CD1FE90FC66" bold="true" box="[199,324,913,939]" pageId="4" pageNumber="370">Diagnosis:</emphasis>
Separated from other species of
<taxonomicName id="4C2B4D34B939E50A38EE8CD2FD23FC66" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[701,759,914,939]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A38EE8CD2FD23FC66" box="[701,759,914,939]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
by the following combination of characters: only first connexival segment (=second tergite) armed posterolaterally with spine or tubercle; discal armature of anterior pronotal lobe medium-sized, cylindrical, blunt; corium and clavus basally with large patches of dense tomenta; anterior pronotal lobe lacking dense whitish tomenta and that of posterior pronotal lobe limited to eight small spots; and median process of pygophore with distinct apical fold.
</paragraph>
</subSubSection>
<subSubSection id="C331653CB939E50B3A948B7EFE71FC68" lastPageId="5" lastPageNumber="371" pageId="4" pageNumber="370" type="description">
<paragraph id="8B9436B7B939E50A3A948B7EFEB5FB0C" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948B7EFD0DFB95" bold="true" box="[199,729,1086,1113]" pageId="4" pageNumber="370">
Description: Macropterous male (
<typeStatus id="54908815B939E50A38338B7EFD18FB95" box="[608,716,1086,1112]" pageId="4" pageNumber="370" type="holotype">holotype</typeStatus>
).
</emphasis>
<emphasis id="B95FEAA5B939E50A38B18B7FFCB2FB94" box="[738,870,1087,1113]" italics="true" pageId="4" pageNumber="370">Coloration:</emphasis>
Rusty red-brown, except apices of head and rostrum, most of profemur to apex of foreleg, meso- and metatrochanter to apex of respective leg, and venter pale yellowish. Setal patches of basal half of hemelytra whitish. Veins of hemelytral membrane black. Pleura and venter suffused with red.
</paragraph>
<paragraph id="8B9436B7B939E50A3A948B8AFE19FAE4" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948B8AFE50FB2E" box="[199,388,1225,1252]" italics="true" pageId="4" pageNumber="370">Structure: Head.</emphasis>
Postantennal spines medium, robust, straight, sharp, otherwise head as per description under
<taxonomicName id="4C2B4D34B939E50A3AC48BADFF05FACB" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[151,209,1261,1286]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A3AC48BADFF05FACB" box="[151,209,1261,1286]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
. Antennae with scape largely glabrous, other antennomeres missing in
<typeStatus id="54908815B939E50A39A78BACFB8EFACB" box="[1012,1114,1260,1286]" pageId="4" pageNumber="370" type="holotype">holotype</typeStatus>
. Eyes, ocelli, and rostrum as per description under
<taxonomicName id="4C2B4D34B939E50A3BDF8A4FFE12FAE5" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[396,454,1295,1320]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A3BDF8A4FFE12FAE5" box="[396,454,1295,1320]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436B7B939E50A3A948A71FB08F94E" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948A71FEF4FA86" box="[199,288,1329,1355]" italics="true" pageId="4" pageNumber="370">Thorax.</emphasis>
Pronotum with anterior pronotal lobe covered with short appressed pale setae but with a few glabrous channels, with collar armed with obliquely-directed, somewhat sharp conical tubercle, disc armed with two medium, robust, apically-blunt spines. Posterior pronotal lobe disc noticeably convex, with faint, short, paired carinae emanating from transverse constriction behind tubercles of anterior pronotal lobe, covered with sparse long erect setae (more conspicuous laterally) and with paired submedial and paired sublateral dense golden setal spots near transverse constriction, disc armed with two medium, thin, essentially erect, sharp spines constricted at middle, humeral anglei with medium, sharp spine over humerus directed laterally and somewhat dorsad, posterior margin slightly convex in front of scutellum and sinuate between humeri and posterior angles, posterior angles roundly obtuse. Scutellum with short appressed white setae and sparse erect longish golden setae. Pleura and sterna as per description under
<taxonomicName id="4C2B4D34B939E50A3AC4892AFF1BF94E" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[151,207,1642,1667]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A3AC4892AFF1BF94E" box="[151,207,1642,1667]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
. Hemelytra with corium covered with dense, white, waxy setal patches and spots in basal half.
</paragraph>
<paragraph id="8B9436B7B939E50A3A9489CDFC90F904" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A9489CDFED0F96B" box="[199,260,1677,1702]" italics="true" pageId="4" pageNumber="370">Legs.</emphasis>
Forelegs with profemur straight, largely glabrous on dorsal and lateral surfaces. Protibia slightly curved apically. Middle and hind legs as per description under
<taxonomicName id="4C2B4D34B939E50A395089EFFCE9F905" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[771,829,1711,1736]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A395089EFFCE9F905" box="[771,829,1711,1736]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436B7B939E50A3A948991FD4DF921" blockId="4.[151,1437,740,1981]" box="[199,665,1745,1772]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A948991FEEDF926" box="[199,313,1745,1771]" italics="true" pageId="4" pageNumber="370">Abdomen.</emphasis>
As per description under
<taxonomicName id="4C2B4D34B939E50A380B8992FD46F926" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[600,658,1746,1771]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B939E50A380B8992FD46F926" box="[600,658,1746,1771]" italics="true" pageId="4" pageNumber="370">Heza</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436B7B939E50A3A9489B4FD9DF870" blockId="4.[151,1437,740,1981]" pageId="4" pageNumber="370">
<emphasis id="B95FEAA5B939E50A3A9489B4FE49F8C3" box="[199,413,1780,1806]" italics="true" pageId="4" pageNumber="370">External genitalia.</emphasis>
Pygophore with integument smooth, essentially glabrous except sparse moderately long erect setae, with posterior margin setose. Median apical process robust, erect, approximately 1.5 times taller than apical width, lateral margins converging from base to middle and diverging slightly from middle to apex, apex strongly convex, margin robustly “hooked” or folded caudad, fold wider than shaft. Parameres moderately-sized, almost reaching median process at rest, more or less cylindrical but slightly sinuate at middle in dorsal plane, densely setose on dorsal margin and near apex.
</paragraph>
<paragraph id="8B9436B7B938E50B3A948FD7FBB7FD3C" blockId="5.[151,1437,150,1437]" pageId="5" pageNumber="371">
<emphasis id="B95FEAA5B938E50B3A948FD7FE04FF7C" box="[199,464,151,177]" italics="true" pageId="5" pageNumber="371">Measurements (in mm).</emphasis>
Total length (apex of head to apex of abdomen): HT: 16.4, PT: 15.7; total length (apex of head to apex of hemelytra): HT: 16.9, PT: 16.8; head (+ neck) length: HT: 2.6, PT: (2.1 + 0.5); head width (across eyes): HT: 1.4, PT: 1.4; anteocular length: HT: 0.7, PT: 0.7; postocular (+ neck) length: HT: 1.3, PT: (0.8 + 0.5); scape length: HT: 6.5, PT: 7.1; pedicel length: HT:?, PT: 1.5; basiflagellum length: HT:?, PT: 8.0; distiflagellum length: HT:?, PT: 1.8; antennal segment ratio: approx. 1.0: 0.2: 1.1: 0.3; eye length: HT: 0.6, PT: 0.6; eye width: HT: 0.3, PT: 0.3; rostral segment 1 length: HT: 1.6, PT: 1.6; rostral segment 2 length: HT: 1.1, PT: 1.2; rostral segment 3 length: HT: 0.5, PT: 0.4; rostral segment ratio: approx. 1.0: 0.7: 0.3; pronotum length: HT: 4.1, PT: 3.8; pronotum width (across humeri, without spines): HT: 4.2, PT: 3.7; pronotum width (across humeri, with spines): HT: 5.4, PT: 4.9; anterior pronotal lobe length: HT: 1.5, PT: 1.4; posterior pronotal lobe length: HT: 2.6, PT: 2.4; scutellum length: HT: 1.1, PT: 1.1; scutellum width (at base): HT: 1.3, PT: 1.4; hemelytra length: HT: 11.1, PT: 11.1; procoxa length: HT: 0.9, PT: 0.7; protrochanter length: HT: 0.9, PT: 0.8; profemur length: HT: 6.0, PT: 6.1; protibia length: HT: 5.5, PT: 5.6; protarsus length: HT: 0.7, PT: 0.7; protarsal segment ratio: approx. 1.0: 2.0: 2.5; mesocoxa length: HT: 0.8, PT: 0.7; mesotrochanter length: HT: 0.7, PT: 0.8; mesofemur length: HT: 4.7, PT: 5.0; mesotibia length: HT: 5.0, PT: 5.2; mesotarsus length: HT: 0.6, PT: 0.6; mesotarsal segment ratio: approx. 1.0: 2.0: 2.5; metacoxa length: HT: 1.0, PT: 0.7; metatrochanter length: HT: 0.7, PT: 0.7; metafemur length: HT: 7.0, PT: 7.1; metatibia length: HT: 8.0, PT: 8.0; metatarsus length: HT: 0.8, PT: 0.6; metatarsal segment ratio: approx. 1.0: 2.0: 2.5; abdomen length: HT: 9.2, PT: 8.8; abdomen (widest) width: HT: 2.9, PT: 2.9.
</paragraph>
<paragraph id="8B9436B7B938E50B3A948DBAFCE2FC4C" blockId="5.[151,1437,150,1437]" pageId="5" pageNumber="371">
<emphasis id="B95FEAA5B938E50B3A948DBAFE95FCD9" bold="true" box="[199,321,762,788]" pageId="5" pageNumber="371">Variation:</emphasis>
Pedicel and flagellomeres of
<typeStatus id="54908815B938E50B38DA8DBBFD3DFCD8" box="[649,745,763,789]" pageId="5" pageNumber="371" type="paratype">paratype</typeStatus>
as per description under
<taxonomicName id="4C2B4D34B938E50B39AD8DBBFBECFCD9" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" box="[1022,1080,763,788]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEAA5B938E50B39AD8DBBFBECFCD9" box="[1022,1080,763,788]" italics="true" pageId="5" pageNumber="371">Heza</emphasis>
</taxonomicName>
. The
<typeStatus id="54908815B938E50B3E268DBBFB01FCD8" box="[1141,1237,763,789]" pageId="5" pageNumber="371" type="paratype">paratype</typeStatus>
has a slightly less swollen pronotum and at least vestiges of two more pairs of tomentose spots near the basal margin of the posterior pronotal lobe, much in the same arrangement as
<taxonomicName id="4C2B4D34B938E50B38FD8C03FCEDFC90" authority="Swanson, 2021" authorityName="Swanson" authorityYear="2021" box="[686,825,835,861]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="phthinica" status="sp. nov.">
<emphasis id="B95FEAA5B938E50B38FD8C03FCEDFC90" box="[686,825,835,861]" italics="true" pageId="5" pageNumber="371">H. phthinica</emphasis>
</taxonomicName>
<emphasis id="B95FEAA5B938E50B396C8C03FC43FC90" bold="true" box="[831,919,835,861]" pageId="5" pageNumber="371">
<taxonomicNameLabel id="A26C57DEB938E50B396C8C03FC43FC90" box="[831,919,835,861]" pageId="5" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
The tomentose patches and spots of the clavus and corium are a little more discrete in the
<typeStatus id="54908815B938E50B382A8C27FD0DFC4C" box="[633,729,871,897]" pageId="5" pageNumber="371" type="paratype">paratype</typeStatus>
as well.
</paragraph>
<paragraph id="8B9436B7B938E50B3A948CCBFE71FC68" blockId="5.[151,1437,150,1437]" box="[199,421,907,933]" pageId="5" pageNumber="371">
<emphasis id="B95FEAA5B938E50B3A948CCBFEFCFC68" bold="true" box="[199,296,907,933]" pageId="5" pageNumber="371">Female:</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C331653CB938E50B3A948CEEFDB4FC04" box="[199,608,942,969]" pageId="5" pageNumber="371" type="distribution">
<paragraph id="8B9436B7B938E50B3A948CEEFDB4FC04" blockId="5.[151,1437,150,1437]" box="[199,608,942,969]" pageId="5" pageNumber="371">
<emphasis id="B95FEAA5B938E50B3A948CEEFEB7FC05" bold="true" box="[199,355,942,968]" pageId="5" pageNumber="371">Distribution:</emphasis>
<collectingCountry id="F33C7627B938E50B3B398CEFFE74FC04" box="[362,416,943,969]" name="Peru" pageId="5" pageNumber="371">Peru</collectingCountry>
,
<collectingCountry id="F33C7627B938E50B3BF88CEFFE3BFC04" box="[427,495,943,969]" name="Brazil" pageId="5" pageNumber="371">Brazil</collectingCountry>
(
<figureCitation id="13102A32B938E50B3BAC8CEFFD86FC04" box="[511,594,943,969]" captionStart="FIGURE 21" captionStartId="11.[151,250,1328,1352]" captionTargetBox="[151,1436,345,1304]" captionTargetId="figure-102@11.[151,1436,345,1304]" captionTargetPageId="11" captionText="FIGURE 21. Type localities of Heza nebrias sp. nov., Heza phthinica sp. nov., and Heza viridipes sp. nov. Filled symbols represent the type locality, open symbols represent localities of paratypic material." figureDoi="http://doi.org/10.5281/zenodo.4692102" httpUri="https://zenodo.org/record/4692102/files/figure.png" pageId="5" pageNumber="371">Fig. 21</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C331653CB938E50B3A948C93FB96FA50" pageId="5" pageNumber="371" type="discussion">
<paragraph id="8B9436B7B938E50B3A948C93FE4AFB6C" blockId="5.[151,1437,150,1437]" pageId="5" pageNumber="371">
<emphasis id="B95FEAA5B938E50B3A948C93FEE9FC20" bold="true" box="[199,317,979,1005]" pageId="5" pageNumber="371">Remarks:</emphasis>
Owing to the distinctly tomentose clavus and base of the corium,
<taxonomicName id="4C2B4D34B938E50B3E488C93FB44FC20" authority="Swanson, 2021" authorityName="Swanson" authorityYear="2021" box="[1051,1168,979,1005]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="nebrias" status="sp. nov.">
<emphasis id="B95FEAA5B938E50B3E488C93FB44FC20" box="[1051,1168,979,1005]" italics="true" pageId="5" pageNumber="371">H. nebrias</emphasis>
</taxonomicName>
<emphasis id="B95FEAA5B938E50B3EC58C93FB3BFC20" bold="true" box="[1174,1263,979,1005]" pageId="5" pageNumber="371">
<taxonomicNameLabel id="A26C57DEB938E50B3EC58C93FB3BFC20" box="[1174,1263,979,1005]" pageId="5" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
will reach couplet #
<quantity id="4CD39B52B938E50B3A848CB7FEDAFBDC" box="[215,270,1015,1041]" metricMagnitude="-1" metricUnit="m" metricValue="4.572" pageId="5" pageNumber="371" unit="in" value="18.0">18 in</quantity>
<bibRefCitation id="EFBA4B46B938E50B3B478CB7FE23FBDF" author="Maldonado" box="[276,503,1015,1042]" pageId="5" pageNumber="371" pagination="403 - 433" refId="ref8630" refString="Maldonado C., J. (1976) The genus Heza (Hemiptera: Reduviidae). Journal of Agriculture of the University of Puerto Rico, 60 (3), 403 - 433. https: // doi. org / 10.46429 / jaupr. v 60 i 3.10533" type="journal article" year="1976">Maldonado’s (1976)</bibRefCitation>
key. However, it differs from
<taxonomicName id="4C2B4D34B938E50B39168CB7FB19FBDC" authority="Champion, 1899" authorityName="Champion" authorityYear="1899" box="[837,1229,1015,1041]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="multiguttata">
<emphasis id="B95FEAA5B938E50B39168CB7FBD9FBDC" box="[837,1037,1015,1041]" italics="true" pageId="5" pageNumber="371">Heza multiguttata</emphasis>
<bibRefCitation id="EFBA4B46B938E50B3E478CB7FB19FBDC" author="Champion, G. C." box="[1044,1229,1015,1041]" pageId="5" pageNumber="371" pagination="193 - 304" refId="ref8217" refString="Champion, G. C. (1899) Insecta: Rhynchota (Hemiptera-Heteroptera). Volume II. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali-Americana (Prospectus): Zoology, Botany and Archaeology. B. Quaritch, London, pp. 193 - 304." type="book chapter" year="1899">
Champion,
<quantity id="4CD39B52B938E50B3EC68CB7FB19FBDC" box="[1173,1229,1015,1041]" metricMagnitude="1" metricUnit="m" metricValue="4.82346" pageId="5" pageNumber="371" unit="in" value="1899.0">1899</quantity>
</bibRefCitation>
</taxonomicName>
in median process of the pygophore lacking a distinct recurved apical fold.
<taxonomicName id="4C2B4D34B938E50B39438B5BFC70FBF8" authority="Swanson, 2021" authorityName="Swanson" authorityYear="2021" box="[784,932,1051,1077]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="nebrias" status="sp. nov.">
<emphasis id="B95FEAA5B938E50B39438B5BFC70FBF8" box="[784,932,1051,1077]" italics="true" pageId="5" pageNumber="371">Heza nebrias</emphasis>
</taxonomicName>
<emphasis id="B95FEAA5B938E50B39F88B5BFBD0FBF8" bold="true" box="[939,1028,1051,1077]" pageId="5" pageNumber="371">
<taxonomicNameLabel id="A26C57DEB938E50B39F88B5BFBD0FBF8" box="[939,1028,1051,1077]" pageId="5" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
also differs from
<taxonomicName id="4C2B4D34B938E50B3E988B5BFF1BFB94" authority="Stal, 1859" authorityName="Stal" authorityYear="1859" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="insignis">
<emphasis id="B95FEAA5B938E50B3E988B5BFAB7FBF9" box="[1227,1379,1051,1076]" italics="true" pageId="5" pageNumber="371">Heza insignis</emphasis>
<bibRefCitation id="EFBA4B46B938E50B3F398B5AFF1BFB94" author="Stal, C." pageId="5" pageNumber="371" pagination="175 - 204" refId="ref8807" refString="Stal, C. (1859) Till kannedomen om Reduvini. Ofversigt af Kongliga Vetenskaps-Akademiens Forhandlingar, 16 (10), 175 - 204." type="journal article" year="1859">
Stål,
<quantity id="4CD39B52B938E50B3AC48B7EFF1BFB94" box="[151,207,1086,1113]" metricMagnitude="1" metricUnit="m" metricValue="4.7218599999999995" pageId="5" pageNumber="371" unit="in" value="1859.0">1859</quantity>
</bibRefCitation>
</taxonomicName>
in the thin, less convex, and wider apical fold on the median process of the pygophore. Both of these species differ from
<taxonomicName id="4C2B4D34B938E50B3B498B23FE7AFBB0" authority="Swanson, 2021" authorityName="Swanson" authorityYear="2021" box="[282,430,1123,1149]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="nebrias" status="sp. nov.">
<emphasis id="B95FEAA5B938E50B3B498B23FE7AFBB0" box="[282,430,1123,1149]" italics="true" pageId="5" pageNumber="371">Heza nebrias</emphasis>
</taxonomicName>
<emphasis id="B95FEAA5B938E50B3BE68B23FDDBFBB0" bold="true" box="[437,527,1123,1149]" pageId="5" pageNumber="371">
<taxonomicNameLabel id="A26C57DEB938E50B3BE68B23FDDBFBB0" box="[437,527,1123,1149]" pageId="5" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
further in having more extensive tomentose spots near the anterior margin of the posterior pronotal lobe.
</paragraph>
<paragraph id="8B9436B7B938E50B3A948BEBFB96FA50" blockId="5.[151,1437,150,1437]" pageId="5" pageNumber="371">
The median process of the pygophore most closely matches
<taxonomicName id="4C2B4D34B938E50B39268BEBFB11FB08" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[885,1221,1195,1221]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B95FEAA5B938E50B39268BEBFC22FB08" box="[885,1014,1195,1221]" italics="true" pageId="5" pageNumber="371">Heza rubra</emphasis>
<bibRefCitation id="EFBA4B46B938E50B39AD8BEBFB11FB08" author="Maldonado" box="[1022,1221,1195,1221]" pageId="5" pageNumber="371" pagination="403 - 433" refId="ref8630" refString="Maldonado C., J. (1976) The genus Heza (Hemiptera: Reduviidae). Journal of Agriculture of the University of Puerto Rico, 60 (3), 403 - 433. https: // doi. org / 10.46429 / jaupr. v 60 i 3.10533" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C2B4D34B938E50B3EAE8BEBFDE9FB24" authority="(Lepeletier &amp; Audinet-Serville, 1825)" baseAuthorityName="Lepeletier &amp; Audinet-Serville" baseAuthorityYear="1825" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="binotata">
<emphasis id="B95FEAA5B938E50B3EAE8BEBFA48FB08" box="[1277,1436,1195,1221]" italics="true" pageId="5" pageNumber="371">Heza binotata</emphasis>
(
<bibRefCitation id="EFBA4B46B938E50B3ACC8B8FFDE1FB24" author="Lepeletier, L. - M. &amp; Audinet-Serville, J. - G." box="[159,565,1230,1257]" pageId="5" pageNumber="371" pagination="273 - 280" refId="ref8501" refString="Lepeletier, L. - M. &amp; Audinet-Serville, J. - G. (1825) Reduve. In: Latreille, P. A., Lepeletier, L. - M., Audinet-Serville, J. - G. &amp; Guerin-Meneville, F. E. (Eds.), Histoire naturelle. Entomologie, ou Histoire naturelle des Crustaces, des Arachnides, et des insectes. Tome Dixieme. Mme veuve Agasse, Paris, pp. 273 - 280." type="book chapter" year="1825">Lepeletier &amp; Audinet-Serville, 1825</bibRefCitation>
)
</taxonomicName>
; however, it differs from the former in possessing a strongly tomentose clavus and in the reddish-brown legs (rather than green faded to yellow in
<taxonomicName id="4C2B4D34B938E50B39C38BB3FC04FAC0" authorityName="Maldonado" authorityYear="1976" box="[912,976,1267,1293]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="rubra">
<emphasis id="B95FEAA5B938E50B39C38BB3FC04FAC0" box="[912,976,1267,1293]" italics="true" pageId="5" pageNumber="371">rubra</emphasis>
</taxonomicName>
) and from the latter in the relative thickness of the recurved apical fold of the median process of the pygophore, as well as lacking the large tomentose spots on the posterior lobe of the pronotum.
<taxonomicName id="4C2B4D34B938E50B381B8A7BFC66FA98" authority="Maldonado, 1976" authorityName="Maldonado" authorityYear="1976" box="[584,946,1339,1365]" class="Insecta" family="Reduviidae" genus="Heza" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="ocellata">
<emphasis id="B95FEAA5B938E50B381B8A7BFD36FA98" box="[584,738,1339,1365]" italics="true" pageId="5" pageNumber="371">Heza ocellata</emphasis>
<bibRefCitation id="EFBA4B46B938E50B38B98A7BFC66FA98" author="Maldonado" box="[746,946,1339,1365]" pageId="5" pageNumber="371" pagination="403 - 433" refId="ref8630" refString="Maldonado C., J. (1976) The genus Heza (Hemiptera: Reduviidae). Journal of Agriculture of the University of Puerto Rico, 60 (3), 403 - 433. https: // doi. org / 10.46429 / jaupr. v 60 i 3.10533" type="journal article" year="1976">Maldonado, 1976</bibRefCitation>
</taxonomicName>
, a species which shares a similar dispersion pattern of small tomentose spots on the posterior pronotal lobe, differs in having a truncate or slightly concave apex on the median process of the pygophore and also lacks a strongly tomentose clavus.
</paragraph>
</subSubSection>
</treatment>
</document>
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