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<document id="4C3B489332B4F7FCCADA41E31AF1D21B" ID-CLB-Dataset="27951" ID-DOI="10.11646/zootaxa.4521.3.9" ID-GBIF-Dataset="027053c2-d1ce-4699-b6a2-a5a5033878de" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610011" ID-ZooBank="BAA9C380-9FB7-442B-B439-554F748F842C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553624191022" checkinUser="plazi" docAuthor="Buchsbaum, Ulf, Hsu, Li-Peng, Chen, Ding-Jia &amp; Grehan, John R." docDate="2018" docId="03B387AC6A2D6712FF58FEB0FD819528" docLanguage="en" docName="zootaxa.4521.3.9.pdf" docOrigin="Zootaxa 4521 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Endoclita atayala Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan 2018, sp. n." docType="treatment" docVersion="8" lastPageNumber="446" masterDocId="FF8AFFD46A2C6717FFCFFB77FFB29437" masterDocTitle="A new Endoclita (C. &amp;. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae)" masterLastPageNumber="450" masterPageNumber="441" pageNumber="442" updateTime="1698713771925" updateUser="ExternalLinkService">
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<mods:title id="53E2BE7C14C00F2AED723971E525A168">A new Endoclita (C. &amp;. R. Felder, 1874) from the High Mountains of Taiwan with notes about its evolutionary origins (Lepidoptera: Hepialidae)</mods:title>
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<mods:namePart id="D9FDEEF92E1B312BE0829526F0335966">Buchsbaum, Ulf</mods:namePart>
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<mods:namePart id="3F1DA2C6FFE34E610FBF8A22C0C8EB24">Hsu, Li-Peng</mods:namePart>
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<mods:namePart id="D002A8E9C16E2627EEA0F7FD8AEBE4CD">Chen, Ding-Jia</mods:namePart>
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<mods:namePart id="692DB0DFD0C7BFBB6B769B30FE62A0DD">Grehan, John R.</mods:namePart>
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<treatment id="03B387AC6A2D6712FF58FEB0FD819528" ID-DOI="http://doi.org/10.5281/zenodo.5962007" ID-GBIF-Taxon="154498900" ID-Zenodo-Dep="5962007" LSID="urn:lsid:plazi:treatment:03B387AC6A2D6712FF58FEB0FD819528" httpUri="http://treatment.plazi.org/id/03B387AC6A2D6712FF58FEB0FD819528" lastPageId="5" lastPageNumber="446" pageId="1" pageNumber="442">
<subSubSection id="C30065316A2D6716FF58FEB0FD1B91D5" box="[151,681,1479,1506]" pageId="1" pageNumber="442" type="nomenclature">
<paragraph id="8BA536BA6A2D6716FF58FEB0FD1B91D5" blockId="1.[151,681,1479,1539]" box="[151,681,1479,1506]" pageId="1" pageNumber="442">
<heading id="D0ED81D66A2D6716FF58FEB0FD1B91D5" bold="true" box="[151,681,1479,1506]" fontSize="11" level="1" pageId="1" pageNumber="442" reason="1">
<emphasis id="B96EEAA86A2D6716FF58FEB0FD1B91D5" bold="true" box="[151,681,1479,1506]" pageId="1" pageNumber="442">
<taxonomicName id="4C1A4D396A2D6716FF58FEB0FDD291D5" authority="Buchsbaum &amp; Hsu" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[151,608,1479,1506]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2D6716FF58FEB0FED991D6" bold="true" box="[151,363,1479,1505]" italics="true" pageId="1" pageNumber="442">Endoclita atayala</emphasis>
Buchsbaum &amp; Hsu
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2D6716FDA9FEBFFD1B91D5" box="[614,681,1480,1506]" pageId="1" pageNumber="442" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C30065316A2D6716FF58FE9DFEC99234" box="[151,379,1514,1539]" pageId="1" pageNumber="442" type="description">
<paragraph id="8BA536BA6A2D6716FF58FE9DFEC99234" blockId="1.[151,681,1479,1539]" box="[151,379,1514,1539]" pageId="1" pageNumber="442">
(
<figureCitation id="13212A3F6A2D6716FF50FE9DFEB09234" box="[159,258,1514,1539]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="1" pageNumber="442">Figs 1, 3</figureCitation>
¯6a, 7¯11)
</paragraph>
</subSubSection>
<subSubSection id="C30065316A2D6716FF58FD45FBF19314" pageId="1" pageNumber="442" type="diagnosis">
<paragraph id="8BA536BA6A2D6716FF58FD45FBF19314" blockId="1.[151,1436,1586,2007]" pageId="1" pageNumber="442">
<emphasis id="B96EEAA86A2D6716FF58FD45FEA3927C" bold="true" box="[151,273,1586,1611]" pageId="1" pageNumber="442">Diagnosis.</emphasis>
In
<collectingCountry id="F30D762A6A2D6716FEF5FD45FE3E927C" box="[314,396,1586,1611]" name="Taiwan" pageId="1" pageNumber="442">Taiwan</collectingCountry>
the presence of a FW costal lobe is shared by
<taxonomicName id="4C1A4D396A2D6716FC64FD45FBC6927C" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[939,1140,1586,1611]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2D6716FC64FD45FBC6927C" box="[939,1140,1586,1611]" italics="true" pageId="1" pageNumber="442">Endoclita atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2D6716FBB1FD45FB0D927D" box="[1150,1215,1586,1611]" pageId="1" pageNumber="442" rank="species">
<emphasis id="B96EEAA86A2D6716FBB1FD45FB2A927C" bold="true" box="[1150,1176,1586,1611]" pageId="1" pageNumber="442">sp</emphasis>
.
<emphasis id="B96EEAA86A2D6716FB67FD45FB05927C" bold="true" box="[1192,1207,1586,1611]" pageId="1" pageNumber="442">n</emphasis>
.
</taxonomicNameLabel>
and
<taxonomicName id="4C1A4D396A2D6716FB36FD43FAD2927C" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[1273,1376,1586,1611]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B96EEAA86A2D6716FB36FD43FAD2927C" box="[1273,1376,1586,1611]" italics="true" pageId="1" pageNumber="442">E. davidi</emphasis>
</taxonomicName>
. The scattered and irregular dark brownish-black transverse lines and the crenulated margin of the FW separates
<taxonomicName id="4C1A4D396A2D6716FA4BFD20FF5992A4" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2D6716FA4BFD20FF5992A4" italics="true" pageId="1" pageNumber="442">E. atayala</emphasis>
</taxonomicName>
<emphasis id="B96EEAA86A2D6716FF3AFD0DFE8A92A4" bold="true" box="[245,312,1658,1683]" pageId="1" pageNumber="442">
<taxonomicNameLabel id="A25D57D36A2D6716FF3AFD0DFE8A92A4" box="[245,312,1658,1683]" pageId="1" pageNumber="442" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from all other Taiwanese
<taxonomicName id="4C1A4D396A2D6716FDA2FD0DFD6892A4" authorityName="Felder" authorityYear="1874" box="[621,730,1658,1683]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA86A2D6716FDA2FD0DFD6892A4" box="[621,730,1658,1683]" italics="true" pageId="1" pageNumber="442">Endoclita</emphasis>
</taxonomicName>
, including
<taxonomicName id="4C1A4D396A2D6716FC90FD0BFC7592A4" baseAuthorityName="Poujade" baseAuthorityYear="1886" box="[863,967,1658,1683]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="davidi">
<emphasis id="B96EEAA86A2D6716FC90FD0BFC7592A4" box="[863,967,1658,1683]" italics="true" pageId="1" pageNumber="442">E. davidi</emphasis>
</taxonomicName>
, and all other
<taxonomicName id="4C1A4D396A2D6716FBB9FD0DFB5192A4" authorityName="Felder" authorityYear="1874" box="[1142,1251,1658,1683]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA86A2D6716FBB9FD0DFB5192A4" box="[1142,1251,1658,1683]" italics="true" pageId="1" pageNumber="442">Endoclita</emphasis>
</taxonomicName>
species in Asia except two Chinese species—
<taxonomicName id="4C1A4D396A2D6716FE27FDE8FCFC9280" authority="(Le Cerf, 1919)" authorityName="Le Cerf" authorityYear="1919" baseAuthorityName="Le Cerf" baseAuthorityYear="1919" box="[488,846,1694,1719]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="crenilimbata">
<emphasis id="B96EEAA86A2D6716FE27FDE8FD249280" box="[488,662,1694,1719]" italics="true" pageId="1" pageNumber="442">E. crenilimbata</emphasis>
(
<bibRefCitation id="EF8B4B4B6A2D6716FD69FDE8FCF49280" author="Le Cerf, F." box="[678,838,1694,1719]" pageId="1" pageNumber="442" pagination="471" refId="ref5695" refString="Le Cerf, F. (1919) Observations sur le genre Phassus Wlkr.; diagnoses de genres nouveaux et description d'une espece nouvelle (Lepidopt. Hepialidae). Bulletin du Museum National d'histoire Naturelle, 25, 469 ‾ 471." type="journal article" year="1919">Le Cerf, 1919</bibRefCitation>
)
</taxonomicName>
and
<emphasis id="B96EEAA86A2D6716FC49FDE8FC5B9280" box="[902,1001,1694,1719]" italics="true" pageId="1" pageNumber="442">E. nodus</emphasis>
(
<bibRefCitation id="EF8B4B4B6A2D6716FC37FDE9FB5B9280" author="Chu, H. - F. &amp; Wang, L. - Y." box="[1016,1257,1694,1719]" pageId="1" pageNumber="442" pagination="302" refId="ref5027" refString="Chu, H. - F. &amp; Wang, L. - Y. (1985) On the stem-borers of Chinese Hepialidae (Lepidoptera: Hepialidae). Acta Entomologica sinica, 28, 293 ‾ 302." type="journal article" year="1985">Chu and Wang, 1985</bibRefCitation>
). The FW of
<taxonomicName id="4C1A4D396A2D6716FA4BFDE8FF5992EC" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="442" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2D6716FA4BFDE8FF5992EC" italics="true" pageId="1" pageNumber="442">E. atayala</emphasis>
</taxonomicName>
<emphasis id="B96EEAA86A2D6716FF38FDB5FE8F92EC" bold="true" box="[247,317,1730,1755]" pageId="1" pageNumber="442">
<taxonomicNameLabel id="A25D57D36A2D6716FF38FDB5FE8F92EC" box="[247,317,1730,1755]" pageId="1" pageNumber="442" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from these two Chinese species by having a slightly triangular costal lobe (vs evenly rounded), a stronger curve on the outer costa between costal lobe and apex (vs shallow, evenly curved margin) and the presence of a submarginal transverse marking between Rs3 and Rs4 (vs absent).
</paragraph>
</subSubSection>
<subSubSection id="C30065316A2D6714FF08FC59FB9A934E" lastPageId="3" lastPageNumber="444" pageId="1" pageNumber="442" type="description">
<paragraph id="8BA536BA6A2D6716FF08FC59FC81935C" blockId="1.[151,1436,1586,2007]" pageId="1" pageNumber="442">
<emphasis id="B96EEAA86A2D6716FF08FC59FE0B9370" bold="true" box="[199,441,1838,1863]" pageId="1" pageNumber="442">Description. Female</emphasis>
(
<figureCitation id="13212A3F6A2D6716FE05FC59FDC99370" box="[458,635,1838,1863]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="1" pageNumber="442">Figs 1, 35, 6a</figureCitation>
,
<figureCitation id="13212A3F6A2D6716FD46FC58FD769370" box="[649,708,1839,1863]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="1" pageNumber="442">711</figureCitation>
). Wingspan:
<quantity id="4CE29B5F6A2D6716FC93FC59FC0F9371" box="[860,957,1838,1863]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="442" unit="mm" value="150.0">150 mm</quantity>
, forewing length:
<quantity id="4CE29B5F6A2D6716FB5EFC58FB579371" box="[1169,1253,1839,1863]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="1" pageNumber="442" unit="mm" value="79.0">79 mm</quantity>
, width:
<quantity id="4CE29B5F6A2D6716FA8BFC58FA2A9371" box="[1348,1432,1839,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="442" unit="mm" value="32.0">32 mm</quantity>
, ratio 2.5: 1; HW length:
<quantity id="4CE29B5F6A2D6716FE65FC24FE49935D" box="[426,507,1875,1899]" metricMagnitude="-2" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="442" unit="mm" value="71.0">71 mm</quantity>
, width:
<quantity id="4CE29B5F6A2D6716FD9AFC25FD14935D" box="[597,678,1874,1899]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="442" unit="mm" value="26.0">26 mm</quantity>
, ratio 2.7: 1.
</paragraph>
<paragraph id="8BA536BA6A2D6716FF08FC01FAA993B8" blockId="1.[151,1436,1586,2007]" box="[199,1307,1910,1935]" pageId="1" pageNumber="442">
<emphasis id="B96EEAA86A2D6716FF08FC01FEB693B8" box="[199,260,1910,1935]" italics="true" pageId="1" pageNumber="442">Head</emphasis>
. Orangish-brown, antennae missing [broken off]. Eyes prominent. Mouthparts not examined.
</paragraph>
<paragraph id="8BA536BA6A2D6715FF08FCEDFCB0976A" blockId="1.[151,1436,1586,2007]" lastBlockId="2.[151,1437,151,861]" lastPageId="2" lastPageNumber="443" pageId="1" pageNumber="442">
<emphasis id="B96EEAA86A2D6716FF08FCEDFE929384" box="[199,288,1946,1971]" italics="true" pageId="1" pageNumber="442">Thorax.</emphasis>
Orangish-brown, dorsal scales short except for dorsal metathorax, anterior scutum III free of scales, iridescent dark chocolate brown. FW costal margin with prominent lobate extension opposite the common stem of radial sector veins (
<figureCitation id="13212A3F6A2E6715FEB7FBE0FE0C9487" box="[376,446,151,176]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 4</figureCitation>
), immediately distal to lobe slightly concave, then convex, sloping posteriorly to apex (
<figureCitation id="13212A3F6A2E6715FF50FBCBFF5294E3" box="[159,224,188,213]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 1</figureCitation>
). Termen slightly crenulated, concave between veins, tornus not discernable. Venation hepialine with close approximation of the common root of Rs1+2 and Rs3 (
<figureCitation id="13212A3F6A2E6715FCDDFBA8FCEA94CF" box="[786,856,223,248]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 3</figureCitation>
). Width between costal margin and Sc broad from wing base to outer edge of costal lobe, then sharply narrow (
<figureCitation id="13212A3F6A2E6715FCF3FA73FC33952B" box="[828,897,260,285]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 4</figureCitation>
). Sc1 well developed, subtending apex of costal lobe. Distance between Sc and R narrows beyond costal lobe. Presence of four basal cells not confirmed. Dorsal ground colour pale orangish-brown to pale yellowish brown or pale gray brown distally, with scattered, mostly transverse dark brownish-black lines or partial circles between veins, particularly in the form of two oblique transverse rows in the post discal region; termen with ovoid dark spot centred on each vein, and two curved, submarginal dark curved lines sub-parallel to margin enclosing greyish brown line and central line of pale greyish brown between Rs2 and Rs3 (
<figureCitation id="13212A3F6A2E6715FE3FFAABFD8495C2" box="[496,566,476,501]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 5</figureCitation>
); outer anterior discal cell with curved longitudinal lines near M2 (
<figureCitation id="13212A3F6A2E6715FAF5FAABFA3C95C2" box="[1338,1422,476,501]" captionStart="FIGURES 16" captionStartId="2.[151,264,1878,1901]" captionTargetBox="[281,1305,916,1856]" captionTargetId="figure@2.[281,1305,916,1856]" captionTargetPageId="2" captionText="FIGURES 16. Voucher specimens: Endoclita atayala sp. n., female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2, E. crenilimbata female HT (MNHN), dorsal view. Photo by Joël Minet. E. atayala sp. n., morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a) E. atayala sp. n., (b) E. crenilimbata, (c) E. nodus (from Zhu et al. 2004)." httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443">Fig. 6a</figureCitation>
); outer half of FW beyond cross veins with irregular, dark, oval to ovoid lines; area between costal margin and Sc with two irregular pale markings edged with black from wing base and costal lobe, and three or four similar markings, much smaller between costal lobe and apex; shallow curved pale yellowish band mottled with pale greyish-brown extends basal to 1A and across the central region between 1A and CuA2. FW ventral ground colour greyish brown to pale yellowish brown anteriorly, especially where piliform scales have been rubbed from specimen. Some dorsal ornamentation present between costal margin and Sc, and along termen and inner margins. HW venation similar to FW except for absence of Sc1; Rs1 almost straight rather than convex relative to costal margin as in FW; dorsal and ventral ground colour greyish brown except for dorsal ornamentation along costal and outer margins. Legs (
<figureCitation id="13212A3F6A2E6715FE58F868FE50970F" box="[407,482,799,824]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="2" pageNumber="443">Fig. 7</figureCitation>
): foreleg with shorter femur and tibia than middle-leg; (hind-legs missing from specimen after removal of abdomen); epiphysis absent.
</paragraph>
<caption id="DF6566326A2E6715FF58FC21FD2293DF" httpUri="https://zenodo.org/record/2610013/files/figure.png" pageId="2" pageNumber="443" startId="2.[151,264,1878,1901]" targetBox="[281,1305,916,1856]" targetPageId="2">
<paragraph id="8BA536BA6A2E6715FF58FC21FD2293DF" blockId="2.[151,1437,1878,2024]" pageId="2" pageNumber="443">
<emphasis id="B96EEAA86A2E6715FF58FC21FE8C935B" bold="true" box="[151,318,1878,1901]" pageId="2" pageNumber="443">FIGURES 16.</emphasis>
Voucher specimens:
<taxonomicName id="4C1A4D396A2E6715FDEEFC20FD68935A" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[545,730,1879,1901]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2E6715FDEEFC20FD68935A" box="[545,730,1879,1901]" italics="true" pageId="2" pageNumber="443">Endoclita atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2E6715FD2BFC20FC93935A" box="[740,801,1879,1901]" pageId="2" pageNumber="443" rank="species">
<emphasis id="B96EEAA86A2E6715FD2BFC20FD4E935B" bold="true" box="[740,764,1879,1900]" pageId="2" pageNumber="443">sp</emphasis>
.
<emphasis id="B96EEAA86A2E6715FCC2FC20FCA9935B" bold="true" box="[781,795,1879,1900]" pageId="2" pageNumber="443">n</emphasis>
.
</taxonomicNameLabel>
, female HT (NMNS). 1a, dorsal; 1b, ventral. Photo by Ulf Buchsbaum. 2,
<taxonomicName id="4C1A4D396A2E6715FEFBFC00FE6093BB" baseAuthorityName="Le Cerf" baseAuthorityYear="1919" box="[308,466,1910,1932]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="crenilimbata">
<emphasis id="B96EEAA86A2E6715FEFBFC00FE6093BB" box="[308,466,1910,1932]" italics="true" pageId="2" pageNumber="443">E. crenilimbata</emphasis>
</taxonomicName>
female HT (MNHN), dorsal view. Photo by Joël Minet.
<taxonomicName id="4C1A4D396A2E6715FBDDFC00FBC893BB" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[1042,1146,1910,1932]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2E6715FBDDFC00FBC893BB" box="[1042,1146,1910,1932]" italics="true" pageId="2" pageNumber="443">E. atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2E6715FB4FFC01FB0B93BB" box="[1152,1209,1910,1932]" pageId="2" pageNumber="443" rank="species">
<emphasis id="B96EEAA86A2E6715FB4FFC01FB2A93BC" bold="true" box="[1152,1176,1910,1931]" pageId="2" pageNumber="443">sp</emphasis>
.
<emphasis id="B96EEAA86A2E6715FB6AFC01FB0193BC" bold="true" box="[1189,1203,1910,1931]" pageId="2" pageNumber="443">n</emphasis>
.
</taxonomicNameLabel>
, morphology: 3, wing venation. Vein connections in basal regions unresolved. 4, costal lobe detail. 5, submarginal transverse band on outer margin of forewing. 6, boundaries of anterior discal cell (blue line) enclosing longitudinal markings: (a)
<taxonomicName id="4C1A4D396A2E6715FB5BFCC3FAB393FE" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[1172,1281,1971,1993]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2E6715FB5BFCC3FAB393FE" box="[1172,1281,1971,1993]" italics="true" pageId="2" pageNumber="443">E. atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2E6715FAC2FCC3FAF993FE" box="[1293,1355,1972,1993]" pageId="2" pageNumber="443" rank="species">
<emphasis id="B96EEAA86A2E6715FAC2FCC3FA9793FE" bold="true" box="[1293,1317,1972,1993]" pageId="2" pageNumber="443">sp</emphasis>
.
<emphasis id="B96EEAA86A2E6715FAF8FCC3FAF793FE" bold="true" box="[1335,1349,1972,1993]" pageId="2" pageNumber="443">n</emphasis>
.
</taxonomicNameLabel>
, (b)
<taxonomicName id="4C1A4D396A2E6715FA49FCC3FEA593DF" baseAuthorityName="Le Cerf" baseAuthorityYear="1919" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="443" phylum="Arthropoda" rank="species" species="crenilimbata">
<emphasis id="B96EEAA86A2E6715FA49FCC3FEA593DF" italics="true" pageId="2" pageNumber="443">E. crenilimbata</emphasis>
</taxonomicName>
, (c)
<emphasis id="B96EEAA86A2E6715FE89FCA4FE2C93DF" box="[326,414,2002,2024]" italics="true" pageId="2" pageNumber="443">E. nodus</emphasis>
(from Zhu
<emphasis id="B96EEAA86A2E6715FDDBFCA4FDF393DF" box="[532,577,2002,2024]" italics="true" pageId="2" pageNumber="443">et al</emphasis>
. 2004).
</paragraph>
</caption>
<caption id="DF6566326A2F6714FF58FF8FFC8D919F" httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444" startId="3.[151,264,1272,1295]" targetBox="[313,1273,193,1251]" targetPageId="3">
<paragraph id="8BA536BA6A2F6714FF58FF8FFC8D919F" blockId="3.[151,1436,1272,1448]" pageId="3" pageNumber="444">
<emphasis id="B96EEAA86A2F6714FF58FF8FFEFE913A" bold="true" box="[151,332,1272,1295]" pageId="3" pageNumber="444">FIGURES 711.</emphasis>
<taxonomicName id="4C1A4D396A2F6714FE98FF8FFDA09139" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[343,530,1272,1294]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="444" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A2F6714FE98FF8FFDA09139" box="[343,530,1272,1294]" italics="true" pageId="3" pageNumber="444">Endoclita atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A2F6714FDD1FF8EFDEF9139" box="[542,605,1273,1294]" pageId="3" pageNumber="444" rank="species">
<emphasis id="B96EEAA86A2F6714FDD1FF8EFD849139" bold="true" box="[542,566,1273,1294]" pageId="3" pageNumber="444">sp</emphasis>
.
<emphasis id="B96EEAA86A2F6714FD86FF8EFDE59139" bold="true" box="[585,599,1273,1294]" pageId="3" pageNumber="444">n</emphasis>
.
</taxonomicNameLabel>
, Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules.
</paragraph>
</caption>
<paragraph id="8BA536BA6A2F6714FF08FEB8FC2E924F" blockId="3.[151,1437,1487,2022]" pageId="3" pageNumber="444">
<emphasis id="B96EEAA86A2F6714FF08FEB8FE8391DF" box="[199,305,1487,1512]" italics="true" pageId="3" pageNumber="444">Abdomen</emphasis>
(
<figureCitation id="13212A3F6A2F6714FE8FFEB8FE1691D0" box="[320,420,1487,1512]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Figs 8, 9</figureCitation>
). Covered with greyish brown scales with pale yellowish brown scales posteriorly on each segment [inferred due to poor condition of abdominal scales]. Tergosternal bar fused with intermediate zone of tergal brace, tergal brace right angled between lateral and dorsal arms, intermediate zone without tergal knob (
<figureCitation id="13212A3F6A2F6714FAA0FD60FF159263" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig. 8</figureCitation>
). Sternite II rectangular with shallow V-shaped anterior margin, anterior lateral arms and anterior margin strongly sclerotized (
<figureCitation id="13212A3F6A2F6714FED0FD28FED69240" box="[287,356,1631,1656]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig. 9</figureCitation>
), apex of lateral arms with sub apical ridge (
<figureCitation id="13212A3F6A2F6714FC98FD28FC21924F" box="[855,915,1631,1656]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig.8</figureCitation>
)
</paragraph>
<paragraph id="8BA536BA6A2F6714FF08FDF4FB9A934E" blockId="3.[151,1437,1487,2022]" pageId="3" pageNumber="444">
<emphasis id="B96EEAA86A2F6714FF08FDF4FE8392AB" box="[199,305,1667,1692]" italics="true" pageId="3" pageNumber="444">Genitalia</emphasis>
(
<figureCitation id="13212A3F6A2F6714FE8FFDF4FE2892AB" box="[320,410,1667,1692]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Figs 10</figureCitation>
¯11). External genitalia (
<figureCitation id="13212A3F6A2F6714FD7FFDF4FCB192AB" box="[688,771,1667,1692]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig. 10</figureCitation>
) roughly shaped as a vertical isosceles triangle; tergum IX (dorsal plate) strongly sclerotized, narrowly fused dorsally, posterior dorsal margin forming an inverted U that ends ventrally at a triangular lobe (= anal papilla of
<bibRefCitation id="EF8B4B4B6A2F6714FD76FDBCFC4092D3" author="Nielsen, E. S. &amp; Kristensen, N. P." box="[697,1010,1739,1764]" pageId="3" pageNumber="444" pagination="206" refId="ref5919" refString="Nielsen, E. S. &amp; Kristensen, N. P. (1989) Primitive ghost moths. Morphology and taxonomy of the Australian genus Fraus Walker (Lepidoptera: Hepialidae s. lat.). Monographs on Australian Lepidoptera, 1, 1 ‾ 206." type="journal article" year="1989">Nielsen &amp; Kristensen 1989</bibRefCitation>
) without setae; lamella antevaginalis with each lateral lobe projecting dorsally and a central lobe strongly sclerotized with dorsally triangular projection; sub anal plates weakly sclerotized, vertically oriented. Internal genitalia (
<figureCitation id="13212A3F6A2F6714FC02FC64FB98931B" box="[973,1066,1811,1836]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig. 11a</figureCitation>
) with ductus bursae
<quantity id="4CE29B5F6A2F6714FADBFC64FAD1931C" box="[1300,1379,1811,1836]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="444" unit="mm" value="15.0">15 mm</quantity>
long and 1.5 mm wide, initially narrow, then slightly wider for remaining
<superScript id="7C6F9BF26A2F6714FC1BFC41FC6E9373" attach="right" box="[980,988,1846,1860]" fontSize="6" pageId="3" pageNumber="444">2</superScript>
/
<subScript id="179E34FF6A2F6714FC2BFC3EFC5E9360" attach="left" box="[996,1004,1865,1879]" fontSize="6" pageId="3" pageNumber="444">3</subScript>
distance, distal end with sclerotized spicules (
<figureCitation id="13212A3F6A2F6714FECFFC17FED2934E" box="[256,352,1888,1913]" captionStart="FIGURES 711" captionStartId="3.[151,264,1272,1295]" captionTargetBox="[313,1273,193,1251]" captionTargetId="figure@3.[313,1273,193,1251]" captionTargetPageId="3" captionText="FIGURES 711. Endoclita atayala sp. n., Appendages and abdomen: 7, foreleg (left), mesoleg (right). 8, tergosternal connection: db—dorsal brace, lb—lateral brace, iz—intermediate zone, tsb—tergosternal bar, ar—apical ridge of sternum II lateral lobe. 9, anterior margin of sternum II with tergum II overlapping as dotted outline: ala—anterior lateral lobe of sternum II. 10, external female genitalia, posterior view: dpl—dorsal plate, ap—anal papilla, pa—paranal lobe, lldp, lateral lobe dorsal projection, mldp—median lobe dorsal projection, l.a-v—lamellar antivaginalis (ventral edge marked as dotted line). 11, internal genitalia. 11a, bursa copulatrix, 11b, ductus bursa spicules." httpUri="https://zenodo.org/record/2610015/files/figure.png" pageId="3" pageNumber="444">Fig. 11b</figureCitation>
); corpus bursae cylindrical,
<quantity id="4CE29B5F6A2F6714FD52FC16FD5E934F" box="[669,748,1889,1913]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="444" unit="mm" value="17.0">17 mm</quantity>
long and
<quantity id="4CE29B5F6A2F6714FC94FC17FC2E934F" box="[859,924,1888,1913]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="444" unit="mm" value="5.0">5 mm</quantity>
in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C30065316A2F6713FF08FCF3FB7795BF" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444" type="distribution">
<paragraph id="8BA536BA6A2F6713FF08FCF3FB7795BF" blockId="3.[151,1437,1487,2022]" lastBlockId="4.[151,1436,151,392]" lastPageId="4" lastPageNumber="445" pageId="3" pageNumber="444">
<emphasis id="B96EEAA86A2F6714FF08FCF3FED293AA" bold="true" box="[199,352,1924,1949]" pageId="3" pageNumber="444">Distribution.</emphasis>
The
<typeStatus id="54A188186A2F6714FE6EFCF1FE6393A9" box="[417,465,1926,1950]" pageId="3" pageNumber="444">type</typeStatus>
locality is on the south slope facing an open valley created by the Madala Stream (Madalaxi) of Kuan Wu in a mountainous area of North
<collectingCountry id="F30D762A6A2F6714FCD4FCDFFCDF93F6" box="[795,877,1960,1985]" name="Taiwan" pageId="3" pageNumber="444">Taiwan</collectingCountry>
(
<figureCitation id="13212A3F6A2F6714FCB2FCDFFC6093F6" box="[893,978,1960,1985]" captionStart="FIGURES 1217" captionStartId="4.[151,264,1615,1638]" captionTargetBox="[313,1273,433,1604]" captionTargetId="figure@4.[313,1273,433,1604]" captionTargetPageId="4" captionText="FIGURES 1217. Distribution and diversity. 12, type locality of Endoclita atayala sp. n., Miaoli County, Taiwan. 13, Biotope of Endoclita atayala sp. n., Miaoli County, Taiwan. 14, Species numbers of Hepialidae recorded from different countries and regions (data from Nielsen et al. (2000). 15, Regional comparison of Endoclita species numbers by major land units. 16, Grid of species distribution densities for Endoclita. Grid units 15 x 15°. 17, distribution records for E. atayala sp. n. (red circle, Taiwan), E. crenilimbata (blue circles, Ping-fa, Giuzhou and Mupin, Sichuan) and E. nodus (white circles, Yuexi, Anhui, and Lushan, Jiangxi)." httpUri="https://zenodo.org/record/2610017/files/figure.png" pageId="3" pageNumber="444">Fig. 12</figureCitation>
). This area ranges between 2,000~2,500 meters a.s.l and is constantly enveloped by fog and clouds. Its climate is cool in winter and warm in summer with a mean annual temperature of 13°C. The vegetation is warm temperate and cold temperate zone forest, a mixed forest composed of coniferous and broadleaf trees (
<figureCitation id="13212A3F6A286713FD21FBCBFCF694E3" box="[750,836,188,213]" captionStart="FIGURES 1217" captionStartId="4.[151,264,1615,1638]" captionTargetBox="[313,1273,433,1604]" captionTargetId="figure@4.[313,1273,433,1604]" captionTargetPageId="4" captionText="FIGURES 1217. Distribution and diversity. 12, type locality of Endoclita atayala sp. n., Miaoli County, Taiwan. 13, Biotope of Endoclita atayala sp. n., Miaoli County, Taiwan. 14, Species numbers of Hepialidae recorded from different countries and regions (data from Nielsen et al. (2000). 15, Regional comparison of Endoclita species numbers by major land units. 16, Grid of species distribution densities for Endoclita. Grid units 15 x 15°. 17, distribution records for E. atayala sp. n. (red circle, Taiwan), E. crenilimbata (blue circles, Ping-fa, Giuzhou and Mupin, Sichuan) and E. nodus (white circles, Yuexi, Anhui, and Lushan, Jiangxi)." httpUri="https://zenodo.org/record/2610017/files/figure.png" pageId="4" pageNumber="445">Fig. 13</figureCitation>
). Japanese Cedar and Formosan Red Cypress grow extensively in a vast area of natural forest, and along with Taiwanese Yellow Cypress, Hemlock, Taiwanese Red Pine, Formosan Alder,
<taxonomicName id="4C1A4D396A286713FE55FA72FDCF952A" box="[410,637,260,285]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="445" phylum="Tracheophyta" rank="species" species="carlesii">
<emphasis id="B96EEAA86A286713FE55FA72FDCF952A" box="[410,637,260,285]" italics="true" pageId="4" pageNumber="445">Castanopsis carlesii</emphasis>
</taxonomicName>
, and Wheel Trees, make up a rich forest ecosystem. It also includes the largest forest of purely Taiwanese Sassafras which is the host plant of the endemic Broad-tailed Swallowtail (
<emphasis id="B96EEAA86A286713FF6FFA3BFEB69552" box="[160,260,332,357]" italics="true" pageId="4" pageNumber="445">Agehana</emphasis>
<taxonomicName id="4C1A4D396A286713FEDEFA3BFD3D9553" authority="Shiraki &amp; Sonan, 1934" authorityName="Shiraki &amp; Sonan" authorityYear="1934" box="[273,655,332,357]" class="Insecta" family="Papilionidae" genus="Papilio" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="maraho">
<emphasis id="B96EEAA86A286713FEDEFA3BFEDA9552" box="[273,360,332,357]" italics="true" pageId="4" pageNumber="445">maraho</emphasis>
<bibRefCitation id="EF8B4B4B6A286713FEB9FA3BFD3D9553" author="Shiraki, T. &amp; Sonan, J." box="[374,655,332,357]" pageId="4" pageNumber="445" pagination="182" refId="ref6255" refString="Shiraki, T. &amp; Sonan, J. (1934) On a new species of Papilionidae, Papilio maraho Shiraki and Sonan. Zephyrus, 5, 177 ‾ 182." type="journal article" year="1934">Shiraki &amp; Sonan, 1934</bibRefCitation>
</taxonomicName>
). Among animals this region also includes the endemic Guanwu Formosan Salamander (
<emphasis id="B96EEAA86A286713FE6DFA18FDF795BF" box="[418,581,367,392]" italics="true" pageId="4" pageNumber="445">Hynobius fuca</emphasis>
<bibRefCitation id="EF8B4B4B6A286713FD83FA18FCBA95BF" author="Lai, J. - S. &amp; Lue, K. - Y." box="[588,776,367,392]" pageId="4" pageNumber="445" pagination="63" refId="ref5645" refString="Lai, J. - S. &amp; Lue, K. - Y. (2008) Two new Hynobius (Caudata: Hynobiidae) salamanders from Taiwan. Herpetologica, 64, 63 ‾ 80. https: // doi. org / 10.1655 / 06 - 065.1" type="journal article" year="2008">Lai &amp; Lue, 2008</bibRefCitation>
) that lives in nearby small, cold creeks.
</paragraph>
</subSubSection>
<caption id="DF6566326A286713FF58FD38FEF592C8" httpUri="https://zenodo.org/record/2610017/files/figure.png" pageId="4" pageNumber="445" startId="4.[151,264,1615,1638]" targetBox="[313,1273,433,1604]" targetPageId="4">
<paragraph id="8BA536BA6A286713FF58FD38FEF592C8" blockId="4.[151,1436,1615,1791]" pageId="4" pageNumber="445">
<emphasis id="B96EEAA86A286713FF58FD38FEE19253" bold="true" box="[151,339,1615,1638]" pageId="4" pageNumber="445">FIGURES 1217.</emphasis>
Distribution and diversity. 12, type locality of
<taxonomicName id="4C1A4D396A286713FCE1FD38FC519252" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[814,995,1615,1637]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A286713FCE1FD38FC519252" box="[814,995,1615,1637]" italics="true" pageId="4" pageNumber="445">Endoclita atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A286713FC25FD27FB909252" box="[1002,1058,1616,1637]" pageId="4" pageNumber="445" rank="species">
<emphasis id="B96EEAA86A286713FC25FD27FBB09252" bold="true" box="[1002,1026,1616,1637]" pageId="4" pageNumber="445">sp</emphasis>
.
<emphasis id="B96EEAA86A286713FBC1FD27FBAE9252" bold="true" box="[1038,1052,1616,1637]" pageId="4" pageNumber="445">n</emphasis>
.
</taxonomicNameLabel>
, Miaoli County, Taiwan. 13, Biotope of
<taxonomicName id="4C1A4D396A286713FF7CFD19FEDB92B3" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[179,361,1646,1668]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A286713FF7CFD19FEDB92B3" box="[179,361,1646,1668]" italics="true" pageId="4" pageNumber="445">Endoclita atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A286713FEBFFD18FE1892B3" box="[368,426,1647,1668]" pageId="4" pageNumber="445" rank="species">
<emphasis id="B96EEAA86A286713FEBFFD18FE3A92B3" bold="true" box="[368,392,1647,1668]" pageId="4" pageNumber="445">sp</emphasis>
.
<emphasis id="B96EEAA86A286713FE59FD18FE1692B3" bold="true" box="[406,420,1647,1668]" pageId="4" pageNumber="445">n</emphasis>
.
</taxonomicNameLabel>
, Miaoli County, Taiwan. 14, Species numbers of
<taxonomicName id="4C1A4D396A286713FC67FD19FBA492B3" authorityName="Stephens" authorityYear="1829" box="[936,1046,1646,1668]" class="Insecta" family="Hepialidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
recorded from different countries and regions (data from Nielsen
<emphasis id="B96EEAA86A286713FE61FDF9FE6E9294" box="[430,476,1677,1699]" italics="true" pageId="4" pageNumber="445">et al</emphasis>
. (2000). 15, Regional comparison of
<taxonomicName id="4C1A4D396A286713FC93FDFAFC0D9294" authorityName="Felder" authorityYear="1874" box="[860,959,1677,1699]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA86A286713FC93FDFAFC0D9294" box="[860,959,1677,1699]" italics="true" pageId="4" pageNumber="445">Endoclita</emphasis>
</taxonomicName>
species numbers by major land units. 16, Grid of species distribution densities for
<taxonomicName id="4C1A4D396A286713FDC4FDDBFDDF92F5" authorityName="Felder" authorityYear="1874" box="[523,621,1708,1730]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="genus">
<emphasis id="B96EEAA86A286713FDC4FDDBFDDF92F5" box="[523,621,1708,1730]" italics="true" pageId="4" pageNumber="445">Endoclita</emphasis>
</taxonomicName>
. Grid units 15 x 15°. 17, distribution records for
<taxonomicName id="4C1A4D396A286713FBBAFDDAFB5292F5" authority="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan, 2018" authorityName="Buchsbaum &amp; Hsu &amp; Chen &amp; Grehan" authorityYear="2018" box="[1141,1248,1708,1730]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="atayala" status="sp. nov.">
<emphasis id="B96EEAA86A286713FBBAFDDAFB5292F5" box="[1141,1248,1708,1730]" italics="true" pageId="4" pageNumber="445">E. atayala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25D57D36A286713FB27FDDBFA9192F5" box="[1256,1315,1708,1730]" pageId="4" pageNumber="445" rank="species">
<emphasis id="B96EEAA86A286713FB27FDDBFAB292F6" bold="true" box="[1256,1280,1708,1729]" pageId="4" pageNumber="445">sp</emphasis>
.
<emphasis id="B96EEAA86A286713FAC0FDDBFAAF92F6" bold="true" box="[1295,1309,1708,1729]" pageId="4" pageNumber="445">n</emphasis>
.
</taxonomicNameLabel>
(red circle, Taiwan),
<taxonomicName id="4C1A4D396A286713FF38FDBCFE2792D7" baseAuthorityName="Le Cerf" baseAuthorityYear="1919" box="[247,405,1738,1760]" class="Insecta" family="Hepialidae" genus="Endoclita" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="445" phylum="Arthropoda" rank="species" species="crenilimbata">
<emphasis id="B96EEAA86A286713FF38FDBCFE2792D7" box="[247,405,1738,1760]" italics="true" pageId="4" pageNumber="445">E. crenilimbata</emphasis>
</taxonomicName>
(blue circles, Ping-fa, Giuzhou and Mupin, Sichuan) and
<emphasis id="B96EEAA86A286713FC26FDBCFBF092D7" box="[1001,1090,1738,1760]" italics="true" pageId="4" pageNumber="445">E. nodus</emphasis>
(white circles, Yuexi, Anhui, and Lushan, Jiangxi).
</paragraph>
</caption>
<subSubSection id="C30065316A286713FF08FC51FD329354" pageId="4" pageNumber="445" type="biology_ecology">
<paragraph id="8BA536BA6A286713FF08FC51FD329354" blockId="4.[151,1437,1830,2035]" pageId="4" pageNumber="445">
<emphasis id="B96EEAA86A286713FF08FC51FE959308" bold="true" box="[199,295,1830,1855]" pageId="4" pageNumber="445">Biology.</emphasis>
The July record for the
<typeStatus id="54A188186A286713FDFBFC50FDD69308" box="[564,612,1831,1855]" pageId="4" pageNumber="445">type</typeStatus>
specimen coincides with midsummer and season with the highest rainfall (Tourism Bureau,
<collectingCountry id="F30D762A6A286713FEACFC3DFD829354" box="[355,560,1866,1891]" name="Taiwan" pageId="4" pageNumber="445">
Republic of
<bibRefCitation id="EF8B4B4B6A286713FE22FC3DFD829354" author="Tourism Bureau &amp; Republic of China" box="[493,560,1866,1891]" pageId="4" pageNumber="445" refId="ref6562" refString="Tourism Bureau, Republic of China (2016) Climate. Available from: http: // eng. taiwan. net. tw / m 1. aspx? sNo = 0002005 (accessed 13 August 2018)" type="url" year="2016">China</bibRefCitation>
</collectingCountry>
2016).
</paragraph>
</subSubSection>
<subSubSection id="C30065316A286713FF08FC19FD0693C4" pageId="4" pageNumber="445" type="etymology">
<paragraph id="8BA536BA6A286713FF08FC19FD0693C4" blockId="4.[151,1437,1830,2035]" pageId="4" pageNumber="445">
<emphasis id="B96EEAA86A286713FF08FC19FEFC93B0" bold="true" box="[199,334,1902,1927]" pageId="4" pageNumber="445">Etymology.</emphasis>
The species name is of feminine gender in agreement with the generic name. The new species is named after the Taiwanese indigenous tribe living in the region around the
<typeStatus id="54A188186A286713FC38FCE4FB95939C" box="[1015,1063,1939,1963]" pageId="4" pageNumber="445">type</typeStatus>
locality. The Atayal (also called Tayal or Tayan) comprise the second largest indigenous population in
<collectingCountry id="F30D762A6A286713FC65FCC1FC4E93F8" box="[938,1020,1974,1999]" name="Taiwan" pageId="4" pageNumber="445">Taiwan</collectingCountry>
after the Amis in the east of
<collectingCountry id="F30D762A6A286713FA8CFCC1FA2B93F8" box="[1347,1433,1974,1999]" name="Taiwan" pageId="4" pageNumber="445">Taiwan</collectingCountry>
. The name means “real human” or “strong man”.
</paragraph>
</subSubSection>
<subSubSection id="C30065316A296712FF08FBE0FD819528" pageId="5" pageNumber="446" type="materials_examined">
<paragraph id="8BA536BA6A296712FF08FBE0FD819528" blockId="5.[151,1436,151,287]" pageId="5" pageNumber="446">
<emphasis id="B96EEAA86A296712FF08FBE0FECB9487" bold="true" box="[199,377,151,176]" pageId="5" pageNumber="446">Type material.</emphasis>
<materialsCitation id="3B723CE76A296712FE4BFBE0FD0094CF" ID-GBIF-Occurrence="2028521081" collectingDate="2008-07-24" collectorName="Mercury Lamp &amp; Li-Peng Hsu" country="Taiwan" county="Meiyun Village" elevation="2360" location="KuanWu Dalu Forest" municipality="KuanWu Dalu Forest Road East" pageId="5" pageNumber="446" specimenCount="1" stateProvince="Miaoli" typeStatus="holotype">
<typeStatus id="54A188186A296712FE4BFBE0FE599487" box="[388,491,151,176]" pageId="5" pageNumber="446" type="holotype">
<emphasis id="B96EEAA86A296712FE4BFBE0FE599487" box="[388,491,151,176]" italics="true" pageId="5" pageNumber="446">Holotype</emphasis>
</typeStatus>
female (with the following labels separated by forward slashes): /
<date id="FFA4107A6A296712FB33FBEFFA2A9487" box="[1276,1432,151,176]" pageId="5" pageNumber="446" value="2008-07-24">
<collectingDate id="EFE0E9926A296712FB33FBEFFA2A9487" box="[1276,1432,151,176]" pageId="5" pageNumber="446" value="2008-07-24">24 July 2008</collectingDate>
</date>
,
<collectingCountry id="F30D762A6A296712FF58FBCBFF5E94E2" box="[151,236,188,213]" name="Taiwan" pageId="5" pageNumber="446">Taiwan</collectingCountry>
,
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,
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,
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KuanWu
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Road
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Branch, 6 K,
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a.s.l.
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, 121°0514 E/ 24°2959 N,
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, leg.
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<emphasis id="B96EEAA86A296712FCF0FBA8FBB994CF" box="[831,1035,223,248]" italics="true" pageId="5" pageNumber="446">Endoclita atayala</emphasis>
Ulf Buchsbaum &amp; Hsu
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des 2016/
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0 1. (
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)
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.
</paragraph>
</subSubSection>
</treatment>
</document>