treatments-xml/data/03/B3/87/03B387C9EF7E3C1CFE960B0AFBCEF510.xml

<document id="563EB15603B28DEA059180FB276EE404" ID-DOI="10.5281/zenodo.170880" ID-GBIF-Dataset="4302257a-8b19-4c5e-ac2e-0256b9294ca1" ID-ISSN="1175­5326" ID-Zenodo-Dep="170880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459716131336" checkinUser="plazi" docAuthor="Dugdale, John S." docDate="2005" docId="03B387C9EF7E3C1CFE960B0AFBCEF510" docLanguage="en" docName="zt00870.pdf" docOrigin="Zootaxa 870" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tracholena liparodes Dugdale, 2005, sp. nov." docType="treatment" docVersion="9" lastPageNumber="9" masterDocId="FF8AFFB1EF783C14FF9E0F1EFF92F765" masterDocTitle="Three new species of Tracholena Common, 1965 (Lepidoptera: Tortricidae: Tortricinae: Schoenotenini) from New Caledonia associated with Araucariaceae" masterLastPageNumber="16" masterPageNumber="1" pageNumber="7" updateTime="1698207980742" updateUser="plazi">
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<mods:title id="FD6EC174DE2B41674BE35026FC190476">Three new species of Tracholena Common, 1965 (Lepidoptera: Tortricidae: Tortricinae: Schoenotenini) from New Caledonia associated with Araucariaceae</mods:title>
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<mods:namePart id="0250132513BC66D6A1EFC413C53736D9">Dugdale, John S.</mods:namePart>
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<mods:date id="C1B75A9B81C5A9BB35E4C31B1AF99663">2005</mods:date>
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<treatment id="03B387C9EF7E3C1CFE960B0AFBCEF510" ID-DOI="http://doi.org/10.5281/zenodo.6267711" ID-GBIF-Taxon="119334650" ID-Zenodo-Dep="6267711" LSID="urn:lsid:plazi:treatment:03B387C9EF7E3C1CFE960B0AFBCEF510" httpUri="http://treatment.plazi.org/id/03B387C9EF7E3C1CFE960B0AFBCEF510" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="C3006554EF7E3C12FE960B0AFCFFF34B" box="[264,877,1044,1070]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BA536DFEF7E3C12FE960B0AFCFFF34B" blockId="6.[264,877,1044,1070]" box="[264,877,1044,1070]" pageId="6" pageNumber="7">
<heading id="D0ED81B3EF7E3C12FE960B0AFCFFF34B" bold="true" box="[264,877,1044,1070]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B96EEACDEF7E3C12FE960B0AFCFFF34B" bold="true" box="[264,877,1044,1070]" pageId="6" pageNumber="7">
<taxonomicName id="4C1A4D5CEF7E3C12FE960B0AFD9AF34B" ID-CoL="B5G3T" box="[264,520,1044,1070]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="liparodes" status="sp. nov.">
<emphasis id="B96EEACDEF7E3C12FE960B0AFD9AF34B" bold="true" box="[264,520,1044,1070]" italics="true" pageId="6" pageNumber="7">Tracholena liparodes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57B6EF7E3C12FD8E0B0AFDFCF34B" box="[528,622,1044,1070]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13212A5AEF7E3C12FDE00B0AFD40F34B" box="[638,722,1044,1070]" captionStart="FIGURES 1 – 3" captionStartId="13.[264,383,1796,1820]" captionTargetBox="[292,1271,293,1778]" captionTargetId="figure@13.[291,1305,280,1786]" captionTargetPageId="13" captionText="FIGURES 1 – 3. Fig. 1. Tracholena nigrilinea sp. nov. Male, Ponerihouen, habitus. (LH wings omitted). Fig. 2. T. liparodes sp. nov. PT male, Mt Panié, habitus. Fig. 3. T. paniense sp. nov. PT male, Mt Panié, habitus. (Scales represent 0.5 mm)" httpUri="https://zenodo.org/record/170881/files/figure.png" pageId="6" pageNumber="7">Figs. 2</figureCitation>
, 
<figureCitation id="13212A5AEF7E3C12FD410B0AFCAAF34B" box="[735,824,1044,1070]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="6" pageNumber="7">6, 9, 10</figureCitation>
, 
<figureCitation id="13212A5AEF7E3C12FCD80B0AFCF6F34B" box="[838,868,1044,1070]" captionStart="FIGURES 11 – 15" captionStartId="15.[264,383,1360,1384]" captionTargetBox="[306,1263,328,1302]" captionTargetId="figure@15.[286,1300,326,1335]" captionTargetPageId="15" captionText="FIGURES 11 – 15. Fig. 11. T. paniense, PT male, genitalia, Mt Panié [NZAC prep. L 878]. Fig. 12. T. nigrilinea, female genitalia, Baie des Tortues [NZAC prep. 866]. Fig. 13. T. liparodes, PT female genitalia, Mt Panié [NZAC prep. L 873, 874]. Fig. 14. T. paniense PT female genitalia: ductus and corpus bursae, Mt Panié [NZAC prep. 877]. Fig. 15. T. paniense PT female genitalia: ovipositor, sterigmal and limen structure [NZAC prep. L 877]. (All scales represent 0.5 mm.)" httpUri="https://zenodo.org/record/170883/files/figure.png" pageId="6" pageNumber="7">13</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3006554EF7E3C12FE960B40FB43F27C" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BA536DFEF7E3C12FE960B40FDB5F3AC" blockId="6.[264,1323,1118,1905]" pageId="6" pageNumber="7">
<emphasis id="B96EEACDEF7E3C12FE960B40FE7CF31D" bold="true" box="[264,494,1118,1144]" italics="true" pageId="6" pageNumber="7">Material examined</emphasis>
: 
<emphasis id="B96EEACDEF7E3C12FD9E0B41FDE1F31D" box="[512,627,1119,1144]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="54A1887DEF7E3C12FD9E0B41FDFDF31D" box="[512,623,1119,1144]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male, “NOUVELLE CALEDONIE: Mt Panié, 
<quantity id="4CE29B3AEF7E3C12FB2E0B41FA99F31C" box="[1200,1291,1119,1145]" metricMagnitude="3" metricUnit="m" metricValue="1.38" pageId="6" pageNumber="7" unit="m" value="1380.0">1380 m</quantity>
to light, 
<date id="FFA4101FEF7E3C12FED10B99FDA7F3C4" box="[335,565,1159,1185]" pageId="6" pageNumber="7" value="1990-12-15">15 December 1990</date>
, J.S. Dugdale.” “ 
<typeStatus id="54A1887DEF7E3C12FC9D0B99FC33F3C4" box="[771,929,1159,1185]" pageId="6" pageNumber="7" type="holotype">HOLOTYPE</typeStatus>
male/ 
<taxonomicName id="4C1A4D5CEF7E3C12FC720B99FEA9F3AC" authority="Dugdale" authorityName="Dugdale" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="liparodes">Tracholena liparodes Dugdale</taxonomicName>
” [red card], 
<collectionCode id="ED0BAE1AEF7E3C12FE500BB1FDB5F3AC" LSID="urn:lsid:biocol.org:col:34988" box="[462,551,1199,1225]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="6" pageNumber="7">MNHN</collectionCode>
</paragraph>
<paragraph id="8BA536DFEF7E3C12FEA60BC9FB43F27C" blockId="6.[264,1323,1118,1905]" pageId="6" pageNumber="7">
<emphasis id="B96EEACDEF7E3C12FEA60BC9FE24F395" box="[312,438,1239,1264]" italics="true" pageId="6" pageNumber="7">
<typeStatus id="54A1887DEF7E3C12FEA60BC9FE21F395" box="[312,435,1239,1264]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<emphasis id="B96EEACDEF7E3C12FE210BC8FDEDF395" bold="true" box="[447,639,1238,1264]" pageId="6" pageNumber="7">
<collectingCountry id="F30D764FEF7E3C12FE210BC8FDEDF395" box="[447,639,1238,1264]" name="New Caledonia" pageId="6" pageNumber="7">New Caledonia</collectingCountry>
</emphasis>
: 
<specimenCount id="9D1CFD56EF7E3C12FD0F0BC9FD77F394" box="[657,741,1239,1265]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
, 
<specimenCount id="9D1CFD56EF7E3C12FD6D0BC9FCCCF394" box="[755,862,1239,1265]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
, same data as 
<typeStatus id="54A1887DEF7E3C12FB900BC9FBE7F394" box="[1038,1141,1239,1265]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
except capture dates span 
<date id="FFA4101FEF7E3C12FE170BE1FD15F27C" box="[393,647,1279,1305]" pageId="6" pageNumber="7" value="1990-12-06" valueMax="1990-12-16" valueMin="1990-12-06">6–16 December 1990</date>
, 
<collectionCode id="ED0BAE1AEF7E3C12FD0A0BE1FD75F27C" box="[660,743,1279,1305]" httpUri="http://grbio.org/cool/crvj-94ys" name="Office de la Recherche Scientifique et Technique d'Outre-Mer" pageId="6" pageNumber="7">ONNC</collectionCode>
Noumea; 
<specimenCount id="9D1CFD56EF7E3C12FCFC0BE1FC26F27C" box="[866,948,1279,1305]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
, 
<specimenCount id="9D1CFD56EF7E3C12FC5E0BE1FBBAF27C" box="[960,1064,1279,1305]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
, ditto, 
<collectionCode id="ED0BAE1AEF7E3C12FBE70BE1FB5FF27C" LSID="urn:lsid:biocol.org:col:34070" box="[1145,1229,1279,1305]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="6" pageNumber="7">NZAC</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3006554EF7E3C13FEA60A38FDECF393" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="8BA536DFEF7E3C13FEA60A38FB1BF6E3" blockId="6.[264,1323,1118,1905]" lastBlockId="7.[264,1324,284,1870]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B96EEACDEF7E3C12FEA60A38FE5CF225" bold="true" box="[312,462,1318,1344]" pageId="6" pageNumber="7">
<emphasis id="B96EEACDEF7E3C12FEA60A38FE56F225" bold="true" box="[312,452,1318,1344]" italics="true" pageId="6" pageNumber="7">Description</emphasis>
:
</emphasis>
<emphasis id="B96EEACDEF7E3C12FE470A39FDE4F225" box="[473,630,1319,1344]" italics="true" pageId="6" pageNumber="7">Male, female</emphasis>
(
<figureCitation id="13212A5AEF7E3C12FD140A39FD44F224" box="[650,726,1319,1345]" captionStart="FIGURES 1 – 3" captionStartId="13.[264,383,1796,1820]" captionTargetBox="[292,1271,293,1778]" captionTargetId="figure@13.[291,1305,280,1786]" captionTargetPageId="13" captionText="FIGURES 1 – 3. Fig. 1. Tracholena nigrilinea sp. nov. Male, Ponerihouen, habitus. (LH wings omitted). Fig. 2. T. liparodes sp. nov. PT male, Mt Panié, habitus. Fig. 3. T. paniense sp. nov. PT male, Mt Panié, habitus. (Scales represent 0.5 mm)" httpUri="https://zenodo.org/record/170881/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
): Head with vertex tufts grey brown, scape grey brown; with vom Raths organ less than half length of labial palp apical segment; maxillary palp one­segmented on an inwardly spinulose pedicel, apical palp segment with 1–2 scales and apically with 4 short stiff sensilla. Flagellomeres longer than wide, 1–3 cylindrical, rest tapering apically, and each with a basal whorl of trichomes longest on the basal flagellomeres and at most only slightly longer than flagellomere length, flagellomeres with the paired sensilla coeloconica contiguous, ventro­apical; labial palpus second segment scaling evenly grey­brown speckled outwardly; prosternal midline scale tuft pale buff contrasting with the brown speckled fore coxae. Forewing: Shape more or less as in 
<taxonomicName id="4C1A4D5CEF7E3C12FB5C0979FAB8F1E5" box="[1218,1322,1639,1664]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="paniense">
<emphasis id="B96EEACDEF7E3C12FB5C0979FAB8F1E5" box="[1218,1322,1639,1664]" italics="true" pageId="6" pageNumber="7">paniense</emphasis>
</taxonomicName>
(
<figureCitation id="13212A5AEF7E3C12FE8F0991FECAF1CC" box="[273,344,1679,1705]" captionStart="FIGURES 1 – 3" captionStartId="13.[264,383,1796,1820]" captionTargetBox="[292,1271,293,1778]" captionTargetId="figure@13.[291,1305,280,1786]" captionTargetPageId="13" captionText="FIGURES 1 – 3. Fig. 1. Tracholena nigrilinea sp. nov. Male, Ponerihouen, habitus. (LH wings omitted). Fig. 2. T. liparodes sp. nov. PT male, Mt Panié, habitus. Fig. 3. T. paniense sp. nov. PT male, Mt Panié, habitus. (Scales represent 0.5 mm)" httpUri="https://zenodo.org/record/170881/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
), length ca. 2.3X width (females), ca. 2.8X width (males); forewing underside with costal cell area dark grey­scaled, contrasting with the rest of the wing; forewing upperside (and mesonotum) ochreous grey, with indistinct blackish markings at about ¼, ½, and a series marking apices of veins R1, R2, and R3; an interrupted serrate dark brown­scaled line at ¼, and another, even less complete at about ½ wing length; Sc, R and branches with scattered small groups of dark brown scales, 1A+2A; discally with scattered irregular patches of more ochreous scales; areas of raised scales as in 
<taxonomicName id="4C1A4D5CEF7F3C13FC7A0E02FBCAF650" box="[996,1112,284,309]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigrilinea">
<emphasis id="B96EEACDEF7F3C13FC7A0E02FBCAF650" box="[996,1112,284,309]" italics="true" pageId="7" pageNumber="8">nigrilinea</emphasis>
</taxonomicName>
. Hindwing: Grey buff; the anal tuft over half the length of anal wing margin; venation: veins M3 and Cu1A coincident at base (i.e., “bifid”) and distant from vein M2 (
<figureCitation id="13212A5AEF7F3C13FC270E72FB93F6E3" box="[953,1025,364,390]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="7" pageNumber="8">Fig. 6</figureCitation>
, cf. 
<figureCitation id="13212A5AEF7F3C13FBAD0E72FBE8F6E3" box="[1075,1146,364,390]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="7" pageNumber="8">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BA536DFEF7F3C13FEA60E8AFCB1F6B3" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
<emphasis id="B96EEACDEF7F3C13FEA60E8AFE39F6C8" box="[312,427,404,429]" italics="true" pageId="7" pageNumber="8">Wingspan</emphasis>
: 
<typeStatus id="54A1887DEF7F3C13FE230E8AFDB8F6CB" box="[445,554,404,430]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
male: 
<quantity id="4CE29B3AEF7F3C13FDE10E8AFD7EF6CB" box="[639,748,404,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="7" pageNumber="8" unit="mm" value="15.2">15.2 mm</quantity>
; 2 
<typeStatus id="54A1887DEF7F3C13FC8D0E8AFCEAF6CB" box="[787,888,404,430]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
males: 14.8 and 
<quantity id="4CE29B3AEF7F3C13FBD70E8AFB25F6CB" box="[1097,1207,404,430]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="7" pageNumber="8" unit="mm" value="15.2">15.2 mm</quantity>
; 
<typeStatus id="54A1887DEF7F3C13FB580E8AFAB9F6CB" box="[1222,1323,404,430]" pageId="7" pageNumber="8" type="paratype">paratype</typeStatus>
females 15.8, 16.0 and 
<quantity id="4CE29B3AEF7F3C13FD860EA2FD16F6B3" box="[536,644,444,470]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="7" pageNumber="8" unit="mm" value="18.2">18.2 mm</quantity>
, respectively.
</paragraph>
<paragraph id="8BA536DFEF7F3C13FEA60EFAFDD0F4EB" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
<emphasis id="B96EEACDEF7F3C13FEA60EFAFE77F698" box="[312,485,484,509]" italics="true" pageId="7" pageNumber="8">Male genitalia</emphasis>
(
<figureCitation id="13212A5AEF7F3C13FE680EFAFDE8F69B" box="[502,634,484,510]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="7" pageNumber="8">Figs. 9, 10</figureCitation>
): Uncus long, slender, decurved; uncus base broad, mesal part as in 
<taxonomicName id="4C1A4D5CEF7F3C13FE1C0D12FE64F540" box="[386,502,524,549]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigrilinea">
<emphasis id="B96EEACDEF7F3C13FE1C0D12FE64F540" box="[386,502,524,549]" italics="true" pageId="7" pageNumber="8">nigrilinea</emphasis>
</taxonomicName>
, produced anteriorly between the posteriorly extended tegumen apices; socii lobate; gnathos arms forming a U thickened towards apex, with lateral spinules outwardly, and thickened part with dorsal spinules, apex with a pair of ventral “wings; transtilla dorsally with a pair of dorsally and outwardly­directed spinose clavate arms; valva apex oblique (costal margin shorter than saccular margin), and no sclerotised strut below the M2 muscle attachment (
<taxonomicName id="4C1A4D5CEF7F3C13FD3D0DCAFCADF588" box="[675,831,724,750]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigrilinea">
cf. 
<emphasis id="B96EEACDEF7F3C13FD550DCAFCADF588" box="[715,831,724,749]" italics="true" pageId="7" pageNumber="8">nigrilinea</emphasis>
</taxonomicName>
); aedeagus (
<figureCitation id="13212A5AEF7F3C13FC4B0DCAFB8EF58B" box="[981,1052,724,750]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
) stout, not longer than valva; with a large apical dextral hook (appressed to the vesica in 
<figureCitation id="13212A5AEF7F3C13FB980DE2FBDEF473" box="[1030,1100,764,790]" captionStart="FIGURES 1 – 10" captionStartId="14.[264,382,1544,1568]" captionTargetBox="[318,1303,344,1492]" captionTargetId="figure@14.[285,1328,316,1522]" captionTargetPageId="14" captionText="FIGURES 1 – 10. Fig. 4. T. paniense. PT male fore and hind wing venation [NZAC prep. 875]. Fig. 5. T. nigrilinea. Fore and hind wing venation, male [NZAC prep. L 862]. (Outer scale tuft across vein CuA 2 omitted.) Fig. 6. T. liparodes sp. nov. Hind wing M 2 ­ M 3 ­ CuA 1 disposition. Fig. 7. T. paniense, male flagellomere 10, ventral, showing widely separated sensillae coeloconica. [NZAC prep. L 879]. Fig. 8. T. nigrilinea, PT male, genitalia, Ponerihouen [NZAC prep. L 859]. Fig. 9. T. liparodes, PT male, aedeagus and bulbus ejaculatorius, Mt Panié [NZAC prep. L 872]. Fig. 10. T. liparodes PT male, genitalia, Mt Panié [NZAC prep. L 880]. (Scales by Figs 4 &amp; 5 repesent 1.0 mm; those by Figs 6, 8 – 10, 0.5 mm.)" httpUri="https://zenodo.org/record/170882/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
); bulbus ejaculatorius entering at ca. half aedeagus length, vesica with a basal spinulose patch and 14 nondeciduous plate­like cornuti arranged in 2 arcs; bulbus ejaculatorius tube stout, sinuous, hood not as long as sleeve.
</paragraph>
<paragraph id="8BA536DFEF7F3C13FEA60C82FDECF393" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
<emphasis id="B96EEACDEF7F3C13FEA60C82FD90F4D0" box="[312,514,924,949]" italics="true" pageId="7" pageNumber="8">Female genitalia</emphasis>
(
<figureCitation id="13212A5AEF7F3C13FD8B0C82FDE2F4D3" box="[533,624,924,950]" captionStart="FIGURES 11 – 15" captionStartId="15.[264,383,1360,1384]" captionTargetBox="[306,1263,328,1302]" captionTargetId="figure@15.[286,1300,326,1335]" captionTargetPageId="15" captionText="FIGURES 11 – 15. Fig. 11. T. paniense, PT male, genitalia, Mt Panié [NZAC prep. L 878]. Fig. 12. T. nigrilinea, female genitalia, Baie des Tortues [NZAC prep. 866]. Fig. 13. T. liparodes, PT female genitalia, Mt Panié [NZAC prep. L 873, 874]. Fig. 14. T. paniense PT female genitalia: ductus and corpus bursae, Mt Panié [NZAC prep. 877]. Fig. 15. T. paniense PT female genitalia: ovipositor, sterigmal and limen structure [NZAC prep. L 877]. (All scales represent 0.5 mm.)" httpUri="https://zenodo.org/record/170883/files/figure.png" pageId="7" pageNumber="8">Fig. 13</figureCitation>
): Ovipositor lobes axes more or less parallel, unmodified anteriorly. Ovipore on a level with anterior third of lobes, and minutely spinulose; sternite 8 with a raised pair of transverse curved ridges; lamella postvaginalis a broad scobinate field with a narrow, sclerotised, scobinate strip bordering the ostium bursae; lamella antevaginalis a thin sinuous sclerotised bar, ostium bursae laterally produced anteriorly (but no ligament discernible); colliculum short, ventrally split; ductus seminalis arising on a latero­ventral papilla; ductus bursae and cestum sinuous, forming one long complete spiral; corpus bursae with a broad­margined, thorn­like signum, corpus bursae covered in minute pit­like micro­sculpture.
</paragraph>
</subSubSection>
<subSubSection id="C3006554EF7F3C1CFEA60A1AFD58F52B" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="diagnosis">
<paragraph id="8BA536DFEF7F3C13FEA60A1AFD27F283" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
<emphasis id="B96EEACDEF7F3C13FEA60A1AFE2AF27B" box="[312,440,1284,1310]" italics="true" pageId="7" pageNumber="8">Diagnosis:</emphasis>
This species resembles 
<taxonomicName id="4C1A4D5CEF7F3C13FD6C0A1AFCEEF278" box="[754,892,1284,1309]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="paniense">
<emphasis id="B96EEACDEF7F3C13FD6C0A1AFCEEF278" box="[754,892,1284,1309]" italics="true" pageId="7" pageNumber="8">T. paniense</emphasis>
</taxonomicName>
and 
<emphasis id="B96EEACDEF7F3C13FC200A1AFBBEF278" box="[958,1068,1284,1309]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C1A4D5CEF7F3C13FC200A1AFBB7F278" box="[958,1061,1284,1309]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lipara">T. lipara</taxonomicName>
,
</emphasis>
being broad­winged (forewing length &lt;2.9X width). It can be distinguished from those two by the forewing with costal cell area dark­grey scaled ventrally (not so in the others); hindwing veins M3 and CuA1 contiguous basally but not stalked (stalked in 
<taxonomicName id="4C1A4D5CEF7F3C13FC3E0A62FC7AF2F0" box="[928,1000,1404,1429]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B96EEACDEF7F3C13FC3E0A62FC7AF2F0" box="[928,1000,1404,1429]" italics="true" pageId="7" pageNumber="8">lipara</emphasis>
</taxonomicName>
, distant in 
<taxonomicName id="4C1A4D5CEF7F3C13FBF40A62FB40F2F0" box="[1130,1234,1404,1429]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="paniense">
<emphasis id="B96EEACDEF7F3C13FBF40A62FB40F2F0" box="[1130,1234,1404,1429]" italics="true" pageId="7" pageNumber="8">paniense</emphasis>
</taxonomicName>
); aedeagus with an apical hooked process; and vesica with 14 chunky, non­deciduous, basal cornuti (a possible species apomorphy).
</paragraph>
<paragraph id="8BA536DFEF7F3C13FEA60AEAFA8AF13B" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
The unmodified ovipositor lobes, the sinuous cestum/ductus bursae (a single, elongate spiral), and presence of a strong hollow signum are character states shared with the Queensland 
<taxonomicName id="4C1A4D5CEF7F3C13FED3095AFE7BF138" box="[333,489,1604,1629]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="micropolia">
<emphasis id="B96EEACDEF7F3C13FED3095AFE7BF138" box="[333,489,1604,1629]" italics="true" pageId="7" pageNumber="8">T. micropolia</emphasis>
</taxonomicName>
. Other differences are noted in the “Diagnosis” of 
<taxonomicName id="4C1A4D5CEF7F3C13FBA0095AFB51F138" box="[1086,1219,1604,1629]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="paniense">
<emphasis id="B96EEACDEF7F3C13FBA0095AFB51F138" box="[1086,1219,1604,1629]" italics="true" pageId="7" pageNumber="8">T. paniense</emphasis>
</taxonomicName>
below.
</paragraph>
<paragraph id="8BA536DFEF7F3C13FEA60972FDC9F02B" blockId="7.[264,1324,284,1870]" pageId="7" pageNumber="8">
<emphasis id="B96EEACDEF7F3C13FEA60972FE4DF1E0" box="[312,479,1644,1669]" italics="true" pageId="7" pageNumber="8">General notes</emphasis>
: While most specimens examined have a wingspan of 
<quantity id="4CE29B3AEF7F3C13FBEF0972FB61F1E3" box="[1137,1267,1644,1670]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="7" pageNumber="8" unit="mm" value="15.0" valueMax="16.0" valueMin="14.0">14–16 mm</quantity>
, one female of this species exceeds 
<quantity id="4CE29B3AEF7F3C13FDED098AFD5BF1CB" box="[627,713,1684,1710]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="8" unit="mm" value="18.0">18 mm</quantity>
, but it has the same venation and genital conformation as the smaller specimens of 
<emphasis id="B96EEACDEF7F3C13FD1709A2FD6CF1B0" box="[649,766,1724,1749]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C1A4D5CEF7F3C13FD1709A2FD68F1B0" box="[649,762,1724,1749]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="liparodes">liparodes</taxonomicName>
.
</emphasis>
Similar variation in wingspan was reported for the female­strobilus/seed­eating 
<taxonomicName id="4C1A4D5CEF7F3C13FD1809FAFC7DF19B" authority="Common 1973: 302" authorityName="Common" authorityPageNumber="302" authorityYear="1973" box="[646,1007,1764,1790]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B96EEACDEF7F3C13FD1809FAFD7AF198" box="[646,744,1764,1789]" italics="true" pageId="7" pageNumber="8">T. lipara</emphasis>
(
<bibRefCitation id="EF8B4B2EEF7F3C13FD6709FAFC7AF19B" author="Common" box="[761,1000,1764,1790]" pageId="7" pageNumber="12" refString="Common, I. F. B. (1973) A new species of Tracholena (Lepidoptera: Tortricidae) attacking klinkii pine in Papua New Guinea, with a description of the larva and pupa of T. homopolia from Australia. Journal of the Australian Entomological Society, 12, 301 - 306." type="journal article" year="1973">Common 1973: 302</bibRefCitation>
)
</taxonomicName>
but not for the foliage­eating 
<taxonomicName id="4C1A4D5CEF7F3C13FEA80812FE2BF040" box="[310,441,1804,1829]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="dialeuca">
<emphasis id="B96EEACDEF7F3C13FEA80812FE2BF040" box="[310,441,1804,1829]" italics="true" pageId="7" pageNumber="8">T. dialeuca</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1A4D5CEF7F3C13FE6A0812FDE0F040" box="[500,626,1804,1829]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hedraea">
<emphasis id="B96EEACDEF7F3C13FE6A0812FDE0F040" box="[500,626,1804,1829]" italics="true" pageId="7" pageNumber="8">T. hedraea</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8B4B2EEF7F3C13FD1D0812FC23F043" author="Common" box="[643,945,1804,1830]" pageId="7" pageNumber="12" refString="Common, I. F. B. (1982) Two new species of Tracholena Common (Lepidoptera: Tortricidae) feeding on Araucaria in Queensland and Norfolk Island. Journal of the Australian Entomological Society, 21, 221 - 224." type="journal article" year="1982">Common 1982: 222, 223</bibRefCitation>
), nor for the male­strobilus­eating 
<taxonomicName id="4C1A4D5CEF7F3C13FEAB082AFE54F028" box="[309,454,1844,1869]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigrilinea">
<emphasis id="B96EEACDEF7F3C13FEAB082AFE54F028" box="[309,454,1844,1869]" italics="true" pageId="7" pageNumber="8">T. nigrilinea</emphasis>
</taxonomicName>
(see above).
</paragraph>
<paragraph id="8BA536DFEF703C1CFEA60E02FD58F52B" blockId="8.[264,1324,284,630]" pageId="8" pageNumber="9">
Specimens were collected at light at the same time and at the same site as 
<taxonomicName id="4C1A4D5CEF703C1CFB000E02FAB1F650" box="[1182,1315,284,309]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="paniense">
<emphasis id="B96EEACDEF703C1CFB000E02FAB1F650" box="[1182,1315,284,309]" italics="true" pageId="8" pageNumber="9">T. paniense</emphasis>
</taxonomicName>
, under a stand of 
<taxonomicName id="4C1A4D5CEF703C1CFE520E5AFD01F638" box="[460,659,324,349]" class="Insecta" family="Braconidae" genus="Agathis" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="B96EEACDEF703C1CFE520E5AFD01F638" box="[460,659,324,349]" italics="true" pageId="8" pageNumber="9">Agathis montana</emphasis>
</taxonomicName>
. No feeding site on any host could be found. Most specimens are greasy, indicating that larvae may be seed­eaters. 
<bibRefCitation id="EF8B4B2EEF703C1CFC420E72FB72F6E3" author="Common" box="[988,1248,364,390]" pageId="8" pageNumber="12" refString="Common, I. F. B. (1973) A new species of Tracholena (Lepidoptera: Tortricidae) attacking klinkii pine in Papua New Guinea, with a description of the larva and pupa of T. homopolia from Australia. Journal of the Australian Entomological Society, 12, 301 - 306." type="journal article" year="1973">Common (1973: 303)</bibRefCitation>
noted that 
<taxonomicName id="4C1A4D5CEF703C1CFEA30E8AFE36F6C8" box="[317,420,404,429]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B96EEACDEF703C1CFEA30E8AFE36F6C8" box="[317,420,404,429]" italics="true" pageId="8" pageNumber="9">T. lipara</emphasis>
</taxonomicName>
specimens, reared from damaged klinkii pine (
<taxonomicName id="4C1A4D5CEF703C1CFC450E8AFB42F6C8" box="[987,1232,404,429]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="9" phylum="Tracheophyta" rank="species" species="hunsteinii">
<emphasis id="B96EEACDEF703C1CFC450E8AFB42F6C8" box="[987,1232,404,429]" italics="true" pageId="8" pageNumber="9">Araucaria hunsteinii</emphasis>
</taxonomicName>
) seeds, can become greasy, and most 
<taxonomicName id="4C1A4D5CEF703C1CFDE70EA2FC9FF6B0" box="[633,781,444,469]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="nigrilinea">
<emphasis id="B96EEACDEF703C1CFDE70EA2FC9FF6B0" box="[633,781,444,469]" italics="true" pageId="8" pageNumber="9">T. nigrilinea</emphasis>
</taxonomicName>
specimens, all from male cones, are greasy. Superficially, this species resembles 2 unset 
<taxonomicName id="4C1A4D5CEF703C1CFC860EFAFCECF698" box="[792,894,484,509]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lipara">
<emphasis id="B96EEACDEF703C1CFC860EFAFCECF698" box="[792,894,484,509]" italics="true" pageId="8" pageNumber="9">T. lipara</emphasis>
</taxonomicName>
specimens (in ANIC, CSIRO) from Bulolo in the 
<collectingCountry id="F30D764FEF703C1CFE280D12FE6CF543" box="[438,510,524,550]" name="Papua New Guinea" pageId="8" pageNumber="9">Papua</collectingCountry>
New Guinean Highlands, but lacks the transverse lines of orange scales so characteristic of that species.
</paragraph>
</subSubSection>
<subSubSection id="C3006554EF703C1CFEA60D42FBCEF510" box="[312,1116,604,630]" pageId="8" pageNumber="9" type="etymology">
<paragraph id="8BA536DFEF703C1CFEA60D42FBCEF510" blockId="8.[264,1324,284,630]" box="[312,1116,604,630]" pageId="8" pageNumber="9">
<emphasis id="B96EEACDEF703C1CFEA60D42FE2CF510" box="[312,446,604,629]" italics="true" pageId="8" pageNumber="9">Etymology:</emphasis>
The name refers to the external resemblance to 
<emphasis id="B96EEACDEF703C1CFC6D0D42FBCEF510" box="[1011,1116,604,629]" italics="true" pageId="8" pageNumber="9">
<taxonomicName id="4C1A4D5CEF703C1CFC6D0D42FBC6F510" box="[1011,1108,604,629]" class="Insecta" family="Tortricidae" genus="Tracholena" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="lipara">T. lipara</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>