1879 lines
288 KiB
XML
1879 lines
288 KiB
XML
<document id="C95F36D793B8893DEF0D1934D9A70D9F" ID-CLB-Dataset="25919" ID-DOI="10.11646/zootaxa.4660.1.1" ID-GBIF-Dataset="2334efd2-e346-48ff-85ff-0bc8661538b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3377134" ID-ZooBank="DA70C158-7FB0-4B46-9A1D-EA32D24B19B6" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566808464076" checkinUser="plazi" docAuthor="Moreno-González, Jairo A., O, Ranulfo González & D, Eduardo Flórez" docDate="2019" docId="03B387E632401011FF3EFA0EDBED3B96" docLanguage="en" docName="zootaxa.4660.1.1.imd" docOrigin="Zootaxa 4660 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Tityus columbianus" docType="treatment" docVersion="12" lastPageNumber="38" masterDocId="FF8AFF9E325C1034FFA9FFD5D8493D28" masterDocTitle="Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species" masterLastPageNumber="94" masterPageNumber="1" pageNumber="29" updateTime="1698752490567" updateUser="ExternalLinkService">
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<mods:title id="1DE406C70DFB047DF7C1590819F67A07">Taxonomic revision of the Colombian Tityus (Archaeotityus) (Scorpiones, Buthidae) species: a morphological and morphometric approach, with a description of a new species</mods:title>
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<mods:namePart id="983C641DD78510E926D6AAE1CC167B86">Moreno-González, Jairo A.</mods:namePart>
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<mods:namePart id="217D187178664973FC892091C5F5EE33">O, Ranulfo González</mods:namePart>
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<mods:namePart id="15F920C03237BC25DFE3BB4D5BEA900D">D, Eduardo Flórez</mods:namePart>
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<treatment id="03B387E632401011FF3EFA0EDBED3B96" ID-DOI="http://doi.org/10.5281/zenodo.4648584" ID-GBIF-Taxon="159243654" ID-Zenodo-Dep="4648584" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96" httpUri="http://treatment.plazi.org/id/03B387E632401011FF3EFA0EDBED3B96" lastPageId="37" lastPageNumber="38" pageId="28" pageNumber="29">
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<paragraph id="8BA536F032401028FF3EFA0EDA0C38DD" blockId="28.[151,1436,1499,1597]" box="[151,581,1499,1526]" pageId="28" pageNumber="29">
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<heading id="D0ED819C32401028FF3EFA0EDA0C38DD" bold="true" box="[151,581,1499,1526]" fontSize="11" level="1" pageId="28" pageNumber="29" reason="1">
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<taxonomicName id="4C1A4D7332401028FF3EFA0EDA0C38DD" ID-CoL="57994" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[151,581,1499,1526]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="columbianus">
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<emphasis id="B96EEAE232401028FF3EFA0EDA0C38DD" bold="true" box="[151,581,1499,1526]" pageId="28" pageNumber="29">
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<emphasis id="B96EEAE232401028FF3EFA0ED9CD38DE" bold="true" box="[151,388,1499,1526]" italics="true" pageId="28" pageNumber="29">Tityus columbianus</emphasis>
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(
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<bibRefCitation id="EF8B4B0132401028FE3DFA0EDA7438DD" author="Thorell, T." box="[404,573,1499,1525]" pageId="28" pageNumber="29" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C300657B32401028FF3EFA2ADA773B15" pageId="28" pageNumber="29" type="description">
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<paragraph id="8BA536F032401028FF3EFA2ADA773B15" blockId="28.[151,1436,1499,1597]" pageId="28" pageNumber="29">
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<figureCitation id="13212A7532401028FF3EFA2AD9713B31" box="[151,312,1534,1561]" captionStart="FIGURE 5" captionStartId="13.[151,250,1940,1965]" captionTargetBox="[199,1390,186,1902]" captionTargetId="figure@13.[179,1409,166,1921]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Archaeotityus) Lourenço, 2006, carapace coloration, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377145/files/figure.png" pageId="28" pageNumber="29">Figures 5C, D</figureCitation>
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,
|
||
<figureCitation id="13212A7532401028FEEDFA2AD9CF3B31" box="[324,390,1535,1561]" captionStart="FIGURE 6" captionStartId="14.[151,250,1837,1862]" captionTargetBox="[151,1415,186,1776]" captionTargetId="figure@14.[151,1436,181,1776]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377147/files/figure.png" pageId="28" pageNumber="29">6B, E</figureCitation>
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,
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<figureCitation id="13212A7532401028FE3BFA2AD9913B31" box="[402,472,1535,1561]" captionStart="FIGURE 7" captionStartId="15.[151,250,1942,1967]" captionTargetBox="[181,1406,194,1913]" captionTargetId="figure@15.[172,1415,166,1922]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Archaeotityus) Lourenço, 2006, chelicerae, dorsal aspect, showing reticulation pattern. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377149/files/figure.png" pageId="28" pageNumber="29">7C, D</figureCitation>
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,
|
||
<figureCitation id="13212A7532401028FE4AFA2ADA603B31" box="[483,553,1535,1561]" captionStart="FIGURE 8" captionStartId="16.[151,250,1940,1965]" captionTargetBox="[195,1393,166,1921]" captionTargetId="figure@16.[195,1393,166,1921]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Archaeotityus) Lourenço, 2006, carapace sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984. A. Female (ICN-As-740). B. Male (ICN-As-741). C, D. Tityus columbianus (Thorell, 1876). C. Female (ICN-As-739). D. Male (ICN-As-524). E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype. Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377151/files/figure.png" pageId="28" pageNumber="29">8C, D</figureCitation>
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,
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||
<figureCitation id="13212A7532401028FD9CFA2ADA323B31" box="[565,635,1535,1561]" captionStart="FIGURE 9" captionStartId="17.[151,250,1761,1786]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure@17.[151,1436,181,1741]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377153/files/figure.png" pageId="28" pageNumber="29">9C, D</figureCitation>
|
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,
|
||
<figureCitation id="13212A7532401028FD2FFA2ADA933B31" box="[646,730,1535,1561]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="28" pageNumber="29">10C, D</figureCitation>
|
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,
|
||
<figureCitation id="13212A7532401028FD4CFA2ADAB43B31" box="[741,765,1535,1561]" captionStart="FIGURE 11" captionStartId="21.[151,250,1944,1969]" captionTargetBox="[272,1316,166,1924]" captionTargetId="figure@21.[272,1316,166,1924]" captionTargetPageId="21" captionText="FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. A–C. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377157/files/figure.png" pageId="28" pageNumber="29">11</figureCitation>
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D–F, 14E, F, 15B, 16C, D, 17B, 18B, 19B, 20B, 21–23, 30, 46B, 47B, C, 48, 49;
|
||
<tableCitation id="C698034B32401028FE2CF9F6DA483B15" box="[389,513,1571,1597]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStart-2="TABLE 3" captionStartId-0="6.[151,239,550,574]" captionStartId-1="8.[151,239,478,502]" captionStartId-2="28.[151,239,730,754]" captionText-0="TABLE 1. Updated checklist of the subgenus Tityus (Archaeotityus)." captionText-1="TABLE 2. Pectinal tooth counts of Colombian Tityus (Archaeotityus) species. Note: Both pectines of each examined specimen were counted. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." captionText-2="TABLE 3. Number of movable finger oblique rows of denticles among Colombian Tityus (Archaeotityus) species. Note: Both movable fingers of each examined specimen were included. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="28" pageNumber="29">Tables 1–3</tableCitation>
|
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,
|
||
<tableCitation id="C698034B32401028FDA4F9F6DA733B15" box="[525,570,1571,1597]" captionStart-0="TABLE 6" captionStart-1="TABLE 7" captionStart-2="TABLE 8" captionStart-3="TABLE 9" captionStartId-0="81.[151,239,153,177]" captionStartId-1="83.[151,239,153,177]" captionStartId-2="84.[151,239,153,177]" captionStartId-3="85.[151,239,488,514]" captionTargetBox-1="[159,1409,261,1598]" captionTargetBox-2="[159,1384,261,1111]" captionTargetBox-3="[155,2014,561,1666]" captionTargetPageId-1="83" captionText-0="TABLE 6. Eigenvalues of a Principal Component Analysis (PCA) analysis obtained from 34 morphometric ratios of males and females of Tityus bastosi, T. columbianus, T. guane sp. nov., T. parvulus and T. tayrona, showing the values of the first five principal components. Note: Highest values are highlighted in bold. Abbreviation: PC, principal component." captionText-1="TABLE 7. Correlations matrix derived from the Principal Component Analysis (PCA) of 34 morphometric ratios from males and females of T. bastosi, T. columbianus, T. guane, T. parvulus and T. tayrona. Note: Informative correlation values are highlighted in bold." captionText-2="TABLE 8. Morphometric ratios ranges of the most influential ratios derived from the PC1 (females and males) and PC3 (males) from Tityus bastosi, T. columbianus, T. guane sp. nov., T. parvulus and T. tayrona. Note: Ranges highlighted in bold seem to be useful." captionText-3="TABLE 9. Measurements (mm) of Tityus (Archaeotityus) bastosi Lourenço, 1991, and Tityus (Archaeotityus) columbianus (Thorell, 1876). Abbreviations: F, females; M, males; SD, standard deviation." httpUri-1="http://table.plazi.org/id/DF656678320F1067FF3EFF4CD9F23DD2" httpUri-2="http://table.plazi.org/id/DF65667832081060FF3EFF4CD9C63DD2" httpUri-3="http://table.plazi.org/id/DF65667832091061FF3EFE3DD9D93F0E" pageId="28" pageNumber="29" tableUuid-1="DF656678320F1067FF3EFF4CD9F23DD2" tableUuid-2="DF65667832081060FF3EFF4CD9C63DD2" tableUuid-3="DF65667832091061FF3EFE3DD9D93F0E">6–9</tableCitation>
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.
|
||
</paragraph>
|
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</subSubSection>
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<subSubSection id="C300657B32401028FF3EF9B5DB6F3A65" pageId="28" pageNumber="29" type="reference_group">
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<paragraph id="8BA536F032401028FF3EF9B5DA6B3B51" blockId="28.[151,546,1632,1657]" box="[151,546,1632,1657]" pageId="28" pageNumber="29">
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<treatmentCitationGroup id="AB0A11DE32401028FF3EF9B5DA6B3B51" box="[151,546,1632,1657]" pageId="28" pageNumber="29">
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<taxonomicName id="4C1A4D7332401028FF3EF9B5DA563B51" authority="Thorell, 1876: 8" authorityName="Thorell" authorityPageNumber="8" authorityYear="1876" box="[151,543,1632,1657]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="columbianus">
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<emphasis id="B96EEAE232401028FF3EF9B5D93B3B50" box="[151,370,1632,1656]" italics="true" pageId="28" pageNumber="29">Phassus columbianus</emphasis>
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<treatmentCitation id="0ABB10E132401028FED1F9B5DA563B51" author="Thorell, T." box="[376,543,1632,1657]" page="8" pageId="28" pageNumber="29" year="1876">
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<bibRefCitation id="EF8B4B0132401028FED1F9B5DA563B51" author="Thorell, T." box="[376,543,1632,1657]" pageId="28" pageNumber="29" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876: 8</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
|
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.
|
||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BA536F032401028FF3EF949D91C3A39" blockId="28.[151,1437,1692,1809]" pageId="28" pageNumber="29">
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<emphasis id="B96EEAE232401028FF3EF949DA173B9D" bold="true" box="[151,606,1692,1717]" pageId="28" pageNumber="29">
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References after
|
||
<bibRefCitation id="EF8B4B0132401028FEF8F948DA103B9D" author="Fet, V. & Lowe, G." box="[337,601,1692,1717]" pageId="28" pageNumber="29" pagination="54 - 286" refId="ref48817" refString="Fet, V. & Lowe, G. (2000) Family Buthidae C. L. Koch, 1837. In: Fet, V., Sissom, W. D., Lowe, G. & Braunwalder, M. E. (Eds.), Catalog of the scorpions of the world. The New York Entomological Society: New York, pp. 54 - 286." type="book chapter" year="2000">Fet & Lowe (2000: 239)</bibRefCitation>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C1A4D7332401028FDCFF948DC463B9D" ID-CoL="57994" authority=": Lourenco 2000: 456" authorityName=": Lourenco" authorityPageNumber="456" authorityYear="2000" box="[614,1039,1692,1717]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE232401028FDCFF948DB653B9C" box="[614,812,1692,1716]" italics="true" pageId="28" pageNumber="29">Tityus columbianus</emphasis>
|
||
:
|
||
<bibRefCitation id="EF8B4B0132401028FC95F948DC463B9D" author="Lourenco, W. R." box="[828,1039,1692,1717]" pageId="28" pageNumber="29" pagination="449 - 461" refId="ref49925" refString="Lourenco, W. R. (2000) Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34 (3), 449 - 461. https: // doi. org / 10.1080 / 00222930029956 1" type="journal article" year="2000">Lourenço 2000: 456</bibRefCitation>
|
||
</taxonomicName>
|
||
, 458, figure 13;
|
||
<bibRefCitation id="EF8B4B0132401028FB17F949DD263B9D" author="Florez, E." box="[1214,1391,1692,1717]" pageId="28" pageNumber="29" refId="ref48892" refString="Florez, E. (2001 a) Sinopsis de los escorpiones de la familia Buthidae en Colombia. Master Thesis. Universidad Nacional de Colombia: Bogota, 108 pp." type="book" year="2001">Flórez 2001a: 58</bibRefCitation>
|
||
, 59;
|
||
<bibRefCitation id="EF8B4B0132401028FF6EF96FD9303BFB" author="Florez, E." box="[199,377,1722,1747]" pageId="28" pageNumber="29" pagination="25 - 30" refId="ref48924" refString="Florez, E. (2001 b) Escorpiones de la Familia Buthidae (Chelicerata: Scorpiones) de Colombia. Biota Colombiana, 2 (1), 25 - 30." type="journal article" year="2001">Flórez 2001b: 28</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8B4B0132401028FE2CF969DA2D3BFB" author="Lourenco, W. R." box="[389,612,1723,1747]" pageId="28" pageNumber="29" pagination="127 - 141" refId="ref49982" refString="Lourenco, W. R. (2002 a) Nouvelles additions a la faune de scorpions neotropicaux (Arachnida). Revue Suisse de Zoologie, 109 (1), 127 - 141. https: // doi. org / 10.5962 / bhl. part. 7958 2" type="journal article" year="2002">Lourenço 2002a: 133</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8B4B0132401028FDC6F969DB7C3BFB" author="Lourenco, W. R. & Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F." box="[623,821,1722,1747]" pageId="28" pageNumber="29" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh " o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço 2006: 56</bibRefCitation>
|
||
, 58, 60;
|
||
<bibRefCitation id="EF8B4B0132401028FC39F96FDCB53BFB" author="Botero-Trujillo, R. & Fagua, G." box="[912,1276,1722,1748]" pageId="28" pageNumber="29" pagination="129 - 134" refId="ref48464" refString="Botero-Trujillo, R. & Fagua, G. (2006) Additions to the knowledge of the geographical distribution of some Colombian Scorpions (Buthidae: Ananteris, Rhopalurus, Tityus). Revista Iberica de Aracnologia, 14, 129 - 134." type="journal article" year="2006">Botero-Trujillo & Fagua 2006: 130</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8B4B0132401028FAA1F96FD89F3BDA" author="Kovarik, F." pageId="28" pageNumber="29" pagination="1 - 7" refId="ref49333" refString="Kovarik, F. (2007) Description of Tityus mraceki sp. n. from Colombia and synonymization of T. meridanus Gonzalez-Sponga with T. nematochirus Mello-Leit " o (Scorpiones: Buthidae). Euscorpius, 54, 1 - 7. https: // doi. org / 10.18590 / euscorpius. 2007. vol 2007. iss 54. 1" type="journal article" year="2007">Kovarik 2007: 3</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8B4B0132401028FF48F90FD9F83BDA" author="Lourenco, W. R." box="[225,433,1754,1778]" pageId="28" pageNumber="29" pagination="195 - 209" refId="ref50142" refString="Lourenco, W. R. (2008) Nouvelles considerations sur le statut taxonomique de quelques especes du genre Tityus C. L. Koch, 1836. (Scorpiones, Buthidae) decrites de la Colombie. Acta Biologica Paranaense, Curitiba, 37, 195 - 209. https: // doi. org / 10.5380 / abpr. v 37 i 0.1320 6" type="journal article" year="2008">Lourenço 2008: 197</bibRefCitation>
|
||
, 199;
|
||
<bibRefCitation id="EF8B4B0132401028FE46F90CDAAE3BDA" author="Teruel, R. & Garcia, L. F." box="[495,743,1753,1778]" pageId="28" pageNumber="29" pagination="1 - 14" refId="ref51868" refString="Teruel, R. & Garcia, L. F. (2008) Rare or poorly known scorpions from Colombia. II. Redescription of Tityus columbianus (Thorell, 1876) (Scorpiones: Buthidae). Euscorpius, 64, 1 - 14. https: // doi. org / 10.18590 / euscorpius. 2008. vol 2008. iss 64. 1" type="journal article" year="2008">Teruel & García 2008: 1</bibRefCitation>
|
||
, 2–10, figures 1–6, table 1–4;
|
||
<bibRefCitation id="EF8B4B0132401028FBB7F90CDD6E3BD9" author="Teruel, R. & Kovarik, F." box="[1054,1319,1753,1778]" pageId="28" pageNumber="29" pagination="1 - 9" refId="ref51935" refString="Teruel, R. & Kovarik, F. (2011) Redescription and taxonomic position of Tityus atriventer Pocock, 1897 (Scorpiones: Buthidae). Euscorpius, 115, 1 - 9. https: // doi. org / 10.18590 / euscorpius. 2011. vol 2011. iss 115. 1" type="journal article" year="2011">Teruel & Kovarik 2011: 7</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF8B4B0132401028FA9BF90CD9183A39" author="Francke, O. F. & Teruel, R. & Santibanez-Lopez, C. E." pageId="28" pageNumber="29" pagination="220 - 232" refId="ref49045" refString="Francke, O. F., Teruel, R. & Santibanez-Lopez, C. E. (2014) A new genus and a new species of scorpion (Scorpiones: Buthidae) from southeastern Mexico. Journal of Arachnology, 42 (3), 220 - 232. https: // doi. org / 10.1636 / Ha 13 - 33. 1" type="journal article" year="2014">
|
||
Francke
|
||
<emphasis id="B96EEAE232401028FA23F90ED8AB3A38" italics="true" pageId="28" pageNumber="29">et al.</emphasis>
|
||
2014: 232
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA536F032401028FF3EF8E0DB6F3A65" blockId="28.[151,806,1844,1869]" box="[151,806,1844,1869]" pageId="28" pageNumber="29">
|
||
<treatmentCitationGroup id="AB0A11DE32401028FF3EF8E0DB6F3A65" box="[151,806,1844,1869]" pageId="28" pageNumber="29">
|
||
<taxonomicName id="4C1A4D7332401028FF3EF8E0D9503A64" authorityName="Lourenco" authorityYear="1999" box="[151,281,1844,1868]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="erikae">
|
||
<emphasis id="B96EEAE232401028FF3EF8E0D9503A64" box="[151,281,1844,1868]" italics="true" pageId="28" pageNumber="29">Tityus erikae</emphasis>
|
||
</taxonomicName>
|
||
(misidentification):
|
||
<treatmentCitation id="0ABB10E132401028FE43F8E0DAF03A65" author="Lourenco, W. R." box="[490,697,1844,1869]" page="456" pageId="28" pageNumber="29" year="2000">
|
||
<bibRefCitation id="EF8B4B0132401028FE43F8E0DAF03A65" author="Lourenco, W. R." box="[490,697,1844,1869]" pageId="28" pageNumber="29" pagination="449 - 461" refId="ref49925" refString="Lourenco, W. R. (2000) Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34 (3), 449 - 461. https: // doi. org / 10.1080 / 00222930029956 1" type="journal article" year="2000">Lourenço 2000: 456</bibRefCitation>
|
||
</treatmentCitation>
|
||
, figure 13.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300657B32401029FF3EF8A1DBE13BCB" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="8BA536F032401028FF3EF8A1DBA63A9B" blockId="28.[151,1436,1907,2043]" pageId="28" pageNumber="29">
|
||
<emphasis id="B96EEAE232401028FF3EF8A1D90D3AA6" bold="true" box="[151,324,1907,1934]" pageId="28" pageNumber="29">Type material.</emphasis>
|
||
<materialsCitation id="3B723CAD32401028FEE3F8A0DBA63A9B" ID-GBIF-Occurrence="2351005733" collectionCode="NMG" country="Colombia" location="America merid., Columbia" municipality="Bogota" pageId="28" pageNumber="29" specimenCode="NMG-Scorp-42" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Cundinamarca department" typeStatus="holotype">
|
||
<taxonomicName id="4C1A4D7332401028FEE3F8A0DA6A3AA6" authorityName=": Lourenco" authorityYear="2000" box="[330,547,1908,1934]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE232401028FEE3F8A0DA6A3AA6" box="[330,547,1908,1934]" italics="true" pageId="28" pageNumber="29">Tityus columbianus</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54A1885232401028FD98F8A1DAD73AA6" box="[561,670,1908,1934]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F30D766032401028FD0EF8A1DB083AA6" box="[679,833,1908,1934]" name="Colombia" pageId="28" pageNumber="29">COLOMBIA</collectingCountry>
|
||
:
|
||
<collectingRegion id="49DEF81232401028FCE3F8A1DC3E3AA6" box="[842,1143,1908,1934]" country="Colombia" name="Cundinamarca" pageId="28" pageNumber="29">
|
||
<emphasis id="B96EEAE232401028FCE3F8A1DC3E3AA6" box="[842,1143,1908,1934]" italics="true" pageId="28" pageNumber="29">Cundinamarca department</emphasis>
|
||
</collectingRegion>
|
||
: one
|
||
<specimenCount id="9D1CFD7932401028FB1DF8A1DCA53AA6" box="[1204,1260,1908,1934]" pageId="28" pageNumber="29" type="adult">adult</specimenCount>
|
||
<specimenCount id="9D1CFD7932401028FB5BF8A1DD083AA6" box="[1266,1345,1908,1934]" pageId="28" pageNumber="29" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32401028FAE2F8A1DDD53AA6" box="[1355,1436,1908,1934]" pageId="28" pageNumber="29">Bogotá</collectingMunicipality>
|
||
(‘’
|
||
<location id="8EC5602B32401028FF01F84DD9AB3A9A" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32401028FF01F84DD9AB3A9A" box="[168,482,1944,1970]" country="Colombia" municipality="Bogota" name="America merid., Columbia" pageId="28" pageNumber="29" stateProvince="Cundinamarca department">America merid., Columbia</location>
|
||
, Bogotá?’’) (
|
||
<specimenCode id="DBBC9E8B32401028FDDAF84DDB613A9A" box="[627,808,1944,1970]" pageId="28" pageNumber="29">NMG-Scorp-42</specimenCode>
|
||
) (
|
||
<location id="8EC5602B32401028FC97F84DDB233A9A" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32401028FC97F84DDB233A9A" box="[830,874,1944,1970]" country="Colombia" municipality="Bogota" name="Not" pageId="28" pageNumber="29" stateProvince="Cundinamarca department">Not</location>
|
||
examined).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA536F032401029FF6EF86EDBE13BCB" blockId="28.[151,1436,1907,2043]" lastBlockId="29.[151,1437,153,2015]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
<emphasis id="B96EEAE232401028FF6EF86ED9FB3AFE" bold="true" box="[199,434,1979,2006]" pageId="28" pageNumber="29">Examined material.</emphasis>
|
||
<materialsCitation id="3B723CAD32401029FE11F869DB103D9C" ID-GBIF-Occurrence="2365685692" collectedFrom="inside a dry forest" collectingDate="1996-07" collectorName="F. Escobar & W. R. Lourenco" country="Colombia" elevation="390" lastPageId="29" lastPageNumber="30" latitude="9.956667" location="Santuario de Flora y Fauna Los Colorados" longLatPrecision="21" longitude="-75.08667" municipality="Zambrano" pageId="28" pageNumber="29" specimenCode="ICN-As- 188" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Bolivar department">
|
||
<collectingCountry id="F30D766032401028FE11F869DA1B3AFE" box="[440,594,1980,2006]" name="Colombia" pageId="28" pageNumber="29" value="Colombia">COLOMBIA</collectingCountry>
|
||
:
|
||
<emphasis id="B96EEAE232401028FDF2F869DB743AFE" box="[603,829,1980,2006]" italics="true" pageId="28" pageNumber="29">
|
||
<collectingRegion id="49DEF81232401028FDF2F869DB713AFE" box="[603,824,1980,2006]" country="Colombia" name="Bolivar" pageId="28" pageNumber="29">Bolívar department</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
one adult
|
||
<specimenCount id="9D1CFD7932401028FC06F869DBA13AFE" box="[943,1000,1980,2006]" pageId="28" pageNumber="29" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32401028FC5BF869DC213AFE" box="[1010,1128,1980,2006]" pageId="28" pageNumber="29">Zambrano</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32401028FBDBF869D9083AD2" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32401028FBDBF869D9083AD2" country="Colombia" latitude="9.956667" longLatPrecision="21" longitude="-75.08667" municipality="Zambrano" name="Santuario de Flora y Fauna Los Colorados" pageId="28" pageNumber="29" stateProvince="Bolivar department">Santuario de Flora y Fauna Los Colorados</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732401028FEE7F80AD99C3AD3" box="[334,469,2015,2043]" degrees="9" direction="north" minutes="57" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="24" value="9.956667">9°57’24’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732401028FE76F80ADA253AD3" box="[479,620,2015,2043]" degrees="75" direction="west" minutes="5" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="12" value="-75.08667">75°5’12’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532401028FDD0F835DAAC3AD2" box="[633,741,2016,2042]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="28" pageNumber="29" unit="m" value="390.0">
|
||
<elevation id="0037D1C332401028FDD0F835DAAC3AD2" box="[633,741,2016,2042]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="28" pageNumber="29" unit="m" value="390.0">390 masl</elevation>
|
||
</quantity>
|
||
, collected
|
||
<collectedFrom id="3806750D32401028FCC8F835DC7C3AD2" box="[865,1077,2016,2043]" pageId="28" pageNumber="29">inside a dry forest</collectedFrom>
|
||
,
|
||
<date id="FFA4103032401028FBEBF835DCEB3AD2" box="[1090,1186,2016,2042]" pageId="28" pageNumber="29" value="1996-07">
|
||
<collectingDate id="EFE0E9D832401028FBEBF835DCEB3AD2" box="[1090,1186,2016,2042]" pageId="28" pageNumber="29" value="1996-07">vii.1996</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632401028FB06F834DD6E3AD2" box="[1199,1319,2016,2043]" pageId="28" pageNumber="29">F. Escobar</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32401029FA91F835D88B3D9B" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">ICN-As- 188</specimenCode>
|
||
) (misidentified as
|
||
<taxonomicName id="4C1A4D7332411029FE33FF4FD9B43D9B" authorityName="Lourenco" authorityYear="1999" box="[410,509,153,179]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="erikae">
|
||
<emphasis id="B96EEAE232411029FE33FF4FD9B43D9B" box="[410,509,153,179]" italics="true" pageId="29" pageNumber="30">T. erikae</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<collectorName id="26EF532632411029FD82FF4CDAAA3D9B" box="[555,739,153,179]" pageId="29" pageNumber="30">W. R. Lourenço</collectorName>
|
||
,
|
||
<date id="FFA4103032411029FD59FF4CDB183D9C" box="[752,849,153,180]" pageId="29" pageNumber="30" value="1998-12">
|
||
<collectingDate id="EFE0E9D832411029FD59FF4CDB183D9C" box="[752,849,153,180]" pageId="29" pageNumber="30" value="1998-12">xii.1998</collectingDate>
|
||
</date>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B723CAD32411029FCCFFF4CDB323DD4" ID-GBIF-Occurrence="2365685891" collectedFrom="under rock near an oak forest" collectingDate="2000-08-01" collectingDateMax="2000-08-03" collectingDateMin="2000-08-01" collectorName="A. Sabogal" country="Colombia" elevation="2500" latitude="5.75" location="Vereda Montesuarez" longLatPrecision="21" longitude="-73.433334" municipality="Arcabuco" pageId="29" pageNumber="30" specimenCode="ICN-As-364" specimenCount="12" specimenCount-adult="3" specimenCount-female="11" specimenCount-juvenile="9" specimenCount-male="1" stateProvince="Boyaca department">
|
||
<collectingRegion id="49DEF81232411029FCCFFF4CDC0A3D9B" box="[870,1091,153,179]" country="Colombia" name="Boyaca" pageId="29" pageNumber="30">
|
||
<emphasis id="B96EEAE232411029FCCFFF4CDC0A3D9B" box="[870,1091,153,179]" italics="true" pageId="29" pageNumber="30">Boyacá department</emphasis>
|
||
</collectingRegion>
|
||
: three adult
|
||
<specimenCount id="9D1CFD7932411029FB7CFF4CDD663D9B" box="[1237,1327,153,179]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
, eight juvenile
|
||
<specimenCount id="9D1CFD7932411029FF4DFF68D9723DFF" box="[228,315,189,215]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one juvenile
|
||
<specimenCount id="9D1CFD7932411029FDA1FF68DA083DFF" box="[520,577,189,215]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FDE4FF68DAF43DFF" box="[589,701,189,215]" pageId="29" pageNumber="30">Arcabuco</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FD61FF68DBFA3DFF" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FD61FF68DBFA3DFF" box="[712,947,189,215]" country="Colombia" latitude="5.75" longLatPrecision="21" longitude="-73.433334" municipality="Arcabuco" name="Vereda Montesuarez" pageId="29" pageNumber="30" stateProvince="Boyaca department">Vereda Montesuárez</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC16FF69DC7F3DFF" box="[959,1078,188,215]" degrees="5" direction="north" minutes="45" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="5.75">5°45’0’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB96FF68DC823DFF" box="[1087,1227,189,215]" degrees="73" direction="west" minutes="26" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="-73.433334">73°26’0’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FB71FF69DD193DFF" box="[1240,1360,188,215]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="30" unit="m" value="2500.0">
|
||
<elevation id="0037D1C332411029FB71FF69DD193DFF" box="[1240,1360,188,215]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="30" unit="m" value="2500.0">2500 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FAF5FF68D9EB3DD3" pageId="29" pageNumber="30">under rock near an oak forest</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FE07FF34DA0F3DD3" box="[430,582,225,251]" pageId="29" pageNumber="30" value="2000-08-01" valueMax="2000-08-03" valueMin="2000-08-01">
|
||
<collectingDate id="EFE0E9D832411029FE07FF34DA0F3DD3" box="[430,582,225,251]" pageId="29" pageNumber="30" value="2000-08-01" valueMax="2000-08-03" valueMin="2000-08-01">1–3.viii.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FDFBFF34DA993DD3" box="[594,720,225,251]" pageId="29" pageNumber="30">A. Sabogal</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FD49FF34DB3A3DD4" box="[736,883,225,252]" pageId="29" pageNumber="30">ICN-As-364</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FC21FF34DCFF3C37" ID-GBIF-Occurrence="2365685940" collectedFrom="under rock inside a crop" collectingDate="1991-04" collectorName="L. G. Baptiste" country="Colombia" elevation="2700" latitude="6.1894445" location="Vereda Chipa" longLatPrecision="21" longitude="-72.47667" municipality="Chita" pageId="29" pageNumber="30" specimenCode="ICN-As-130" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FC50FF34DC7B3DD3" box="[1017,1074,225,251]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FB96FF34DC363DD3" box="[1087,1151,225,251]" pageId="29" pageNumber="30">Chita</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FB22FF34DD613DD3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FB22FF34DD613DD3" box="[1163,1320,225,251]" country="Colombia" latitude="6.1894445" longLatPrecision="21" longitude="-72.47667" municipality="Chita" name="Vereda Chipa" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Vereda Chipa</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FA9DFF34D8E23C37" degrees="6" direction="north" minutes="11" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="22" value="6.1894445">6°11’22’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FF1DFED0D9063C37" box="[180,335,261,287]" degrees="72" direction="west" minutes="28" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="36" value="-72.47667">72°28’36’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FEF2FED0D99D3C37" box="[347,468,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="29" pageNumber="30" unit="m" value="2700.0">
|
||
<elevation id="0037D1C332411029FEF2FED0D99D3C37" box="[347,468,261,287]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="29" pageNumber="30" unit="m" value="2700.0">2700 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FE48FED0DAB23C37" box="[481,763,261,287]" pageId="29" pageNumber="30">under rock inside a crop</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FCAEFED0DB143C37" box="[775,861,261,287]" pageId="29" pageNumber="30" value="1991-04">
|
||
<collectingDate id="EFE0E9D832411029FCAEFED0DB143C37" box="[775,861,261,287]" pageId="29" pageNumber="30" value="1991-04">iv.1991</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FCC3FED0DC423C37" box="[874,1035,261,287]" pageId="29" pageNumber="30">L. G. Baptiste</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FBB5FED0DCE73C37" box="[1052,1198,261,287]" pageId="29" pageNumber="30">ICN-As-130</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FB6AFED0D9733C4F" ID-GBIF-Occurrence="2365685751" collectingDate="2000-01-07" collectorName="R. Alvarado" country="Colombia" elevation="2500" latitude="6.1894445" location="Vereda El Alcaparral" longLatPrecision="21" longitude="-72.47667" municipality="Firavitoba" pageId="29" pageNumber="30" specimenCode="ICN-As-288" specimenCount="4" specimenCount-adult="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
three adult
|
||
<specimenCount id="9D1CFD7932411029FAECFED0DDD53C37" box="[1349,1436,261,287]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one adult
|
||
<specimenCount id="9D1CFD7932411029FE9EFEFCD9393C6B" box="[311,368,297,323]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FED5FEFCD9BB3C6B" box="[380,498,297,323]" pageId="29" pageNumber="30">Firavitoba</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FE54FEFCDAA23C6B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE54FEFCDAA23C6B" box="[509,747,297,323]" country="Colombia" latitude="6.1894445" longLatPrecision="21" longitude="-72.47667" municipality="Firavitoba" name="Vereda El Alcaparral" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Vereda El Alcaparral</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FD5FFEFCDB333C6B" box="[758,890,297,323]" degrees="6" direction="north" minutes="11" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="22" value="6.1894445">6°11’22’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FC2BFEFCDC523C6B" box="[898,1051,297,323]" degrees="72" direction="west" minutes="28" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="36" value="-72.47667">72°28’36’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FB8EFEFDDCD73C6B" box="[1063,1182,296,323]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="30" unit="m" value="2500.0">
|
||
<elevation id="0037D1C332411029FB8EFEFDDCD73C6B" box="[1063,1182,296,323]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="29" pageNumber="30" unit="m" value="2500.0">2500 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FB03FEFCDD413C6B" box="[1194,1288,297,323]" pageId="29" pageNumber="30" value="2000-01-07">
|
||
<collectingDate id="EFE0E9D832411029FB03FEFCDD413C6B" box="[1194,1288,297,323]" pageId="29" pageNumber="30" value="2000-01-07">7.i.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FABDFEFCDDD53C6B" box="[1300,1436,297,323]" pageId="29" pageNumber="30">R. Alvarado</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FF36FE98D97B3C4F" box="[159,306,333,359]" pageId="29" pageNumber="30">ICN-As-288</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FEEDFE98DA893CA3" ID-GBIF-Occurrence="2365685647" collectingDate="2000-07-10" collectorName="E. Florez" country="Colombia" elevation="2590" latitude="5.613333" location="Iza" longLatPrecision="15" longitude="-72.981094" pageId="29" pageNumber="30" specimenCode="ICN-As-388" specimenCount="10" specimenCount-adult="2" specimenCount-female="5" specimenCount-juvenile="8" specimenCount-male="5" stateProvince="Boyaca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FE1AFE98DA443C4F" box="[435,525,333,359]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
, five juvenile
|
||
<specimenCount id="9D1CFD7932411029FD03FE98DAA23C4F" box="[682,747,333,359]" pageId="29" pageNumber="30" type="male">males</specimenCount>
|
||
and three juvenile
|
||
<specimenCount id="9D1CFD7932411029FC16FE98DC503C4F" box="[959,1049,333,359]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FB8DFE98DC003C4F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FB8DFE98DC003C4F" box="[1060,1097,333,359]" country="Colombia" latitude="5.613333" longLatPrecision="15" longitude="-72.981094" name="Iza" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Iza</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FBFAFE99DC9E3C4F" box="[1107,1239,332,359]" degrees="5" direction="north" minutes="36" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="48" value="5.613333">5°36’48’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB74FE99DDD13C4F" box="[1245,1432,332,359]" degrees="72" direction="west" minutes="58" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="51.93" value="-72.981094">72°58’51.93’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EFEA5D9593CA3" box="[151,272,368,395]" metricMagnitude="3" metricUnit="m" metricValue="2.59" pageId="29" pageNumber="30" unit="m" value="2590.0">
|
||
<elevation id="0037D1C332411029FF3EFEA5D9593CA3" box="[151,272,368,395]" metricMagnitude="3" metricUnit="m" metricValue="2.59" pageId="29" pageNumber="30" unit="m" value="2590.0">2590 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FEB4FEA4D9D63CA3" box="[285,415,369,395]" pageId="29" pageNumber="30" value="2000-07-10">
|
||
<collectingDate id="EFE0E9D832411029FEB4FEA4D9D63CA3" box="[285,415,369,395]" pageId="29" pageNumber="30" value="2000-07-10">10.vii.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE05FEA4DA5C3CA3" box="[428,533,369,395]" pageId="29" pageNumber="30">E. Flórez</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FD8CFEA4DAF13CA3" box="[549,696,369,395]" pageId="29" pageNumber="30">ICN-As-388</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD64FEA4DC6F3C98" ID-GBIF-Occurrence="2365686108" collectedFrom="inside a rotten log" collectingDate="2010-05-21" collectorName="H. Gasca" country="Colombia" elevation="2315" latitude="6.2328334" location="Vereda San Ignacio" longLatPrecision="1" longitude="-72.70006" municipality="Susacon" pageId="29" pageNumber="30" specimenCode="ICN-As-846" specimenCount="2" specimenCount-adult="2" specimenCount-female="2" stateProvince="Boyaca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FCE9FEA4DBD33CA3" box="[832,922,369,395]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FC0EFEA4DC4E3CA3" box="[935,1031,369,395]" pageId="29" pageNumber="30">Susacón</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FBBDFEA4DCBD3CA3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FBBDFEA4DCBD3CA3" box="[1044,1268,369,395]" country="Colombia" latitude="6.2328334" longLatPrecision="1" longitude="-72.70006" municipality="Susacon" name="Vereda San Ignacio" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Vereda San Ignacio</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FAA8FEA5DDD53CA3" box="[1281,1436,368,395]" degrees="6" direction="north" minutes="13" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="58.2" value="6.2328334">6°13’58.2’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FF3EFE40D90A3C87" box="[151,323,405,431]" degrees="72" direction="west" minutes="42" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="0.23" value="-72.70006">72°42’0.23’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FEE4FE41D98A3C87" box="[333,451,404,431]" metricMagnitude="3" metricUnit="m" metricValue="2.315" pageId="29" pageNumber="30" unit="m" value="2315.0">
|
||
<elevation id="0037D1C332411029FEE4FE41D98A3C87" box="[333,451,404,431]" metricMagnitude="3" metricUnit="m" metricValue="2.315" pageId="29" pageNumber="30" unit="m" value="2315.0">2315 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FE65FE40DADD3C87" box="[460,660,405,431]" pageId="29" pageNumber="30">inside a rotten log</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FD37FE40DB463C87" box="[670,783,405,431]" pageId="29" pageNumber="30" value="2010-05-21">
|
||
<collectingDate id="EFE0E9D832411029FD37FE40DB463C87" box="[670,783,405,431]" pageId="29" pageNumber="30" value="2010-05-21">21.v.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FCB0FE40DB373C87" box="[793,894,405,431]" pageId="29" pageNumber="30">H. Gasca</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FC25FE40DC573C98" box="[908,1054,405,432]" pageId="29" pageNumber="30">ICN-As-846</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FB99FE40DD6F3CFB" ID-GBIF-Occurrence="2365686013" collectingDate="2002-08-25" collectorName="E. Gonzalez" country="Colombia" elevation="2128" latitude="5.633333" location="Villa de Leyva" longLatPrecision="15" longitude="-73.53333" municipality="Villa de Leyva" pageId="29" pageNumber="30" specimenCode="ICN-As-739" specimenCount="4" specimenCount-adult="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
three adult
|
||
<specimenCount id="9D1CFD7932411029FB02FE40DD4B3C87" box="[1195,1282,405,431]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one adult
|
||
<specimenCount id="9D1CFD7932411029FF3EFE6CD8993CFB" box="[151,208,441,467]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FF75FE6CD9CC3CFB" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FF75FE6CD9CC3CFB" box="[220,389,441,467]" country="Colombia" latitude="5.633333" longLatPrecision="15" longitude="-73.53333" name="Villa de Leyva" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Villa de Leyva</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FE38FE6DDA413CFB" box="[401,520,440,467]" degrees="5" direction="north" minutes="38" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="5.633333">5°38’0’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FDB9FE6DDA843CFB" box="[528,717,440,467]" degrees="73" direction="west" minutes="31" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.99" value="-73.53333">73°31’59.99’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD70FE6CDB193CFB" box="[729,848,441,467]" metricMagnitude="3" metricUnit="m" metricValue="2.128" pageId="29" pageNumber="30" unit="m" value="2128.0">
|
||
<elevation id="0037D1C332411029FD70FE6CDB193CFB" box="[729,848,441,467]" metricMagnitude="3" metricUnit="m" metricValue="2.128" pageId="29" pageNumber="30" unit="m" value="2128.0">2128 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FCF5FE6DDBAF3CFB" box="[860,998,440,467]" pageId="29" pageNumber="30" value="2002-08-25">
|
||
<collectingDate id="EFE0E9D832411029FCF5FE6DDBAF3CFB" box="[860,998,440,467]" pageId="29" pageNumber="30" value="2002-08-25">25.viii.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FC5BFE6CDC353CFB" box="[1010,1148,441,467]" pageId="29" pageNumber="30">E. González</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FB25FE6CDD563CFB" box="[1164,1311,441,467]" pageId="29" pageNumber="30">ICN-As-739</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FA9BFE6CD8BF3F17" ID-GBIF-Occurrence="2365685927" collectedFrom="under rocks and logs inside a paddock" collectingDate="1991-06-09" collectionCode="ICN" collectorName="F. Quevedo" country="Colombia" elevation="2600" latitude="5.6833334" location="Santuario de Flora y Fauna de Iguaque" longLatPrecision="21" longitude="-73.48333" municipality="Villa de Leyva" pageId="29" pageNumber="30" specimenCode="ICN- As-524" specimenCount="5" specimenCount-adult="4" specimenCount-female="3" specimenCount-juvenile="1" specimenCount-male="2" stateProvince="Boyaca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FF3EFE08D8B83CDF" box="[151,241,477,503]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
, two adult
|
||
<specimenCount id="9D1CFD7932411029FED9FE08D9F83CDF" box="[368,433,477,503]" pageId="29" pageNumber="30" type="male">males</specimenCount>
|
||
and one juvenile
|
||
<specimenCount id="9D1CFD7932411029FDD5FE08DA823CDF" box="[636,715,477,503]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FD7EFE08DBC83CDF" box="[727,897,477,503]" pageId="29" pageNumber="30">Villa de Leyva</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FC24FE08DD013CDF" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC24FE08DD013CDF" box="[909,1352,477,503]" country="Colombia" latitude="5.6833334" longLatPrecision="21" longitude="-73.48333" municipality="Villa de Leyva" name="Santuario de Flora y Fauna de Iguaque" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Santuario de Flora y Fauna de Iguaque</location>
|
||
,
|
||
<location id="8EC5602B32411029FAFCFE08D9433F33" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FAFCFE08D9433F33" country="Colombia" latitude="5.6833334" longLatPrecision="21" longitude="-73.48333" municipality="Villa de Leyva" name="Sector de Chaina" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Sector de Chaina</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FEBCFDD5D9C33F33" box="[277,394,512,539]" degrees="5" direction="north" minutes="41" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="5.6833334">5°41’0’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FE38FDD4DA523F33" box="[401,539,513,539]" degrees="73" direction="west" minutes="29" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="-73.48333">73°29’0’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD8CFDD4DAD53F33" box="[549,668,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FD8CFDD4DAD53F33" box="[549,668,513,539]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FD0FFDD4DC183F33" box="[678,1105,513,539]" pageId="29" pageNumber="30">under rocks and logs inside a paddock</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FBF5FDD4DC813F33" box="[1116,1224,513,539]" pageId="29" pageNumber="30" value="1991-06-09">
|
||
<collectingDate id="EFE0E9D832411029FBF5FDD4DC813F33" box="[1116,1224,513,539]" pageId="29" pageNumber="30" value="1991-06-09">9.vi.1991</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FB7AFDD4DD1A3F33" box="[1235,1363,513,539]" pageId="29" pageNumber="30">F. Quevedo</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FAC8FDD4D8A63F17" pageId="29" pageNumber="30">ICN- As-524</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FEAAFDF0DA6E3F4B" ID-GBIF-Occurrence="2365686372" collectingDate="1985-10" collectorName="R. Gonzalez" country="Colombia" elevation="2200" latitude="5.638611" location="El Roble" longLatPrecision="21" longitude="-73.528336" municipality="Villa de Leyva" pageId="29" pageNumber="30" specimenCode="ICN-As-352" specimenCount="32" specimenCount-adult="32" specimenCount-female="24" specimenCount-male="8" stateProvince="Boyaca Department">
|
||
24 adult
|
||
<specimenCount id="9D1CFD7932411029FECEFDF0D9F73F17" box="[359,446,549,575]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and eight adult
|
||
<specimenCount id="9D1CFD7932411029FDDFFDF0DAF33F17" box="[630,698,549,575]" pageId="29" pageNumber="30" type="male">males</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FD6FFDF0DB263F17" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FD6FFDF0DB263F17" box="[710,879,549,575]" country="Colombia" latitude="5.638611" longLatPrecision="21" longitude="-73.528336" name="Villa de Leyva" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Villa de Leyva</location>
|
||
,
|
||
<location id="8EC5602B32411029FCD2FDF0DBAB3F17" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FCD2FDF0DBAB3F17" box="[891,994,549,575]" country="Colombia" latitude="5.638611" longLatPrecision="21" longitude="-73.528336" name="El Roble" pageId="29" pageNumber="30" stateProvince="Boyaca Department">El Roble</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC47FDF1DC3A3F17" box="[1006,1139,548,575]" degrees="5" direction="north" minutes="38" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="19" value="5.638611">5°38’19’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FBD2FDF0DD5C3F17" box="[1147,1301,549,575]" degrees="73" direction="west" minutes="31" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="42" value="-73.528336">73°31’42’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FA89FDF0DDD13F17" box="[1312,1432,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">
|
||
<elevation id="0037D1C332411029FA89FDF0DDD13F17" box="[1312,1432,549,575]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FF3EFD9DD8AF3F4B" box="[151,230,584,611]" pageId="29" pageNumber="30" value="1985-10">
|
||
<collectingDate id="EFE0E9D832411029FF3EFD9DD8AF3F4B" box="[151,230,584,611]" pageId="29" pageNumber="30" value="1985-10">x.1985</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FF5AFD9CD9373F4B" box="[243,382,585,611]" pageId="29" pageNumber="30">R. González</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE24FD9DDA563F4B" box="[397,543,584,611]" pageId="29" pageNumber="30">ICN-As-352</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD9AFD9CDCAB3FAF" ID-GBIF-Occurrence="2365686150" collectedFrom="inside a pastureland" collectingDate="2000-04" collectorName="E. Hernandez" country="Colombia" elevation="2200" latitude="5.638611" location="near river Cebada and river Cene" longLatPrecision="21" longitude="-73.528336" pageId="29" pageNumber="30" specimenCode="ICN-As-314" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Boyaca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FD0BFD9CDAB83F4B" box="[674,753,585,611]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FD55FD9CDBEC3F4B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FD55FD9CDBEC3F4B" box="[764,933,585,611]" country="Colombia" latitude="5.638611" longLatPrecision="21" longitude="-73.528336" name="Villa de Leyva" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Villa de Leyva</location>
|
||
, near river
|
||
<location id="8EC5602B32411029FB8DFD9CDD6E3F4B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FB8DFD9CDD6E3F4B" box="[1060,1319,585,611]" country="Colombia" latitude="5.638611" longLatPrecision="21" longitude="-73.528336" name="Cebada and river Cene" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Cebada and river Cene</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FA9AFD9DD8E23FAF" degrees="5" direction="north" minutes="38" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="19" value="5.638611">5°38’19’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FF1DFDB8D9043FAF" box="[180,333,621,647]" degrees="73" direction="west" minutes="31" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="42" value="-73.528336">73°31’42’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FEF0FDB8D99B3FAF" box="[345,466,621,647]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">
|
||
<elevation id="0037D1C332411029FEF0FDB8D99B3FAF" box="[345,466,621,647]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200 masl</elevation>
|
||
</quantity>
|
||
, collected
|
||
<collectedFrom id="3806750D32411029FDE2FDB8DB663FAF" box="[587,815,621,647]" pageId="29" pageNumber="30">inside a pastureland</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FC92FDB8DBD83FAF" box="[827,913,621,647]" pageId="29" pageNumber="30" value="2000-04">
|
||
<collectingDate id="EFE0E9D832411029FC92FDB8DBD83FAF" box="[827,913,621,647]" pageId="29" pageNumber="30" value="2000-04">iv.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FC34FDB8DC7E3FAF" box="[925,1079,621,647]" pageId="29" pageNumber="30">E. Hernández</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FBEEFDB8DC903FAF" box="[1095,1241,621,647]" pageId="29" pageNumber="30">ICN-As-314</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FB47FDB8DD233F83" ID-GBIF-Occurrence="2365686424" collectedFrom="inside a house" collectingDate="2000-08-01" collectorName="H. Schultz" country="Colombia" elevation="2200" latitude="5.633861" location="Villa de Leyva" longLatPrecision="1" longitude="-73.52352" municipality="Villa de Leyva" pageId="29" pageNumber="30" specimenCode="ICN-As-419" specimenCount="1" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FAF6FDB8DDD13FAF" box="[1375,1432,621,647]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FF3EFD44D9743F83" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FF3EFD44D9743F83" box="[151,317,657,683]" country="Colombia" latitude="5.633861" longLatPrecision="1" longitude="-73.52352" name="Villa de Leyva" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Villa de Leyva</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FEEEFD45D9983F83" box="[327,465,656,683]" degrees="5" direction="north" minutes="38" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="1.9" value="5.633861">5°38’1.9’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FE7EFD44DADB3F83" box="[471,658,657,683]" degrees="73" direction="west" minutes="31" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="24.67" value="-73.52352">73°31’24.67’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD35FD44DB5B3F83" box="[668,786,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">
|
||
<elevation id="0037D1C332411029FD35FD44DB5B3F83" box="[668,786,657,683]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FCB5FD44DBF23F83" box="[796,955,657,683]" pageId="29" pageNumber="30">inside a house</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FC6FFD44DC0B3F83" box="[966,1090,657,683]" pageId="29" pageNumber="30" value="2000-08-01">
|
||
<collectingDate id="EFE0E9D832411029FC6FFD44DC0B3F83" box="[966,1090,657,683]" pageId="29" pageNumber="30" value="2000-08-01">1.viii.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FBE5FD44DC883F83" box="[1100,1217,657,683]" pageId="29" pageNumber="30">H. Schultz</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FB66FD44DD2B3F83" box="[1231,1378,657,683]" pageId="29" pageNumber="30">ICN-As-419</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FADDFD44D9FB3FDB" ID-GBIF-Occurrence="2365685741" collectedFrom="inside leaf litter" collectingDate="2004-02-19" collectingMethod="manual capture" collectorName="M. Porras" country="Colombia" elevation="2200" latitude="5.633333" location="Villa de Leyva" longLatPrecision="15" longitude="-73.53333" pageId="29" pageNumber="30" specimenCode="ICN-As-570" specimenCount="1" specimenCount-juvenile="1" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
one juvenile
|
||
<specimenCount id="9D1CFD7932411029FF51FD60D9783FE7" box="[248,305,693,719]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FE95FD60D9AD3FE7" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE95FD60D9AD3FE7" box="[316,484,693,719]" country="Colombia" latitude="5.633333" longLatPrecision="15" longitude="-73.53333" name="Villa de Leyva" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Villa de Leyva</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FE46FD61DA2C3FE7" box="[495,613,692,719]" degrees="5" direction="north" minutes="38" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="5.633333">5°38’0’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FDC5FD61DB613FE7" box="[620,808,692,719]" degrees="73" direction="west" minutes="31" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.99" value="-73.53333">73°31’59.99’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FC9AFD60DBE33FE7" box="[819,938,693,719]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">
|
||
<elevation id="0037D1C332411029FC9AFD60DBE33FE7" box="[819,938,693,719]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="m" value="2200.0">2200 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FC1DFD60DC293FE7" box="[948,1120,693,719]" pageId="29" pageNumber="30">manual capture</collectingMethod>
|
||
<collectedFrom id="3806750D32411029FBCEFD60DD503FE7" box="[1127,1305,693,719]" pageId="29" pageNumber="30">inside leaf litter</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FA8DFD60DDD13FE7" box="[1316,1432,693,719]" pageId="29" pageNumber="30" value="2004-02-19">
|
||
<collectingDate id="EFE0E9D832411029FA8DFD60DDD13FE7" box="[1316,1432,693,719]" pageId="29" pageNumber="30" value="2004-02-19">19.ii.2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FF3EFD0CD9413FDB" box="[151,264,729,755]" pageId="29" pageNumber="30">M. Porras</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FEB1FD0DD9E23FDB" box="[280,427,728,755]" pageId="29" pageNumber="30">ICN-As-570</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FE16FD0CDA923E3F" ID-GBIF-Occurrence="2365686429" collectingDate="1979-05-24" collectorName="C. Hernandez" country="Colombia" elevation="2580" latitude="5.616892" location="Monte Batallon Sucre" longLatPrecision="1" longitude="-73.81652" municipality="Chiquinquira" pageId="29" pageNumber="30" specimenCode="ICN-As-129" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Boyaca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FD99FD0CDA363FDB" box="[560,639,729,755]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FD25FD0CDB6A3FDB" box="[652,803,729,755]" pageId="29" pageNumber="30">Chiquinquirá</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FC86FD0CDC653FDB" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC86FD0CDC653FDB" box="[815,1068,729,755]" country="Colombia" latitude="5.616892" longLatPrecision="1" longitude="-73.81652" municipality="Chiquinquira" name="Monte Batallon Sucre" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Monte Batallón Sucre</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FB91FD0DDC9A3FDB" box="[1080,1235,728,755]" degrees="5" direction="north" minutes="37" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="0.81" value="5.616892">5°37’0.81’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB72FD0DDDD13FDB" box="[1243,1432,728,755]" degrees="73" direction="west" minutes="48" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.47" value="-73.81652">73°48’59.47’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EFD29D9463E3F" box="[151,271,764,791]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="29" pageNumber="30" unit="m" value="2580.0">
|
||
<elevation id="0037D1C332411029FF3EFD29D9463E3F" box="[151,271,764,791]" metricMagnitude="3" metricUnit="m" metricValue="2.58" pageId="29" pageNumber="30" unit="m" value="2580.0">2580 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FEB3FD28D9C23E3F" box="[282,395,765,791]" pageId="29" pageNumber="30" value="1979-05-24">
|
||
<collectingDate id="EFE0E9D832411029FEB3FD28D9C23E3F" box="[282,395,765,791]" pageId="29" pageNumber="30" value="1979-05-24">24.v.1979</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE3EFD28DA783E3F" box="[407,561,765,791]" pageId="29" pageNumber="30">C. Hernández</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FDE9FD28DA9B3E3F" box="[576,722,765,791]" pageId="29" pageNumber="30">ICN-As-129</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD4FFD28DAC43E13" ID-GBIF-Occurrence="2365685802" collectingDate="2010-08-20" collectorName="J. Iglesias" country="Colombia" elevation="2100" latitude="5.566703" location="Tinjaca" longLatPrecision="1" longitude="-73.64997" pageId="29" pageNumber="30" specimenCode="ICN-As-851" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Boyaca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FCFFFD28DBC63E3F" box="[854,911,765,791]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FC33FD28DBB93E3F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC33FD28DBB93E3F" box="[922,1008,765,791]" country="Colombia" latitude="5.566703" longLatPrecision="1" longitude="-73.64997" name="Tinjaca" pageId="29" pageNumber="30" stateProvince="Boyaca Department">Tinjacá</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC55FD29DCDC3E3F" box="[1020,1173,764,791]" degrees="5" direction="north" minutes="34" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="0.13" value="5.566703">5°34’0.13’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB34FD29DD103E3F" box="[1181,1369,764,791]" degrees="73" direction="west" minutes="38" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.89" value="-73.64997">73°38’59.89’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FACDFD28D8863E13" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="m" value="2100.0">
|
||
<elevation id="0037D1C332411029FACDFD28D8863E13" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="29" pageNumber="30" unit="m" value="2100.0">2100 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FF72FCF4D92C3E13" box="[219,357,801,827]" pageId="29" pageNumber="30" value="2010-08-20">
|
||
<collectingDate id="EFE0E9D832411029FF72FCF4D92C3E13" box="[219,357,801,827]" pageId="29" pageNumber="30" value="2010-08-20">20.viii.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FEDBFCF4D9AB3E13" box="[370,482,801,827]" pageId="29" pageNumber="30">J. Iglesias</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE5BFCF5DACD3E13" box="[498,644,800,827]" pageId="29" pageNumber="30">ICN-As-851</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B723CAD32411029FD31FCF4DD6C3E77" ID-GBIF-Occurrence="2365686312" collectingDate="1961-05-07" collectorName="J. M. Idroba" country="Colombia" elevation="2600" latitude="4.7536473" location="Suba" longLatPrecision="1" longitude="-74.088715" municipality="Bogota" pageId="29" pageNumber="30" specimenCode="ICN-As-088" specimenCount="4" specimenCount-adult="3" specimenCount-female="4" specimenCount-juvenile="1" stateProvince="Cundinamarca department">
|
||
<emphasis id="B96EEAE232411029FD31FCF4DB993E13" box="[664,976,801,827]" italics="true" pageId="29" pageNumber="30">
|
||
<collectingRegion id="49DEF81232411029FD31FCF4DB823E13" box="[664,971,801,827]" country="Colombia" name="Cundinamarca" pageId="29" pageNumber="30">Cundinamarca department</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
three adult
|
||
<specimenCount id="9D1CFD7932411029FBF3FCF4DCF83E13" box="[1114,1201,801,827]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one juvenile
|
||
<specimenCount id="9D1CFD7932411029FAD4FCF4D8993E77" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FF77FC90D97B3E77" box="[222,306,837,863]" pageId="29" pageNumber="30">Bogotá</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FE96FC90D9333E77" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE96FC90D9333E77" box="[319,378,837,863]" country="Colombia" latitude="4.7536473" longLatPrecision="1" longitude="-74.088715" municipality="Bogota" name="Suba" pageId="29" pageNumber="30" stateProvince="Cundinamarca department">Suba</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FE2EFC91DA793E77" box="[391,560,836,863]" degrees="4" direction="north" minutes="45" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="13.13" value="4.7536473">4°45’13.13’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FD93FC91DAA03E77" box="[570,745,836,863]" degrees="74" direction="west" minutes="5" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="19.36" value="-74.088715">74°5’19.36’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD5FFC90DB263E77" box="[758,879,837,863]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FD5FFC90DB263E77" box="[758,879,837,863]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FCD5FC90DB963E77" box="[892,991,837,863]" pageId="29" pageNumber="30" value="1961-05-07">
|
||
<collectingDate id="EFE0E9D832411029FCD5FC90DB963E77" box="[892,991,837,863]" pageId="29" pageNumber="30" value="1961-05-07">7.v.1961</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FC44FC90DC333E77" box="[1005,1146,837,863]" pageId="29" pageNumber="30">J. M. Idroba</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FB22FC90DD543E77" box="[1163,1309,837,863]" pageId="29" pageNumber="30">ICN-As-088</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FA9AFC90DA4A3E8F" ID-GBIF-Occurrence="2365685812" collectedFrom="under a rock" collectingDate="2000-09" collectorName="D. Tobar" country="Colombia" elevation="2850" latitude="4.6183057" location="Parque de San Rafael" longLatPrecision="1" longitude="-74.05566" municipality="Bogota" pageId="29" pageNumber="30" specimenCode="ICN-As-432" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Cundinamarca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FF3EFCBCD8AF3EAB" box="[151,230,873,899]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FF5AFCBCD90E3EAB" box="[243,327,873,899]" pageId="29" pageNumber="30">Bogotá</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FEFDFCBCDA053EAB" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FEFDFCBCDA053EAB" box="[340,588,873,899]" country="Colombia" latitude="4.6183057" longLatPrecision="1" longitude="-74.05566" municipality="Bogota" name="Parque de San Rafael" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Parque de San Rafael</location>
|
||
,
|
||
<location id="8EC5602B32411029FDF0FCBCDB6B3EAB" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FDF0FCBCDB6B3EAB" box="[601,802,873,899]" country="Colombia" latitude="4.6183057" longLatPrecision="1" longitude="-74.05566" municipality="Bogota" name="Cerros Orientales" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Cerros Orientales</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC86FCBDDBF53EAB" box="[815,956,872,899]" degrees="4" direction="north" minutes="37" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="5.9" value="4.6183057">4°37’5.9’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FC6CFCBCDC3D3EAB" box="[965,1140,873,899]" degrees="74" direction="west" minutes="3" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="20.36" value="-74.05566">74°3’20.36’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FB28FCBDDCB03EAB" box="[1153,1273,872,899]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="29" pageNumber="30" unit="m" value="2850.0">
|
||
<elevation id="0037D1C332411029FB28FCBDDCB03EAB" box="[1153,1273,872,899]" metricMagnitude="3" metricUnit="m" metricValue="2.85" pageId="29" pageNumber="30" unit="m" value="2850.0">2850 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FAAFFCBCDDD13EAB" box="[1286,1432,873,899]" pageId="29" pageNumber="30">under a rock</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FF3EFC58D8A63E8F" box="[151,239,909,935]" pageId="29" pageNumber="30" value="2000-09">
|
||
<collectingDate id="EFE0E9D832411029FF3EFC58D8A63E8F" box="[151,239,909,935]" pageId="29" pageNumber="30" value="2000-09">ix.2000</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FF50FC58D9123E8F" box="[249,347,909,935]" pageId="29" pageNumber="30">D. Tobar</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FEC1FC58D9B23E8F" box="[360,507,909,935]" pageId="29" pageNumber="30">ICN-As-432</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FDA4FC58DB6F3EE3" ID-GBIF-Occurrence="2365685822" collectedFrom="under rocks" collectingDate="2001-04-27" collectorName="M. Ramirez" country="Colombia" elevation="2600" latitude="4.4730697" location="Usme" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" pageId="29" pageNumber="30" specimenCode="ICN-As-409" specimenCount="2" specimenCount-female="2" stateProvince="Cundinamarca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FDD2FC58DA9C3E8F" box="[635,725,909,935]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FD76FC58DB7A3E8F" box="[735,819,909,935]" pageId="29" pageNumber="30">Bogotá</collectingMunicipality>
|
||
, rural zone
|
||
<location id="8EC5602B32411029FC1AFC58DBBE3E8F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC1AFC58DBBE3E8F" box="[947,1015,909,935]" country="Colombia" latitude="4.4730697" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" name="Usme" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Usme</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FBA8FC59DCE13E8F" box="[1025,1192,908,935]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="23.05" value="4.4730697">4°28’23.05’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB07FC59DD133E8F" box="[1198,1370,908,935]" degrees="74" direction="west" minutes="6" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="57.93" value="-74.11609">74°6’57.93’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FACDFC58D8863EE3" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FACDFC58D8863EE3" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FF72FC64D92B3EE3" box="[219,354,945,971]" pageId="29" pageNumber="30">under rocks</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FEC6FC64D9AE3EE3" box="[367,487,945,971]" pageId="29" pageNumber="30" value="2001-04-27">
|
||
<collectingDate id="EFE0E9D832411029FEC6FC64D9AE3EE3" box="[367,487,945,971]" pageId="29" pageNumber="30" value="2001-04-27">27.iv.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE5DFC64DA323EE3" box="[500,635,945,971]" pageId="29" pageNumber="30">M. Ramírez</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FD22FC64DB573EE3" box="[651,798,945,971]" pageId="29" pageNumber="30">ICN-As-409</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FC9BFC64D884393B" ID-GBIF-Occurrence="2365686411" collectingDate="2003-05-04" collectingMethod="night manual capture" collectorName="T. Vivas" country="Colombia" elevation="2600" latitude="4.4730697" location="Cerro Juan Rey" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" pageId="29" pageNumber="30" specimenCode="ICN-As- 525" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FC0AFC64DBBB3EE3" box="[931,1010,945,971]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FC57FC64DC1B3EE3" box="[1022,1106,945,971]" pageId="29" pageNumber="30">Bogotá</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FBF7FC64DD593EE3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FBF7FC64DD593EE3" box="[1118,1296,945,971]" country="Colombia" latitude="4.4730697" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" name="Cerro Juan Rey" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Cerro Juan Rey</location>
|
||
, near
|
||
<location id="8EC5602B32411029FAFCFC64D9423EC7" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FAFCFC64D9423EC7" country="Colombia" latitude="4.4730697" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" name="Barrio La Fiscala" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Barrio La Fiscala</location>
|
||
in
|
||
<location id="8EC5602B32411029FE86FC00D93A3EC7" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE86FC00D93A3EC7" box="[303,371,981,1007]" country="Colombia" latitude="4.4730697" longLatPrecision="1" longitude="-74.11609" municipality="Bogota" name="Usme" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Usme</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FED7FC01DA6C3EC7" box="[382,549,980,1007]" degrees="4" direction="north" minutes="28" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="23.05" value="4.4730697">4°28’23.05’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FD85FC01DA933EC7" box="[556,730,980,1007]" degrees="74" direction="west" minutes="6" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="57.93" value="-74.11609">74°6’57.93’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD4DFC00DB123EC7" box="[740,859,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FD4DFC00DB123EC7" box="[740,859,981,1007]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FCCFFC00DC1C3EC7" box="[870,1109,981,1007]" pageId="29" pageNumber="30">night manual capture</collectingMethod>
|
||
,
|
||
<date id="FFA4103032411029FBC9FC00DC8A3EC7" box="[1120,1219,981,1007]" pageId="29" pageNumber="30" value="2003-05-04">
|
||
<collectingDate id="EFE0E9D832411029FBC9FC00DC8A3EC7" box="[1120,1219,981,1007]" pageId="29" pageNumber="30" value="2003-05-04">4.v.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FB67FC00DD633EC7" box="[1230,1322,981,1007]" pageId="29" pageNumber="30">T. Vivas</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FA91FC00D88C393B" pageId="29" pageNumber="30">ICN-As- 525</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FF70FC2CDC00391F" ID-GBIF-Occurrence="2365686009" collectedFrom="under rocks" collectingDate="2011-06-25" collectingMethod="diurnal manual capture" collectorName="J. Cabra" country="Colombia" elevation="2971" latitude="5.3502803" location="Vereda San Jose" longLatPrecision="1" longitude="-73.9053" municipality="Carmen de Carupa, Vereda San José" pageId="29" pageNumber="30" specimenCode="ICN-As-995" specimenCount="5" specimenCount-adult="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Cundinamarca Department">
|
||
four adult
|
||
<specimenCount id="9D1CFD7932411029FEE4FC2CD9ED393B" box="[333,420,1017,1043]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one adult
|
||
<specimenCount id="9D1CFD7932411029FDE0FC2CDACB393B" box="[585,642,1017,1043]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FD24FC2CDB2B393B" box="[653,866,1017,1043]" pageId="29" pageNumber="30">Carmen de Carupa</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FCC5FC2CDC6D393B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FCC5FC2CDC6D393B" box="[876,1060,1017,1043]" country="Colombia" latitude="5.3502803" longLatPrecision="1" longitude="-73.9053" municipality="Carmen de Carupa" name="Vereda San Jose" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Vereda San José</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FB86FC2DDC9F393B" box="[1071,1238,1016,1043]" degrees="5" direction="north" minutes="21" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="01.01" value="5.3502803">5°21’01.01’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB74FC2DDDD1393B" box="[1245,1432,1016,1043]" degrees="73" direction="west" minutes="54" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="19.06" value="-73.9053">73°54’19.06’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EFBC8D946391F" box="[151,271,1053,1079]" metricMagnitude="3" metricUnit="m" metricValue="2.971" pageId="29" pageNumber="30" unit="m" value="2971.0">
|
||
<elevation id="0037D1C332411029FF3EFBC8D946391F" box="[151,271,1053,1079]" metricMagnitude="3" metricUnit="m" metricValue="2.971" pageId="29" pageNumber="30" unit="m" value="2971.0">2971 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FEB3FBC8DA69391F" box="[282,544,1053,1079]" pageId="29" pageNumber="30">diurnal manual capture</collectingMethod>
|
||
,
|
||
<collectedFrom id="3806750D32411029FD85FBC8DAFB391F" box="[556,690,1053,1079]" pageId="29" pageNumber="30">under rocks</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FD17FBC9DB7E391F" box="[702,823,1052,1079]" pageId="29" pageNumber="30" value="2011-06-25">
|
||
<collectingDate id="EFE0E9D832411029FD17FBC9DB7E391F" box="[702,823,1052,1079]" pageId="29" pageNumber="30" value="2011-06-25">25.vi.2011</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FCEAFBC8DBD6391F" box="[835,927,1053,1079]" pageId="29" pageNumber="30">J. Cabra</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FC07FBC9DC08391F" box="[942,1089,1052,1079]" pageId="29" pageNumber="30">ICN-As-995</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FBFCFBC8D9D639A8" ID-GBIF-Occurrence="2365685957" collectedFrom="under rocks" collectingDate="2010-04" collectingMethod="diurnal manual capture" collectorName="J. Cabra" country="Colombia" elevation="2971" latitude="5.3502803" location="Carmen de Carupa, Vereda San Jose" longLatPrecision="1" longitude="-73.9053" municipality="Carmen de Carupa" pageId="29" pageNumber="30" specimenCode="ICN-As-996" specimenCount="6" specimenCount-adult="6" specimenCount-female="6" stateProvince="Cundinamarca Department">
|
||
six adult
|
||
<specimenCount id="9D1CFD7932411029FB15FBC8DD5F391F" box="[1212,1302,1053,1079]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FA8BFBC8D8A23973" pageId="29" pageNumber="30">Carmen de Carupa</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FF5CFB94D9E23973" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FF5CFB94D9E23973" box="[245,427,1089,1115]" country="Colombia" latitude="5.3502803" longLatPrecision="1" longitude="-73.9053" municipality="Carmen de Carupa" name="Vereda San Jose" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Vereda San José</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FE1CFB95DA123973" box="[437,603,1088,1115]" degrees="5" direction="north" minutes="21" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="01.01" value="5.3502803">5°21’01.01’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FDCBFB95DB553973" box="[610,796,1088,1115]" degrees="73" direction="west" minutes="54" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="19.06" value="-73.9053">73°54’19.06’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FC8FFB94DBD53973" box="[806,924,1089,1115]" metricMagnitude="3" metricUnit="m" metricValue="2.971" pageId="29" pageNumber="30" unit="m" value="2971.0">
|
||
<elevation id="0037D1C332411029FC8FFB94DBD53973" box="[806,924,1089,1115]" metricMagnitude="3" metricUnit="m" metricValue="2.971" pageId="29" pageNumber="30" unit="m" value="2971.0">2971 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FC0FFB94DCE03973" box="[934,1193,1089,1115]" pageId="29" pageNumber="30">diurnal manual capture</collectingMethod>
|
||
,
|
||
<collectedFrom id="3806750D32411029FB1AFB94DD713973" box="[1203,1336,1089,1115]" pageId="29" pageNumber="30">under rocks</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FAEBFB94DDD13973" box="[1346,1432,1089,1115]" pageId="29" pageNumber="30" value="2010-04">
|
||
<collectingDate id="EFE0E9D832411029FAEBFB94DDD13973" box="[1346,1432,1089,1115]" pageId="29" pageNumber="30" value="2010-04">iv.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FF3EFBB0D8BD3957" box="[151,244,1125,1151]" pageId="29" pageNumber="30">J. Cabra</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FEADFBB0D9DF39A8" box="[260,406,1125,1152]" pageId="29" pageNumber="30">ICN-As-996</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FE02FBB0DC26398B" ID-GBIF-Occurrence="2365685688" collectedFrom="on a hill with bushes" collectingDate="2003-08-24" collectorName="F. Fernandez" country="Colombia" elevation="2700" latitude="4.814986" location="El Rosal-Faca" longLatPrecision="1" longitude="-74.356476" municipality="Facatativa" pageId="29" pageNumber="30" specimenCode="ICN-As-512" specimenCount="6" specimenCount-adult="6" specimenCount-female="6" stateProvince="Cundinamarca Department">
|
||
six adult
|
||
<specimenCount id="9D1CFD7932411029FDBDFBB0DA273957" box="[532,622,1125,1151]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FDD3FBB0DAB93957" box="[634,752,1125,1151]" pageId="29" pageNumber="30">Facatativá</collectingMunicipality>
|
||
, highway from
|
||
<location id="8EC5602B32411029FC0AFBB0DC0C3957" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC0AFBB0DC0C3957" box="[931,1093,1125,1151]" country="Colombia" latitude="4.814986" longLatPrecision="1" longitude="-74.356476" municipality="Facatativa" name="El Rosal-Faca" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">El Rosal-Faca</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FBFBFBB1DCB33957" box="[1106,1274,1124,1151]" degrees="4" direction="north" minutes="48" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="53.95" value="4.814986">4°48’53.95’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FAABFBB0D8FB398B" degrees="74" direction="west" minutes="21" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="23.31" value="-74.356476">74°21’23.31’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF14FB5CD97D398B" box="[189,308,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="29" pageNumber="30" unit="m" value="2700.0">
|
||
<elevation id="0037D1C332411029FF14FB5CD97D398B" box="[189,308,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="29" pageNumber="30" unit="m" value="2700.0">2700 masl</elevation>
|
||
</quantity>
|
||
, collected
|
||
<collectedFrom id="3806750D32411029FE02FB5CDADC398B" box="[427,661,1161,1187]" pageId="29" pageNumber="30">on a hill with bushes</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FD08FB5CDB63398B" box="[673,810,1161,1187]" pageId="29" pageNumber="30" value="2003-08-24">
|
||
<collectingDate id="EFE0E9D832411029FD08FB5CDB63398B" box="[673,810,1161,1187]" pageId="29" pageNumber="30" value="2003-08-24">24.viii.2003</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FC9FFB5CDB8F398B" box="[822,966,1161,1187]" pageId="29" pageNumber="30">F. Fernández</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FC7CFB5DDC21398B" box="[981,1128,1160,1187]" pageId="29" pageNumber="30">ICN-As-512</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FBD3FB5CDCE239EF" ID-GBIF-Occurrence="2365685501" country="Colombia" elevation="2600" latitude="4.865797" location="Guasca" longLatPrecision="1" longitude="-73.88328" municipality="Guasca" pageId="29" pageNumber="30" specimenCode="ICN-As-134" specimenCount="1" specimenCount-female="1" stateProvince="Cundinamarca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FB40FB5CDD71398B" box="[1257,1336,1161,1187]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FAEAFB5CDDD1398B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FAEAFB5CDDD1398B" box="[1347,1432,1161,1187]" country="Colombia" latitude="4.865797" longLatPrecision="1" longitude="-73.88328" name="Guasca" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Guasca</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FF3EFB79D97639EF" box="[151,319,1196,1223]" degrees="4" direction="north" minutes="51" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="56.87" value="4.865797">4°51’56.87’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FEE1FB79DA4D39EF" box="[328,516,1196,1223]" degrees="73" direction="west" minutes="52" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.78" value="-73.88328">73°52’59.78’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FDB9FB78DAC039EF" box="[528,649,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FDB9FB78DAC039EF" box="[528,649,1197,1223]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
, without any other data available (
|
||
<specimenCode id="DBBC9E8B32411029FBB8FB78DCEA39EF" box="[1041,1187,1197,1223]" pageId="29" pageNumber="30">ICN-As-134</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FB1EFB78DBDC3838" ID-GBIF-Occurrence="2365686118" collectingDate="1997-06" collectingMethod="manual capture" collectionCode="ICN" collectorName="F. Escobar" country="Colombia" elevation="1500" latitude="4.6093163" location="Vereda Miralindo" longLatPrecision="1" longitude="-73.421844" municipality="Medina" pageId="29" pageNumber="30" specimenCode="ICN- As-194" specimenCount="1" specimenCount-adult="1" specimenCount-male="1" stateProvince="Cundinamarca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FA81FB78DD2839EF" box="[1320,1377,1197,1223]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FAC7FB78D88339C3" pageId="29" pageNumber="30">Medina</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FF7EFB04D9E839C3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FF7EFB04D9E839C3" box="[215,417,1233,1259]" country="Colombia" latitude="4.6093163" longLatPrecision="1" longitude="-73.421844" municipality="Medina" name="Vereda Miralindo" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Vereda Miralindo</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FE06FB05DA1139C3" box="[431,600,1232,1259]" degrees="4" direction="north" minutes="36" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="33.54" value="4.6093163">4°36’33.54’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FDCBFB05DB6939C3" box="[610,800,1232,1259]" degrees="73" direction="west" minutes="25" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="18.65" value="-73.421844">73°25’18.65’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FC84FB05DBEE39C3" box="[813,935,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">
|
||
<elevation id="0037D1C332411029FC84FB05DBEE39C3" box="[813,935,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="29" pageNumber="30" unit="m" value="1500.0">1500 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FC1DFB04DC2C39C3" box="[948,1125,1233,1259]" pageId="29" pageNumber="30">manual capture</collectingMethod>
|
||
,
|
||
<date id="FFA4103032411029FBDAFB04DC8339C3" box="[1139,1226,1233,1259]" pageId="29" pageNumber="30" value="1997-06">
|
||
<collectingDate id="EFE0E9D832411029FBDAFB04DC8339C3" box="[1139,1226,1233,1259]" pageId="29" pageNumber="30" value="1997-06">vi.1997</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FB71FB04DD1939C3" box="[1240,1360,1233,1259]" pageId="29" pageNumber="30">F. Escobar</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FAC8FB04D8A23827" pageId="29" pageNumber="30">ICN- As-194</specimenCode>
|
||
) (misidentified as
|
||
<taxonomicName id="4C1A4D7332411029FE60FB23DA673827" authorityName="Lourenco" authorityYear="1999" box="[457,558,1269,1295]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="erikae">
|
||
<emphasis id="B96EEAE232411029FE60FB23DA673827" box="[457,558,1269,1295]" italics="true" pageId="29" pageNumber="30">T. erikae</emphasis>
|
||
</taxonomicName>
|
||
by W. R. Lourenço,
|
||
<date id="FFA4103032411029FC85FB20DBC43838" box="[812,909,1269,1296]" pageId="29" pageNumber="30" value="1998-12">xii.1998</date>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FC0CFB20DA0F387F" ID-GBIF-Occurrence="2365685757" collectingDate="ii-iv. 2002" collectingMethod="pitfall" collectorName="C. Nino & G. Mora & J. Martinez" country="Colombia" elevation="2600" latitude="4.668044" location="Mondonedo" longLatPrecision="1" longitude="-74.262764" municipality="Mosquera" pageId="29" pageNumber="30" specimenCode="ICN-As-499" specimenCount="13" specimenCount-adult="2" specimenCount-female="13" specimenCount-juvenile="11" stateProvince="Cundinamarca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FBB4FB20DC3D3827" box="[1053,1140,1269,1295]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and 11 juvenile
|
||
<specimenCount id="9D1CFD7932411029FA97FB20DDD13827" box="[1342,1432,1269,1295]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FF3EFACCD943381B" box="[151,266,1305,1331]" pageId="29" pageNumber="30">Mosquera</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FEBEFACCD9E8381B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FEBEFACCD9E8381B" box="[279,417,1305,1331]" country="Colombia" latitude="4.668044" longLatPrecision="1" longitude="-74.262764" municipality="Mosquera" name="Mondonedo" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Mondoñedo</location>
|
||
,
|
||
<location id="8EC5602B32411029FE07FACCDA69381B" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE07FACCDA69381B" box="[430,544,1305,1331]" country="Colombia" latitude="4.668044" longLatPrecision="1" longitude="-74.262764" municipality="Mosquera" name="Zabrinsky" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Zabrinsky</location>
|
||
desert,
|
||
<geoCoordinate id="EE2E503732411029FDD3FACCDB5C381B" box="[634,789,1305,1331]" degrees="4" direction="north" minutes="40" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="4.96" value="4.668044">4°40’4.96’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FCB7FACDDB92381B" box="[798,987,1304,1331]" degrees="74" direction="west" minutes="15" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="45.95" value="-74.262764">74°15’45.95’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FC41FACCDC29381B" box="[1000,1120,1305,1331]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FC41FACCDC29381B" box="[1000,1120,1305,1331]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectingMethod id="525B4EE732411029FBC4FACCDCFA381B" box="[1133,1203,1305,1331]" pageId="29" pageNumber="30">pitfall</collectingMethod>
|
||
,
|
||
<collectingDate id="EFE0E9D832411029FB16FACCDD67381B" box="[1215,1326,1305,1331]" pageId="29" pageNumber="30">ii-iv.2002</collectingDate>
|
||
,
|
||
<collectorName id="26EF532632411029FA92FACCDDD1381B" box="[1339,1432,1305,1331]" pageId="29" pageNumber="30">C. Niño</collectorName>
|
||
,
|
||
<collectorName id="26EF532632411029FF3EFAE8D9D2387F" box="[151,411,1341,1367]" pageId="29" pageNumber="30">G. Mora & J. Martínez</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE02FAE8DA77387F" box="[427,574,1341,1367]" pageId="29" pageNumber="30">ICN-As-499</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FDFBFAE8DAD83854" ID-GBIF-Occurrence="2365686104" collectingDate="1988-02-19" collectorName="P. Ruiz" country="Colombia" elevation="2600" latitude="4.598889" location="Km7- road to Cota" longLatPrecision="21" longitude="-74.08083" municipality="Mosquera" pageId="29" pageNumber="30" specimenCode="ICN-As-016" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Cundinamarca Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FD6BFAE8DB58387F" box="[706,785,1341,1367]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FCB4FAE8DBD9387F" box="[797,912,1341,1367]" pageId="29" pageNumber="30">Mosquera</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FC35FAE8DB80387F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC35FAE8DB80387F" box="[924,969,1341,1367]" country="Colombia" latitude="4.598889" longLatPrecision="21" longitude="-74.08083" municipality="Mosquera" name="Km" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Km</location>
|
||
7- road to
|
||
<location id="8EC5602B32411029FB95FAE8DC3D387F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FB95FAE8DC3D387F" box="[1084,1140,1341,1367]" country="Colombia" latitude="4.598889" longLatPrecision="21" longitude="-74.08083" municipality="Mosquera" name="Cota" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Cota</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FB29FAE9DD4C387F" box="[1152,1285,1340,1367]" degrees="4" direction="north" minutes="35" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="56" value="4.598889">4°35’56’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FAA4FAE9DDD1387F" box="[1293,1432,1340,1367]" degrees="74" direction="west" minutes="4" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="51" value="-74.08083">74°4’51’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EFAB4D9473853" box="[151,270,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FF3EFAB4D9473853" box="[151,270,1377,1403]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FEB0FAB4D9C43853" box="[281,397,1377,1403]" pageId="29" pageNumber="30" value="1988-02-19">
|
||
<collectingDate id="EFE0E9D832411029FEB0FAB4D9C43853" box="[281,397,1377,1403]" pageId="29" pageNumber="30" value="1988-02-19">19.ii.1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE31FAB4D9A13853" box="[408,488,1377,1403]" pageId="29" pageNumber="30">P. Ruiz</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE5FFAB4DAC03854" box="[502,649,1377,1404]" pageId="29" pageNumber="30">ICN-As-016</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD35FAB4DAC738B7" ID-GBIF-Occurrence="2365685701" collectedFrom="inside a house" collectingDate="1998-11-01" collectorName="R. Jaramillo" country="Colombia" elevation="263" latitude="4.4005556" location="Narino" longLatPrecision="21" longitude="-74.83584" pageId="29" pageNumber="30" specimenCode="ICN-As-347" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Narino Department">
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FCA3FAB4DB103853" box="[778,857,1377,1403]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FCCDFAB4DBFC3853" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FCCDFAB4DBFC3853" box="[868,949,1377,1403]" country="Colombia" latitude="4.4005556" longLatPrecision="21" longitude="-74.83584" name="Narino" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Nariño</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC69FAB4DC7F3853" box="[960,1078,1377,1403]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="2" value="4.4005556">4°24’2’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB94FAB5DC8E3853" box="[1085,1223,1376,1403]" degrees="74" direction="west" minutes="50" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="9" value="-74.83584">74°50’9’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FB7BFAB4DD723853" box="[1234,1339,1377,1403]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="29" pageNumber="30" unit="m" value="263.0">
|
||
<elevation id="0037D1C332411029FB7BFAB4DD723853" box="[1234,1339,1377,1403]" metricMagnitude="2" metricUnit="m" metricValue="2.63" pageId="29" pageNumber="30" unit="m" value="263.0">263 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<collectedFrom id="3806750D32411029FAEFFAB4D89238B7" pageId="29" pageNumber="30">inside a house</collectedFrom>
|
||
,
|
||
<date id="FFA4103032411029FF4FFA50D91B38B7" box="[230,338,1413,1439]" pageId="29" pageNumber="30" value="1998-11-01">
|
||
<collectingDate id="EFE0E9D832411029FF4FFA50D91B38B7" box="[230,338,1413,1439]" pageId="29" pageNumber="30" value="1998-11-01">1.xi.1998</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FEF4FA50D9AC38B7" box="[349,485,1413,1439]" pageId="29" pageNumber="30">R. Jaramillo</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE5AFA50DACF38B7" box="[499,646,1413,1439]" pageId="29" pageNumber="30">ICN-As-347</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD31FA50DC8C38EB" ID-GBIF-Occurrence="2365686344" collectingDate="1961-05-13" collectorName="J. Hernandez" country="Colombia" elevation="2600" latitude="4.5499053" location="Hacienda Terrenos, Hoya de la Quebrada Tibanica" longLatPrecision="1" longitude="-74.18328" municipality="Soacha" pageId="29" pageNumber="30" specimenCode="ICN-As-083" specimenCount="5" specimenCount-adult="5" specimenCount-female="5" stateProvince="Cundinamarca Department">
|
||
five adult
|
||
<specimenCount id="9D1CFD7932411029FCA0FA50DB2A38B7" box="[777,867,1413,1439]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FCC4FA50DB8838B7" box="[877,961,1413,1439]" pageId="29" pageNumber="30">Soacha</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FC62FA50DCE938B7" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC62FA50DCE938B7" box="[971,1184,1413,1439]" country="Colombia" latitude="4.5499053" longLatPrecision="1" longitude="-74.18328" municipality="Soacha" name="Hacienda Terrenos" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Hacienda Terrenos</location>
|
||
,
|
||
<location id="8EC5602B32411029FB03FA50DDD138B7" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FB03FA50DDD138B7" box="[1194,1432,1413,1439]" country="Colombia" latitude="4.5499053" longLatPrecision="1" longitude="-74.18328" municipality="Soacha" name="Hoya de la Quebrada" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Hoya de la Quebrada</location>
|
||
,
|
||
<location id="8EC5602B32411029FF3EFA7CD8B238EB" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FF3EFA7CD8B238EB" box="[151,251,1449,1475]" country="Colombia" latitude="4.5499053" longLatPrecision="1" longitude="-74.18328" municipality="Soacha" name="Tibanica" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Tibanica</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FEA1FA7DD9F838EB" box="[264,433,1448,1475]" degrees="4" direction="north" minutes="32" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="59.66" value="4.5499053">4°32’59.66’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FE10FA7DDA3E38EB" box="[441,631,1448,1475]" degrees="74" direction="west" minutes="10" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="59.83" value="-74.18328">74°10’59.83’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FD2AFA7CDAB538EB" box="[643,764,1449,1475]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FD2AFA7CDAB538EB" box="[643,764,1449,1475]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FCA1FA7CDB3038EB" box="[776,889,1449,1475]" pageId="29" pageNumber="30" value="1961-05-13">
|
||
<collectingDate id="EFE0E9D832411029FCA1FA7CDB3038EB" box="[776,889,1449,1475]" pageId="29" pageNumber="30" value="1961-05-13">13.v.1961</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FC2FFA7CDC5338EB" box="[902,1050,1449,1475]" pageId="29" pageNumber="30">J. Hernández</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FB82FA7CDCF438EB" box="[1067,1213,1449,1475]" pageId="29" pageNumber="30">ICN-As-083</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FB7BFA7CDA1A3B23" ID-GBIF-Occurrence="2365685718" collectingDate="2006-12-30" collectorName="J. Pinzon & G. Alarcon" country="Colombia" elevation="2710" latitude="5.4166665" location="Finca Bermejales, Vereda La Fragua" longLatPrecision="21" longitude="-73.816666" municipality="Susa" pageId="29" pageNumber="30" specimenCode="ICN-As-727" specimenCount="3" specimenCount-adult="3" specimenCount-female="1" specimenCount-male="1" stateProvince="Cundinamarca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FAECFA7CDDD538EB" box="[1349,1436,1449,1475]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one adult
|
||
<specimenCount id="9D1CFD7932411029FE92FA18D93D38CF" box="[315,372,1485,1511]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FE28FA18D9F038CF" box="[385,441,1485,1511]" pageId="29" pageNumber="30">Susa</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FE6FFA18DAC738CF" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FE6FFA18DAC738CF" box="[454,654,1485,1511]" country="Colombia" latitude="5.4166665" longLatPrecision="21" longitude="-73.816666" municipality="Susa" name="Finca Bermejales" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Finca Bermejales</location>
|
||
,
|
||
<location id="8EC5602B32411029FD32FA18DB2038CF" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FD32FA18DB2038CF" box="[667,873,1485,1511]" country="Colombia" latitude="5.4166665" longLatPrecision="21" longitude="-73.816666" municipality="Susa" name="Vereda La Fragua" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Vereda La Fragua</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FCDCFA19DBA438CF" box="[885,1005,1484,1511]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="5.4166665">5°25’0’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FC5EFA18DCCA38CF" box="[1015,1155,1485,1511]" degrees="73" direction="west" minutes="49" orientation="longitude" pageId="29" pageNumber="30" precision="15" seconds="0" value="-73.816666">73°49’0’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FB39FA18DD4038CF" box="[1168,1289,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="29" pageNumber="30" unit="m" value="2710.0">
|
||
<elevation id="0037D1C332411029FB39FA18DD4038CF" box="[1168,1289,1485,1511]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="29" pageNumber="30" unit="m" value="2710.0">2710 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FABFFA18DDD138CF" box="[1302,1432,1485,1511]" pageId="29" pageNumber="30" value="2006-12-30">
|
||
<collectingDate id="EFE0E9D832411029FABFFA18DDD138CF" box="[1302,1432,1485,1511]" pageId="29" pageNumber="30" value="2006-12-30">30.xii.2006</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FF3EFA24D9EF3B23" box="[151,422,1521,1547]" pageId="29" pageNumber="30">J. Pinzón & G. Alarcón</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE11FA24DA033B23" box="[440,586,1521,1547]" pageId="29" pageNumber="30">ICN-As-727</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FDC9FA24DA6D3B07" ID-GBIF-Occurrence="2365685954" collectingDate="2002-07-01" collectorName="G. Stiles" country="Colombia" elevation="2690" latitude="4.7536473" location="Tabio" longLatPrecision="1" longitude="-74.088715" municipality="Tabio" pageId="29" pageNumber="30" specimenCode="ICN-As-498" specimenCount="2" specimenCount-female="2" specimenCount-juvenile="2" stateProvince="Cundinamarca Department">
|
||
two juvenile
|
||
<specimenCount id="9D1CFD7932411029FD5EFA24DB183B23" box="[759,849,1521,1547]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FCF7FA24DBE93B23" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FCF7FA24DBE93B23" box="[862,928,1521,1547]" country="Colombia" latitude="4.7536473" longLatPrecision="1" longitude="-74.088715" name="Tabio" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Tabio</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FC07FA25DC113B23" box="[942,1112,1520,1547]" degrees="4" direction="north" minutes="45" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="13.13" value="4.7536473">4°45’13.13’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FBC8FA25DD5B3B23" box="[1121,1298,1520,1547]" degrees="74" direction="west" minutes="5" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="19.36" value="-74.088715">74°5’19.36’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FAB6FA24DDD13B23" box="[1311,1432,1521,1547]" metricMagnitude="3" metricUnit="m" metricValue="2.69" pageId="29" pageNumber="30" unit="m" value="2690.0">
|
||
<elevation id="0037D1C332411029FAB6FA24DDD13B23" box="[1311,1432,1521,1547]" metricMagnitude="3" metricUnit="m" metricValue="2.69" pageId="29" pageNumber="30" unit="m" value="2690.0">2690 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FF3EF9C0D9423B07" box="[151,267,1557,1583]" pageId="29" pageNumber="30" value="2002-07-01">
|
||
<collectingDate id="EFE0E9D832411029FF3EF9C0D9423B07" box="[151,267,1557,1583]" pageId="29" pageNumber="30" value="2002-07-01">1.vii.2002</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FEB1F9C0D9303B07" box="[280,377,1557,1583]" pageId="29" pageNumber="30">G. Stiles</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE20F9C0DA523B07" box="[393,539,1557,1583]" pageId="29" pageNumber="30">ICN-As-498</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD99F9C0DAD33B7B" ID-GBIF-Occurrence="2365686101" collectingDate="2001-12" collectorName="A. Uribe" country="Colombia" elevation="2600" latitude="5.311906" location="Ubate" longLatPrecision="1" longitude="-73.81821" pageId="29" pageNumber="30" specimenCode="ICN-As-100" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Cundinamarca Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FD0BF9C0DAB03B07" box="[674,761,1557,1583]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
and one adult
|
||
<specimenCount id="9D1CFD7932411029FC0BF9C0DB923B07" box="[930,987,1557,1583]" pageId="29" pageNumber="30" type="male">male</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FC41F9C0DC673B07" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC41F9C0DC673B07" box="[1000,1070,1557,1583]" country="Colombia" latitude="5.311906" longLatPrecision="1" longitude="-73.81821" name="Ubate" pageId="29" pageNumber="30" stateProvince="Cundinamarca Department">Ubaté</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FB90F9C1DCAB3B07" box="[1081,1250,1556,1583]" degrees="5" direction="north" minutes="18" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="42.86" value="5.311906">5°18’42.86’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB43F9C1DDD13B07" box="[1258,1432,1556,1583]" degrees="73" direction="west" minutes="49" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="5.56" value="-73.81821">73°49’5.56’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EF9ECD9583B7B" box="[151,273,1593,1619]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">
|
||
<elevation id="0037D1C332411029FF3EF9ECD9583B7B" box="[151,273,1593,1619]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="29" pageNumber="30" unit="m" value="2600.0">2600 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FEB7F9ECD9373B7B" box="[286,382,1593,1619]" pageId="29" pageNumber="30" value="2001-12">
|
||
<collectingDate id="EFE0E9D832411029FEB7F9ECD9373B7B" box="[286,382,1593,1619]" pageId="29" pageNumber="30" value="2001-12">xii.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE20F9ECD9A73B7B" box="[393,494,1593,1619]" pageId="29" pageNumber="30">A. Uribe</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FE56F9ECDADB3B7B" box="[511,658,1593,1619]" pageId="29" pageNumber="30">ICN-As-100</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B723CAD32411029FD0FF9EFD8BE3BB3" ID-GBIF-Occurrence="2365685921" collectingDate="1992-06" collectionCode="ICN" collectorName="A. Calixto" country="Colombia" elevation="1800" latitude="1.2195889" location="Reserva Natural La Planada" longLatPrecision="1" longitude="-77.984055" pageId="29" pageNumber="30" specimenCode="ICN- As-712" specimenCount="10" specimenCount-adult="9" specimenCount-female="8" specimenCount-juvenile="1" specimenCount-male="2" stateProvince="Narino department">
|
||
<collectingRegion id="49DEF81232411029FD0FF9EFDB363B7B" box="[678,895,1593,1619]" country="Colombia" name="Narino" pageId="29" pageNumber="30">
|
||
<emphasis id="B96EEAE232411029FD0FF9EFDB363B7B" box="[678,895,1593,1619]" italics="true" pageId="29" pageNumber="30">Nariño department</emphasis>
|
||
</collectingRegion>
|
||
: seven adult
|
||
<specimenCount id="9D1CFD7932411029FBB2F9ECDC3C3B7B" box="[1051,1141,1593,1619]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
, two adult
|
||
<specimenCount id="9D1CFD7932411029FB5EF9ECDD713B7B" box="[1271,1336,1593,1619]" pageId="29" pageNumber="30" type="male">males</specimenCount>
|
||
and one juvenile
|
||
<specimenCount id="9D1CFD7932411029FF50F988D9013B5F" box="[249,328,1629,1655]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<location id="8EC5602B32411029FEFDF988DADB3B5F" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FEFDF988DADB3B5F" box="[340,658,1629,1655]" country="Colombia" latitude="1.2195889" longLatPrecision="1" longitude="-77.984055" name="Reserva Natural La Planada" pageId="29" pageNumber="30" stateProvince="Narino department">Reserva Natural La Planada</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FD34F989DB0C3B5F" box="[669,837,1628,1655]" degrees="1" direction="north" minutes="13" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="10.52" value="1.2195889">1°13’10.52’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FCE5F989DBA53B5F" box="[844,1004,1628,1655]" degrees="77" direction="west" minutes="59" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="2.6" value="-77.984055">77°59’2.6’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FC51F988DC263B5F" box="[1016,1135,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">
|
||
<elevation id="0037D1C332411029FC51F988DC263B5F" box="[1016,1135,1629,1655]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="29" pageNumber="30" unit="m" value="1800.0">1800 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FBD2F988DC9B3B5F" box="[1147,1234,1629,1655]" pageId="29" pageNumber="30" value="1992-06">
|
||
<collectingDate id="EFE0E9D832411029FBD2F988DC9B3B5F" box="[1147,1234,1629,1655]" pageId="29" pageNumber="30" value="1992-06">vi.1992</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FB74F988DD183B5F" box="[1245,1361,1629,1655]" pageId="29" pageNumber="30">A. Calixto</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FAC8F988D8A73BB3" pageId="29" pageNumber="30">ICN- As-712</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B723CAD32411029FEAAF954DAEB3B97" ID-GBIF-Occurrence="2365686102" collectingDate="1972-07-06" collectorName="P. Garcia" country="Colombia" elevation="1490" latitude="4.407453" location="Boqueron, Tigre Cave" longLatPrecision="1" longitude="-75.264984" municipality="Ibague" pageId="29" pageNumber="30" specimenCode="ICN-As-131" specimenCount="1" specimenCount-adult="1" specimenCount-female="1" stateProvince="Tolima department">
|
||
<emphasis id="B96EEAE232411029FEAAF954D9AA3BB3" box="[259,483,1665,1691]" italics="true" pageId="29" pageNumber="30">
|
||
<collectingRegion id="49DEF81232411029FEAAF954D9943BB3" box="[259,477,1665,1691]" country="Colombia" name="Tolima" pageId="29" pageNumber="30">Tolima department</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
one adult
|
||
<specimenCount id="9D1CFD7932411029FDC9F954DAE63BB3" box="[608,687,1665,1691]" pageId="29" pageNumber="30" type="female">female</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FD15F954DB423BB3" box="[700,779,1665,1691]" pageId="29" pageNumber="30">Ibagué</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FCB0F954DBC33BB3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FCB0F954DBC33BB3" box="[793,906,1665,1691]" country="Colombia" latitude="4.407453" longLatPrecision="1" longitude="-75.264984" municipality="Ibague" name="Boqueron" pageId="29" pageNumber="30" stateProvince="Tolima department">Boquerón</location>
|
||
,
|
||
<location id="8EC5602B32411029FC31F954DC503BB3" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC31F954DC503BB3" box="[920,1049,1665,1691]" country="Colombia" latitude="4.407453" longLatPrecision="1" longitude="-75.264984" municipality="Ibague" name="Tigre Cave" pageId="29" pageNumber="30" stateProvince="Tolima department">Tigre Cave</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FB8EF954DC983BB3" box="[1063,1233,1665,1691]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="26.83" value="4.407453">4°24’26.83’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FB73F955DDD13BB3" box="[1242,1432,1664,1691]" degrees="75" direction="west" minutes="15" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="53.94" value="-75.264984">75°15’53.94’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FF3EF970D9443B97" box="[151,269,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="29" pageNumber="30" unit="m" value="1490.0">
|
||
<elevation id="0037D1C332411029FF3EF970D9443B97" box="[151,269,1701,1727]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="29" pageNumber="30" unit="m" value="1490.0">1490 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FEBEF970D9C53B97" box="[279,396,1701,1727]" pageId="29" pageNumber="30" value="1972-07-06">
|
||
<collectingDate id="EFE0E9D832411029FEBEF970D9C53B97" box="[279,396,1701,1727]" pageId="29" pageNumber="30" value="1972-07-06">6.vii.1972</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FE3FF970D9B23B97" box="[406,507,1701,1727]" pageId="29" pageNumber="30">P. García</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FDA1F970DAD23B97" box="[520,667,1701,1727]" pageId="29" pageNumber="30">ICN-As-131</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B723CAD32411029FD04F970DBEC3BCB" ID-GBIF-Occurrence="2365685869" collectingDate="2007-05-20" collectorName="C. Samudio" country="Colombia" elevation="1200" latitude="4.4406557" location="Botanical Garden San Jose" longLatPrecision="1" longitude="-75.24414" municipality="Ibague" pageId="29" pageNumber="30" specimenCode="ICN-As-743" specimenCount="2" specimenCount-adult="2" specimenCount-female="2" stateProvince="Tolima Department">
|
||
two adult
|
||
<specimenCount id="9D1CFD7932411029FCB2F970DB3C3B97" box="[795,885,1701,1727]" pageId="29" pageNumber="30" type="female">females</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BC1AC8A32411029FCD6F970DB873B97" box="[895,974,1701,1727]" pageId="29" pageNumber="30">Ibagué</collectingMunicipality>
|
||
,
|
||
<location id="8EC5602B32411029FC70F970DD4F3B97" LSID="urn:lsid:plazi:treatment:03B387E632401011FF3EFA0EDBED3B96:8EC5602B32411029FC70F970DD4F3B97" box="[985,1286,1701,1727]" country="Colombia" latitude="4.4406557" longLatPrecision="1" longitude="-75.24414" municipality="Ibague" name="Botanical Garden San Jose" pageId="29" pageNumber="30" stateProvince="Tolima Department">Botanical Garden San José</location>
|
||
,
|
||
<geoCoordinate id="EE2E503732411029FAB9F970D8E23BCB" degrees="4" direction="north" minutes="26" orientation="latitude" pageId="29" pageNumber="30" precision="1" seconds="26.36" value="4.4406557">4°26’26.36’’ N</geoCoordinate>
|
||
<geoCoordinate id="EE2E503732411029FF1BF91DD9273BCB" box="[178,366,1736,1763]" degrees="75" direction="west" minutes="14" orientation="longitude" pageId="29" pageNumber="30" precision="1" seconds="38.91" value="-75.24414">75°14’38.91’’ W</geoCoordinate>
|
||
,
|
||
<quantity id="4CE29B1532411029FED0F91CD9B93BCB" box="[377,496,1737,1763]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="m" value="1200.0">
|
||
<elevation id="0037D1C332411029FED0F91CD9B93BCB" box="[377,496,1737,1763]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="m" value="1200.0">1200 masl</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFA4103032411029FE52F91CDA223BCB" box="[507,619,1737,1763]" pageId="29" pageNumber="30" value="2007-05-20">
|
||
<collectingDate id="EFE0E9D832411029FE52F91CDA223BCB" box="[507,619,1737,1763]" pageId="29" pageNumber="30" value="2007-05-20">20.v.2007</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26EF532632411029FDDEF91CDAB53BCB" box="[631,764,1737,1763]" pageId="29" pageNumber="30">C. Samudio</collectorName>
|
||
(
|
||
<specimenCode id="DBBC9E8B32411029FCA3F91CDBD43BCB" box="[778,925,1737,1763]" pageId="29" pageNumber="30">ICN-As-743</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300657B3241102AFF6EF939DA5B3CDF" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="diagnosis">
|
||
<paragraph id="8BA536F03241102AFF6EF939DA5B3CDF" blockId="29.[151,1437,153,2015]" lastBlockId="30.[151,1437,153,503]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
<emphasis id="B96EEAE232411029FF6EF939D90B3A2F" bold="true" box="[199,322,1772,1799]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
|
||
This species can be readily recognized from other
|
||
<taxonomicName id="4C1A4D7332411029FC2EF93BDC363A2F" authority="" authorityName="Lourenco" authorityYear="2006" box="[903,1151,1773,1799]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE232411029FC2EF93BDB833A2F" box="[903,970,1774,1799]" italics="true" pageId="29" pageNumber="30">Tityus</emphasis>
|
||
(
|
||
<emphasis id="B96EEAE232411029FC75F938DC3F3A2F" box="[988,1142,1773,1799]" italics="true" pageId="29" pageNumber="30">Archaeotityus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species by the following combination of character states: Total length:
|
||
<quantity id="4CE29B1532411029FD0EF8C4DB2E3A03" box="[679,871,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.3634999999999997" metricValueMax="3.911" metricValueMin="2.816" pageId="29" pageNumber="30" unit="mm" value="33.635" valueMax="39.11" valueMin="28.16">28.16–39.11 mm</quantity>
|
||
(males) and
|
||
<quantity id="4CE29B1532411029FBA9F8C4DC893A03" box="[1024,1216,1809,1835]" metricMagnitude="-2" metricUnit="m" metricValue="3.3634999999999997" metricValueMax="3.911" metricValueMin="2.816" pageId="29" pageNumber="30" unit="mm" value="33.635" valueMax="39.11" valueMin="28.16">28.16–39.11 mm</quantity>
|
||
(females). General body coloration (
|
||
<figureCitation id="13212A7532411029FEFEF8E0D98E3A67" box="[343,455,1845,1871]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="29" pageNumber="30">Figure 21</figureCitation>
|
||
) dark yellow background moderately covered with dark brown variegated spots; metasomal segment V completely dark reddish-brown with three ventral black stripes and with a dorsomedian area without spots (
|
||
<figureCitation id="13212A7532411029FEEBF8A8D9943ABF" box="[322,477,1917,1943]" captionStart="FIGURE 6" captionStartId="14.[151,250,1837,1862]" captionTargetBox="[151,1415,186,1776]" captionTargetId="figure@14.[151,1436,181,1776]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377147/files/figure.png" pageId="29" pageNumber="30">Figure 6B, E</figureCitation>
|
||
). Metasomal segments II–III (
|
||
<figureCitation id="13212A7532411029FC90F8A8DB833ABF" box="[825,970,1916,1943]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="29" pageNumber="30">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A7532411029FC71F8A8DC643ABF" box="[984,1069,1917,1943]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="29" pageNumber="30">16C, D</figureCitation>
|
||
) each has a distoterminal small acute granule on the DL carinae. Metasomal segment IV (
|
||
<figureCitation id="13212A7532411029FCB4F874DBE33A93" box="[797,938,1952,1979]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="29" pageNumber="30">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A7532411029FC1FF874DC413A93" box="[950,1032,1953,1979]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="29" pageNumber="30">16C, D</figureCitation>
|
||
) with a distoterminal medium-sized blunt, and coarse granule on the DL carinae. Metasomal segment V with the VSM2 macrosetae slightly distal or at the same level of the VL2 macrosetae (
|
||
<figureCitation id="13212A753242102AFDD1FF4CDB413D9B" box="[632,776,153,179]" captionStart="FIGURE 17" captionStartId="27.[151,250,1559,1584]" captionTargetBox="[151,1436,181,1539]" captionTargetId="figure@27.[151,1436,181,1539]" captionTargetPageId="27" captionText="FIGURE 17. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377169/files/figure.png" pageId="30" pageNumber="31">Figures 17B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753242102AFCBFFF4CDB0E3D9B" box="[790,839,153,179]" captionStart="FIGURE 18" captionStartId="30.[151,250,1899,1924]" captionTargetBox="[151,1422,525,1880]" captionTargetId="figure@30.[151,1422,525,1880]" captionTargetPageId="30" captionText="FIGURE 18. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As- 181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377171/files/figure.png" pageId="30" pageNumber="31">18B</figureCitation>
|
||
). Subaculear tubercle medium-sized pyramidal, flat, and triangle-like with an acute and coarse apex in lateral view (
|
||
<figureCitation id="13212A753242102AFCF1FF68DBAF3DFF" box="[856,998,189,215]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="30" pageNumber="31">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753242102AFC58FF68DC6D3DFF" box="[1009,1060,189,215]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="30" pageNumber="31">20B</figureCitation>
|
||
). Dorsal margin of the subaculear tubercle with a pair of ventrodistal small rounded granules with apices pointing to the middle portion of the aculeus. Vesicle and subaculear tubercle with VM carina composed of coarse granules and strongly crenulate in lateral view (
|
||
<figureCitation id="13212A753242102AFF36FEFCD9643C6B" box="[159,301,297,323]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="30" pageNumber="31">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753242102AFE93FEFCD9253C6B" box="[314,364,297,323]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="30" pageNumber="31">20B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1A4D733242102AFE29FEFCD9833C6B" box="[384,458,297,323]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="30" pageNumber="84" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
|
||
with well-marked (female) (
|
||
<figureCitation id="13212A753242102AFCA7FEFCDBD93C6B" box="[782,912,297,323]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="30" pageNumber="31">Figure 19B</figureCitation>
|
||
) or moderately marked (male) carinae (
|
||
<figureCitation id="13212A753242102AFAF8FEFCD8833C4F" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="30" pageNumber="31">Figure 20B</figureCitation>
|
||
). Metasomal intercarinal areas densely covered with coarse granules (
|
||
<figureCitation id="13212A753242102AFC5BFE98DCCB3C4F" box="[1010,1154,332,359]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="30" pageNumber="31">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753242102AFB38FE98DCA13C4F" box="[1169,1256,333,359]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="30" pageNumber="31">16C, D</figureCitation>
|
||
). Pectinal tooth counts and movable finger oblique rows of denticles: 11–15/ 11–13 (females) and 13–15/ 12–14 (males) (
|
||
<tableCitation id="C698034B3242102AFAE9FEA4DDDE3CA3" box="[1344,1431,369,395]" captionStart="TABLE 2" captionStartId="8.[151,239,478,502]" captionText="TABLE 2. Pectinal tooth counts of Colombian Tityus (Archaeotityus) species. Note: Both pectines of each examined specimen were counted. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="30" pageNumber="31">Table 2</tableCitation>
|
||
,
|
||
<tableCitation id="C698034B3242102AFF3EFE40D8EE3C87" box="[151,167,405,431]" captionStart="TABLE 3" captionStartId="28.[151,239,730,754]" captionText="TABLE 3. Number of movable finger oblique rows of denticles among Colombian Tityus (Archaeotityus) species. Note: Both movable fingers of each examined specimen were included. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="30" pageNumber="31">3</tableCitation>
|
||
). Metasomal segments II–V (
|
||
<figureCitation id="13212A753242102AFE50FE40DAC13C87" box="[505,648,404,431]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="30" pageNumber="31">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753242102AFD3FFE40DAA23C87" box="[662,747,405,431]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="30" pageNumber="31">16C, D</figureCitation>
|
||
), female pedipalp (
|
||
<figureCitation id="13212A753242102AFC6EFE40DC7A3C87" box="[967,1075,405,431]" captionStart="FIGURE 11" captionStartId="21.[151,250,1944,1969]" captionTargetBox="[272,1316,166,1924]" captionTargetId="figure@21.[272,1316,166,1924]" captionTargetPageId="21" captionText="FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. A–C. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377157/files/figure.png" pageId="30" pageNumber="31">Figure 11</figureCitation>
|
||
D–F), and male pedipalp femur and
|
||
<taxonomicName id="4C1A4D733242102AFF6FFE6CD9583CFB" box="[198,273,441,467]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="30" pageNumber="71" phylum="Mollusca" rank="genus">patella</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A753242102AFEB7FE6CD9893CFB" box="[286,448,441,467]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="30" pageNumber="31">Figure 10C, D</figureCitation>
|
||
) short and of a medium-width.
|
||
<taxonomicName id="4C1A4D733242102AFCB7FE6CDB163CFB" box="[798,863,441,467]" class="Actinopterygii" family="Cyprinidae" genus="Chela" kingdom="Animalia" order="Cypriniformes" pageId="30" pageNumber="76" phylum="Chordata" rank="genus">Chela</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A753242102AFCC4FE6CDB933CFB" box="[877,986,441,467]" captionStart="FIGURE 22" captionStartId="36.[151,250,1928,1953]" captionTargetBox="[316,1271,166,1909]" captionTargetId="figure@36.[316,1271,166,1909]" captionTargetPageId="36" captionText="FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. A–C. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="30" pageNumber="31">Figure 22</figureCitation>
|
||
A–C) and metasomal segment V (
|
||
<figureCitation id="13212A753242102AFAF8FE6CD8A03CDF" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="30" pageNumber="31">Figure 16C, D</figureCitation>
|
||
) incrassate in large males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6566783242102AFF3EF8BEDCF53AD8" httpUri="https://zenodo.org/record/3377171/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1899,1924]" targetBox="[151,1422,525,1880]" targetPageId="30">
|
||
<paragraph id="8BA536F03242102AFF3EF8BEDCF53AD8" blockId="30.[151,1436,1899,2032]" pageId="30" pageNumber="31">
|
||
<emphasis id="B96EEAE23242102AFF3EF8BED96A3AAC" bold="true" box="[151,291,1899,1924]" pageId="30" pageNumber="31">FIGURE 18.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFE85F8B8DAF43AAC" authority="Lourenco, 2006" authorityName="Lourenco" authorityYear="2006" box="[300,701,1899,1924]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE23242102AFE85F8B8DA473AAC" box="[300,526,1900,1924]" italics="true" pageId="30" pageNumber="31">Tityus (Archaeotityus)</emphasis>
|
||
<bibRefCitation id="EF8B4B013242102AFDBEF8B8DAF43AAC" author="Lourenco, W. R. & Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F." box="[535,701,1899,1924]" pageId="30" pageNumber="31" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh " o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, male metasomal segment V sculpturing, showing macrosetae, ventral aspect.
|
||
<emphasis id="B96EEAE23242102AFF4BF85AD8B23A80" bold="true" box="[226,251,1935,1960]" pageId="30" pageNumber="31">A.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFEA8F844DA7A3A80" authority="Lourenco, 1984" authorityName="Lourenco" authorityYear="1984" box="[257,563,1936,1960]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bastosi">
|
||
<emphasis id="B96EEAE23242102AFEA8F844D9C53A80" box="[257,396,1936,1960]" italics="true" pageId="30" pageNumber="31">Tityus bastosi</emphasis>
|
||
<bibRefCitation id="EF8B4B013242102AFE3BF844DA7A3A80" author="Lourenco, W. R." box="[402,563,1936,1960]" pageId="30" pageNumber="31" pagination="349 - 360" refId="ref49629" refString="Lourenco, W. R. (1984) Analyse taxonomique des Scorpions du groupe Tityus clathratus Koch, 1845 (Scorpiones, Buthidae). Bulletin Museum National d' Histoire Naturelle, Paris, 4 e series, 6, 349 - 360." type="journal article" year="1984">Lourenço, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-741).
|
||
<emphasis id="B96EEAE23242102AFD7FF845DAA43A80" bold="true" box="[726,749,1936,1960]" pageId="30" pageNumber="31">B.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFD5AF844DC103A80" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[755,1113,1935,1960]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE23242102AFD5AF844DBFE3A80" box="[755,951,1936,1960]" italics="true" pageId="30" pageNumber="31">Tityus columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B013242102AFC6CF85ADC1B3A80" author="Thorell, T." box="[965,1106,1935,1960]" pageId="30" pageNumber="31" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(ICN-As-524).
|
||
<emphasis id="B96EEAE23242102AFB55F85ADD5C3A80" bold="true" box="[1276,1301,1935,1960]" pageId="30" pageNumber="31">C.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFAB2F844DDD53A80" authorityName="Moreno-González & O & D" authorityYear="2019" box="[1307,1436,1937,1960]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="guane">
|
||
<emphasis id="B96EEAE23242102AFAB2F844DDD53A80" box="[1307,1436,1937,1960]" italics="true" pageId="30" pageNumber="31">Tityus guane</emphasis>
|
||
</taxonomicName>
|
||
(holotype) (ICN-As-1003).
|
||
<emphasis id="B96EEAE23242102AFE1DF861D9843AE4" bold="true" box="[436,461,1972,1996]" pageId="30" pageNumber="31">D.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFE7CF860DB683AE4" authority="Kraepelin, 1914" authorityName="Kraepelin" authorityYear="1914" box="[469,801,1971,1996]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B96EEAE23242102AFE7CF860DA3D3AE4" box="[469,628,1972,1996]" italics="true" pageId="30" pageNumber="31">Tityus parvulus</emphasis>
|
||
<bibRefCitation id="EF8B4B013242102AFDD5F866DB683AE4" author="Kraepelin, K." box="[636,801,1971,1996]" pageId="30" pageNumber="31" pagination="15 - 28" refId="ref49461" refString="Kraepelin, K. (1914) Beitrag zur Kenntnis der Skorpione und Pedipalpen Columbiens. Memoires de la Societe des Sciences naturelles, Neuchatel, 5, 15 - 28." type="journal article" year="1914">Kraepelin, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-1004).
|
||
<emphasis id="B96EEAE23242102AFC7CF861DBA53AE4" bold="true" box="[981,1004,1972,1996]" pageId="30" pageNumber="31">E.</emphasis>
|
||
<taxonomicName id="4C1A4D733242102AFC5CF860DD7A3AE4" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[1013,1331,1972,1996]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="tayrona">
|
||
<emphasis id="B96EEAE23242102AFC5CF860DCC13AE4" box="[1013,1160,1973,1996]" italics="true" pageId="30" pageNumber="31">Tityus tayrona</emphasis>
|
||
<bibRefCitation id="EF8B4B013242102AFB39F860DD7A3AE4" author="Lourenco, W. R." box="[1168,1331,1972,1996]" pageId="30" pageNumber="31" pagination="275 - 288" refId="ref49674" refString="Lourenco, W. R. (1991) Les scorpions de Colombie, II. Les faunes des regions de Santa Marta et de la. Cordillere Orientale. Approche biogeographique. Senckenbergiana Biologica, 71 (4), 275 - 288." type="journal article" year="1991">Lourenço, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As- 181).
|
||
<emphasis id="B96EEAE23242102AFF7BF80DD9163AD8" box="[210,351,2008,2032]" italics="true" pageId="30" pageNumber="31">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96EEAE23242102AFEC4F80CD9D93AD8" box="[365,400,2009,2032]" italics="true" pageId="30" pageNumber="31">ML</emphasis>
|
||
, median lateral;
|
||
<emphasis id="B96EEAE23242102AFD93F80CDA113AD8" box="[570,600,2009,2032]" italics="true" pageId="30" pageNumber="31">VL</emphasis>
|
||
, ventrolateral;
|
||
<emphasis id="B96EEAE23242102AFD5BF80CDB6A3AD8" box="[754,803,2009,2032]" italics="true" pageId="30" pageNumber="31">VSM</emphasis>
|
||
, ventrosubmedian. Scale bars= 1.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6566783243102BFF3EFBACDD1D39F1" httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,250,1145,1170]" targetBox="[151,1436,181,1126]" targetPageId="31">
|
||
<paragraph id="8BA536F03243102BFF3EFBACDD1D39F1" blockId="31.[151,1437,1145,1242]" pageId="31" pageNumber="32">
|
||
<emphasis id="B96EEAE23243102BFF3EFBACD95639BA" bold="true" box="[151,287,1145,1170]" pageId="31" pageNumber="32">FIGURE 19.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFE8CFBAEDAE239BA" authority="Lourenco, 2006" authorityName="Lourenco" authorityYear="2006" box="[293,683,1145,1170]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE23243102BFE8CFBAEDA4A39BA" box="[293,515,1146,1170]" italics="true" pageId="31" pageNumber="32">Tityus (Archaeotityus)</emphasis>
|
||
<bibRefCitation id="EF8B4B013243102BFDA0FBAFDAE239BA" author="Lourenco, W. R. & Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F." box="[521,683,1145,1170]" pageId="31" pageNumber="32" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh " o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, female
|
||
<taxonomicName id="4C1A4D733243102BFD56FBACDB7539BA" box="[767,828,1145,1170]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="31" pageNumber="84" phylum="Arthropoda" rank="genus">telson</taxonomicName>
|
||
sculpturing, lateral aspect.
|
||
<emphasis id="B96EEAE23243102BFBE6FBACDC2139BA" bold="true" box="[1103,1128,1145,1170]" pageId="31" pageNumber="32">A.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFBC4FBAEDDD439BA" authority="Lourenco, 1984" authorityName="Lourenco" authorityYear="1984" box="[1133,1437,1146,1170]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bastosi">
|
||
<emphasis id="B96EEAE23243102BFBC4FBAEDCBE39BA" box="[1133,1271,1146,1170]" italics="true" pageId="31" pageNumber="32">Tityus bastosi</emphasis>
|
||
<bibRefCitation id="EF8B4B013243102BFB54FBAFDDD439BA" author="Lourenco, W. R." box="[1277,1437,1146,1170]" pageId="31" pageNumber="32" pagination="349 - 360" refId="ref49629" refString="Lourenco, W. R. (1984) Analyse taxonomique des Scorpions du groupe Tityus clathratus Koch, 1845 (Scorpiones, Buthidae). Bulletin Museum National d' Histoire Naturelle, Paris, 4 e series, 6, 349 - 360." type="journal article" year="1984">Lourenço, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-740).
|
||
<emphasis id="B96EEAE23243102BFE9CFB4BD905399E" bold="true" box="[309,332,1182,1206]" pageId="31" pageNumber="32">B.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFEFDFB4ADAF7399E" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[340,702,1181,1206]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE23243102BFEFDFB4ADA53399E" box="[340,538,1182,1206]" italics="true" pageId="31" pageNumber="32">Tityus columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B013243102BFD80FB48DAFE399E" author="Thorell, T." box="[553,695,1181,1206]" pageId="31" pageNumber="32" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(ICN-As-739).
|
||
<emphasis id="B96EEAE23243102BFCCDFB48DB35399E" bold="true" box="[868,892,1181,1206]" pageId="31" pageNumber="32">C.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFC2DFB4ADC4E399E" authority="Moreno-González & O & D, 2019" authorityName="Moreno-González & O & D" authorityYear="2019" box="[900,1031,1183,1206]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="guane" status="sp. nov.">
|
||
<emphasis id="B96EEAE23243102BFC2DFB4ADC4E399E" box="[900,1031,1183,1206]" italics="true" pageId="31" pageNumber="32">Tityus guane</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25D57993243102BFBA6FB4BDC28399E" box="[1039,1121,1182,1206]" pageId="31" pageNumber="32" rank="species">
|
||
<emphasis id="B96EEAE23243102BFBA6FB4BDC28399E" bold="true" box="[1039,1121,1182,1206]" pageId="31" pageNumber="32">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(paratype) (ICN-As-1003).
|
||
<emphasis id="B96EEAE23243102BFA2DFB4BDDD5399E" bold="true" box="[1412,1436,1182,1206]" pageId="31" pageNumber="32">D.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFF3EFB16D99439F2" authority="Kraepelin, 1914" authorityName="Kraepelin" authorityYear="1914" box="[151,477,1217,1242]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B96EEAE23243102BFF3EFB16D97A39F2" box="[151,307,1218,1242]" italics="true" pageId="31" pageNumber="32">Tityus parvulus</emphasis>
|
||
<bibRefCitation id="EF8B4B013243102BFE90FB14D99439F2" author="Kraepelin, K." box="[313,477,1217,1242]" pageId="31" pageNumber="32" pagination="15 - 28" refId="ref49461" refString="Kraepelin, K. (1914) Beitrag zur Kenntnis der Skorpione und Pedipalpen Columbiens. Memoires de la Societe des Sciences naturelles, Neuchatel, 5, 15 - 28." type="journal article" year="1914">Kraepelin, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-1004).
|
||
<emphasis id="B96EEAE23243102BFD22FB17DAEB39F2" bold="true" box="[651,674,1218,1242]" pageId="31" pageNumber="32">E.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFD00FB16DBA839F2" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[681,993,1218,1242]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tayrona">
|
||
<emphasis id="B96EEAE23243102BFD00FB16DB7339F2" box="[681,826,1219,1242]" italics="true" pageId="31" pageNumber="32">Tityus tayrona</emphasis>
|
||
<bibRefCitation id="EF8B4B013243102BFCE9FB17DBA839F2" author="Lourenco, W. R." box="[832,993,1218,1242]" pageId="31" pageNumber="32" pagination="275 - 288" refId="ref49674" refString="Lourenco, W. R. (1991) Les scorpions de Colombie, II. Les faunes des regions de Santa Marta et de la. Cordillere Orientale. Approche biogeographique. Senckenbergiana Biologica, 71 (4), 275 - 288." type="journal article" year="1991">Lourenço, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-181). Scale bars= 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C300657B3243102BFF6EFADDDC4C3A83" pageId="31" pageNumber="32" type="biology_ecology">
|
||
<paragraph id="8BA536F03243102BFF6EFADDDB563BA3" blockId="31.[151,1437,1288,2035]" pageId="31" pageNumber="32">
|
||
<emphasis id="B96EEAE23243102BFF6EFADDDAFB380B" bold="true" box="[199,690,1288,1315]" pageId="31" pageNumber="32">Affinities with other Colombian species.</emphasis>
|
||
<taxonomicName id="4C1A4D733243102BFD15FADCDBD0380A" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[700,921,1288,1314]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE23243102BFD15FADCDBD0380A" box="[700,921,1288,1314]" italics="true" pageId="31" pageNumber="32">Tityus columbianus</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C1A4D733243102BFBD2FADCDCB5380A" authorityName="Kraepelin" authorityYear="1914" box="[1147,1276,1288,1314]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B96EEAE23243102BFBD2FADCDCB5380A" box="[1147,1276,1288,1314]" italics="true" pageId="31" pageNumber="32">T. parvulus</emphasis>
|
||
</taxonomicName>
|
||
, sharing with it the presence of a subaculear tubercle that is medium-sized pyramidal, flat, and triangle-like with an acute and coarse apex in lateral view (
|
||
<figureCitation id="13212A753243102BFE79FA85DA173843" box="[464,606,1360,1387]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="31" pageNumber="32">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFDC3FA84DAD53843" box="[618,668,1361,1387]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="31" pageNumber="32">20B</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1A4D733243102BFD19FA84DBC13842" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[688,904,1360,1386]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE23243102BFD19FA84DBC13842" box="[688,904,1360,1386]" italics="true" pageId="31" pageNumber="32">Tityus columbianus</emphasis>
|
||
</taxonomicName>
|
||
can be readily distinguished by the presence of a general body coloration with a dark yellow background moderately covered with dark brown spots (
|
||
<figureCitation id="13212A753243102BFAB5FAA1DDC738A7" box="[1308,1422,1396,1423]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="31" pageNumber="32">Figure 21</figureCitation>
|
||
); metasomal segment V completely dark reddish-brown with three ventral black stripes, a dorsomedian area without spots (
|
||
<figureCitation id="13212A753243102BFF77FA69D92738FF" box="[222,366,1468,1495]" captionStart="FIGURE 6" captionStartId="14.[151,250,1837,1862]" captionTargetBox="[151,1415,186,1776]" captionTargetId="figure@14.[151,1436,181,1776]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377147/files/figure.png" pageId="31" pageNumber="32">Figure 6B, E</figureCitation>
|
||
) and with the VSM2 macrosetae; metasomal segments II–III each with a distoterminal small and acute granule on the DL carinae (
|
||
<figureCitation id="13212A753243102BFDA7FA35DAD838D3" box="[526,657,1504,1531]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="31" pageNumber="32">Figure 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFD34FA35DAA638D3" box="[669,751,1504,1531]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="31" pageNumber="32">16C, D</figureCitation>
|
||
) and metasomal segment IV with a medium-sized blunt, and coarse granule on the DL carinae (
|
||
<figureCitation id="13212A753243102BFDBEF9D1DAED3B37" box="[535,676,1540,1567]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="31" pageNumber="32">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFD19F9D1DB4D3B37" box="[688,772,1540,1567]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="31" pageNumber="32">16C, D</figureCitation>
|
||
); metasomal intercarinal areas densely covered with coarse granules (
|
||
<figureCitation id="13212A753243102BFEAEF9FDD9DE3B6B" box="[263,407,1576,1603]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="31" pageNumber="32">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFE0FF9FDD9B53B6B" box="[422,508,1576,1603]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="31" pageNumber="32">16C, D</figureCitation>
|
||
); subaculear tubercle with VM carina composed of coarse granules and strongly crenulate in lateral view (
|
||
<figureCitation id="13212A753243102BFE1BF999DA763B4F" box="[434,575,1612,1639]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="31" pageNumber="32">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFDE3F998DA343B4F" box="[586,637,1613,1639]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="31" pageNumber="32">20B</figureCitation>
|
||
); metasomal segments (
|
||
<figureCitation id="13212A753243102BFC2EF999DC5D3B4F" box="[903,1044,1612,1639]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="31" pageNumber="32">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFBB6F999DC393B4F" box="[1055,1136,1612,1639]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="31" pageNumber="32">16C, D</figureCitation>
|
||
) and male pedipalp femur and
|
||
<taxonomicName id="4C1A4D733243102BFF6EF9A5D95B3BA2" box="[199,274,1648,1674]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="31" pageNumber="71" phylum="Mollusca" rank="genus">patella</taxonomicName>
|
||
short and of a medium-width (
|
||
<figureCitation id="13212A753243102BFDC5F9A5DB5B3BA3" box="[620,786,1648,1675]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="31" pageNumber="32">Figure 10C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA536F03243102BFF6EF941DC4C3A83" blockId="31.[151,1437,1288,2035]" pageId="31" pageNumber="32">
|
||
Conversely, in
|
||
<taxonomicName id="4C1A4D733243102BFED9F940D9A73B86" authorityName="Kraepelin" authorityYear="1914" box="[368,494,1684,1710]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B96EEAE23243102BFED9F940D9A73B86" box="[368,494,1684,1710]" italics="true" pageId="31" pageNumber="32">T. parvulus</emphasis>
|
||
</taxonomicName>
|
||
the general body coloration is with a dark yellow background slightly covered with dark brown variegated spots (
|
||
<figureCitation id="13212A753243102BFE76F96DDA063BFB" box="[479,591,1720,1747]" captionStart="FIGURE 31" captionStartId="54.[151,250,1932,1957]" captionTargetBox="[172,1391,176,1901]" captionTargetId="figure@54.[162,1425,166,1912]" captionTargetPageId="54" captionText="FIGURE 31. Tityus (Archaeotityus) parvulus Kraepelin, 1914, habitus. A, B. Female (ICN-As-1004). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Chinchiná, Caldas) (ICN-As-1004). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Chinchiná, Caldas) (ICN-As-1004). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377199/files/figure.png" pageId="31" pageNumber="32">Figure 31</figureCitation>
|
||
); the metasomal segment V is dark yellow slightly covered with dark brown variegated spots and a pair of distal VL dark brown spots, but does not have dorsal spots neither the VSM macrosetae (
|
||
<figureCitation id="13212A753243102BFF76F8D5D9C03A33" box="[223,393,1792,1819]" captionStart="FIGURE 33" captionStartId="56.[151,250,1937,1962]" captionTargetBox="[338,1190,173,1909]" captionTargetId="figure@56.[338,1249,166,1918]" captionTargetPageId="56" captionText="FIGURE 33. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, C. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). A. Female. C. Male. B, D. Tityus tayrona Lourenço, 1991 (ICN-As-181). B. Female. D. Male." httpUri="https://zenodo.org/record/3377203/files/figure.png" pageId="31" pageNumber="32">Figure 33A, C</figureCitation>
|
||
); the metasomal segments II–IV do not have conspicuous distoterminal granules on the DL carinae (
|
||
<figureCitation id="13212A753243102BFF50F8F1D9C33A17" box="[249,394,1828,1855]" captionStart="FIGURE 39" captionStartId="63.[151,250,1975,2000]" captionTargetBox="[473,1115,166,1956]" captionTargetId="figure@63.[473,1115,166,1956]" captionTargetPageId="63" captionText="FIGURE 39. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus parvulus Kraepelin, 1914 (ICN-As-104). B. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bar= 3.0 mm." httpUri="https://zenodo.org/record/3377215/files/figure.png" pageId="31" pageNumber="32">Figures 39A</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFE3EF8F0D9A43A17" box="[407,493,1829,1855]" captionStart="FIGURE 40" captionStartId="64.[151,250,1896,1921]" captionTargetBox="[323,1265,166,1876]" captionTargetId="figure@64.[323,1265,166,1876]" captionTargetPageId="64" captionText="FIGURE 40. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus parvulus Kraepelin, 1914. A. Male (Chinchiná, Caldas) (ICN-As-1004). B. Male (Chinchiná, Caldas) (ICN-As-1004). C, D. Tityus tayrona Lourenço, 1991. C. Male (Zambrano, Bolívar) (ICN-As-181). D. Male (Zambrano, Bolívar) (ICN-As-181). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377217/files/figure.png" pageId="31" pageNumber="32">40A, B</figureCitation>
|
||
); the metasomal intercarinal areas are densely covered with fine granules (
|
||
<figureCitation id="13212A753243102BFAEEF8F1D8833A4B" captionStart="FIGURE 39" captionStartId="63.[151,250,1975,2000]" captionTargetBox="[473,1115,166,1956]" captionTargetId="figure@63.[473,1115,166,1956]" captionTargetPageId="63" captionText="FIGURE 39. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus parvulus Kraepelin, 1914 (ICN-As-104). B. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bar= 3.0 mm." httpUri="https://zenodo.org/record/3377215/files/figure.png" pageId="31" pageNumber="32">Figures 39A</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFF7EF89CD9453A4B" box="[215,268,1865,1891]" captionStart="FIGURE 40" captionStartId="64.[151,250,1896,1921]" captionTargetBox="[323,1265,166,1876]" captionTargetId="figure@64.[323,1265,166,1876]" captionTargetPageId="64" captionText="FIGURE 40. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus parvulus Kraepelin, 1914. A. Male (Chinchiná, Caldas) (ICN-As-1004). B. Male (Chinchiná, Caldas) (ICN-As-1004). C, D. Tityus tayrona Lourenço, 1991. C. Male (Zambrano, Bolívar) (ICN-As-181). D. Male (Zambrano, Bolívar) (ICN-As-181). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377217/files/figure.png" pageId="31" pageNumber="32">40A</figureCitation>
|
||
); the subaculear tubercle has a VM carina composed of fine granules and is feebly crenulate in lateral view (
|
||
<figureCitation id="13212A753243102BFF74F8B9D9273AAF" box="[221,366,1900,1927]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="31" pageNumber="32">Figures 19D</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFED2F8B8D9F93AAF" box="[379,432,1901,1927]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="31" pageNumber="32">20D</figureCitation>
|
||
); the metasomal segments are slender in both sexes (
|
||
<figureCitation id="13212A753243102BFBA0F8B9DCD33AAF" box="[1033,1178,1900,1927]" captionStart="FIGURE 39" captionStartId="63.[151,250,1975,2000]" captionTargetBox="[473,1115,166,1956]" captionTargetId="figure@63.[473,1115,166,1956]" captionTargetPageId="63" captionText="FIGURE 39. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus parvulus Kraepelin, 1914 (ICN-As-104). B. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bar= 3.0 mm." httpUri="https://zenodo.org/record/3377215/files/figure.png" pageId="31" pageNumber="32">Figures 39A</figureCitation>
|
||
,
|
||
<figureCitation id="13212A753243102BFB0EF8B8DCB03AAF" box="[1191,1273,1901,1927]" captionStart="FIGURE 40" captionStartId="64.[151,250,1896,1921]" captionTargetBox="[323,1265,166,1876]" captionTargetId="figure@64.[323,1265,166,1876]" captionTargetPageId="64" captionText="FIGURE 40. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus parvulus Kraepelin, 1914. A. Male (Chinchiná, Caldas) (ICN-As-1004). B. Male (Chinchiná, Caldas) (ICN-As-1004). C, D. Tityus tayrona Lourenço, 1991. C. Male (Zambrano, Bolívar) (ICN-As-181). D. Male (Zambrano, Bolívar) (ICN-As-181). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377217/files/figure.png" pageId="31" pageNumber="32">40A, B</figureCitation>
|
||
) and the male pedipalp femur and
|
||
<taxonomicName id="4C1A4D733243102BFED1F845D98A3A82" box="[376,451,1936,1962]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="31" pageNumber="71" phylum="Mollusca" rank="genus">patella</taxonomicName>
|
||
are slightly elongated and slender (
|
||
<figureCitation id="13212A753243102BFCF9F845DBB03A83" box="[848,1017,1936,1963]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="31" pageNumber="32">Figure 10G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300657B32431014FF6EF860DBE93CA3" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="discussion">
|
||
<paragraph id="8BA536F032431014FF6EF860DBAD3DD3" blockId="31.[151,1437,1288,2035]" lastBlockId="32.[151,1437,153,503]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
|
||
<emphasis id="B96EEAE23243102BFF6EF860D9723AE7" bold="true" box="[199,315,1973,1999]" pageId="31" pageNumber="32">Remarks.</emphasis>
|
||
<bibRefCitation id="EF8B4B013243102BFEEBF860D9B63AE7" author="Lourenco, W. R." box="[322,511,1973,1999]" pageId="31" pageNumber="32" pagination="449 - 461" refId="ref49925" refString="Lourenco, W. R. (2000) Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34 (3), 449 - 461. https: // doi. org / 10.1080 / 00222930029956 1" type="journal article" year="2000">Lourenço (2000)</bibRefCitation>
|
||
provided records for
|
||
<taxonomicName id="4C1A4D733243102BFD59F860DBC93AE6" authorityName="Lourenco" authorityYear="1999" box="[752,896,1972,1998]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="erikae">
|
||
<emphasis id="B96EEAE23243102BFD59F860DBC93AE6" box="[752,896,1972,1998]" italics="true" pageId="31" pageNumber="32">Tityus erikae</emphasis>
|
||
</taxonomicName>
|
||
from Vereda Miralindo, Medina,
|
||
<collectingRegion id="49DEF8123243102BFB50F861D95C3ADA" country="Colombia" name="Cundinamarca" pageId="31" pageNumber="32">Cundinamarca department</collectingRegion>
|
||
(ICN-As-194) and Santuario de Flora y Fauna Los Colorados,
|
||
<collectingRegion id="49DEF8123243102BFC79F80DDCE13ADA" box="[976,1192,2008,2034]" country="Colombia" name="Bolivar" pageId="31" pageNumber="32">Bolívar department</collectingRegion>
|
||
(ICN-As-188). These specimens were examined and we consider that they correspond to adult males of
|
||
<taxonomicName id="4C1A4D73327C1014FBE5FF4FDD613D9B" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[1100,1320,153,179]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE2327C1014FBE5FF4FDD613D9B" box="[1100,1320,153,179]" italics="true" pageId="32" pageNumber="33">Tityus columbianus</emphasis>
|
||
</taxonomicName>
|
||
. Also, we examined two adult males, seven adult females and one juvenile female from Reserva Natural La Planada,
|
||
<collectingRegion id="49DEF812327C1014FAE7FF68D95C3DD3" country="Colombia" name="Narino" pageId="32" pageNumber="33">Nariño department</collectingRegion>
|
||
(ICN-As-712); and they also correspond to
|
||
<taxonomicName id="4C1A4D73327C1014FCADFF37DB943DD3" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[772,989,225,251]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE2327C1014FCADFF37DB943DD3" box="[772,989,225,251]" italics="true" pageId="32" pageNumber="33">Tityus columbianus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327C1014FF6EFED0DBE93CA3" blockId="32.[151,1437,153,503]" pageId="32" pageNumber="33">
|
||
The records from Santuario de Flora y Fauna Los Colorados,
|
||
<collectingRegion id="49DEF812327C1014FCD1FED0DB873C37" box="[888,974,261,287]" country="Venezuela" name="Bolivar" pageId="32" pageNumber="33">Bolívar</collectingRegion>
|
||
, and Reserva Natural La Planada,
|
||
<collectingRegion id="49DEF812327C1014FAE7FED0DDD53C37" box="[1358,1436,261,287]" country="Colombia" name="Narino" pageId="32" pageNumber="33">Nariño</collectingRegion>
|
||
are unusually far away from the known distribution range of
|
||
<taxonomicName id="4C1A4D73327C1014FCE4FEFFDBB03C6B" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[845,1017,297,323]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE2327C1014FCE4FEFFDBB03C6B" box="[845,1017,297,323]" italics="true" pageId="32" pageNumber="33">T. columbianus</emphasis>
|
||
</taxonomicName>
|
||
which is in the Eastern Cordillera in
|
||
<collectingRegion id="49DEF812327C1014FF3EFE98D9733C4F" box="[151,314,333,359]" country="Colombia" name="Cundinamarca" pageId="32" pageNumber="33">Cundinamarca</collectingRegion>
|
||
and
|
||
<collectingRegion id="49DEF812327C1014FEC6FE98D98A3C4F" box="[367,451,333,359]" country="Colombia" name="Boyaca" pageId="32" pageNumber="33">Boyacá</collectingRegion>
|
||
departments. However, only new collections from these localities could explain whether these records are due to mislabeling errors or introduced populations.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300657B327C1011FF6EFE41DA8738CE" lastPageId="37" lastPageNumber="38" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="8BA536F0327C1014FF6EFE41D9E93CFB" blockId="32.[151,1437,153,503]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96EEAE2327C1014FF6EFE41D93C3C87" bold="true" box="[199,373,404,431]" pageId="32" pageNumber="33">Redescription.</emphasis>
|
||
Based on an adult female (ICN-As-739) and an adult male (ICN-As-524). For female live habitus see
|
||
<figureCitation id="13212A75327C1014FEB3FE6CD9D53CFB" box="[282,412,441,467]" captionStart="FIGURE 46" captionStartId="77.[151,250,1653,1678]" captionTargetBox="[151,1436,202,1599]" captionTargetId="figure@77.[151,1436,202,1599]" captionTargetPageId="77" captionText="FIGURE 46. Tityus (Archaeotityus) Lourenço, 2006, habitus in life. A. Tityus bastosi Lourenço, 1984, female from El Carmen, Villavicencio, Meta, Colombia. B. Tityus columbianus (Thorell, 1876), female from Soacha, Cundinamarca, Colombia. C. Tityus guane sp. nov., male from Vereda Pasochico, Finca La Ceiba, Los Santos, Santander. D, E. Tityus parvulus Kraepelin, 1914, females. D. Female from Regional Natural Reserve El Vínculo [Natural Regional Park El Vínculo], Buga, Valle del Cauca, Colombia. E. Female from the Botanical Garden Maria Cespedes [Jardín Botánico Maria Cespedes], Tulúa, Valle del Cauca, Colombia. F. Tityus tayrona Lourenço, 1991, female from National Natural Park Tayrona [Parque Nacional Natural Tayrona], Santa Marta, Magdalena, Colombia. Photo B, courtesy of N. Herreño." httpUri="https://zenodo.org/record/3377229/files/figure.png" pageId="32" pageNumber="33">Figure 46B</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327C1014FF6EFE08DAB23CDF" blockId="32.[151,1437,153,503]" box="[199,763,476,503]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96EEAE2327C1014FF6EFE08D91D3CDF" box="[199,340,477,503]" italics="true" pageId="32" pageNumber="33">Total length.</emphasis>
|
||
Female:
|
||
<quantity id="4CE29B15327C1014FE15FE09DA783CDF" box="[444,561,476,503]" metricMagnitude="-2" metricUnit="m" metricValue="3.522" pageId="32" pageNumber="33" unit="mm" value="35.22">35.22 mm</quantity>
|
||
; male:
|
||
<quantity id="4CE29B15327C1014FD2BFE08DABE3CDF" box="[642,759,477,503]" metricMagnitude="-2" metricUnit="m" metricValue="4.029" pageId="32" pageNumber="33" unit="mm" value="40.29">40.29 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF656678327C1014FF3EFA1BDD1D3B07" httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1486,1511]" targetBox="[151,1436,525,1467]" targetPageId="32">
|
||
<paragraph id="8BA536F0327C1014FF3EFA1BDD1D3B07" blockId="32.[151,1437,1486,1583]" pageId="32" pageNumber="33">
|
||
<emphasis id="B96EEAE2327C1014FF3EFA1BD96838CF" bold="true" box="[151,289,1486,1511]" pageId="32" pageNumber="33">FIGURE 20.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FE8EFA05DAF838CF" authority="Lourenco, 2006" authorityName="Lourenco" authorityYear="2006" box="[295,689,1486,1511]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE2327C1014FE8EFA05DA4E38CF" box="[295,519,1487,1511]" italics="true" pageId="32" pageNumber="33">Tityus (Archaeotityus)</emphasis>
|
||
<bibRefCitation id="EF8B4B01327C1014FDA7FA05DAF838CF" author="Lourenco, W. R. & Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F." box="[526,689,1486,1511]" pageId="32" pageNumber="33" pagination="117 - 120" refId="ref50272" refString="Lourenco, W. R., Jesus-Junior, M. M. B. & Limeira-De-Oliveira, F. (2006) A new species of Tityus C. L. Koch, 1836 (Scorpiones, Buthidae) from the state of Maranh " o in Brazil. Boletin Sociedad Entomologica Aragonesa, 38, 117 - 120." type="journal article" year="2006">Lourenço, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, male
|
||
<taxonomicName id="4C1A4D73327C1014FD5DFA1BDB7838CF" box="[756,817,1486,1511]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="32" pageNumber="84" phylum="Arthropoda" rank="genus">telson</taxonomicName>
|
||
sculpturing, lateral aspect.
|
||
<emphasis id="B96EEAE2327C1014FBE3FA1BDC2B38CF" bold="true" box="[1098,1122,1486,1511]" pageId="32" pageNumber="33">A.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FBC0FA05DDD438CF" authority="Lourenco, 1984" authorityName="Lourenco" authorityYear="1984" box="[1129,1437,1487,1511]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bastosi">
|
||
<emphasis id="B96EEAE2327C1014FBC0FA05DCBC38CF" box="[1129,1269,1487,1511]" italics="true" pageId="32" pageNumber="33">Tityus bastosi</emphasis>
|
||
<bibRefCitation id="EF8B4B01327C1014FB52FA05DDD438CF" author="Lourenco, W. R." box="[1275,1437,1487,1511]" pageId="32" pageNumber="33" pagination="349 - 360" refId="ref49629" refString="Lourenco, W. R. (1984) Analyse taxonomique des Scorpions du groupe Tityus clathratus Koch, 1845 (Scorpiones, Buthidae). Bulletin Museum National d' Histoire Naturelle, Paris, 4 e series, 6, 349 - 360." type="journal article" year="1984">Lourenço, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-741).
|
||
<emphasis id="B96EEAE2327C1014FE9CFA26D9053B23" bold="true" box="[309,332,1523,1547]" pageId="32" pageNumber="33">B.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FEFDFA21DAF73B23" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[340,702,1522,1547]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="columbianus">
|
||
<emphasis id="B96EEAE2327C1014FEFDFA21DA503B23" box="[340,537,1523,1547]" italics="true" pageId="32" pageNumber="33">Tityus columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B01327C1014FD80FA27DAFE3B23" author="Thorell, T." box="[553,695,1522,1547]" pageId="32" pageNumber="33" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(ICN-As-524).
|
||
<emphasis id="B96EEAE2327C1014FCCAFA27DB353B23" bold="true" box="[867,892,1522,1547]" pageId="32" pageNumber="33">C.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FC2DFA21DC4F3B23" authority="Moreno-González & O & D, 2019" authorityName="Moreno-González & O & D" authorityYear="2019" box="[900,1030,1524,1547]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="guane" status="sp. nov.">
|
||
<emphasis id="B96EEAE2327C1014FC2DFA21DC4F3B23" box="[900,1030,1524,1547]" italics="true" pageId="32" pageNumber="33">Tityus guane</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25D5799327C1014FBA7FA26DC293B23" box="[1038,1120,1523,1547]" pageId="32" pageNumber="33" rank="species">
|
||
<emphasis id="B96EEAE2327C1014FBA7FA26DC293B23" bold="true" box="[1038,1120,1523,1547]" pageId="32" pageNumber="33">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype) (ICN-As-1003).
|
||
<emphasis id="B96EEAE2327C1014FA2DFA26DDD53B23" bold="true" box="[1412,1436,1523,1547]" pageId="32" pageNumber="33">D.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FF3EF9CDD9943B07" authority="Kraepelin, 1914" authorityName="Kraepelin" authorityYear="1914" box="[151,477,1558,1583]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="B96EEAE2327C1014FF3EF9CDD97A3B07" box="[151,307,1559,1583]" italics="true" pageId="32" pageNumber="33">Tityus parvulus</emphasis>
|
||
<bibRefCitation id="EF8B4B01327C1014FE90F9C3D9943B07" author="Kraepelin, K." box="[313,477,1558,1583]" pageId="32" pageNumber="33" pagination="15 - 28" refId="ref49461" refString="Kraepelin, K. (1914) Beitrag zur Kenntnis der Skorpione und Pedipalpen Columbiens. Memoires de la Societe des Sciences naturelles, Neuchatel, 5, 15 - 28." type="journal article" year="1914">Kraepelin, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-1004).
|
||
<emphasis id="B96EEAE2327C1014FD22F9C2DAEB3B07" bold="true" box="[651,674,1559,1583]" pageId="32" pageNumber="33">E.</emphasis>
|
||
<taxonomicName id="4C1A4D73327C1014FD00F9CDDBA83B07" authority="Lourenco, 1991" authorityName="Lourenco" authorityYear="1991" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[681,993,1559,1583]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="tayrona">
|
||
<emphasis id="B96EEAE2327C1014FD00F9CDDB733B07" box="[681,826,1560,1583]" italics="true" pageId="32" pageNumber="33">Tityus tayrona</emphasis>
|
||
<bibRefCitation id="EF8B4B01327C1014FCE9F9CDDBA83B07" author="Lourenco, W. R." box="[832,993,1559,1583]" pageId="32" pageNumber="33" pagination="275 - 288" refId="ref49674" refString="Lourenco, W. R. (1991) Les scorpions de Colombie, II. Les faunes des regions de Santa Marta et de la. Cordillere Orientale. Approche biogeographique. Senckenbergiana Biologica, 71 (4), 275 - 288." type="journal article" year="1991">Lourenço, 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
(ICN-As-181). Scale bars= 1.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA536F0327C1015FF6EF988DC5E3F33" blockId="32.[151,1437,1629,2016]" lastBlockId="33.[151,1437,153,2015]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
|
||
<emphasis id="B96EEAE2327C1014FF6EF988D9063B50" bold="true" box="[199,335,1629,1656]" pageId="32" pageNumber="33">Coloration.</emphasis>
|
||
General body coloration (in ethanol 70%) (
|
||
<figureCitation id="13212A75327C1014FC93F988DBE33B50" box="[826,938,1629,1656]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="32" pageNumber="33">Figure 21</figureCitation>
|
||
) with a dark yellow background moderately covered with dark brown variegated spots.
|
||
<emphasis id="B96EEAE2327C1014FDD9F957DA973BB4" box="[624,734,1666,1692]" italics="true" pageId="32" pageNumber="33">Carapace</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327C1014FD42F954DB343BB4" box="[747,893,1665,1692]" captionStart="FIGURE 5" captionStartId="13.[151,250,1940,1965]" captionTargetBox="[199,1390,186,1902]" captionTargetId="figure@13.[179,1409,166,1921]" captionTargetPageId="13" captionText="FIGURE 5. Tityus (Archaeotityus) Lourenço, 2006, carapace coloration, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377145/files/figure.png" pageId="32" pageNumber="33">Figure 5C, D</figureCitation>
|
||
) moderately covered with dark brown variegated spots, except for the anterior margin that is dark yellow; lateral and median eyes surrounded by black spots; median ocular tubercle surrounded by a lateral black stripe on each side; posterior area to the median ocular tubercle with an inverted ‘’Y’’-shaped spot.
|
||
<emphasis id="B96EEAE2327C1014FE45F93BDA2E3A20" box="[492,615,1774,1800]" italics="true" pageId="32" pageNumber="33">Chelicerae</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327C1014FDDFF938DB423A20" box="[630,779,1773,1800]" captionStart="FIGURE 7" captionStartId="15.[151,250,1942,1967]" captionTargetBox="[181,1406,194,1913]" captionTargetId="figure@15.[172,1415,166,1922]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Archaeotityus) Lourenço, 2006, chelicerae, dorsal aspect, showing reticulation pattern. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377149/files/figure.png" pageId="32" pageNumber="33">Figure 7C, D</figureCitation>
|
||
) coxa and hand with a light yellow background; hand with dark brown reticulated spots on the anterior half and a small areas without spots on the adjacent area to the movable finger articulation; movable and fixed fingers with dark brown spots along their lengths; teeth dark reddish-brown.
|
||
<emphasis id="B96EEAE2327C1014FF3EF88FD94E3A5C" box="[151,263,1882,1908]" italics="true" pageId="32" pageNumber="33">Pedipalps</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327C1014FEBEF88CD9C13A5C" box="[279,392,1881,1908]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="32" pageNumber="33">Figure 21</figureCitation>
|
||
) moderately covered with dark brown variegated spots and yellow spots; trochanter, femur, and
|
||
<taxonomicName id="4C1A4D73327C1014FF6CF8A8D9593ABF" box="[197,272,1917,1943]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="32" pageNumber="71" phylum="Mollusca" rank="genus">patella</taxonomicName>
|
||
ventrally yellow; trichobothrial pits yellow.
|
||
<emphasis id="B96EEAE2327C1014FD52F8ABDB793ABF" box="[763,816,1918,1943]" italics="true" pageId="32" pageNumber="33">Legs</emphasis>
|
||
With all the segments having a light yellow background moderately covered with dark brown variegated spots on their prolateral surfaces.
|
||
<emphasis id="B96EEAE2327C1014FB86F877DD4E3A93" box="[1071,1287,1954,1980]" italics="true" pageId="32" pageNumber="33">Coxosternal region</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327C1014FABCF874D88F3AC8" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="32" pageNumber="33">Figure 21B, E, F</figureCitation>
|
||
) coxae I–IV, sternum, genital operculum, and pectines (teeth) light yellow; basal pectinal piece with an anterior white area (female); coxaphophyses I–II, pectines (marginal lamellae, middle lamellae, fulcra), and basal pectinal piece with a light yellow background moderately covered with dark brown variegated spots.
|
||
<taxonomicName id="4C1A4D73327D1015FB61FF6BDD093DFF" box="[1224,1344,190,215]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96EEAE2327D1015FB61FF6BDD093DFF" box="[1224,1344,190,215]" italics="true" pageId="33" pageNumber="34">Mesosoma</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A75327D1015FAF8FF68D8FD3DD3" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="33" pageNumber="34">Figure 21</figureCitation>
|
||
) tergites I–VII each with a dark yellow background moderately covered with dark brown variegated spots and with four posterior dark brown spots (two submedian and two lateral); sternites III–VII each densely covered with dark reddish-brown variegated spots; spiracles yellow.
|
||
<emphasis id="B96EEAE2327D1015FCA2FEFFDBC93C6B" box="[779,896,298,323]" italics="true" pageId="33" pageNumber="34">Metasoma</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327D1015FC39FEFCDC6E3C6B" box="[912,1063,297,323]" captionStart="FIGURE 6" captionStartId="14.[151,250,1837,1862]" captionTargetBox="[151,1415,186,1776]" captionTargetId="figure@14.[151,1436,181,1776]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377147/files/figure.png" pageId="33" pageNumber="34">Figure 6B, E</figureCitation>
|
||
) with a dark yellow background moderately covered with dark brown variegated spots and with ventral yellow spots; each segment progressively darker towards segment V; segment V completely dark reddish-brown with three ventral black stripes and with a dorsomedian area without spots (
|
||
<figureCitation id="13212A75327D1015FDADFE40DADE3C87" box="[516,663,405,431]" captionStart="FIGURE 6" captionStartId="14.[151,250,1837,1862]" captionTargetBox="[151,1415,186,1776]" captionTargetId="figure@14.[151,1436,181,1776]" captionTargetPageId="14" captionText="FIGURE 6. Tityus (Archaeotityus) Lourenço, 2006, metasoma coloration pattern, dorsal (left) and ventral (right). A, D. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. D. Male. B, E. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). B. Female. E. Male. C, F. Tityus guane sp. nov. (ICN-As-1003). C. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377147/files/figure.png" pageId="33" pageNumber="34">Figure 6B, E</figureCitation>
|
||
). DL intercarinal areas of segments I–IV each with an anterior median arrow-shaped brown spot and a posterior median brown spot; ML, VSM, and VM intercarinal areas of segments I–IV each with a pair of distal brown spots.
|
||
<taxonomicName id="4C1A4D73327D1015FD29FE08DA813CDF" box="[640,712,477,503]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B96EEAE2327D1015FD29FE08DA813CDF" box="[640,712,477,503]" italics="true" pageId="33" pageNumber="34">Telson</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A75327D1015FD7EFE08DB0C3CDF" box="[727,837,477,503]" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="33" pageNumber="34">Figure 21</figureCitation>
|
||
) almost completely dark reddish-brown with a lateral light reddish-brown stripe; subaculear tubercle and aculeus dark reddish-brown.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFDF0DC5A3FDC" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFDF0D9283F17" bold="true" box="[199,353,549,575]" pageId="33" pageNumber="34">Morphology.</emphasis>
|
||
<emphasis id="B96EEAE2327D1015FECEFDF0D99C3F17" box="[359,469,549,575]" italics="true" pageId="33" pageNumber="34">Carapace</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327D1015FE4DFDF0DA333F17" box="[484,634,549,575]" captionStart="FIGURE 8" captionStartId="16.[151,250,1940,1965]" captionTargetBox="[195,1393,166,1921]" captionTargetId="figure@16.[195,1393,166,1921]" captionTargetPageId="16" captionText="FIGURE 8. Tityus (Archaeotityus) Lourenço, 2006, carapace sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984. A. Female (ICN-As-740). B. Male (ICN-As-741). C, D. Tityus columbianus (Thorell, 1876). C. Female (ICN-As-739). D. Male (ICN-As-524). E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype. Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377151/files/figure.png" pageId="33" pageNumber="34">Figure 8C, D</figureCitation>
|
||
) densely covered with coarse granules and a few fine granules; anterior margin with a shallow median notch; carinae (anterior median, lateral, and posterior median) and furrows (anterior, lateral, median ocular, posterior median, posterior marginal, and posterior lateral) well-marked (female) or feebly marked (male); median ocular tubercle well-marked and located on the anterior half of the carapace; median eyes separated by about 1.6 (female) and 1.3 (male) ocular diameters. Lateral eye pattern
|
||
<typeStatus id="54A18852327D1015FBFDFD60DCCD3FE7" box="[1108,1156,693,719]" pageId="33" pageNumber="34">type</typeStatus>
|
||
4A: three pairs of major ocelli present (PLMa, MLMa and
|
||
<taxonomicName id="4C1A4D73327D1015FDBFFD0CDA2A3FDB" authorityName="Grube" authorityYear="1855" box="[534,611,729,755]" class="Clitellata" family="Glossoscolecidae" genus="Alma" kingdom="Animalia" order="Haplotaxida" pageId="33" pageNumber="6" phylum="Annelida" rank="genus">ALMa</taxonomicName>
|
||
) and one pair of minor ocelli (ADMi).
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFD28DB7B3E13" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFD28D90B3E3F" box="[199,322,765,791]" italics="true" pageId="33" pageNumber="34">Chelicerae</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327D1015FEFDFD28D9B93E3F" box="[340,496,765,791]" captionStart="FIGURE 7" captionStartId="15.[151,250,1942,1967]" captionTargetBox="[181,1406,194,1913]" captionTargetId="figure@15.[172,1415,166,1922]" captionTargetPageId="15" captionText="FIGURE 7. Tityus (Archaeotityus) Lourenço, 2006, chelicerae, dorsal aspect, showing reticulation pattern. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740, 741). A. Female. B. Male. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739, 524). C. Female. D. Male. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Female paratype. F. Male holotype." httpUri="https://zenodo.org/record/3377149/files/figure.png" pageId="33" pageNumber="34">Figure 7C, D</figureCitation>
|
||
) dentition characteristic of the family
|
||
<taxonomicName id="4C1A4D73327D1015FC18FD28DC853E30" authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[945,1228,765,792]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="33" pageNumber="34" phylum="Arthropoda" rank="family">
|
||
Buthidae (
|
||
<bibRefCitation id="EF8B4B01327D1015FB81FD28DC8D3E30" author="Vachon, M." box="[1064,1220,765,792]" pageId="33" pageNumber="34" pagination="161 - 166" refId="ref52210" refString="Vachon, M. (1963) De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin Museum National d' Histoire Naturelle, Paris, 2 e series, 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
; hand and fingers densely covered with setae on internal and ventral surfaces.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFC90DD5039EF" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFC90D9763E77" box="[199,319,837,863]" italics="true" pageId="33" pageNumber="34">Pedipalps.</emphasis>
|
||
<taxonomicName id="4C1A4D73327D1015FEE3FC90D9C23E77" box="[330,395,837,863]" class="Actinopterygii" family="Cyprinidae" genus="Chela" kingdom="Animalia" order="Cypriniformes" pageId="33" pageNumber="76" phylum="Chordata" rank="genus">Chela</taxonomicName>
|
||
short and of a medium-width (female) (Cl/Cw= 4.9) or incrassate (male) (Cl/Cw= 3.1). Trichobothriotaxic pattern
|
||
<typeStatus id="54A18852327D1015FE62FCBCDA4A3EAB" box="[459,515,873,899]" pageId="33" pageNumber="34">Type</typeStatus>
|
||
A with alfa configuration (hand: Eb3:Eb2:Eb1:Esb:Est:Et, fixed finger: eb:esb: est:et:db:dt:it) (e.g.,
|
||
<figureCitation id="13212A75327D1015FED4FC58D9973E8F" box="[381,478,909,935]" captionStart="FIGURE 3" captionStartId="11.[151,250,1912,1937]" captionTargetBox="[302,1285,166,1892]" captionTargetId="figure@11.[302,1285,166,1892]" captionTargetPageId="11" captionText="FIGURE 3. Tityus (Archaeotityus) Lourenço, 2006, right pedipalp, showing trichobothrial pattern. A–C. Pedipalp chela. A. External aspect. B. Dorsal aspect. C. Ventral aspect. D. Femur, dorsal aspect. E. Patella, dorsal aspect. Abbreviations: d, dorsal; db, dorsal basal; dt, dorsal terminal; e, external; eb, Eb, external basal; esb, Esb, external subbasal; est, Est, external subterminal; et, Et, external terminal; i, internal; it, internal terminal; v, ventral. Scale bars= A–C= 2.0 mm; D, E= 1.0 mm." httpUri="https://zenodo.org/record/3377140/files/figure.png" pageId="33" pageNumber="34">Figure 3</figureCitation>
|
||
).
|
||
<emphasis id="B96EEAE2327D1015FE5DFC5BDA763E8F" box="[500,575,910,935]" italics="true" pageId="33" pageNumber="34">Femur</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327D1015FDE7FC58DA873E8F" box="[590,718,909,935]" captionStart="FIGURE 9" captionStartId="17.[151,250,1761,1786]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure@17.[151,1436,181,1741]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377153/files/figure.png" pageId="33" pageNumber="34">Figures 9C</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327D1015FD72FC58DB423E8F" box="[731,779,909,935]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="33" pageNumber="34">10C</figureCitation>
|
||
) with five well-marked and complete carinae: VI, DI, and DE crenulate; VE and IM serratocrenulate; intercarinal areas moderately covered with coarse granules and densely covered with fine granules (
|
||
<figureCitation id="13212A75327D1015FE7BFC00DA1B3EC7" box="[466,594,981,1007]" captionStart="FIGURE 9" captionStartId="17.[151,250,1761,1786]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure@17.[151,1436,181,1741]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377153/files/figure.png" pageId="33" pageNumber="34">Figures 9C</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327D1015FDF7FC00DAD83EC7" box="[606,657,981,1007]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="33" pageNumber="34">10C</figureCitation>
|
||
).
|
||
<taxonomicName id="4C1A4D73327D1015FD0CFC00DABF3EC7" box="[677,758,981,1007]" class="Gastropoda" family="Patellidae" genus="Patella" kingdom="Animalia" order="Pezizales" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B96EEAE2327D1015FD0CFC00DABF3EC7" box="[677,758,981,1007]" italics="true" pageId="33" pageNumber="34">Patella</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A75327D1015FCACFC00DBCF3EC7" box="[773,902,981,1007]" captionStart="FIGURE 9" captionStartId="17.[151,250,1761,1786]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure@17.[151,1436,181,1741]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377153/files/figure.png" pageId="33" pageNumber="34">Figures 9D</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327D1015FC3AFC00DB8C3EC7" box="[915,965,981,1007]" captionStart="FIGURE 10" captionStartId="20.[151,250,1754,1779]" captionTargetBox="[151,1436,181,1734]" captionTargetId="figure@20.[151,1436,181,1734]" captionTargetPageId="20" captionText="FIGURE 10. Tityus (Archaeotityus) Lourenço, 2006, male pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-741). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN-As- 1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377155/files/figure.png" pageId="33" pageNumber="34">10D</figureCitation>
|
||
) with seven carinae: VI, VE, DI, DE, and EM complete and crenulate; DM incomplete and crenulate; IM complete and serratocrenulate with a spur near the segment base; intercarinal areas densely covered with fine granules.
|
||
<taxonomicName id="4C1A4D73327D1015FC0EFBC8DC77391F" authority="(Tibia)" baseAuthorityName="Tibia" box="[935,1086,1053,1079]" class="Actinopterygii" family="Cyprinidae" genus="Chela" kingdom="Animalia" order="Cypriniformes" pageId="33" pageNumber="34" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96EEAE2327D1015FC0EFBC8DBA3391F" box="[935,1002,1053,1079]" italics="true" pageId="33" pageNumber="34">Chela</emphasis>
|
||
(
|
||
<emphasis id="B96EEAE2327D1015FC55FBC8DC7F391F" box="[1020,1078,1053,1079]" italics="true" pageId="33" pageNumber="34">Tibia</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13212A75327D1015FBE6FBC8DC8C391F" box="[1103,1221,1053,1079]" captionStart="FIGURE 11" captionStartId="21.[151,250,1944,1969]" captionTargetBox="[272,1316,166,1924]" captionTargetId="figure@21.[272,1316,166,1924]" captionTargetPageId="21" captionText="FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. A–C. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377157/files/figure.png" pageId="33" pageNumber="34">Figures 11</figureCitation>
|
||
D–F, 22A–C) with eight carinae: VI, VE, D, DS, DMA, and ES complete and crenulate; DI complete and feebly crenulate (female) (
|
||
<figureCitation id="13212A75327D1015FF36FBB0D9403957" box="[159,265,1125,1151]" captionStart="FIGURE 11" captionStartId="21.[151,250,1944,1969]" captionTargetBox="[272,1316,166,1924]" captionTargetId="figure@21.[272,1316,166,1924]" captionTargetPageId="21" captionText="FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. A–C. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377157/files/figure.png" pageId="33" pageNumber="34">Figure 11</figureCitation>
|
||
D–F) or obsolete (male) (
|
||
<figureCitation id="13212A75327D1015FD80FBB0DAD13957" box="[553,664,1125,1151]" captionStart="FIGURE 22" captionStartId="36.[151,250,1928,1953]" captionTargetBox="[316,1271,166,1909]" captionTargetId="figure@36.[316,1271,166,1909]" captionTargetPageId="36" captionText="FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. A–C. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="33" pageNumber="34">Figure 22</figureCitation>
|
||
A–C); SA incomplete and crenulate, only present on the anterior half of the hand. Pedipalp movable and fixed fingers with an obsolete basal lobe (male) (
|
||
<figureCitation id="13212A75327D1015FBEAFB5CDC8F398B" box="[1091,1222,1161,1187]" captionStart="FIGURE 22" captionStartId="36.[151,250,1928,1953]" captionTargetBox="[316,1271,166,1909]" captionTargetId="figure@36.[316,1271,166,1909]" captionTargetPageId="36" captionText="FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. A–C. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="33" pageNumber="34">Figure 22A</figureCitation>
|
||
) or without a basal lobe (female) (
|
||
<figureCitation id="13212A75327D1015FE95FB78D98939EF" box="[316,448,1197,1223]" captionStart="FIGURE 11" captionStartId="21.[151,250,1944,1969]" captionTargetBox="[272,1316,166,1924]" captionTargetId="figure@21.[272,1316,166,1924]" captionTargetPageId="21" captionText="FIGURE 11. Tityus (Archaeotityus) Lourenço, 2006, right female pedipalp sculpturing. A–C. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Tityus columbianus (Thorell, 1876) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377157/files/figure.png" pageId="33" pageNumber="34">Figure 11D</figureCitation>
|
||
); dorsal surface of the movable finger with 13–13 oblique rows of denticles.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFB04DB6B381B" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFB04D9E139C3" box="[199,424,1233,1259]" italics="true" pageId="33" pageNumber="34">Coxosternal region.</emphasis>
|
||
Covered with fine granules and a few setae, except for the coxapophyses I–II which are smooth. Sternum with posterior depression, outer ridge, and apical button well-marked. Genital operculum longitudinally divided and composed of two subtriangular plates.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFAEBD92038CF" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFAEBD96F387F" box="[199,294,1342,1367]" italics="true" pageId="33" pageNumber="34">Pectines</emphasis>
|
||
. Basal piece sexually dimorphic (e.g.,
|
||
<figureCitation id="13212A75327D1015FD5BFAE8DB1C387F" box="[754,853,1341,1367]" captionStart="FIGURE 2" captionStartId="10.[151,250,1977,2002]" captionTargetBox="[236,1348,184,1943]" captionTargetId="figure@10.[219,1368,166,1958]" captionTargetPageId="10" captionText="FIGURE 2. Tityus (Archaeotityus) guane sp. nov., sternum and pectines, showing the shape of the basal pectinal piece. A. Female paratype (ICN-As-1003). B. Male holotype (ICN-As-1003). Abbreviations: A, anterior region; P, posterior region." httpUri="https://zenodo.org/record/3377138/files/figure.png" pageId="33" pageNumber="34">Figure 2</figureCitation>
|
||
) sub rectangular-shaped with an anterior elevated glandular area (female) (e.g.,
|
||
<figureCitation id="13212A75327D1015FE4EFAB4DA153853" box="[487,604,1377,1403]" captionStart="FIGURE 2" captionStartId="10.[151,250,1977,2002]" captionTargetBox="[236,1348,184,1943]" captionTargetId="figure@10.[219,1368,166,1958]" captionTargetPageId="10" captionText="FIGURE 2. Tityus (Archaeotityus) guane sp. nov., sternum and pectines, showing the shape of the basal pectinal piece. A. Female paratype (ICN-As-1003). B. Male holotype (ICN-As-1003). Abbreviations: A, anterior region; P, posterior region." httpUri="https://zenodo.org/record/3377138/files/figure.png" pageId="33" pageNumber="34">Figure 2A</figureCitation>
|
||
) or flat, shield-shaped with an anterior median notch well-marked (male) (e.g.,
|
||
<figureCitation id="13212A75327D1015FF71FA50D90438B7" box="[216,333,1413,1439]" captionStart="FIGURE 2" captionStartId="10.[151,250,1977,2002]" captionTargetBox="[236,1348,184,1943]" captionTargetId="figure@10.[219,1368,166,1958]" captionTargetPageId="10" captionText="FIGURE 2. Tityus (Archaeotityus) guane sp. nov., sternum and pectines, showing the shape of the basal pectinal piece. A. Female paratype (ICN-As-1003). B. Male holotype (ICN-As-1003). Abbreviations: A, anterior region; P, posterior region." httpUri="https://zenodo.org/record/3377138/files/figure.png" pageId="33" pageNumber="34">Figure 2B</figureCitation>
|
||
); pectinal tooth counts: 13–14 (female) and 14–14 (male). Intermediate plate, marginal plate, and fulcra moderately covered with setae (
|
||
<figureCitation id="13212A75327D1015FDE5FA7CDA9838EB" box="[588,721,1449,1475]" captionStart="FIGURE 14" captionStartId="24.[151,250,1868,1893]" captionTargetBox="[199,1386,187,1829]" captionTargetId="figure@24.[182,1404,166,1848]" captionTargetPageId="24" captionText="FIGURE 14. Tityus (Archaeotityus) Lourenço, 2006, left pectine with peg sensillae details, ventral aspect. A–D. Tityus bastosi Lourenço, 1984.A, B. Female (MZSP 71740). A. Pectine. B. Peg sensillae shape. C, D. Male (MZSP 71740). C. Pectine. D. Peg sensillae shape. E, F. Tityus columbianus (Thorell, 1876) (ICN-As-739). E. Female pectine. F. Female peg sensillae shape. G–J. Tityus guane sp. nov.. G, H. Female (MZSP 71736). G. Pectine. H. Peg sensillae shape. I, J. Male (MZSP 71736). I. Pectine. J. Peg sensillae shape. Scale bars= A, C, E, G, I= 400 µm; B, D, F, H, J= 10 µm." httpUri="https://zenodo.org/record/3377163/files/figure.png" pageId="33" pageNumber="34">Figure 14E</figureCitation>
|
||
). Pectinal teeth peg sensillae flattened with a slightly spatulate apex (
|
||
<figureCitation id="13212A75327D1015FF72FA18D91538CF" box="[219,348,1485,1511]" captionStart="FIGURE 14" captionStartId="24.[151,250,1868,1893]" captionTargetBox="[199,1386,187,1829]" captionTargetId="figure@24.[182,1404,166,1848]" captionTargetPageId="24" captionText="FIGURE 14. Tityus (Archaeotityus) Lourenço, 2006, left pectine with peg sensillae details, ventral aspect. A–D. Tityus bastosi Lourenço, 1984.A, B. Female (MZSP 71740). A. Pectine. B. Peg sensillae shape. C, D. Male (MZSP 71740). C. Pectine. D. Peg sensillae shape. E, F. Tityus columbianus (Thorell, 1876) (ICN-As-739). E. Female pectine. F. Female peg sensillae shape. G–J. Tityus guane sp. nov.. G, H. Female (MZSP 71736). G. Pectine. H. Peg sensillae shape. I, J. Male (MZSP 71736). I. Pectine. J. Peg sensillae shape. Scale bars= A, C, E, G, I= 400 µm; B, D, F, H, J= 10 µm." httpUri="https://zenodo.org/record/3377163/files/figure.png" pageId="33" pageNumber="34">Figure 14F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EFA27DA363B07" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EFA27D8B53B23" box="[199,252,1522,1547]" italics="true" pageId="33" pageNumber="34">Legs</emphasis>
|
||
. Carinae present; intercarinal areas with a few fine granules; telotarsus covered by numerous ventrosubmedian setae; claws short and symmetrical.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1015FF6EF9EFD9703A5B" blockId="33.[151,1437,153,2015]" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EF9EFD90E3B7B" box="[199,327,1594,1619]" italics="true" pageId="33" pageNumber="34">
|
||
<taxonomicName id="4C1A4D73327D1015FF6EF9EFD90B3B7B" box="[199,322,1594,1619]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Tergites I–VI densely covered with coarse granules and a few fine granules; pre-tergite well-marked with the median carina visible on the posterior margin of the post-tergite; tergite VII with DSM and DL carinae complete and crenulate; median carina of the tergite VII composed of an anterior median crenulate eminence located on the anterior half of the post-tergite. Sternites densely covered with coarse granules and a few fine granules; sternites III–VI each with a longitudinal median hyaline suture and a pair of elliptic spiracles on the posterior half; spiracles progressively larger towards sternite VI; posterior margin of the sternite V with a subtriangular glandular area; sternite VI with VSM carinae crenulate occupying the posterior half of the sternite; sternite VII with VSM carinae crenulate occupying the posterior 2/3 of the sternite and VL carinae crenulate occupying the anterior half of the sternite.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327D1016FF6EF8A8DA663C6B" blockId="33.[151,1437,153,2015]" lastBlockId="34.[151,1437,153,2015]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
<emphasis id="B96EEAE2327D1015FF6EF8A8D9FD3ABF" box="[199,436,1917,1943]" italics="true" pageId="33" pageNumber="34">Hemispermatophore.</emphasis>
|
||
General morphological characteristics as in other
|
||
<taxonomicName id="4C1A4D73327D1015FC4CF8ABDC923ABF" authority="" authorityName="Lourenco" authorityYear="2006" box="[997,1243,1917,1943]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE2327D1015FC4CF8ABDC613ABF" box="[997,1064,1918,1943]" italics="true" pageId="33" pageNumber="34">Tityus</emphasis>
|
||
(
|
||
<emphasis id="B96EEAE2327D1015FB9EF8A8DC983ABF" box="[1079,1233,1917,1943]" italics="true" pageId="33" pageNumber="34">Archaeotityus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
species.
|
||
<emphasis id="B96EEAE2327D1015FAE9F8A8DDD53ABF" box="[1344,1436,1917,1943]" italics="true" pageId="33" pageNumber="34">Capsule</emphasis>
|
||
(
|
||
<figureCitation id="13212A75327D1015FF36F874D9443A93" box="[159,269,1953,1979]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="33" pageNumber="34">Figure 23</figureCitation>
|
||
A–C) with a straight distal carina ending in a 70° angle; sperm duct reduced not overpassing the basal carina level and with a translucent area between the base of the basal hook and the base of the basal carina; trans- lucent area basally wide but progressively narrower towards the distal region (
|
||
<figureCitation id="13212A75327E1016FBB3FF4CDC8A3D9B" box="[1050,1219,153,179]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23B, C</figureCitation>
|
||
); basal carina with a rounded apex (
|
||
<figureCitation id="13212A75327E1016FEFFFF68D9953DFF" box="[342,476,189,215]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23C</figureCitation>
|
||
); basal hook semi ovate-shaped with anterior margin straight in lateral view (
|
||
<figureCitation id="13212A75327E1016FAF8FF68D8803DD3" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23B</figureCitation>
|
||
), and slightly curved in anterior and posterior views; basal hook forming a ‘’U’’-shaped curve with the stem (
|
||
<figureCitation id="13212A75327E1016FF36FED0D90E3C37" box="[159,327,261,287]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23A, C</figureCitation>
|
||
); basal hook elongated with a thick and rounded apex and approximately 1.2 longer than its basal width in anterior view (
|
||
<figureCitation id="13212A75327E1016FE37FEFCDA6B3C6B" box="[414,546,297,323]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327E1016FF6EFE9BDC513FDC" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96EEAE2327E1016FF6EFE9BD90A3C4F" box="[199,323,334,359]" italics="true" pageId="34" pageNumber="35">Metasoma.</emphasis>
|
||
Segments II–V short and of a medium-width in both sexes (Msl/Msw ratio= female/ male: II=1.4/ 1.5; III= 1.6/ 1.7; IV= 1.8/ 1.9; V= 2.0/ 1.7). Segment V short and of a medium-width (female) (
|
||
<figureCitation id="13212A75327E1016FB59FEA4DD3A3CA3" box="[1264,1395,368,395]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="34" pageNumber="35">Figure 15B</figureCitation>
|
||
) or incrassate (male) (
|
||
<figureCitation id="13212A75327E1016FECDFE40D9A03C98" box="[356,489,405,432]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="34" pageNumber="35">Figure 16C</figureCitation>
|
||
). Segments I–II (
|
||
<figureCitation id="13212A75327E1016FD00FE40DB7E3C87" box="[681,823,404,431]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="34" pageNumber="35">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FCEAFE40DB3A3C98" box="[835,883,405,432]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="34" pageNumber="35">16C</figureCitation>
|
||
) with 10 complete, parallel and crenulate carinae (paired DL, ML, LIM, VL, and VSM; DL carinae serratocrenulate on segment II); LIM on segment II composed of coarse granules almost along all its length; intercarinal areas densely covered with coarse granules. Segment III–IV (
|
||
<figureCitation id="13212A75327E1016FF36FDD4D9653F33" box="[159,300,512,539]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="34" pageNumber="35">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FE9EFDD4D92E3F34" box="[311,359,513,540]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="34" pageNumber="35">16C</figureCitation>
|
||
) with eight complete, parallel, and crenulate carinae (paired DL, ML, VL, and VSM; DL carinae serratocrenulate); intercarinal areas densely covered with coarse granules. Segment V with five complete and crenulate carinae (VM, paired DL, and VL); lateral and ventral intercarinal areas densely covered with coarse granules and a few fine granules (
|
||
<figureCitation id="13212A75327E1016FE01FDB8DA7C3FAF" box="[424,565,620,647]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="34" pageNumber="35">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FDE8FDB8DA3A3FA0" box="[577,627,621,648]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="34" pageNumber="35">16C</figureCitation>
|
||
); intercarinal areas with granules irregularly distributed on the middle of each area. Segment II–IV with DL carinae composed of granules which slightly increase in size towards the distal region of each segment; DL carinae ending in a distoterminal acute and small granule (II–III) or a blunt and coarse granule (IV) which increase in size towards the segment IV (
|
||
<figureCitation id="13212A75327E1016FCE9FD0CDB843FDB" box="[832,973,728,755]" captionStart="FIGURE 15" captionStartId="25.[151,250,1936,1961]" captionTargetBox="[283,1305,166,1916]" captionTargetId="figure@25.[283,1305,166,1916]" captionTargetPageId="25" captionText="FIGURE 15. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segments I–V sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377165/files/figure.png" pageId="34" pageNumber="35">Figures 15B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FC70FD0CDC423FDC" box="[985,1035,729,756]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="34" pageNumber="35">16C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327E1016FF6EFD28D90D391F" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96EEAE2327E1016FF6EFD28D9BB3E3F" box="[199,498,765,791]" italics="true" pageId="34" pageNumber="35">Metasomal macrosetation.</emphasis>
|
||
Segments I–IV each with two pairs of VSM macrosetae (2/2) and two pairs of VL macrosetae (2/2); pair VSM1 is located in the anterior third, and pair VSM2 is located near the posterior margin of the segment; pair VL1 is located near the anterior margin of the segment, and pair VL2 is located on the posterior third of the segment (e.g,
|
||
<figureCitation id="13212A75327E1016FE1DFCBCDA583EAB" box="[436,529,873,899]" captionStart="FIGURE 1" captionStartId="9.[151,250,1939,1964]" captionTargetBox="[243,1344,166,1920]" captionTargetId="figure@9.[243,1344,166,1920]" captionTargetPageId="9" captionText="FIGURE 1. Tityus (Archaeotityus) Lourenço, 2006, metasomal segments I–V showing distribution of macrosetae.A. Segments I–V, ventral aspect. B. Segment V, lateral aspect. Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian." httpUri="https://zenodo.org/record/3377136/files/figure.png" pageId="34" pageNumber="35">Figure 1</figureCitation>
|
||
). Segment V (
|
||
<figureCitation id="13212A75327E1016FD1BFCBCDB763EAB" box="[690,831,873,899]" captionStart="FIGURE 17" captionStartId="27.[151,250,1559,1584]" captionTargetBox="[151,1436,181,1539]" captionTargetId="figure@27.[151,1436,181,1539]" captionTargetPageId="27" captionText="FIGURE 17. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377169/files/figure.png" pageId="34" pageNumber="35">Figures 17B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FCE3FCBCDB303EAB" box="[842,889,873,899]" captionStart="FIGURE 18" captionStartId="30.[151,250,1899,1924]" captionTargetBox="[151,1422,525,1880]" captionTargetId="figure@30.[151,1422,525,1880]" captionTargetPageId="30" captionText="FIGURE 18. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As- 181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377171/files/figure.png" pageId="34" pageNumber="35">18B</figureCitation>
|
||
) with three pairs of VSM macrosetae (3/3), three pairs of VL macrosetae (3/3), and a single pair of DL macrosetae; pair VSM1 is located near the anterior margin of the segment; pair VSM2 is located distal or at same level of pair VL2 (
|
||
<figureCitation id="13212A75327E1016FC0DFC64DC783EE3" box="[932,1073,945,971]" captionStart="FIGURE 17" captionStartId="27.[151,250,1559,1584]" captionTargetBox="[151,1436,181,1539]" captionTargetId="figure@27.[151,1436,181,1539]" captionTargetPageId="27" captionText="FIGURE 17. Tityus (Archaeotityus) Lourenço, 2006, female metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377169/files/figure.png" pageId="34" pageNumber="35">Figures 17B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FB95FC64DC243EE3" box="[1084,1133,945,971]" captionStart="FIGURE 18" captionStartId="30.[151,250,1899,1924]" captionTargetBox="[151,1422,525,1880]" captionTargetId="figure@30.[151,1422,525,1880]" captionTargetPageId="30" captionText="FIGURE 18. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segment V sculpturing, showing macrosetae, ventral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As- 181). Abbreviations: ML, median lateral; VL, ventrolateral; VSM, ventrosubmedian. Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377171/files/figure.png" pageId="34" pageNumber="35">18B</figureCitation>
|
||
), and pair VSM3 is located on the anal arch; pair VL1 is located near the anterior margin of the segment; pair VL2 is located on the posterior second third of the segment, and pair VL3 is located on the anal arch; pair DL1 is located near the posterior margin of the segment.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327E1016FF6EFB94DD77387F" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96EEAE2327E1016FF6EFB94D95E3973" box="[199,279,1089,1115]" italics="true" pageId="34" pageNumber="35">
|
||
<taxonomicName id="4C1A4D73327E1016FF6EFB94D95A3973" box="[199,275,1089,1115]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
Vesicle not elongated and suboval (Vl/Vh= 2.0) (
|
||
<figureCitation id="13212A75327E1016FCFCFB94DBAF3973" box="[853,998,1089,1115]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="34" pageNumber="35">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FC5AFB94DC6A3973" box="[1011,1059,1089,1115]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="34" pageNumber="35">20B</figureCitation>
|
||
) with dorsal surface smooth and with a lateral longitudinal furrow on each side; VM, paired VSM, VL, and DL carinae composed of coarse granules; carinae well-marked (female) (
|
||
<figureCitation id="13212A75327E1016FD86FB5CDAF9398B" box="[559,688,1161,1187]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="34" pageNumber="35">Figure 19B</figureCitation>
|
||
) or feebly marked (male) (
|
||
<figureCitation id="13212A75327E1016FC49FB5CDC2C398B" box="[992,1125,1161,1187]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="34" pageNumber="35">Figure 20B</figureCitation>
|
||
); paired DL carinae located dorsally with respect to the lateral longitudinal furrow; VM carina well-marked along its length (
|
||
<figureCitation id="13212A75327E1016FB6FFB78DD1A39EF" box="[1222,1363,1197,1223]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="34" pageNumber="35">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FAF4FB78DDD939EF" box="[1373,1424,1197,1223]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="34" pageNumber="35">20B</figureCitation>
|
||
). Subaculear tubercle medium-sized pyramidal, flat, and triangle-like with an acute and coarse apex in lateral view (
|
||
<figureCitation id="13212A75327E1016FF36FB20D9653827" box="[159,300,1269,1295]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="34" pageNumber="35">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FE9FFB20D9203827" box="[310,361,1269,1295]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="34" pageNumber="35">20B</figureCitation>
|
||
); dorsal margin of the subaculear tubercle with a pair of ventrodistal small rounded granules with apices pointing to the middle portion of the aculeus; VM carina composed of coarse granules and strongly crenulate in lateral view (
|
||
<figureCitation id="13212A75327E1016FEEFFAE8D99D387F" box="[326,468,1341,1367]" captionStart="FIGURE 19" captionStartId="31.[151,250,1145,1170]" captionTargetBox="[151,1436,181,1126]" captionTargetId="figure@31.[151,1436,181,1126]" captionTargetPageId="31" captionText="FIGURE 19. Tityus (Archaeotityus) Lourenço, 2006, female telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-740). B. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Tityus guane sp. nov. (paratype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377173/files/figure.png" pageId="34" pageNumber="35">Figures 19B</figureCitation>
|
||
,
|
||
<figureCitation id="13212A75327E1016FE76FAE8DA5B387F" box="[479,530,1341,1367]" captionStart="FIGURE 20" captionStartId="32.[151,250,1486,1511]" captionTargetBox="[151,1436,525,1467]" captionTargetId="figure@32.[151,1436,525,1467]" captionTargetPageId="32" captionText="FIGURE 20. Tityus (Archaeotityus) Lourenço, 2006, male telson sculpturing, lateral aspect. A. Tityus bastosi Lourenço, 1984 (ICN-As-741). B. Tityus columbianus (Thorell, 1876) (ICN-As-524). C. Tityus guane sp. nov. (holotype) (ICN-As-1003). D. Tityus parvulus Kraepelin, 1914 (ICN-As-1004). E. Tityus tayrona Lourenço, 1991 (ICN-As-181). Scale bars= 1.5 mm." httpUri="https://zenodo.org/record/3377175/files/figure.png" pageId="34" pageNumber="35">20B</figureCitation>
|
||
); aculeus strongly curved shorter than vesicle and with a ventral groove.
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327E1016FF6EFAB5DD5B3A67" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96EEAE2327E1016FF6EFAB5D9043853" bold="true" box="[199,333,1376,1403]" pageId="34" pageNumber="35">Variability.</emphasis>
|
||
<emphasis id="B96EEAE2327E1016FEFDFAB4D9933853" box="[340,474,1377,1403]" italics="true" pageId="34" pageNumber="35">Total length</emphasis>
|
||
(
|
||
<emphasis id="B96EEAE2327E1016FE42FAB4DAD63853" box="[491,671,1377,1403]" italics="true" pageId="34" pageNumber="35">
|
||
including
|
||
<taxonomicName id="4C1A4D73327E1016FDF5FAB4DAD63853" box="[604,671,1377,1403]" class="Maxillopoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Poecilostomatoida" pageId="34" pageNumber="84" phylum="Arthropoda" rank="genus">telson</taxonomicName>
|
||
</emphasis>
|
||
): males
|
||
<quantity id="4CE29B15327E1016FCA9FAB4DBF73853" box="[768,958,1377,1403]" metricMagnitude="-2" metricUnit="m" metricValue="3.3634999999999997" metricValueMax="3.911" metricValueMin="2.816" pageId="34" pageNumber="35" unit="mm" value="33.635" valueMax="39.11" valueMin="28.16">28.16–39.11 mm</quantity>
|
||
(n= 14; mean= 31.83; SD= 3.91); females
|
||
<quantity id="4CE29B15327E1016FF3EFA51D91138B7" box="[151,344,1412,1439]" metricMagnitude="-2" metricUnit="m" metricValue="3.159" metricValueMax="3.754" metricValueMin="2.564" pageId="34" pageNumber="35" unit="mm" value="31.59" valueMax="37.54" valueMin="25.64">25.64–37.54 mm</quantity>
|
||
(n= 57; mean= 33.46; SD= 2.08) (
|
||
<tableCitation id="C698034B327E1016FD41FA50DB0938B7" box="[744,832,1413,1439]" captionStart="TABLE 9" captionStartId="85.[151,239,488,514]" captionTargetBox="[155,2014,561,1666]" captionText="TABLE 9. Measurements (mm) of Tityus (Archaeotityus) bastosi Lourenço, 1991, and Tityus (Archaeotityus) columbianus (Thorell, 1876). Abbreviations: F, females; M, males; SD, standard deviation." httpUri="http://table.plazi.org/id/DF65667832091061FF3EFE3DD9D93F0E" pageId="34" pageNumber="35" tableUuid="DF65667832091061FF3EFE3DD9D93F0E">Table 9</tableCitation>
|
||
).
|
||
<emphasis id="B96EEAE2327E1016FCFEFA50DC0038B7" box="[855,1097,1413,1439]" italics="true" pageId="34" pageNumber="35">Pectinal tooth counts</emphasis>
|
||
: males 13–15 (n= 42; mode= 14); females 11–15 (n= 208; mode= 12) (
|
||
<tableCitation id="C698034B327E1016FDD2FA7CDA9F38EB" box="[635,726,1449,1475]" captionStart="TABLE 2" captionStartId="8.[151,239,478,502]" captionText="TABLE 2. Pectinal tooth counts of Colombian Tityus (Archaeotityus) species. Note: Both pectines of each examined specimen were counted. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="34" pageNumber="35">Table 2</tableCitation>
|
||
).
|
||
<emphasis id="B96EEAE2327E1016FD44FA7CDD0038EB" box="[749,1353,1449,1475]" italics="true" pageId="34" pageNumber="35">Number of movable finger oblique rows of denticles</emphasis>
|
||
: males 12–14 (n= 39; mode= 13); females 11–13 (n= 203; mode= 13) (
|
||
<tableCitation id="C698034B327E1016FCC2FA18DB8A38CF" box="[875,963,1485,1511]" captionStart="TABLE 3" captionStartId="28.[151,239,730,754]" captionText="TABLE 3. Number of movable finger oblique rows of denticles among Colombian Tityus (Archaeotityus) species. Note: Both movable fingers of each examined specimen were included. Abbreviations: F, female; M, male; Juv, juvenile; SD, standard deviation." pageId="34" pageNumber="35">Table 3</tableCitation>
|
||
).
|
||
<emphasis id="B96EEAE2327E1016FC70FA18DD6638CF" box="[985,1327,1485,1511]" italics="true" pageId="34" pageNumber="35">Metasomal macrosetae counts</emphasis>
|
||
: segment I: males (n= 21): 2/2 (n= 10), 2/3 (n= 1), 3/2 (n= 2), 3/3 (n= 7), 3/4 (n= 1) VSM macrosetae on the segment; segments II–IV: males (n= 21) 2/2 VSM on each segment; segment V: males (n= 21): 3/3 VSM and 3/3 VL macrosetae on the segment. Segment I: females (n= 116): 2/2 (n= 57), 2/3 (n= 9), 3/2 (n= 2), 3/3 (n= 46), 3/4 (n= 1), 4/2 (n= 1) VSM macrosetae on the segment; segment II–III: females (n= 116): 2/2 (n= 115), 2/3 (n= 1) VSM macrosetae on each segment; segment IV: females (n= 116): 2/2 (n= 106), 3/2 (n= 2), 3/3 (n= 4) VSM macrosetae on the segment; segment V: females (n= 116): 3/2 (n= 1), 3/3 (n= 114), 3/4 (n= 1) VSM macrosetae; 3/3 VL macrosetae, and 1/1 DL macrosetae on the segment. Segments I–IV: males (n= 21) and females (n= 116): 2/2 VL macrosetae on each segment.
|
||
<emphasis id="B96EEAE2327E1016FF75F938DBA73A2F" box="[220,1006,1773,1799]" italics="true" pageId="34" pageNumber="35">Relative position of the VSM2 macrosetae on the metasomal segment V</emphasis>
|
||
: males (n= 21): distal to VL2 (n= 7) or at the same level of VL2 (n= 14); females (n= 116): distal to VL2 (n= 27); at the same level of VL2 (n= 78); slightly proximal to the level of VL2 (n= 1) or asymmetric with just one seta at the same level of VL2 (n= 10).
|
||
</paragraph>
|
||
<paragraph id="8BA536F0327E1016FF6EF88CD9783AF7" blockId="34.[151,1437,153,2015]" pageId="34" pageNumber="35">
|
||
<emphasis id="B96EEAE2327E1016FF6EF88CDA443A5B" box="[199,525,1881,1907]" italics="true" pageId="34" pageNumber="35">Hemispermatophore capsule</emphasis>
|
||
: the basal hook of the capsule exhibited different shapes in anterior view: (1) elongated with a thick and rounded apex, curved lateral margin, and forming ‘’U’’-shaped curve with the stem (
|
||
<figureCitation id="13212A75327E1016FAC5F8A8D8BA3A93" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23C</figureCitation>
|
||
) and (2) short with an acute apex, curved lateral margin, and forming a ‘’U’’-shaped curve with the stem (
|
||
<figureCitation id="13212A75327E1016FF36F810D96C3AF7" box="[159,293,1989,2015]" captionStart="FIGURE 23" captionStartId="37.[151,250,981,1006]" captionTargetBox="[163,1426,200,952]" captionTargetId="figure@37.[151,1436,181,961]" captionTargetPageId="37" captionText="FIGURE 23. Tityus (Archaeotityus) columbianus (Thorell, 1876), dextral hemispermatophore, showing capsule details, male (ICN-As-739). A. Posterior view. B. Lateral view. C. Anterior view. D. Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524). Abbreviations: Bc, basal carina; Bh, basal hook; Dc, distal carina; Sdm, sperm duct membrane. Scale bar= 1.0 mm." httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="34" pageNumber="35">Figure 23D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF656678327F1017FF3EF84CDC473AD2" httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1945,1970]" targetBox="[366,1227,214,1917]" targetPageId="35">
|
||
<paragraph id="8BA536F0327F1017FF3EF84CDC473AD2" blockId="35.[151,1436,1945,2043]" pageId="35" pageNumber="36">
|
||
<emphasis id="B96EEAE2327F1017FF3EF84CD9683A9A" bold="true" box="[151,289,1945,1970]" pageId="35" pageNumber="36">FIGURE 21.</emphasis>
|
||
<taxonomicName id="4C1A4D73327F1017FE80F84EDB703A9A" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[297,825,1945,1970]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="columbianus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE2327F1017FE80F84EDADA3A9A" box="[297,659,1946,1970]" italics="true" pageId="35" pageNumber="36">Tityus (Archaeotityus) columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B01327F1017FD0BF84CDB7B3A9A" author="Thorell, T." box="[674,818,1945,1970]" pageId="35" pageNumber="36" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, habitus.
|
||
<emphasis id="B96EEAE2327F1017FC35F84CDB9D3A9A" bold="true" box="[924,980,1945,1970]" pageId="35" pageNumber="36">A, B.</emphasis>
|
||
Female (ICN-As-739).
|
||
<emphasis id="B96EEAE2327F1017FB65F84CDCAC3A9A" bold="true" box="[1228,1253,1945,1970]" pageId="35" pageNumber="36">A.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96EEAE2327F1017FA2CF84FDDD53A9A" bold="true" box="[1413,1436,1946,1970]" pageId="35" pageNumber="36">B.</emphasis>
|
||
Ventral aspect.
|
||
<emphasis id="B96EEAE2327F1017FE9CF868D9253AFE" bold="true" box="[309,364,1981,2006]" pageId="35" pageNumber="36">C, E.</emphasis>
|
||
Male (Villa de Leyva, Boyacá) (ICN-As-739).
|
||
<emphasis id="B96EEAE2327F1017FCFEF868DB393AFE" bold="true" box="[855,880,1981,2006]" pageId="35" pageNumber="36">C.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96EEAE2327F1017FBB9F86BDC6E3AFE" bold="true" box="[1040,1063,1982,2006]" pageId="35" pageNumber="36">E.</emphasis>
|
||
Ventral aspect.
|
||
<emphasis id="B96EEAE2327F1017FB65F86BDCB63AFE" bold="true" box="[1228,1279,1982,2006]" pageId="35" pageNumber="36">D, F.</emphasis>
|
||
Male (Villa de Leyva, Boyacá) (ICN-As-524).
|
||
<emphasis id="B96EEAE2327F1017FE73F837D9BA3AD2" bold="true" box="[474,499,2018,2042]" pageId="35" pageNumber="36">D.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96EEAE2327F1017FD26F837DAEB3AD2" bold="true" box="[655,674,2018,2042]" pageId="35" pageNumber="36">F.</emphasis>
|
||
Ventral aspect. Scale bars= 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF65667832781010FF3EF85DDB303AC0" httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,250,1928,1953]" targetBox="[316,1271,166,1909]" targetPageId="36">
|
||
<paragraph id="8BA536F032781010FF3EF85DDB303AC0" blockId="36.[151,1436,1928,2025]" pageId="36" pageNumber="37">
|
||
<emphasis id="B96EEAE232781010FF3EF85DD9683A89" bold="true" box="[151,289,1928,1953]" pageId="36" pageNumber="37">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C1A4D7332781010FE80F85FDB713A89" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[297,824,1928,1953]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="columbianus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE232781010FE80F85FDADB3A89" box="[297,658,1929,1953]" italics="true" pageId="36" pageNumber="37">Tityus (Archaeotityus) columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B0132781010FD08F85DDB783A89" author="Thorell, T." box="[673,817,1928,1953]" pageId="36" pageNumber="37" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, male pedipalp
|
||
<taxonomicName id="4C1A4D7332781010FC75F85DDC5C3A89" box="[988,1045,1928,1953]" class="Actinopterygii" family="Cyprinidae" genus="Chela" kingdom="Animalia" order="Cypriniformes" pageId="36" pageNumber="76" phylum="Chordata" rank="genus">chela</taxonomicName>
|
||
.
|
||
<emphasis id="B96EEAE232781010FBB6F85DDC1E3A89" bold="true" box="[1055,1111,1928,1953]" pageId="36" pageNumber="37">A–C.</emphasis>
|
||
Male (Villa de Leyva, Boyacá) (ICN-As-524).
|
||
<emphasis id="B96EEAE232781010FE9DF879D9043AED" bold="true" box="[308,333,1964,1989]" pageId="36" pageNumber="37">A.</emphasis>
|
||
External aspect.
|
||
<emphasis id="B96EEAE232781010FE55F878DA5A3AED" bold="true" box="[508,531,1965,1989]" pageId="36" pageNumber="37">B.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96EEAE232781010FD18F879DA803AED" bold="true" box="[689,713,1964,1989]" pageId="36" pageNumber="37">C.</emphasis>
|
||
Ventral aspect.
|
||
<emphasis id="B96EEAE232781010FCC5F878DBD73AED" bold="true" box="[876,926,1965,1989]" pageId="36" pageNumber="37">D–F.</emphasis>
|
||
Male (Villa de Leyva, Boyacá) (ICN-As-739).
|
||
<emphasis id="B96EEAE232781010FA2DF878DDD53AED" bold="true" box="[1412,1436,1965,1989]" pageId="36" pageNumber="37">D.</emphasis>
|
||
External aspect.
|
||
<emphasis id="B96EEAE232781010FE96F804D91F3AC1" bold="true" box="[319,342,2001,2025]" pageId="36" pageNumber="37">E.</emphasis>
|
||
Dorsal aspect.
|
||
<emphasis id="B96EEAE232781010FE5AF804DA4F3AC1" bold="true" box="[499,518,2001,2025]" pageId="36" pageNumber="37">F.</emphasis>
|
||
Ventral aspect. Scale bars= 2.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF65667832791011FF3EFC00D9683971" httpUri="https://zenodo.org/record/3377181/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,981,1006]" targetBox="[163,1426,200,952]" targetPageId="37">
|
||
<paragraph id="8BA536F032791011FF3EFC00D9683971" blockId="37.[151,1436,981,1114]" pageId="37" pageNumber="38">
|
||
<emphasis id="B96EEAE232791011FF3EFC00D9683EC6" bold="true" box="[151,289,981,1006]" pageId="37" pageNumber="38">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C1A4D7332791011FE80FC03DB713EC6" authority="(Thorell, 1876)" authorityName="Thorell" authorityYear="1876" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[297,824,981,1006]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="columbianus" subGenus="Archaeotityus">
|
||
<emphasis id="B96EEAE232791011FE80FC03DADB3EC5" box="[297,658,981,1005]" italics="true" pageId="37" pageNumber="38">Tityus (Archaeotityus) columbianus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B0132791011FD08FC00DB783EC6" author="Thorell, T." box="[673,817,981,1006]" pageId="37" pageNumber="38" pagination="1 - 14" refId="ref52173" refString="Thorell, T. (1876) On the classification of scorpions. Annals and Magazine of Natural History, 97, 1 - 14. https: // doi. org / 10.1080 / 0022293760868188 9" type="journal article" year="1876">Thorell, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, dextral hemispermatophore, showing capsule details, male (ICN-As-739).
|
||
<emphasis id="B96EEAE232791011FE9DFC2CD905393A" bold="true" box="[308,332,1017,1042]" pageId="37" pageNumber="38">A.</emphasis>
|
||
Posterior view.
|
||
<emphasis id="B96EEAE232791011FE58FC2CDA413939" bold="true" box="[497,520,1017,1041]" pageId="37" pageNumber="38">B.</emphasis>
|
||
Lateral view.
|
||
<emphasis id="B96EEAE232791011FD33FC2CDAFB393A" bold="true" box="[666,690,1017,1042]" pageId="37" pageNumber="38">C.</emphasis>
|
||
Anterior view.
|
||
<emphasis id="B96EEAE232791011FCF9FC2CDB203939" bold="true" box="[848,873,1017,1041]" pageId="37" pageNumber="38">D.</emphasis>
|
||
Capsule variation, anterior view, male (Villa de Leyva, Boyacá) (ICN-As-524).
|
||
<emphasis id="B96EEAE232791011FE38FBC8DA57391D" box="[401,542,1053,1077]" italics="true" pageId="37" pageNumber="38">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B96EEAE232791011FD85FBCBDA0E391D" box="[556,583,1054,1077]" italics="true" pageId="37" pageNumber="38">Bc</emphasis>
|
||
, basal carina;
|
||
<emphasis id="B96EEAE232791011FD74FBC8DAB0391D" box="[733,761,1053,1077]" italics="true" pageId="37" pageNumber="38">Bh</emphasis>
|
||
, basal hook;
|
||
<emphasis id="B96EEAE232791011FC2DFBCBDBEB391D" box="[900,930,1054,1077]" italics="true" pageId="37" pageNumber="38">Dc</emphasis>
|
||
, distal carina;
|
||
<emphasis id="B96EEAE232791011FB93FBC8DC2F391D" box="[1082,1126,1053,1077]" italics="true" pageId="37" pageNumber="38">Sdm</emphasis>
|
||
, sperm duct membrane. Scale bar= 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA536F032791011FF6EFB5DDA8738CE" blockId="37.[151,1437,1159,1726]" pageId="37" pageNumber="38">
|
||
<emphasis id="B96EEAE232791011FF6EFB5DDA493989" box="[199,512,1160,1186]" italics="true" pageId="37" pageNumber="38">Male intrasexual variations</emphasis>
|
||
: adult males exhibited different body sizes ranging between
|
||
<quantity id="4CE29B1532791011FB05FB52DD25398A" box="[1196,1388,1159,1186]" metricMagnitude="-2" metricUnit="m" metricValue="3.3634999999999997" metricValueMax="3.911" metricValueMin="2.816" pageId="37" pageNumber="38" unit="mm" value="33.635" valueMax="39.11" valueMin="28.16">28.16–39.11 mm</quantity>
|
||
, but despite this they did not show discrete size-classes, but rather a progressive increase in their total lengths (
|
||
<figureCitation id="13212A7532791011FAF8FB79D8F839C2" captionStart="FIGURE 21" captionStartId="35.[151,250,1945,1970]" captionTargetBox="[366,1227,214,1917]" captionTargetId="figure@35.[353,1234,166,1926]" captionTargetPageId="35" captionText="FIGURE 21. Tityus (Archaeotityus) columbianus (Thorell, 1876), habitus. A, B. Female (ICN-As-739). A. Dorsal aspect. B. Ventral aspect. C, E. Male (Villa de Leyva, Boyacá) (ICN-As-739). C. Dorsal aspect. E. Ventral aspect. D, F. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Dorsal aspect. F. Ventral aspect. Scale bars= 10 mm." httpUri="https://zenodo.org/record/3377177/files/figure.png" pageId="37" pageNumber="38">Figure 21</figureCitation>
|
||
C–F,
|
||
<tableCitation id="C698034B32791011FF5BFB05D90339C2" box="[242,330,1232,1258]" captionStart="TABLE 9" captionStartId="85.[151,239,488,514]" captionTargetBox="[155,2014,561,1666]" captionText="TABLE 9. Measurements (mm) of Tityus (Archaeotityus) bastosi Lourenço, 1991, and Tityus (Archaeotityus) columbianus (Thorell, 1876). Abbreviations: F, females; M, males; SD, standard deviation." httpUri="http://table.plazi.org/id/DF65667832091061FF3EFE3DD9D93F0E" pageId="37" pageNumber="38" tableUuid="DF65667832091061FF3EFE3DD9D93F0E">Table 9</tableCitation>
|
||
). However, as the total body size increases among males the following: (1)
|
||
<emphasis id="B96EEAE232791011FB6EFB05D8853825" italics="true" pageId="37" pageNumber="38">Carapace and tergites</emphasis>
|
||
: the number of coarse granules on intercarinal surfaces increases; (2)
|
||
<emphasis id="B96EEAE232791011FC4BFB21DCD93826" box="[994,1168,1268,1294]" italics="true" pageId="37" pageNumber="38">Pedipalp femur</emphasis>
|
||
: intercarinal areas have coarser and finest granules (
|
||
<figureCitation id="13212A7532791011FE70FACDDA18381A" box="[473,593,1304,1330]" captionStart="FIGURE 9" captionStartId="17.[151,250,1761,1786]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure@17.[151,1436,181,1741]" captionTargetPageId="17" captionText="FIGURE 9. Tityus (Archaeotityus) Lourenço, 2006, female pedipalp femur and patella sculpturing, dorsal aspect. A, B. Tityus bastosi Lourenço, 1984 (ICN-As-740). A. Femur. B. Patella. C, D. Tityus columbianus (Thorell, 1876) (ICN-As-739). C. Femur. D. Patella. E, F. Tityus guane sp. nov. (ICN-As-1003). E. Femur. F. Patella. G, H. Tityus parvulus Kraepelin, 1914 (ICN- As-1004). G. Femur. H. Patella. I, J. Tityus tayrona Lourenço, 1991 (ICN-As-181). I. Femur. J. Patella. Scale bars= 1.0 mm." httpUri="https://zenodo.org/record/3377153/files/figure.png" pageId="37" pageNumber="38">Figure 9C</figureCitation>
|
||
); (3)
|
||
<taxonomicName id="4C1A4D7332791011FD38FACDDA9D381A" box="[657,724,1304,1330]" class="Actinopterygii" family="Cyprinidae" genus="Chela" kingdom="Animalia" order="Cypriniformes" pageId="37" pageNumber="38" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96EEAE232791011FD38FACDDA9D381A" box="[657,724,1304,1330]" italics="true" pageId="37" pageNumber="38">Chela</emphasis>
|
||
</taxonomicName>
|
||
: the DI carina is more obsolete (
|
||
<figureCitation id="13212A7532791011FBF9FACDDC88381A" box="[1104,1217,1304,1330]" captionStart="FIGURE 22" captionStartId="36.[151,250,1928,1953]" captionTargetBox="[316,1271,166,1909]" captionTargetId="figure@36.[316,1271,166,1909]" captionTargetPageId="36" captionText="FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. A–C. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="37" pageNumber="38">Figure 22</figureCitation>
|
||
A–F); the manus is more incrassate without apparently exhibiting isometric growth (n= 14; Cl/Cw= 3.1–4.7; Cl/Ch= 3.0–4.0) (
|
||
<figureCitation id="13212A7532791011FAF8FAE9D8FD3852" captionStart="FIGURE 22" captionStartId="36.[151,250,1928,1953]" captionTargetBox="[316,1271,166,1909]" captionTargetId="figure@36.[316,1271,166,1909]" captionTargetPageId="36" captionText="FIGURE 22. Tityus (Archaeotityus) columbianus (Thorell, 1876), male pedipalp chela. A–C. Male (Villa de Leyva, Boyacá) (ICN-As-524). A. External aspect. B. Dorsal aspect. C. Ventral aspect. D–F. Male (Villa de Leyva, Boyacá) (ICN-As-739). D. External aspect. E. Dorsal aspect. F. Ventral aspect. Scale bars= 2.0 mm." httpUri="https://zenodo.org/record/3377179/files/figure.png" pageId="37" pageNumber="38">Figure 22</figureCitation>
|
||
A–F); (4)
|
||
<emphasis id="B96EEAE232791011FE8DFAB5DA063851" box="[292,591,1376,1402]" italics="true" pageId="37" pageNumber="38">Metasomal segments II–IV</emphasis>
|
||
: the DL carinae exhibit coarser distoterminal granules (
|
||
<figureCitation id="13212A7532791011FB13FAB5DD163852" box="[1210,1375,1375,1402]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="37" pageNumber="38">Figure 16C, D</figureCitation>
|
||
); and (5)
|
||
<emphasis id="B96EEAE232791011FF69FA51D9F238B5" box="[192,443,1412,1438]" italics="true" pageId="37" pageNumber="38">Metasomal segment V</emphasis>
|
||
: segment is more incrassate without apparently exhibiting an isometric growth (n= 14; Msl-V/Msw-V= 1.8–2.1; Msl-V/Msh-V= 1.6–2.1) (
|
||
<figureCitation id="13212A7532791011FD48FA7DDBC338EA" box="[737,906,1447,1474]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="37" pageNumber="38">Figure 16C, D</figureCitation>
|
||
), and the granules associated with carinae and intercarinal areas are less marked (
|
||
<figureCitation id="13212A7532791011FDB2FA19DA8838CE" box="[539,705,1483,1510]" captionStart="FIGURE 16" captionStartId="26.[151,250,1905,1930]" captionTargetBox="[253,1334,166,1885]" captionTargetId="figure@26.[253,1334,166,1885]" captionTargetPageId="26" captionText="FIGURE 16. Tityus (Archaeotityus) Lourenço, 2006, male metasomal segments I–V sculpturing, lateral aspect. A, B. Tityus bastosi Lourenço, 1984. A. Male (Puerto López, Meta) (ICN-As-741). B. Male (San Martín, Meta) (ICN-As-738). C, D. Tityus columbianus (Thorell, 1876). C. Male (Villa de Leyva, Boyacá) (ICN-As-524). D. Male (Villa de Leyva, Boyacá) (ICN-As- 739). Scale bars= 3.0 mm." httpUri="https://zenodo.org/record/3377167/files/figure.png" pageId="37" pageNumber="38">Figure 16C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300657B32791011FF6EFA3ADBED3B96" pageId="37" pageNumber="38" type="distribution">
|
||
<paragraph id="8BA536F032791011FF6EFA3ADBED3B96" blockId="37.[151,1437,1159,1726]" pageId="37" pageNumber="38">
|
||
<emphasis id="B96EEAE232791011FF6EFA3AD9A13B22" bold="true" box="[199,488,1519,1546]" pageId="37" pageNumber="38">
|
||
Distribution (
|
||
<figureCitation id="13212A7532791011FECFFA3AD9923B22" box="[358,475,1519,1546]" captionStart="FIGURE 30" captionStartId="52.[151,250,1435,1460]" captionTargetBox="[151,1418,194,1407]" captionTargetId="figure@52.[151,1434,181,1416]" captionTargetPageId="52" captionText="FIGURE 30. Distribution of Tityus (Archaeotityus) columbianus (Thorell, 1876) and Tityus (Archaeotityus) guane sp. nov., showing known records and new locality records." httpUri="https://zenodo.org/record/3377195/files/figure.png" pageId="37" pageNumber="38">Figure 30</figureCitation>
|
||
).
|
||
</emphasis>
|
||
COLOMBIA:
|
||
<emphasis id="B96EEAE232791011FD39FA25DB203B22" box="[656,873,1520,1546]" italics="true" pageId="37" pageNumber="38">Boyacá department</emphasis>
|
||
:Arcabuco, Chiquinquirá, Chita, Cuítiva, Duitama, Iza, Firavitoba, La Uvita, Mongui, Úmbita and Villa de Leyva.
|
||
<emphasis id="B96EEAE232791011FCC4F9C1DCD43B06" box="[877,1181,1556,1582]" italics="true" pageId="37" pageNumber="38">Cundinamarca department</emphasis>
|
||
: Bogotá, Cajíca, Chocontá, Gachancipá, Guasca, Guatavita, La Calera, Mosquera, Pacho, Ubaté, and Soacha (
|
||
<figureCitation id="13212A7532791011FB2AF9EDDD603B7A" box="[1155,1321,1592,1618]" captionStart="FIGURE 47" captionStartId="78.[151,250,1588,1613]" captionTargetBox="[151,1436,181,1568]" captionTargetId="figure@78.[151,1436,181,1568]" captionTargetPageId="78" captionText="FIGURE 47. Tityus (Archaeotityus) Lourenço, 2006, habitat. A. Vereda El Carmen, Villavicencio, Meta, Colombia, habitat of Tityus bastosi Lourenço, 1984. B, C. Soacha, Cundinamarca, Colombia, habitat of Tityus columbianus (Thorell, 1876). D. Botanical Garden Maria Cespedes [Jardín Botánico Maria Cespedes], Tulúa, Valle, Colombia, habitat of Tityus parvulus Kraepelin, 1914. E. Vereda Pasochico, Finca La Ceiba, Los Santos, Santander, habitat of Tityus guane sp. nov. F. Natural National Park Tayrona [Parque Natural Nacional Tayrona], Santa Marta, Magdalena, Colombia, habitat of Tityus tayrona Lourenço, 1991. Photos B and C, courtesy of N. Herreño." httpUri="https://zenodo.org/record/3377231/files/figure.png" pageId="37" pageNumber="38">Figure 47B, C</figureCitation>
|
||
).
|
||
<emphasis id="B96EEAE232791011FA97F9EDD8963B5E" bold="true" pageId="37" pageNumber="38">New records.</emphasis>
|
||
COLOMBIA:
|
||
<emphasis id="B96EEAE232791011FE23F989DA2D3B5E" box="[394,612,1628,1654]" italics="true" pageId="37" pageNumber="38">Bolívar department</emphasis>
|
||
: Santuario de Flora y Fauna Los Colorados.
|
||
<emphasis id="B96EEAE232791011FBFEF989DD783B5E" box="[1111,1329,1628,1654]" italics="true" pageId="37" pageNumber="38">Boyacá department</emphasis>
|
||
: Susacón and Tinjáca.
|
||
<emphasis id="B96EEAE232791011FE80F955DA1E3BB2" box="[297,599,1664,1690]" italics="true" pageId="37" pageNumber="38">Cundinamarca department</emphasis>
|
||
: Carmen de Carupa, Facatativá, Medina, Nariño, Susa, and Tabio.
|
||
<emphasis id="B96EEAE232791011FAE4F955D9693B96" italics="true" pageId="37" pageNumber="38">Nariño department:</emphasis>
|
||
Natural Reserve La Planada.
|
||
<emphasis id="B96EEAE232791011FDC7F971DB0B3B96" box="[622,834,1700,1726]" italics="true" pageId="37" pageNumber="38">Tolima department</emphasis>
|
||
: Ibagué.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |