114 lines
14 KiB
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114 lines
14 KiB
XML
<document id="DADD3875297B3FE6539548A82A9A6437" ID-DOI="10.26879/1220" ID-ISSN="1094-8074" ID-Zenodo-Dep="10962250" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="admin" IM.taxonomicNames_approvedBy="admin" IM.treatments_approvedBy="admin" checkinTime="1712859670786" checkinUser="felipe" docAuthor="Jacisin Iii, John J. & Lawing, A. Michelle" docDate="2024" docId="03B387E8FFA5320484F9F8F89B54FA5B" docLanguage="en" docName="PalaeontolElectron.27.1.a2.pdf" docOrigin="Palaeontologia Electronica (a 2) 27 (1)" docSource="http://dx.doi.org/10.26879/1220" docStyle="DocumentStyle:322FC57CC99045555090E8A84CFB029A.4:PalaeontolElectron.2018-.journal_article" docStyleId="322FC57CC99045555090E8A84CFB029A" docStyleName="PalaeontolElectron.2018-.journal_article" docStyleVersion="4" docTitle="Pantherophis kansensis Gilmore 1938" docType="treatment" docVersion="5" lastPageNumber="16" masterDocId="FF8AFF90FFA8320B8770FFDA9964FFD9" masterDocTitle="Fossil snakes of the Penny Creek Local Fauna from Webster County, Nebraska, USA, and the first record of snakes from the Early Clarendonian (12.5 - 12 Ma) of North America" masterLastPageNumber="42" masterPageNumber="1" pageNumber="14" updateTime="1715215045661" updateUser="admin" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:titleInfo id="B5C164E2A93334E9EA7BD241788E28B4">
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<mods:title id="1031EFE866E7B464DDAF702FC0F0655C">Fossil snakes of the Penny Creek Local Fauna from Webster County, Nebraska, USA, and the first record of snakes from the Early Clarendonian (12.5 - 12 Ma) of North America</mods:title>
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<mods:namePart id="EB38BE05E9E88B2174464BCF0ABC890A">Jacisin Iii, John J.</mods:namePart>
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<mods:name id="EEFEDD83552D0EA7F145653C6ADFDB42" type="personal">
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<mods:namePart id="8B68D79E76778A743F8E75E9F457A459">Lawing, A. Michelle</mods:namePart>
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<mods:typeOfResource id="EAE03CB17E3F371721C7D9B856D7AE3B">text</mods:typeOfResource>
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<mods:title id="F66D24FF433358D37EFAB809B9E7F660">Palaeontologia Electronica</mods:title>
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<mods:part id="E277FB4EBCCE69F25C101265A2FA7616">
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<mods:date id="D9F9361D67F3196A1DDAF7CE06E9CCFA">2024</mods:date>
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<mods:detail id="E9FEA6BB41ABF799E6A1CAAAF8FD1A80" type="series">
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<mods:title id="1940FA4F0229DD49C6369CB3B18201E5">a 2</mods:title>
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<mods:detail id="0C77F2EF0EEE16E878EB6F5A372085A6" type="volume">
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<mods:number id="C471395BDDA40C5E51A2651B402A6956">27</mods:number>
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<mods:number id="AB85C0DDA4BCE6147C181BD123BDBC15">1</mods:number>
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<mods:url id="8DF91FFB668BED9BC44CD38CCBDF85C4">http://dx.doi.org/10.26879/1220</mods:url>
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<mods:classification id="6927184D128D1DEDAD4740CFB1223F77">journal article</mods:classification>
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<mods:identifier id="A2BA0BB8AF1DE54D809B124455E1BAB0" type="DOI">10.26879/1220</mods:identifier>
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<mods:identifier id="B9621FC8596878848BF95DC94510A4E2" type="ISSN">1094-8074</mods:identifier>
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<treatment id="03B387E8FFA5320484F9F8F89B54FA5B" LSID="urn:lsid:plazi:treatment:03B387E8FFA5320484F9F8F89B54FA5B" httpUri="http://treatment.plazi.org/id/03B387E8FFA5320484F9F8F89B54FA5B" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<subSubSection id="C3006575FFA5320684F9F8F89C3BF8EA" box="[905,1375,1823,1852]" pageId="13" pageNumber="14" type="nomenclature">
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<paragraph id="8BA536FEFFA5320684F9F8F89C3BF8EA" blockId="13.[905,1375,1823,1883]" box="[905,1375,1823,1852]" pageId="13" pageNumber="14">
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<heading id="D0ED8192FFA5320684F9F8F89C3BF8EA" box="[905,1375,1823,1852]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="14" reason="4">
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<taxonomicName id="4C1A4D7DFFA5320684F9F8F89C37F8E2" authority="Gilmore, 1938" authorityName="Gilmore" authorityYear="1938" box="[905,1363,1826,1852]" class="Reptilia" family="Colubridae" genus="Pantherophis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="kansensis">
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<emphasis id="B96EEAECFFA5320684F9F8F89DC0F8E2" box="[905,1188,1826,1851]" italics="true" pageId="13" pageNumber="14">Pantherophis kansensis</emphasis>
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<bibRefCitation id="EF8B4B0FFFA5320683DCF8F89C37F8E2" author="Gilmore, C. W." box="[1196,1363,1826,1852]" pageId="13" pageNumber="14" pagination="1 - 96" refId="ref19033" refString="Gilmore, C. W. 1938. Fossil snakes of North America. Geological Society of North America Special Paper, 9: 1 - 96. https: // doi. org / 10.1130 / spe 9 - p 1" type="journal article" year="1938">Gilmore, 1938</bibRefCitation>
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</taxonomicName>
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<superScript id="7C6F9BB6FFA532068223F8C59C3BF8EA" attach="none" box="[1363,1375,1823,1843]" fontSize="8" pageId="13" pageNumber="14">†</superScript>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006575FFA532068333F8989DC1F882" box="[1091,1189,1858,1883]" pageId="13" pageNumber="14" type="description">
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<paragraph id="8BA536FEFFA532068333F8989DC1F882" blockId="13.[905,1375,1823,1883]" box="[1091,1189,1858,1883]" pageId="13" pageNumber="14">
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<figureCitation id="13212A7BFFA532068333F8989DC1F882" box="[1091,1189,1858,1883]" captionStart="FIGURE 7" captionStartId="14.[196,287,1572,1595]" captionTargetBox="[194,1438,164,1556]" captionTargetId="figure-111@14.[194,1438,164,1556]" captionTargetPageId="14" captionText="FIGURE 7. Trunk vertebra of Pantherophis kansensis from the Penny Creek local fauna. From top left: dorsal, ventral, anterior, posterior, and lateral views (anterior to the left). Scale bar equals 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10962266" httpUri="https://zenodo.org/record/10962266/files/figure.png" pageId="13" pageNumber="14">Figure 7</figureCitation>
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</paragraph>
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</subSubSection>
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<caption id="DF656676FFA6320587B4F9FE9D7CF980" ID-DOI="http://doi.org/10.5281/zenodo.10962266" ID-Zenodo-Dep="10962266" httpUri="https://zenodo.org/record/10962266/files/figure.png" pageId="14" pageNumber="15" startId="14.[196,287,1572,1595]" targetBox="[194,1438,164,1556]" targetPageId="14" targetType="figure">
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<paragraph id="8BA536FEFFA6320587B4F9FE9D7CF980" blockId="14.[196,1437,1572,1625]" pageId="14" pageNumber="15">
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<emphasis id="B96EEAECFFA6320587B4F9FE985DF9E5" bold="true" box="[196,313,1572,1596]" pageId="14" pageNumber="15">FIGURE 7.</emphasis>
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Trunk vertebra of
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<taxonomicName id="4C1A4D7DFFA63205868AF9FF9B9DF9E5" authorityName="Gilmore" authorityYear="1938" box="[506,761,1573,1596]" class="Reptilia" family="Colubridae" genus="Pantherophis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="15" phylum="Chordata" rank="species" species="kansensis">
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<emphasis id="B96EEAECFFA63205868AF9FF9B9DF9E5" box="[506,761,1573,1596]" italics="true" pageId="14" pageNumber="15">Pantherophis kansensis</emphasis>
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</taxonomicName>
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from the Penny Creek local fauna. From top left: dorsal, ventral, anterior, posterior, and lateral views (anterior to the left). Scale bar equals 1 mm.
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</paragraph>
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</caption>
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<subSubSection id="C3006575FFA6320587B0F9469879F90C" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="8BA536FEFFA6320587B0F9469879F90C" blockId="14.[192,792,1692,1909]" pageId="14" pageNumber="15">
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<emphasis id="B96EEAECFFA6320587B0F9469848F96C" bold="true" box="[192,300,1692,1717]" pageId="14" pageNumber="15">Material.</emphasis>
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<collectionCode id="ED0BAE3BFFA632058644F94698E7F96F" box="[308,387,1692,1718]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="14" pageNumber="15">UNSM</collectionCode>
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139983 (21 pre-cloacal trunk vertebrae).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3006575FFA6320487B0F9069B09FBBB" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BA536FEFFA6320587B0F9069AB5F8EC" blockId="14.[192,792,1692,1909]" lastBlockId="14.[840,1440,1692,1909]" pageId="14" pageNumber="15">
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<emphasis id="B96EEAECFFA6320587B0F906983EF92C" bold="true" box="[192,346,1756,1781]" pageId="14" pageNumber="15">Description.</emphasis>
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In anterior view, the neural spine is relatively low for a colubrid, the neural arch is moderately vaulted, and the zygosphene is convex dorsally. The zygosphenal articular facets are strongly tilted dorso-ventrally. The neural canal is rounded and slightly smaller than the cotyle, which is also round. The prezygapophyseal articular facets tilt slightly upward, and the diapophyses and parapophyses are rounded, distinct sections of the synapophyses.
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</paragraph>
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<paragraph id="8BA536FEFFA6320484F2F8E6984EFE5B" blockId="14.[840,1440,1692,1909]" lastBlockId="15.[192,793,169,1410]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">In dorsal view, the neural spine is long and located in the middle of the vertebra. The anterior edge of the zygosphene is concave. The prezygapophyseal accessory processes are moderately pointed, the prezygapophyseal articular facets are rounded to oval and tilted slightly upward, and the epizygapophyseal spines are absent. The posterior edge of the neural arch in dorsal view is slightly rounded.</paragraph>
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<paragraph id="8BA536FEFFA73204878AFE539812FD5B" blockId="15.[192,793,169,1410]" pageId="15" pageNumber="16">In lateral view, the neural spine is approximately twice as long as it is high. The condyle is tilted slightly forward. The hemal keel is distinct from the centrum, and the synapophyses can clearly be divided into diapophyseal and parapophyseal sections. The interzygapophyseal ridge is bowed downward, and the subcentral ridge is bowed upward.</paragraph>
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<paragraph id="8BA536FEFFA73204878AFD539A65FCBB" blockId="15.[192,793,169,1410]" pageId="15" pageNumber="16">In posterior view, the neural arch is somewhat vaulted. The condyle is round and approximately the same size as the neural canal, which is an inverted U-shape. The zygosphenal articular facets and the postzygapophyseal articular facets are all tilted sharply upward. The diapophyses and parapophyses are distinct parts of the synapophyses.</paragraph>
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<paragraph id="8BA536FEFFA73204878AFCB39B09FBBB" blockId="15.[192,793,169,1410]" pageId="15" pageNumber="16">In ventral view, the centrum is triangular and bordered by visible subcentral ridges. The hemal keel is well-developed, moderately wide, and constricted in the middle relative to the ends. The prezygapophyseal accessory processes are distinct, oblique, and directed anterolaterally. As in other views, the synapophyses are distinctly divided into the diapophyses and parapophyses.</paragraph>
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</subSubSection>
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<subSubSection id="C3006575FFA7320487B0FBB39B54FA5B" pageId="15" pageNumber="16" type="discussion">
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<paragraph id="8BA536FEFFA7320487B0FBB39B54FA5B" blockId="15.[192,793,169,1410]" pageId="15" pageNumber="16">
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<emphasis id="B96EEAECFFA7320487B0FBB3985CFB5B" bold="true" box="[192,312,1129,1154]" pageId="15" pageNumber="16">Remarks.</emphasis>
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<taxonomicName id="4C1A4D7DFFA732048634FBB39B00FB5B" authorityName="Gilmore" authorityYear="1938" box="[324,612,1129,1154]" class="Reptilia" family="Colubridae" genus="Pantherophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="kansensis">
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<emphasis id="B96EEAECFFA732048634FBB39B00FB5B" box="[324,612,1129,1154]" italics="true" pageId="15" pageNumber="16">Pantherophis kansensis</emphasis>
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</taxonomicName>
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has vertebrae with a lower neural spine and more anterolaterally directed prezygapophyseal accessory processes than any other known species of
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<taxonomicName id="4C1A4D7DFFA732048529FB139B92FB3B" authorityName="Fitzinger" authorityYear="1843" box="[601,758,1225,1250]" class="Reptilia" family="Colubridae" genus="Pantherophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="B96EEAECFFA732048529FB139B92FB3B" box="[601,758,1225,1250]" italics="true" pageId="15" pageNumber="16">Pantherophis</emphasis>
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</taxonomicName>
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or
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<taxonomicName id="4C1A4D7DFFA7320487B0FB33982AFADB" authorityName="Dowling & Price" authorityYear="1988" box="[192,334,1257,1282]" class="Reptilia" family="Colubridae" genus="Bogertophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="genus">
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<emphasis id="B96EEAECFFA7320487B0FB33982AFADB" box="[192,334,1257,1282]" italics="true" pageId="15" pageNumber="16">Bogertophis</emphasis>
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</taxonomicName>
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. This neural spine is relatively taller in some specimens, and is more proportionally more similar to
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<taxonomicName id="4C1A4D7DFFA73204864EFAF39B3EFA9B" baseAuthorityName="Say" baseAuthorityYear="1823" box="[318,602,1321,1346]" class="Reptilia" family="Colubridae" genus="Pantherophis" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="16" phylum="Chordata" rank="species" species="obsoletus">
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<emphasis id="B96EEAECFFA73204864EFAF39B3EFA9B" box="[318,602,1321,1346]" italics="true" pageId="15" pageNumber="16">Pantherophis obsoletus</emphasis>
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</taxonomicName>
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in that regard; these vertebrae appear to be from a more anterior portion of the vertebral column.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |