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<document id="393C520D635D61E66354BB5878303BA7" ID-CLB-Dataset="9462" ID-DOI="10.11646/zootaxa.4890.2.5" ID-GBIF-Dataset="768fbdbe-ad7e-415f-8e22-52c0a52c71f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="4301869" ID-ZooBank="82A5AFC2-8E8D-472D-97B5-9FFD5CB8CC31" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1606905221522" checkinUser="plazi" docAuthor="Scheers, Kevin &amp; Hájek, Jiří" docDate="2020" docId="03B387F5FF9BC16642C6FF65FC57F8DA" docLanguage="en" docName="zootaxa.4890.2.5.pdf" docOrigin="Zootaxa 4890 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoclypeodytes curtulus" docType="treatment" docVersion="8" lastPageNumber="249" masterDocId="FF8AFF8DFF9FC1624251FFF2FF99FFC5" masterDocTitle="Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)" masterLastPageNumber="256" masterPageNumber="245" pageNumber="249" updateTime="1698847084457" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="124CF5177F4784A13A128D804A66B0D8">Neoclypeodytes Young from Panama, with description of two new species (Coleoptera: Dytiscidae: Hydroporinae: Bidessini)</mods:title>
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<mods:namePart id="E19F9EF161E0FA35897C05A23375746B">Scheers, Kevin</mods:namePart>
<mods:affiliation id="77810B847C4C7C69DC08C6F29A1D66F6">Research Institute for Nature and Forest (INBO), Havenlaan 88 bus 73, 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="4AC887FE853D556791F3D10399C394D9">Hájek, Jiří</mods:namePart>
<mods:affiliation id="56B9C9C5E3005D3A232886B0C9F9D400">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. jiri. hajek @ nm. cz; https: // orcid. org / 0000 - 0001 - 5779 - 1542</mods:affiliation>
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<treatment id="03B387F5FF9BC16642C6FF65FC57F8DA" ID-DOI="http://doi.org/10.5281/zenodo.4327973" ID-GBIF-Taxon="170604212" ID-Zenodo-Dep="4327973" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA" httpUri="http://treatment.plazi.org/id/03B387F5FF9BC16642C6FF65FC57F8DA" lastPageNumber="249" pageId="4" pageNumber="249">
<subSubSection id="C3006568FF9BC16642C6FF65FDF7FF74" box="[151,622,151,178]" pageId="4" pageNumber="249" type="nomenclature">
<paragraph id="8BA536E3FF9BC16642C6FF65FDF7FF74" blockId="4.[151,622,151,213]" box="[151,622,151,178]" pageId="4" pageNumber="249">
<heading id="D0ED818FFF9BC16642C6FF65FDF7FF74" bold="true" box="[151,622,151,178]" fontSize="11" level="1" pageId="4" pageNumber="249" reason="1">
<taxonomicName id="4C1A4D60FF9BC16642C6FF65FDF7FF74" ID-CoL="46D87" authority="(Sharp, 1887)" baseAuthorityName="Sharp" baseAuthorityYear="1887" box="[151,622,151,178]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC16642C6FF65FDF7FF74" bold="true" box="[151,622,151,178]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC16642C6FF65FE24FF77" bold="true" box="[151,445,151,178]" italics="true" pageId="4" pageNumber="249">Neoclypeodytes curtulus</emphasis>
(Sharp, 1887)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9BC16642C6FF49FF75FF10" box="[151,236,187,213]" pageId="4" pageNumber="249" type="description">
<paragraph id="8BA536E3FF9BC16642C6FF49FF75FF10" blockId="4.[151,622,151,213]" box="[151,236,187,213]" pageId="4" pageNumber="249">
(
<figureCitation id="13212A66FF9BC16642F1FF49FF7AFF10" box="[160,227,187,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9BC16642C6FEF2FD66FEF3" pageId="4" pageNumber="249" type="reference_group">
<paragraph id="8BA536E3FF9BC16642C6FEF2FD70FEDD" blockId="4.[151,767,255,311]" box="[151,745,255,280]" pageId="4" pageNumber="249">
<taxonomicName id="4C1A4D60FF9BC16642C6FEF2FE64FED2" authority="Sharp, 1887: 753" authorityName="Sharp" authorityPageNumber="753" authorityYear="1887" box="[151,509,255,280]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC16642C6FEF2FED0FEDD" box="[151,329,256,280]" italics="true" pageId="4" pageNumber="249">Bidessus curtulus</emphasis>
Sharp, 1887: 753
</taxonomicName>
(orig. descr.,
<collectingCountry id="F30D7673FF9BC16640D8FEF2FD47FEDD" box="[649,734,256,280]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
).
</paragraph>
<paragraph id="8BA536E3FF9BC16642C6FEECFD66FEF3" blockId="4.[151,767,255,311]" box="[151,767,286,311]" pageId="4" pageNumber="249">
<treatmentCitationGroup id="AB0A11CDFF9BC16642C6FEECFD66FEF3" box="[151,767,286,311]" pageId="4" pageNumber="249">
<treatmentCitation id="0ABB10F2FF9BC16642C6FEECFDABFEF3" author="Young, F. N." box="[151,562,286,311]" page="3" pageId="4" pageNumber="249" year="1969">
<taxonomicName id="4C1A4D60FF9BC16642C6FEECFDABFEF3" ID-CoL="46D87" authority=": Young 1969: 3" authorityName=": Young" authorityPageNumber="3" authorityYear="1969" box="[151,562,286,311]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC16642C6FEECFE09FEF3" box="[151,400,286,310]" italics="true" pageId="4" pageNumber="249">Neoclypeodytes curtulus</emphasis>
:
<bibRefCitation id="EF8B4B12FF9BC16643CDFEEDFDABFEF3" author="Young, F. N." box="[412,562,286,311]" pageId="4" pageNumber="249" pagination="1 - 5" refId="ref5881" refString="Young, F. N. (1969) A checklist of the American Bidessini (Coleoptera: Dytiscidae - Hydroporinae). Smithsonian Contributions to Zoology, 33, 1 - 5. https: // doi. org / 10.5479 / si. 00810282.33" type="journal article" year="1969">Young 1969: 3</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9BC16642C6FE97FB5BFC5E" pageId="4" pageNumber="249" type="materials_examined">
<paragraph id="8BA536E3FF9BC16642C6FE97FBC4FC96" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC16642C6FE97FEDEFEBB" bold="true" box="[151,327,356,383]" pageId="4" pageNumber="249">Type material.</emphasis>
<materialsCitation id="3B723CBEFF9BC1664301FE97FB7FFDCA" ID-GBIF-Occurrence="2986026309" collectorName="B. C. A. Col. &amp; K. B. Miller" country="Panama" location="Taboga island" municipality="Taboga Isl." pageId="4" pageNumber="249" specimenCount="4" typeStatus="lectotype">
<typeStatus id="54A18841FF9BC1664301FE97FE5DFEBA" box="[336,452,357,383]" pageId="4" pageNumber="249" type="lectotype">Lectotype</typeStatus>
: m# (designated by
<bibRefCitation id="EF8B4B12FF9BC16640FDFE96FCAEFEBA" author="Miller, K. B." box="[684,823,356,383]" pageId="4" pageNumber="249" pagination="87 - 123" refId="ref5642" refString="Miller, K. B. (2001) Revision and phylogenetic analysis of the New World genus Neoclypeodytes Young (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Systematic Entomology, 26, 87 - 123. https: // doi. org / 10.1046 / j. 1365 - 3113.2001.00144. x" type="journal article" year="2001">Miller 2001</bibRefCitation>
), “
<taxonomicName id="4C1A4D60FF9BC1664104FE96FC22FEBB" authorityName="Sharp" authorityYear="1882" box="[853,955,356,382]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
cur- / tulus. Types D.S. /
<location id="8EC56038FF9BC16646B6FE96FA11FEBB" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC16646B6FE96FA11FEBB" box="[1255,1416,356,382]" country="Panama" municipality="Taboga Isl." name="Taboga island" pageId="4" pageNumber="249">Taboga island</location>
. / Champion [Sharps hw] //
<collectorName id="26EF5335FF9BC1664392FE7BFDD0FE67" box="[451,585,392,419]" pageId="4" pageNumber="249">B.C.A. Col.</collectorName>
I.2. /
<taxonomicName id="4C1A4D60FF9BC16640DCFE7AFD68FE67" authorityName="Sharp" authorityYear="1882" box="[653,753,392,418]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
/ curtulus, / Sharp. [p] //
<collectingMunicipality id="6BC1AC99FF9BC166465EFE7AFB17FE67" box="[1039,1166,392,418]" pageId="4" pageNumber="249">Taboga Isl.</collectingMunicipality>
, /
<collectingCountry id="F30D7673FF9BC16646FBFE7BFA9FFE66" box="[1194,1286,393,419]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
. / Champion. [p] // Sharp Coll / 1905-313. [p] // Type [p, round label with red frame] //
<typeStatus id="54A18841FF9BC1664197FE5FFBAFFE02" box="[966,1078,429,455]" pageId="4" pageNumber="249" type="lectotype">Lectotype</typeStatus>
specimen / dissected and on red / point. Other specimen / (White point) is a /
<typeStatus id="54A18841FF9BC16640C8FE22FCADFE2F" box="[665,820,464,490]" pageId="4" pageNumber="249" type="paralectotype">paralectotype</typeStatus>
. [p, pink label] KBM [hw] //
<typeStatus id="54A18841FF9BC16646D2FE23FABAFE2E" box="[1155,1315,465,491]" pageId="4" pageNumber="249" type="lectotype">LECTOTYPE</typeStatus>
/
<taxonomicName id="4C1A4D60FF9BC166476BFE22FF6AFDCB" authorityName="Sharp" authorityYear="1887" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC166476BFE22FF6AFDCB" italics="true" pageId="4" pageNumber="249">Bidessus curtulus</emphasis>
</taxonomicName>
/ Sharp 1887 / designated by /
<collectorName id="26EF5335FF9BC1664005FE07FD4FFDCB" box="[596,726,500,527]" pageId="4" pageNumber="249">K.B. Miller</collectorName>
1997 [p, pink label with double black frame]”
</materialsCitation>
.
<materialsCitation id="3B723CBEFF9BC16646A0FE06FD09FDF7" ID-GBIF-Occurrence="2986026305" collectorName="B. C. A. Col. &amp; K. B. Miller" country="Panama" location="Taboga island" municipality="Taboga Isl." pageId="4" pageNumber="249" specimenCount="1" typeStatus="paralectotype">
<typeStatus id="54A18841FF9BC16646A0FE06FA00FDCB" box="[1265,1433,500,526]" pageId="4" pageNumber="249" type="paralectotype">Paralectotypes</typeStatus>
:
<specimenCount id="9D1CFD6AFF9BC16642C6FDEBFE8FFDF7" box="[151,278,536,563]" pageId="4" pageNumber="249" type="generic">1 specimen</specimenCount>
with same label data as lectotype
</materialsCitation>
;
<materialsCitation id="3B723CBEFF9BC16640CDFDEBFBDEFD5A" ID-GBIF-Occurrence="2986026302" collectorName="B. C. A. Col. &amp; K. B. Miller" country="Panama" county="Sharp." location="Taboga island" municipality="Sharp Coll" pageId="4" pageNumber="249" specimenCount="3" typeStatus="paralectotype">
<specimenCount id="9D1CFD6AFF9BC16640CDFDEBFCB1FDF7" box="[668,808,536,563]" pageId="4" pageNumber="249" type="generic">3 specimens</specimenCount>
, “
<taxonomicName id="4C1A4D60FF9BC166416DFDEAFC3BFDF7" authorityName="Sharp" authorityYear="1882" box="[828,930,536,562]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
cur- / tulus D.S. /
<location id="8EC56038FF9BC166463EFDEAFA97FDF7" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC166463EFDEAFA97FDF7" box="[1135,1294,536,562]" country="Panama" county="Sharp." municipality="Sharp Coll" name="Taboga island" pageId="4" pageNumber="249">Taboga island</location>
. /
<location id="8EC56038FF9BC1664779FDEAFA04FDF7" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC1664779FDEAFA04FDF7" box="[1320,1437,536,562]" country="Panama" county="Sharp." municipality="Sharp Coll" name="Champion" pageId="4" pageNumber="249">Champion</location>
[
<location id="8EC56038FF9BC16642CFFDCEFF6CFD93" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC16642CFFDCEFF6CFD93" box="[158,245,572,598]" country="Panama" county="Sharp." municipality="Sharp Coll" name="Sharp's" pageId="4" pageNumber="249">Sharps</location>
hw] // Co- / type [p, round label with yellow frame] //
<location id="8EC56038FF9BC1664106FDCEFC4CFD93" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC1664106FDCEFC4CFD93" box="[855,981,572,598]" country="Panama" county="Sharp." municipality="Sharp Coll" name="Taboga Isl." pageId="4" pageNumber="249">Taboga Isl.</location>
, /
<collectingCountry id="F30D7673FF9BC16641BEFDCFFBD2FD92" box="[1007,1099,573,599]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
. /
<location id="8EC56038FF9BC1664635FDCEFB79FD93" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC1664635FDCEFB79FD93" box="[1124,1248,572,598]" country="Panama" county="Sharp." municipality="Sharp Coll" name="Champion." pageId="4" pageNumber="249">Champion.</location>
[p] //
<collectingMunicipality id="6BC1AC99FF9BC1664775FDCEFA05FD93" box="[1316,1436,572,598]" pageId="4" pageNumber="249">Sharp Coll</collectingMunicipality>
/ 1905-313. [p] //
<collectorName id="26EF5335FF9BC166430BFD93FE46FDBF" box="[346,479,608,635]" pageId="4" pageNumber="249">B.C.A. Col.</collectorName>
I.2. /
<taxonomicName id="4C1A4D60FF9BC166404CFD92FD18FDBF" authorityName="Sharp" authorityYear="1882" box="[541,641,608,634]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
/ curtulus, /
<collectingCounty id="62C44E6FFF9BC166415BFD92FCCAFDBF" box="[778,851,608,634]" pageId="4" pageNumber="249">Sharp.</collectingCounty>
[p] //
<typeStatus id="54A18841FF9BC16641C4FD93FBE4FDBE" box="[917,1149,609,635]" pageId="4" pageNumber="249" type="paralectotype">PARALECTOTYPE</typeStatus>
/
<taxonomicName id="4C1A4D60FF9BC16646C0FD92FA05FDBF" authority="Sharp" authorityName="Sharp" authorityYear="1887" box="[1169,1436,608,634]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC16646C0FD92FACCFDBF" box="[1169,1365,608,634]" italics="true" pageId="4" pageNumber="249">Bidessus curtulus</emphasis>
Sharp
</taxonomicName>
/ 1887, designated by /
<collectorName id="26EF5335FF9BC16643F6FD77FDB2FD5B" box="[423,555,644,671]" pageId="4" pageNumber="249">K.B. Miller</collectorName>
1997 [p, blue label with double black frame]”
</materialsCitation>
;
<materialsCitation id="3B723CBEFF9BC1664605FD77FDAFFCCE" ID-GBIF-Occurrence="2986026303" collectorName="K. B. Miller" country="Panama" county="Sharp Coll" location="Taboga island" municipality="Sharp's" pageId="4" pageNumber="249" specimenCount="1" typeStatus="paralectotype">
<specimenCount id="9D1CFD6AFF9BC1664605FD77FB40FD5B" box="[1108,1241,644,671]" pageId="4" pageNumber="249" type="generic">1 specimen</specimenCount>
, “
<taxonomicName id="4C1A4D60FF9BC16646BEFD76FACCFD5B" authorityName="Sharp" authorityYear="1882" box="[1263,1365,644,670]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
cur- / tulus. /
<location id="8EC56038FF9BC16642BDFD5AFE13FD07" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC16642BDFD5AFE13FD07" box="[236,394,680,706]" country="Panama" county="Sharp Coll" municipality="Sharp's" name="Taboga island" pageId="4" pageNumber="249">Taboga island</location>
/
<collectingCountry id="F30D7673FF9BC16643F3FD5BFE67FD06" box="[418,510,681,707]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
.
<location id="8EC56038FF9BC166405BFD5AFDE6FD07" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC166405BFD5AFDE6FD07" box="[522,639,680,706]" country="Panama" county="Sharp Coll" municipality="Sharp's" name="Champion" pageId="4" pageNumber="249">Champion</location>
[
<collectingMunicipality id="6BC1AC99FF9BC16640DCFD5AFD7DFD07" box="[653,740,680,706]" pageId="4" pageNumber="249">Sharps</collectingMunicipality>
hw] // Co- / type [p, round label with yellow frame] //
<collectingCounty id="62C44E6FFF9BC166470AFD5AFF5EFD23" pageId="4" pageNumber="249">Sharp Coll</collectingCounty>
/ 1905-313. [p] //
<typeStatus id="54A18841FF9BC16643C9FD3FFD19FD22" box="[408,640,717,743]" pageId="4" pageNumber="249" type="paralectotype">PARALECTOTYPE</typeStatus>
/
<taxonomicName id="4C1A4D60FF9BC16640C7FD3EFC3DFD23" authority="Sharp" authorityName="Sharp" authorityYear="1887" box="[662,932,716,742]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC16640C7FD3EFCC5FD23" box="[662,860,716,742]" italics="true" pageId="4" pageNumber="249">Bidessus curtulus</emphasis>
Sharp
</taxonomicName>
/ 1887, designated by /
<collectorName id="26EF5335FF9BC16646E5FD3FFAAEFD23" box="[1204,1335,716,743]" pageId="4" pageNumber="249">K.B. Miller</collectorName>
1997 [p, blue label with double black frame]”
</materialsCitation>
;
<materialsCitation id="3B723CBEFF9BC1664013FD03FB96FCEA" ID-GBIF-Occurrence="2986026304" collectorName="B. C. A. Col." country="Panama" location="Taboga Isl." pageId="4" pageNumber="249" specimenCount="21" typeStatus="paralectotype">
<specimenCount id="9D1CFD6AFF9BC1664013FD03FD44FCCF" box="[578,733,752,779]" pageId="4" pageNumber="249" type="generic">21 specimens</specimenCount>
, “Co- / type [p, round label with yellow frame] //
<location id="8EC56038FF9BC166475BFD02FA11FCCF" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC166475BFD02FA11FCCF" box="[1290,1416,752,778]" country="Panama" name="Taboga Isl." pageId="4" pageNumber="249">Taboga Isl.</location>
, /
<collectingCountry id="F30D7673FF9BC16642C6FCE7FF6AFCEA" box="[151,243,789,815]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
. / Champion. [p] //
<collectorName id="26EF5335FF9BC1664380FCE7FDC1FCEB" box="[465,600,788,815]" pageId="4" pageNumber="249">B.C.A. Col.</collectorName>
I.2. /
<taxonomicName id="4C1A4D60FF9BC16640CDFCE6FC99FCEB" authorityName="Sharp" authorityYear="1882" box="[668,768,788,814]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
/ curtulus, / Sharp. [p]”
</materialsCitation>
;
<materialsCitation id="3B723CBEFF9BC166464DFCE7FBC0FC96" ID-GBIF-Occurrence="2986026307" collectionCode="BMNH" collectorName="B. C. A. Col." country="Panama" county="Champion." location="San Miguel" municipality="Pearl Isl." pageId="4" pageNumber="249" specimenCount="3" typeStatus="paralectotype">
<specimenCount id="9D1CFD6AFF9BC166464DFCE7FB33FCEB" box="[1052,1194,788,815]" pageId="4" pageNumber="249" type="generic">3 specimens</specimenCount>
, “
<location id="8EC56038FF9BC16646EEFCE7FADEFCEB" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC16646EEFCE7FADEFCEB" box="[1215,1351,788,815]" country="Panama" county="Champion." municipality="Pearl Isl." name="San Miguel" pageId="4" pageNumber="249">San Miguel</location>
, /
<collectingMunicipality id="6BC1AC99FF9BC1664732FCE6FF23FC97" pageId="4" pageNumber="249">Pearl Isl.</collectingMunicipality>
/
<collectingCounty id="62C44E6FFF9BC1664281FCCAFED5FC97" box="[208,332,824,850]" pageId="4" pageNumber="249">Champion.</collectingCounty>
[p] //
<collectorName id="26EF5335FF9BC16643C0FCCBFD8FFC97" box="[401,534,824,851]" pageId="4" pageNumber="249">B.C.A. Col.</collectorName>
I.2. /
<taxonomicName id="4C1A4D60FF9BC1664009FCCAFD25FC97" authorityName="Sharp" authorityYear="1882" box="[600,700,824,850]" class="Insecta" family="Dytiscidae" genus="Bidessus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="genus">Bidessus</taxonomicName>
/ curtulus, / Sharp. [p]” (all
<collectionCode id="ED0BAE26FF9BC16641A9FCCBFBC8FC96" box="[1016,1105,825,851]" pageId="4" pageNumber="249">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BA536E3FF9BC1664296FCAEFB5BFC5E" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC1664296FCAEFDA0FCB3" bold="true" box="[199,569,860,886]" pageId="4" pageNumber="249">Additional material examined.</emphasis>
<materialsCitation id="3B723CBEFF9BC1664013FCAFFB26FC5E" ID-GBIF-Occurrence="2986026301" collectingDate="2007-06-19" collectionCode="CKSB, NMPC, ZSMG" country="Panama" elevation="270" location="Campana" municipality="Anam" pageId="4" pageNumber="249" specimenCount="7">
<specimenCount id="9D1CFD6AFF9BC1664013FCAFFD4BFCB3" box="[578,722,860,887]" pageId="4" pageNumber="249" type="generic">7 specimens</specimenCount>
: “
<collectingCountry id="F30D7673FF9BC16640B8FCAFFCD0FCB2" box="[745,841,861,887]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
:
<collectingCountry id="F30D7673FF9BC1664104FCAFFC28FCB2" box="[853,945,861,887]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
,
<location id="8EC56038FF9BC16641EFFCAFFBB2FCB2" LSID="urn:lsid:plazi:treatment:03B387F5FF9BC16642C6FF65FC57F8DA:8EC56038FF9BC16641EFFCAFFBB2FCB2" box="[958,1067,861,887]" country="Panama" municipality="Anam" name="Campana" pageId="4" pageNumber="249">Campana</location>
, / cascade below
<collectingMunicipality id="6BC1AC99FF9BC16646A7FCAFFAA3FCB2" box="[1270,1338,861,887]" pageId="4" pageNumber="249">Anam</collectingMunicipality>
office, /
<quantity id="4CE29B06FF9BC16642C6FC73FF43FC5E" box="[151,218,897,923]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="249" unit="m" value="270.0">
<elevation id="0037D1D0FF9BC16642C6FC73FF43FC5E" box="[151,218,897,923]" metricMagnitude="2" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="249" unit="m" value="270.0">270m</elevation>
</quantity>
,
<date id="FFA41023FF9BC16642B4FC72FEC6FC5F" box="[229,351,896,922]" pageId="4" pageNumber="249" value="2007-06-19">
<collectingDate id="EFE0E9CBFF9BC16642B4FC72FEC6FC5F" box="[229,351,896,922]" pageId="4" pageNumber="249" value="2007-06-19">19.vi.2007</collectingDate>
</date>
, / 08.40.818N 079.54.075W, / Balke (PAN 07) [p]” (
<collectionCode id="ED0BAE26FF9BC16641FFFC73FC62FC5E" box="[942,1019,897,923]" pageId="4" pageNumber="249">CKSB</collectionCode>
,
<collectionCode id="ED0BAE26FF9BC1664657FC73FBC1FC5E" box="[1030,1112,897,923]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="249">NMPC</collectionCode>
,
<collectionCode id="ED0BAE26FF9BC1664635FC73FB2EFC5E" box="[1124,1207,897,923]" pageId="4" pageNumber="249">ZSMG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9BC1664296FC56FF72F912" pageId="4" pageNumber="249" type="diagnosis">
<paragraph id="8BA536E3FF9BC1664296FC56FE43FA56" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC1664296FC56FEDBFC7B" bold="true" box="[199,322,932,958]" pageId="4" pageNumber="249">Diagnosis.</emphasis>
Small, ovoid-shaped species, only slightly attenuated posteriorly; lateral outline nearly continuous between pronotum and elytra in dorsal aspect (
<figureCitation id="13212A66FF9BC16640F3FC3AFD71FC26" box="[674,744,968,995]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Figs 2</figureCitation>
AB). Surface shiny. Elytron rufopiceous with two testaceous maculae; subbasal macula transversely quadrangular, longitudinally extended at lateral margin, narrower basomedially, not reaching suture; posterior macula longitudinal-oblique, expanded medially in apical half, reaching elytral apex. Head finely reticulated and punctate. Pronotum widest at base; lateral bead distinct; finely punctate over entire surface, basal striae sinuated, distinctly curved inwards near base, length about 0.60× length of pronotum measured along same line as stria. Elytra with basal striae straight, slightly shorter than those on pronotum; sutural striae absent; epipleura with moderately oblique humeral carina. Metaventrite sparsely and rather finely punctate over entire surface; metacoxal plates with punctate medial and lateral portions, broadly impunctate along anterior and posterior margins (
<figureCitation id="13212A66FF9BC1664338FB1AFE5BFAC6" box="[361,450,1256,1283]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Fig. 2C</figureCitation>
). Abdominal ventrites I and II coarsely punctate; ventrite VI unmodified. Median lobe in lateral view expanded medially, apical third linear with lateral margins convergent to acute apex (
<figureCitation id="13212A66FF9BC166469BFAFEFAB9FAE2" box="[1226,1312,1292,1319]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Fig. 2E</figureCitation>
); in ventral view with attenuation in apical fifth, then evenly convergent to obtusely pointed apex; ventral surface in apical part with numerous scale-like setae (
<figureCitation id="13212A66FF9BC1664051FAA6FDC2FAAA" box="[512,603,1364,1391]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Fig. 2D</figureCitation>
). Lateral lobe with apical segment almost straight and parallel-sided, apex slightly protruding (
<figureCitation id="13212A66FF9BC1664329FA8AFE54FA56" box="[376,461,1400,1427]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="8BA536E3FF9BC1664296FA6EFDE7F9E6" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<taxonomicName id="4C1A4D60FF9BC1664296FA6EFE40FA73" authorityName=": Young" authorityYear="1969" box="[199,473,1436,1462]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC1664296FA6EFE40FA73" box="[199,473,1436,1462]" italics="true" pageId="4" pageNumber="249">Neoclypeodytes curtulus</emphasis>
</taxonomicName>
is easily distinguished from the other two Panamanian species by the smaller body size (TL:
<quantity id="4CE29B06FF9BC1664280FA32FEEFFA1E" box="[209,374,1472,1499]" metricMagnitude="-3" metricUnit="m" metricValue="1.725" metricValueMax="1.8" metricValueMin="1.65" pageId="4" pageNumber="249" unit="mm" value="1.725" valueMax="1.8" valueMin="1.65">1.651.80 mm</quantity>
; TL-h: 1.451.60; TW: 0.901.00 mm), characteristic elytral color pattern (
<figureCitation id="13212A66FF9BC16646EBFA32FA99FA1E" box="[1210,1280,1472,1499]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Figs 2</figureCitation>
AB), and the male genitalia with the median lobe expanded medially, and the apex of the lateral lobe protruding but less distinctly than in
<taxonomicName id="4C1A4D60FF9BC16642BAF9FBFE11F9E6" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[235,392,1545,1571]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="fortunensis" status="sp. nov.">
<emphasis id="B96EEAF1FF9BC16642BAF9FBFE11F9E6" box="[235,392,1545,1571]" italics="true" pageId="4" pageNumber="249">N. fortunensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9BC16643DEF9FBFE7FF9E6" box="[399,486,1545,1571]" pageId="4" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13212A66FF9BC16643A5F9FAFDA2F9E6" box="[500,571,1544,1571]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Figs 2</figureCitation>
DF).
</paragraph>
<paragraph id="8BA536E3FF9BC1664296F9DFFED6F94A" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC1664296F9DFFE9FF982" bold="true" box="[199,262,1581,1607]" pageId="4" pageNumber="249">Note.</emphasis>
<bibRefCitation id="EF8B4B12FF9BC166435FF9DEFE3EF982" author="Miller, K. B." box="[270,423,1580,1607]" pageId="4" pageNumber="249" pagination="87 - 123" refId="ref5642" refString="Miller, K. B. (2001) Revision and phylogenetic analysis of the New World genus Neoclypeodytes Young (Coleoptera: Dytiscidae: Hydroporinae: Bidessini). Systematic Entomology, 26, 87 - 123. https: // doi. org / 10.1046 / j. 1365 - 3113.2001.00144. x" type="journal article" year="2001">Miller (2001)</bibRefCitation>
describes the subbasal elytral maculae as triangular in
<taxonomicName id="4C1A4D60FF9BC166464AF9DFFB00F983" authorityName=": Young" authorityYear="1969" box="[1051,1177,1580,1606]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="curtulus">
<emphasis id="B96EEAF1FF9BC166464AF9DFFB00F983" box="[1051,1177,1580,1606]" italics="true" pageId="4" pageNumber="249">N. curtulus</emphasis>
</taxonomicName>
, which is however not completely accurate: although the macula is usually somewhat attenuated basomedially, it is never acute triangular (cf.
<figureCitation id="13212A66FF9BC1664295F986FE93F94A" box="[196,266,1652,1679]" captionStart="FIGURE 2" captionStartId="5.[151,250,1914,1938]" captionTargetBox="[164,1419,198,1882]" captionTargetId="figure-17@5.[151,1435,181,1890]" captionTargetPageId="5" captionText="FIGURE 2. Neoclypeodytes curtulus (Sharp, 1887) A. dorsal habitus of the lectotype; B. dorsal habitus of specimen from Campana; C. ventral habitus; D. median lobe of aedeagus in ventral view; E. median lobe of aedeagus in lateral view; F. lateral lobe (paramere). Scale bars: AC = 1 mm; DF = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4301873" httpUri="https://zenodo.org/record/4301873/files/figure.png" pageId="4" pageNumber="249">Figs 2</figureCitation>
AB).
</paragraph>
<paragraph id="8BA536E3FF9BC1664296F96AFF72F912" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC1664296F96AFE61F977" bold="true" box="[199,504,1688,1714]" pageId="4" pageNumber="249">Collection circumstances.</emphasis>
The recently collected specimens were found together with
<taxonomicName id="4C1A4D60FF9BC16646F2F96BFA91F977" authority="Scheers &amp; Hájek, 2020" authorityName="Scheers &amp; Hájek" authorityYear="2020" box="[1187,1288,1688,1714]" class="Insecta" family="Dytiscidae" genus="Neoclypeodytes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="249" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B96EEAF1FF9BC16646F2F96BFA91F977" box="[1187,1288,1688,1714]" italics="true" pageId="4" pageNumber="249">N. balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D578AFF9BC1664740F96BFAF1F976" box="[1297,1384,1689,1715]" pageId="4" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
(see above).
</paragraph>
</subSubSection>
<subSubSection id="C3006568FF9BC1664296F912FC57F8DA" pageId="4" pageNumber="249" type="distribution">
<paragraph id="8BA536E3FF9BC1664296F912FC57F8DA" blockId="4.[151,1437,356,1823]" pageId="4" pageNumber="249">
<emphasis id="B96EEAF1FF9BC1664296F912FEF8F93F" bold="true" box="[199,353,1760,1786]" pageId="4" pageNumber="249">Distribution.</emphasis>
The species was described and known so far only from two islands in Pacific Gulf of
<collectingCountry id="F30D7673FF9BC166476DF913FA01F93E" box="[1340,1432,1761,1787]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
; here we present the first record from continental Central
<collectingCountry id="F30D7673FF9BC1664145F8F7FCF4F8DA" box="[788,877,1797,1823]" name="Panama" pageId="4" pageNumber="249">Panama</collectingCountry>
(
<figureCitation id="13212A66FF9BC166412DF8F6FC59F8DA" box="[892,960,1796,1823]" captionStart="FIGURE 6" captionStartId="10.[151,250,852,876]" captionTargetBox="[151,1435,181,829]" captionTargetId="figure-19@10.[151,1435,181,829]" captionTargetPageId="10" captionText="FIGURE 6. Map of distribution of Neoclypeodytes species in Panama." figureDoi="http://doi.org/10.5281/zenodo.4301881" httpUri="https://zenodo.org/record/4301881/files/figure.png" pageId="4" pageNumber="249">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>