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<document id="FB1A72D874D2E014D1E2CECE07E01A69" ID-DOI="10.5252/geodiversitas2019v41a9" ID-GBIF-Dataset="e9842926-ae93-45e6-813d-248b81b6054d" ID-ISSN="1638-9395" ID-Zenodo-Dep="3700034" ID-ZooBank="urn:lsid:zoobank.org:pub:32E3623C-C47B-4D42-B2EB-E2594D031349" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583515269397" checkinUser="valdenar" docAuthor="Ossó, Àlex &amp; Gagnaison, Cyril" docDate="2019" docId="03D187F1907FFFB1FBB79D230A234199" docLanguage="en" docName="GEODIVERSITAS.41.9.367-383.pdf.imf" docOrigin="Geodiversitas 41 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Liocarcinus sp. Stimpson 1871" docType="treatment" docVersion="6" lastPageNumber="376" masterDocId="FFE8FF899076FFBAFF979B360B3E4674" masterDocTitle="An appraisal of the Middle-Late Miocene fossil decapod crustaceans of the Faluns (Anjou-Touraine, France)" masterLastPageNumber="383" masterPageNumber="367" pageNumber="374" updateTime="1698725925088" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="F26322F31560F6D0B8571EEA03BBD59F">An appraisal of the Middle-Late Miocene fossil decapod crustaceans of the Faluns (Anjou-Touraine, France)</mods:title>
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<mods:namePart id="1C8C9866A740F9103E19FAC1168DB8E3">Ossó, Àlex</mods:namePart>
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<mods:namePart id="1A227ED502D2EED895D006C896ABD511">Gagnaison, Cyril</mods:namePart>
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<mods:title id="C9CA9E6979307C889FE58B0B207D543D">Geodiversitas</mods:title>
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<mods:date id="5EA492E722D52DC7DD1322DD9CE3F2AA">2019</mods:date>
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<mods:number id="D796E9E051A60176C8B39D43B499DE12">2019-04-25</mods:number>
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<treatment id="03D187F1907FFFB1FBB79D230A234199" ID-DOI="http://doi.org/10.5281/zenodo.3705482" ID-GBIF-Taxon="162463503" ID-Zenodo-Dep="3705482" LSID="urn:lsid:plazi:treatment:03D187F1907FFFB1FBB79D230A234199" httpUri="http://treatment.plazi.org/id/03D187F1907FFFB1FBB79D230A234199" lastPageId="11" lastPageNumber="376" pageId="9" pageNumber="374">
<subSubSection id="C362656C907FFFB3FBB79D230F82405B" box="[1056,1212,1557,1583]" pageId="9" pageNumber="374" type="nomenclature">
<paragraph id="8BC736E7907FFFB3FBB79D230F82405B" blockId="9.[1056,1212,1557,1615]" box="[1056,1212,1557,1583]" pageId="9" pageNumber="374">
<heading id="D08F818B907FFFB3FBB79D230F82405B" box="[1056,1212,1557,1583]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="374" reason="2">
<taxonomicName id="4C784D64907FFFB3FBB79D230F82405B" authorityName="Stimpson" authorityYear="1871" box="[1056,1212,1557,1583]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="374" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="B90CEAF5907FFFB3FBB79D230FA8405A" box="[1056,1174,1557,1582]" italics="true" pageId="9" pageNumber="374">Liocarcinus</emphasis>
sp.
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C362656C907FFFB3FBBA9D020F90403B" box="[1069,1198,1588,1615]" pageId="9" pageNumber="374" type="description">
<paragraph id="8BC736E7907FFFB3FBBA9D020F90403B" blockId="9.[1056,1212,1557,1615]" box="[1069,1198,1588,1615]" pageId="9" pageNumber="374">
(
<figureCitation id="13432A62907FFFB3FBA29D020F98403B" box="[1077,1190,1588,1615]" captionStart="FIG" captionStartId="12.[132,143,1605,1623]" captionTargetBox="[132,1455,215,1559]" captionTargetId="figure@12.[132,1455,214,1559]" captionTargetPageId="12" captionText="FIG. 5. — A, B, Liocarcinus sp., MS2012 0 172b: A, dorsal view; B, frontal view; C, D, Necronectes michelini (A. Milne-Edwards, 1861): C, ULB-SO-9, right dactylus; D, ULB-SO-10, left dactylus; E, Paguristes gagnaisoni Gagnaison, 2012, MS2006 FS 134b, right chela; F-I, Xantho cf. moldavicus (Yanakevich, 1977): F, ULB-SO-11, left chela, inner lateral view; G, outer lateral view; H, ULB-SO-12, left chela, outer lateral view; I, frontolateral view; J-M, undetermined chelae: J, ULB-SO-13, lateral view of right chela; K, frontal view; L, ULB-SO-14, lateral view of left chela; M, close-up showing an encrusted colony of bryozoans. Scale bars: A, B, 5 mm; C-M, 10 mm. Photographs by À. Ossó." figureDoi="http://doi.org/10.5281/zenodo.3700044" httpUri="https://zenodo.org/record/3700044/files/figure.png" pageId="9" pageNumber="374">Fig. 5A, B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C362656C907FFFB3FCBA9D5B08D9416B" pageId="9" pageNumber="374" type="materials_examined">
<paragraph id="8BC736E7907FFFB3FCBA9D5B08AD40CE" blockId="9.[813,1456,1645,1722]" pageId="9" pageNumber="374">
MATERIAL EXAMINED AND MEASUREMENTS (in mm). —
<materialsCitation id="3B103CBA907FFFB3FAC99D5B08AD40CE" ID-GBIF-Occurrence="2574969786" pageId="9" pageNumber="374">Partially preserved carapace, MS2012 0 172b: L = 10.5, W = 12, FOW = 8.5, F = 4.</materialsCitation>
</paragraph>
<paragraph id="8BC736E7907FFFB3FCBA9DE708D9416B" blockId="9.[813,1455,1745,1823]" pageId="9" pageNumber="374">LOCALITY AND HORIZON. — La Sonneterie quarry, Meigné-le- Vicomte (Maine-et-Loire). Savignean facies, Langhian-Serravallian (Middle Miocene).</paragraph>
</subSubSection>
<subSubSection id="C362656C907FFFB1FCBA9C650BCC4446" lastPageId="11" lastPageNumber="376" pageId="9" pageNumber="374" type="description">
<paragraph id="8BC736E7907FFFB3FCBA9C650883411F" blockId="9.[813,1455,1875,2030]" box="[813,957,1875,1901]" pageId="9" pageNumber="374">DESCRIPTION</paragraph>
<paragraph id="8BC736E7907FFFB1FCBA9C450BCC4446" blockId="9.[813,1455,1875,2030]" lastBlockId="11.[132,775,217,562]" lastPageId="11" lastPageNumber="376" pageId="9" pageNumber="374">Small sized carapace, subhexagonal, smooth, slightly wider than long. Regions weakly defined. Three longitudinally aligned tubercles at each side of carapace. Front incomplete, straight, but appears to be slightly produced medially. Orbits relatively large; supraorbital margin with two closed fissures. Anterolateral margin slightly arcuate, with four subtriangular teeth (excluding the outer orbital tooth), the last of them (epi- branchial tooth) the sharper. Posterolateral margin straight; reentrant of fifth pereiopod well marked. Posterior margin straight.Frontal region smooth, flattened.Gastric process poorly defined. Epigastric lobes slightly marked. Protogastric lobes slightly swollen with faintly marked transverse ridge at each lobe; marked tubercle near gastrohepatic groove. Mesogastric region weakly marked. Urogastric region depressed. Cardiac region large, slightly swollen. Intestinal region depressed. Hepatic region depressed. Epibranchial region sigmoidal, weakly marked, with medial tubercle. Mesobranchial region slightly swollen, with marked tubercle, Metabranchial region depressed. Cervical groove faintly marked. Branchiocardiac groove weakly marked. Ventral parts and appendages not preserved.</paragraph>
</subSubSection>
<caption id="DF07666F907CFFB0FF139C2609F24112" ID-DOI="http://doi.org/10.5281/zenodo.3700042" ID-Zenodo-Dep="3700042" httpUri="https://zenodo.org/record/3700042/files/figure.png" pageId="10" pageNumber="375" startId="10.[132,143,1808,1826]" targetBox="[132,1455,215,1764]" targetPageId="10">
<paragraph id="8BC736E7907CFFB0FF139C2609F24112" blockId="10.[132,1457,1808,1894]" pageId="10" pageNumber="375">
FIG. 4. —
<emphasis id="B90CEAF5907CFFB0FF449C260BCC4156" bold="true" box="[211,242,1808,1826]" pageId="10" pageNumber="375">A -F</emphasis>
,
<taxonomicName id="4C784D64907CFFB0FF6B9C2609424156" authority="Millet de la Turtaudiere, 1865" authorityName="Millet de la Turtaudiere" authorityYear="1865" box="[252,636,1808,1826]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="375" phylum="Arthropoda" rank="species" species="orbignyana">
<emphasis id="B90CEAF5907CFFB0FF6B9C260ABB4156" box="[252,389,1808,1826]" italics="true" pageId="10" pageNumber="375">Maja orbignyana</emphasis>
<bibRefCitation id="EFE94B16907CFFB0FE1D9C2609424156" author="MILLET DE LA TURTAUDIERE P. A." box="[394,636,1808,1826]" pageId="10" pageNumber="375" refId="ref13332" refString="MILLET DE LA TURTAUDIERE P. A. 1865. - Paleontographie ou Description des fossiles nouveaux du terrain tertiaire marin ou terrain miocene superieur du departement de Maine-et-Loire (Supplement). Indicateur de Maine et Loire. Cosnier et Lachese, Angers, vol. 2, 616 p." type="book" year="1865">Millet de la Turtaudière, 1865</bibRefCitation>
</taxonomicName>
:
<emphasis id="B90CEAF5907CFFB0FD159C2609B14156" bold="true" box="[642,655,1808,1826]" pageId="10" pageNumber="375">A</emphasis>
, ULB-SO-2, frontal view of left orbit;
<emphasis id="B90CEAF5907CFFB0FC559C2608F14156" bold="true" box="[962,975,1808,1826]" pageId="10" pageNumber="375">B</emphasis>
, MS2012 0 172a, dorsal view of anterior part of carapace;
<emphasis id="B90CEAF5907CFFB0FF139C100BAC414C" bold="true" box="[132,146,1830,1848]" pageId="10" pageNumber="375">C</emphasis>
, ULB-SO-3, frontal view of left orbit;
<emphasis id="B90CEAF5907CFFB0FE509C110AEB414D" bold="true" box="[455,469,1831,1849]" pageId="10" pageNumber="375">D</emphasis>
, ULB-SO-3, frontal view of left orbit;
<emphasis id="B90CEAF5907CFFB0FC9D9C110828414D" bold="true" box="[778,790,1831,1849]" pageId="10" pageNumber="375">E</emphasis>
, ULB-SO-4, merus of ambulatory leg;
<emphasis id="B90CEAF5907CFFB0FBC19C110F5F414D" bold="true" box="[1110,1121,1831,1849]" pageId="10" pageNumber="375">F</emphasis>
, ULB-SO-5, dactylus of ambulatory leg;
<emphasis id="B90CEAF5907CFFB0FF139C0B0B9B413B" bold="true" box="[132,165,1853,1871]" pageId="10" pageNumber="375">G -L</emphasis>
,
<taxonomicName id="4C784D64907CFFB0FF399C0B0932413B" authority="(Lorenthey, 1897)" baseAuthorityName="Lorenthey" baseAuthorityYear="1897" box="[174,524,1853,1871]" class="Malacostraca" family="Pilumnidae" genus="Pilumnus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="375" phylum="Arthropoda" rank="species" species="mediterraneus">
<emphasis id="B90CEAF5907CFFB0FF399C0B0A48413B" box="[174,374,1853,1871]" italics="true" pageId="10" pageNumber="375">Pilumnus mediterraneus</emphasis>
(
<bibRefCitation id="EFE94B16907CFFB0FEE89C0B0936413B" author="LORENTHEY I." box="[383,520,1853,1871]" pageId="10" pageNumber="375" pagination="149 - 169" refId="ref12916" refString="LORENTHEY I. 1897. - Adatok Magyarorszag harmadkoru rak faunajahoz. Mathematikai es Termeszettudomanyi Erttesito 15: 149 - 169" type="journal article" year="1897">Lőrenthey, 1897</bibRefCitation>
)
</taxonomicName>
:
<emphasis id="B90CEAF5907CFFB0FD849C0B091F413B" bold="true" box="[531,545,1853,1871]" pageId="10" pageNumber="375">G</emphasis>
, ULB-SO-6, dorsal view;
<emphasis id="B90CEAF5907CFFB0FD649C0B083F413B" bold="true" box="[755,769,1853,1871]" pageId="10" pageNumber="375">H</emphasis>
, frontal view;
<emphasis id="B90CEAF5907CFFB0FCE49C0B0847413B" bold="true" box="[883,889,1853,1871]" pageId="10" pageNumber="375">I</emphasis>
, MS2012 0 170,frontal view;
<emphasis id="B90CEAF5907CFFB0FBFB9C0B0F48413B" bold="true" box="[1132,1142,1853,1871]" pageId="10" pageNumber="375">J</emphasis>
, dorsal view;
<emphasis id="B90CEAF5907CFFB0FB709C0B0FCA413B" bold="true" box="[1255,1268,1853,1871]" pageId="10" pageNumber="375">K</emphasis>
, ULB-SO-7,left chela;
<emphasis id="B90CEAF5907CFFB0FF139C620BB14112" bold="true" box="[132,143,1876,1894]" pageId="10" pageNumber="375">L</emphasis>
, ULB-SO-8, left chela. Scale bars: 10 mm. Photographs by À. Ossó.
</paragraph>
</caption>
<subSubSection id="C362656C907DFFB1FF1399610A234199" pageId="11" pageNumber="376" type="discussion">
<paragraph id="8BC736E7907DFFB1FF1399610BD64404" blockId="11.[132,776,599,2029]" box="[132,232,599,625]" pageId="11" pageNumber="376">REMARKS</paragraph>
<paragraph id="8BC736E7907DFFB1FF13994E08364364" blockId="11.[132,776,599,2029]" pageId="11" pageNumber="376">
The features of the studied specimen fit with those of
<taxonomicName id="4C784D64907DFFB1FD2B994E0B8644C5" authorityName="Stimpson" authorityYear="1871" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5907DFFB1FD2B994E0B8644C5" italics="true" pageId="11" pageNumber="376">Liocarcinus</emphasis>
</taxonomicName>
, and its general outline matches well with the known Miocene
<taxonomicName id="4C784D64907DFFB1FF78998E0A5444A5" authorityName="Stimpson" authorityYear="1871" box="[239,362,696,721]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5907DFFB1FF78998E0A5444A5" box="[239,362,696,721]" italics="true" pageId="11" pageNumber="376">Liocarcinus</emphasis>
</taxonomicName>
species (see
<bibRefCitation id="EFE94B16907DFFB1FE6C998109C844A6" author="SCHWEITZER C. E. &amp; FELDMANN R. M. &amp; GARASSINO A. &amp; KARASAWA H. &amp; SCHWEIGERT G." box="[507,758,695,722]" pageId="11" pageNumber="376" pagination="1 - 1222" refId="ref14815" refString="SCHWEITZER C. E., FELDMANN R. M., GARASSINO A., KARASAWA H. &amp; SCHWEIGERT G. 2010. - Systematic list of fossil decapod crustacean species. Crustaceana Monograph 10: 1 - 1222. https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="journal article" year="2010">
Schweitzer
<emphasis id="B90CEAF5907DFFB1FDEB998E098A44A5" box="[636,692,695,721]" italics="true" pageId="11" pageNumber="376">et al.</emphasis>
2010
</bibRefCitation>
). However, they differ in having a most decorated or rough surface of the carapace, as for instance
<emphasis id="B90CEAF5907DFFB1FD8199CE09AA4565" box="[534,660,759,785]" italics="true" pageId="11" pageNumber="376">L. rakosensis</emphasis>
Lőrenthey
<emphasis id="B90CEAF5907DFFB1FF13982E0BA44545" box="[132,154,792,817]" italics="true" pageId="11" pageNumber="376">in</emphasis>
<bibRefCitation id="EFE94B16907DFFB1FF3498210AE94545" author="LORENTHEY E. &amp; BEURLEN K." box="[163,471,791,817]" pageId="11" pageNumber="376" pagination="1 - 420" refId="ref13030" refString="LORENTHEY E. &amp; BEURLEN K. 1929. - Die fossilen Decapoden der Lander der Ungarischen Krone. Geologica Hungarica, Series Palaeontologica 3: 1 - 420." type="journal article" year="1929">Lőrenthey &amp; Beurlen, 1929</bibRefCitation>
and
<emphasis id="B90CEAF5907DFFB1FD86982E09494545" box="[529,631,791,817]" italics="true" pageId="11" pageNumber="376">L. kuehni</emphasis>
(Bachmayer, 1953), whereas in the studied specimen the carapace surface is completely smooth; or in having more marked and raised regions as in the case of
<emphasis id="B90CEAF5907DFFB1FE01984E0A9B45E5" box="[406,421,888,913]" italics="true" pageId="11" pageNumber="376">L</emphasis>
.
<emphasis id="B90CEAF5907DFFB1FE22984E0AC545E5" box="[437,507,888,913]" italics="true" pageId="11" pageNumber="376">oroszyi</emphasis>
(Bachmayer, 1953) and
<emphasis id="B90CEAF5907DFFB1FF1398A10A1B45C4" box="[132,293,919,944]" italics="true" pageId="11" pageNumber="376">L. praearcuatus</emphasis>
<bibRefCitation id="EFE94B16907DFFB1FEB998A10AFE45C5" author="MULLER P." box="[302,448,919,945]" pageId="11" pageNumber="376" pagination="3 - 14" refId="ref13912" refString="MULLER P. 1996. - Middle Miocene Decapod Crustacea from southern Poland. Prace Muzeum Ziemi, Prace paleozoologiczne 43: 3 - 14." type="journal article" year="1996">Müller, 1996</bibRefCitation>
(see
<bibRefCitation id="EFE94B16907DFFB1FE6D98A1098345C6" author="BACHMAYER F." box="[506,701,919,946]" pageId="11" pageNumber="376" pagination="237 - 262" refId="ref10060" refString="BACHMAYER F. 1953 a. - Die Dekapodenfauna des tortonischen Leithakalkes von Deutsch-Altenburg (Niederosterreich). Mitteilungen der Geologischen Gesellschaft in Wien 44: 237 - 262." type="journal article" year="1953">Bachmayer 1953a</bibRefCitation>
: pl. 2, figs 2, 5-5a;
<bibRefCitation id="EFE94B16907DFFB1FE8698810AC245A5" author="MULLER P." box="[273,508,951,977]" pageId="11" pageNumber="376" pagination="25 - 317" refId="ref13833" refString="MULLER P. 1984 a. - Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica 42: 25 - 317." type="journal article" year="1984">Müller 1984a: 83-84</bibRefCitation>
, pl. 69, figs 2-6, pl. 71, figs 1-4; 1996: 10, pl. 2, figs 2-3).
<emphasis id="B90CEAF5907DFFB1FD9D98E108394584" box="[522,775,983,1008]" italics="true" pageId="11" pageNumber="376">
<taxonomicName id="4C784D64907DFFB1FD9D98E109BF4584" authorityName="Stimpson" authorityYear="1871" box="[522,641,983,1008]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">Liocarcinus</taxonomicName>
ottnangensis
</emphasis>
(Bachmayer, 1953) differs clearly in having a trifid front and a characteristic prominent transverse ridge running across the mesogastric and epibranchial regions, which is absent in the studied specimen (see
<bibRefCitation id="EFE94B16907DFFB1FE2D9F6009444205" author="HYZNY M. &amp; HARZHAUSER M. &amp; DANNINGER W." box="[442,634,1110,1137]" pageId="11" pageNumber="376" pagination="217 - 233" refId="ref12042" refString="HYZNY M., HARZHAUSER M. &amp; DANNINGER W. 2015. - Decapod Crustacea of the Central Paratethyan Ottnangian Stage (middle Burdigalian): implications for systematics and biogeography. Geologica Carpathica 66 (3): 217 - 233. https: // doi. org / 10.1515 / geoca- 2015 - 0021" type="journal article" year="2015">
Hyžný
<emphasis id="B90CEAF5907DFFB1FD9E9F6109024204" box="[521,572,1110,1136]" italics="true" pageId="11" pageNumber="376">et al.</emphasis>
2015
</bibRefCitation>
).
<emphasis id="B90CEAF5907DFFB1FD069F610BC042E4" italics="true" pageId="11" pageNumber="376">
<taxonomicName id="4C784D64907DFFB1FD069F6108394204" authorityName="Stimpson" authorityYear="1871" box="[657,775,1111,1136]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">Liocarcinus</taxonomicName>
oligocenicus
</emphasis>
(
<bibRefCitation id="EFE94B16907DFFB1FE999F400AA742E4" author="PAUCA M." box="[270,409,1142,1168]" pageId="11" pageNumber="376" pagination="40 - 44" refId="ref14309" refString="PAUCA M. 1929. - Zwei neue fossile Decapoden aus den Oligozanen Clupea crenata Schiefern von Suslaneski-Muscel und uber die Bildung der Menilitschiefer. Bulletin de la Section scientifique de l'Academie roumaine 12: 4, 5, 40 - 44." type="journal article" year="1929">Paucă, 1929</bibRefCitation>
) and
<emphasis id="B90CEAF5907DFFB1FE4B9F4109AD42E4" box="[476,659,1142,1168]" italics="true" pageId="11" pageNumber="376">L. lancetidactylus</emphasis>
(
<bibRefCitation id="EFE94B16907DFFB1FD349F400B8342C4" author="SMIRNOV V. P." pageId="11" pageNumber="376" pagination="1 - 49" refId="ref15002" refString="SMIRNOV V. P. 1929. - Decapoda of the fish beds at Black river in the vicinity of Vladikavkaz. (Decapoda iz rybnykplastov na Chernoy rechke v okrastnostyakh Vladikavkaza). Arbeiten der Nord-Kaukasischen AssoziationWissenschaftlicher Institute 59: 1 - 49 (in Russian)." type="journal article" year="1929">Smirnov, 1929</bibRefCitation>
) present a different pattern of anterolateral margins and front, furthermore, their poor preservation makes difficult an accurate comparison (e.g.
<bibRefCitation id="EFE94B16907DFFB1FE029FE009774284" author="JERZMANSKA A." box="[405,585,1238,1264]" pageId="11" pageNumber="376" pagination="539 - 545" refId="ref12210" refString="JERZMANSKA A. 1967. - Crabs of the genus Portunus Weber from the Menilite Series of the Carpathians. Annales de la Societe geologique de Pologne 37: 539 - 545. (in Polish, with English summary)." type="journal article" year="1967">Jerzmańska 1967</bibRefCitation>
: fig. 1;
<bibRefCitation id="EFE94B16907DFFB1FD029FE00BCE4364" author="SCHWEITZER C. E. &amp; FELDMANN R. M. &amp; LAZA R I." pageId="11" pageNumber="376" pagination="1 - 14" refId="ref14872" refString="SCHWEITZER C. E., FELDMANN R. M. &amp; LAZA R I. 2009. - Fossil Crustacea (excluding Cirripedia and Ostracoda) in the University of Bucharest Collections, Romania, including two new species. Bulletin of the Mizunami Fossil Museum 35: 1 - 14." type="journal article" year="2009">
Schweitzer
<emphasis id="B90CEAF5907DFFB1FF139FC00B93437B" box="[132,173,1269,1295]" italics="true" pageId="11" pageNumber="376">et al</emphasis>
. 2009
</bibRefCitation>
: fig. 6; and
<bibRefCitation id="EFE94B16907DFFB1FEFF9FC3094D4364" author="GARASSINO A. &amp; NOVATI M." box="[360,627,1269,1296]" pageId="11" pageNumber="376" pagination="269 - 281" refId="ref11439" refString="GARASSINO A. &amp; NOVATI M. 2001. - Liocarcinus lancetidactylus (Smirnov, 1929) and Platymaia lethaeus (Smirnov, 1929) from the Miocene of N Caucasus (Russia). Atti della Societa iItaliana di Scienze naturali e del Museo Civico di Storia naturale di Milano 141 (2): 269 - 281." type="journal article" year="2001">Garassino &amp; Novati 2001</bibRefCitation>
, respectively).
</paragraph>
<paragraph id="8BC736E7907DFFB1FF0C9E230839401B" blockId="11.[132,776,599,2029]" pageId="11" pageNumber="376">
Regarding the extant species of
<taxonomicName id="4C784D64907DFFB1FE6F9E200950435B" authorityName="Stimpson" authorityYear="1871" box="[504,622,1302,1327]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5907DFFB1FE6F9E200950435B" box="[504,622,1302,1327]" italics="true" pageId="11" pageNumber="376">Liocarcinus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFE94B16907DFFB1FD309E200B82433B" author="NG P. K. L. &amp; GUINOT D. &amp; DAVIE P. J. F." pageId="11" pageNumber="376" pagination="1 - 286" refId="ref13973" refString="NG P. K. L., GUINOT D. &amp; DAVIE P. J. F. 2008. - Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology, Supplement 17: 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B90CEAF5907DFFB1FD459E200839435B" box="[722,775,1301,1327]" italics="true" pageId="11" pageNumber="376">et al.</emphasis>
2008
</bibRefCitation>
), they have usually a more arched anterolateral margins than the studied specimen, and most of them have a trifid front, and/or a rough surface of carapace, which is smooth in the studied specimen.
<taxonomicName id="4C784D64907DFFB1FE0D9EA0083943C4" authority="(Leach, 1815)" baseAuthorityName="Leach" baseAuthorityYear="1815" box="[410,775,1429,1456]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B90CEAF5907DFFB1FE0D9EA0095B43DB" box="[410,613,1429,1455]" italics="true" pageId="11" pageNumber="376">Liocarcinus pusillus</emphasis>
(
<bibRefCitation id="EFE94B16907DFFB1FDE29EA309C143C4" author="LEACH W. E." box="[629,767,1429,1456]" pageId="11" pageNumber="376" pagination="145 - 154" refId="ref12679" refString="LEACH W. E. 1815. - The Zoological Miscellany; being Descriptions of New, or Interesting Animals, Illustrated with Coloured Figures, Drawn from Nature, in NODDER R. P. et al. (eds). E. Nodder &amp; Son, London, vol. 2: 145 - 154. https: // doi. org / 10.5962 / bhl. title. 41372" type="journal article" year="1815">Leach, 1815</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C784D64907DFFB1FF229E800ADA43BB" authority="(Herbst, 1794)" baseAuthorityName="Herbst" baseAuthorityYear="1794" box="[181,484,1461,1487]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="species" species="navigator">
<emphasis id="B90CEAF5907DFFB1FF229E800A0643BB" box="[181,312,1462,1487]" italics="true" pageId="11" pageNumber="376">L. navigator</emphasis>
(Herbst, 1794)
</taxonomicName>
have a similar blunt front medially produced, similar to the studied specimen, but they differ in having a rougher dorsal surface, and by the lack of the conspicuous protogastric ridges and three characteristic tubercles, longitudinally aligned, present in the French specimen possess (e.g.
<bibRefCitation id="EFE94B16907DFFB1FE9D9D620927401A" author="FROGLIA C. &amp; MANNING R. B." box="[266,537,1620,1646]" pageId="11" pageNumber="376" pagination="257 - 266" refId="ref11136" refString="FROGLIA C. &amp; MANNING R. B. 1982. - Notes on Liocarcinus pusillus (Leach) and related species. Quaderni del Laboratorio di Tecnologia della Pesca 3 (2 - 5): 257 - 266." type="journal article" year="1982">Froglia &amp; Manning 1982</bibRefCitation>
; Koch &amp; Ďuriš 2016)
</paragraph>
<paragraph id="8BC736E7907DFFB1FF0C9D4209364139" blockId="11.[132,776,599,2029]" pageId="11" pageNumber="376">
The general outline and dorsal sculpture of the studied specimen also resemble that of
<emphasis id="B90CEAF5907DFFB1FE0F9DA30AC640DA" box="[408,504,1685,1710]" italics="true" pageId="11" pageNumber="376">Miopipus</emphasis>
<taxonomicName id="4C784D64907DFFB1FE6B9DA3083940DA" authority="(Brocchi, 1883)" baseAuthorityName="Brocchi" baseAuthorityYear="1883" box="[508,775,1684,1710]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B90CEAF5907DFFB1FE6B9DA3096540DA" box="[508,603,1685,1710]" italics="true" pageId="11" pageNumber="376">pygmaeus</emphasis>
(
<bibRefCitation id="EFE94B16907DFFB1FDFF9DA209C140DA" author="BROCCHI P." box="[616,767,1684,1710]" pageId="11" pageNumber="376" pagination="1 - 8" refId="ref10341" refString="BROCCHI P. 1883. - Notes sur les crustaces fossiles des terrains tertiaires de la Hongrie. Annales des Sciences geologiques 14 (2): 1 - 8." type="journal article" year="1883">Brocchi, 1883</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EFE94B16907DFFB1FF1A9D820A0440BA" author="BROCCHI P." box="[141,314,1716,1743]" pageId="11" pageNumber="376" pagination="1 - 8" refId="ref10341" refString="BROCCHI P. 1883. - Notes sur les crustaces fossiles des terrains tertiaires de la Hongrie. Annales des Sciences geologiques 14 (2): 1 - 8." type="journal article" year="1883">Brocchi 1883: 1</bibRefCitation>
, pl. 5, figs 4-4a). However,
<bibRefCitation id="EFE94B16907DFFB1FDF49D820B9C409A" author="MULLER P." pageId="11" pageNumber="376" pagination="25 - 317" refId="ref13833" refString="MULLER P. 1984 a. - Decapod Crustacea of the Badenian. Geologica Hungarica, Series Palaeontologica 42: 25 - 317." type="journal article" year="1984">Müller (1984a: 84</bibRefCitation>
, pl. 72, figs 1-4), described the front of
<taxonomicName id="4C784D64907DFFB1FDF69DE309D4409A" baseAuthorityName="Brocchi" baseAuthorityYear="1883" box="[609,746,1749,1774]" class="Malacostraca" family="Majidae" genus="Maja" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="species" species="pygmaeus">
<emphasis id="B90CEAF5907DFFB1FDF69DE309D4409A" box="[609,746,1749,1774]" italics="true" pageId="11" pageNumber="376">M. pygmaeus</emphasis>
</taxonomicName>
as trilobed, narrow and prominent, and the carapace regions as strongly inflated, which precludes a congeneric relationship with the specimen described herein.
</paragraph>
<paragraph id="8BC736E7907DFFB1FF0C9C650A234199" blockId="11.[132,776,599,2029]" pageId="11" pageNumber="376">
Therefore, in spite that the aforementioned differences could lead us to propose a new species of
<taxonomicName id="4C784D64907DFFB1FDA99C42098A41F9" authorityName="Stimpson" authorityYear="1871" box="[574,692,1908,1933]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5907DFFB1FDA99C42098A41F9" box="[574,692,1908,1933]" italics="true" pageId="11" pageNumber="376">Liocarcinus</emphasis>
</taxonomicName>
for this specimen, based on the poorly preserved sole specimen, we report it as
<taxonomicName id="4C784D64907DFFB1FE929C820A4541B9" authorityName="Stimpson" authorityYear="1871" box="[261,379,1972,1997]" class="Malacostraca" family="Polybiidae" genus="Liocarcinus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="376" phylum="Arthropoda" rank="genus">
<emphasis id="B90CEAF5907DFFB1FE929C820A4541B9" box="[261,379,1972,1997]" italics="true" pageId="11" pageNumber="376">Liocarcinus</emphasis>
</taxonomicName>
sp., until more specimens are available for study.
</paragraph>
</subSubSection>
</treatment>
</document>